treatments-xml/data/03/EF/DD/03EFDD6EFFEEFF9144D5FC35FEDCFCDA.xml

<document id="66D7BDFE0FDD1E4ABDDD0A0E993CA469" ID="10.11646/zootaxa.4250.6.6SLASH10765" ID-CLB-Dataset="33262" ID-DOI="10.11646/zootaxa.4250.6.6" ID-GBIF-Dataset="d50fd45f-543b-4273-ab64-21074cc8ceda" ID-ISSN="1175-5326" ID-Zenodo-Dep="495872" ID-ZooBank="C2D552D3-8198-4EBC-8D14-6D1166A81601" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491811635627" checkinUser="plazi" docAuthor="Carrerette, Orlemir, Nogueira, João Miguel De Matos &amp; Hutchings, Pat" docDate="2017" docId="03EFDD6EFFEEFF9144D5FC35FEDCFCDA" docLanguage="en" docName="zootaxa.4250.6.6SLASH10765.pdf" docOrigin="Zootaxa 4250 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thelepus brevitori Carrerette, Nogueira &amp; Hutchings, 2017, n. sp." docType="treatment" docVersion="9" lastPageNumber="598" masterDocId="FFD6A516FFE6FF9A4442FF90FFEDFFCF" masterDocTitle="The genus Thelepus Leuckart, 1849 (Annelida, Thelepodidae) in Brazil, with a redescription of the holotype of T. setosus (Quatrefages, 1866)" masterLastPageNumber="599" masterPageNumber="587" pageNumber="595" updateTime="1698438819292" updateUser="ExternalLinkService">
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<mods:title id="EEDA4A5A5385179832810F6E606655D3">The genus Thelepus Leuckart, 1849 (Annelida, Thelepodidae) in Brazil, with a redescription of the holotype of T. setosus (Quatrefages, 1866)</mods:title>
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<mods:namePart id="6DCFC4CE09A8A01B58B26051B3C90223">Carrerette, Orlemir</mods:namePart>
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<treatment id="03EFDD6EFFEEFF9144D5FC35FEDCFCDA" ID-DOI="http://doi.org/10.5281/zenodo.5628882" ID-GBIF-Taxon="128344888" ID-Zenodo-Dep="5628882" LSID="urn:lsid:plazi:treatment:03EFDD6EFFEEFF9144D5FC35FEDCFCDA" httpUri="http://treatment.plazi.org/id/03EFDD6EFFEEFF9144D5FC35FEDCFCDA" lastPageId="11" lastPageNumber="598" pageId="8" pageNumber="595">
<subSubSection id="C35C3FF3FFEEFF9244D5FC35FE86FC2F" pageId="8" pageNumber="595" type="nomenclature">
<paragraph id="8BF96C78FFEEFF9244D5FC35FE50FC70" blockId="8.[151,445,933,992]" box="[151,445,933,959]" pageId="8" pageNumber="595">
<heading id="D0B1DB14FFEEFF9244D5FC35FE50FC70" bold="true" box="[151,445,933,959]" fontSize="11" level="1" pageId="8" pageNumber="595" reason="1">
<emphasis id="B932B06AFFEEFF9244D5FC35FE50FC70" bold="true" box="[151,445,933,959]" pageId="8" pageNumber="595">
<taxonomicName id="4C4617FBFFEEFF9244D5FC35FE99FC70" ID-CoL="5686F" box="[151,372,933,959]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="8" pageNumber="595" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEEFF9244D5FC35FE99FC70" bold="true" box="[151,372,933,959]" italics="true" pageId="8" pageNumber="595">Thelepus brevitori</emphasis>
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<taxonomicNameLabel id="A2010D11FFEEFF924539FC35FE50FC70" box="[379,445,933,959]" pageId="8" pageNumber="595" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BF96C78FFEEFF9244D5FC57FE86FC2F" blockId="8.[151,445,933,992]" box="[151,363,967,992]" pageId="8" pageNumber="595">
(
<figureCitation id="137D70FDFFEEFF9244DDFC57FEEEFC2F" box="[159,259,967,992]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1886,1908]" captionStartId-1="10.[151,250,1924,1946]" captionTargetBox-0="[157,1432,629,1857]" captionTargetBox-1="[155,1429,197,1895]" captionTargetId-0="figure@9.[151,1436,625,1865]" captionTargetId-1="figure@10.[151,1436,193,1902]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." captionText-1="FIGURE 6. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A – C). Notochaetae, segments 8 and 26 (2), respectively. (D – G), uncini, segments 5, 18, 33 (2), and 26, respectively. b = base of uncinus; c = crest with secondary rows of teeth; db = dorsal button; h = heel; mf = main fang; p = prow. Scale bars: A – C = 60 µm; D – F = 30 µm; F = 10 µm; G – H = 20 µm." httpUri-0="https://zenodo.org/record/495877/files/figure.png" httpUri-1="https://zenodo.org/record/495878/files/figure.png" pageId="8" pageNumber="595">Figs 5–6</figureCitation>
, 
<tableCitation id="C6C459C3FFEEFF92454DFC57FE8DFC2F" box="[271,352,967,992]" captionStart="TABLE 1" captionStartId="4.[151,239,613,636]" captionTargetBox="[159,1419,664,1376]" captionTargetPageId="4" captionText="TABLE 1. Morphological variation within the type-series of Thelepus megalabiatum n. sp. and Thelepus brevitori n. sp." httpUri="http://table.plazi.org/id/DF393CF0FFE2FF9E44D5FDF5FA71FDB3" pageId="8" pageNumber="595" tableUuid="DF393CF0FFE2FF9E44D5FDF5FA71FDB3">Table 1</tableCitation>
)
</paragraph>
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<paragraph id="8BF96C78FFEEFF9244D5FB9AFC7DFBEF" blockId="8.[151,912,1034,1056]" box="[151,912,1034,1056]" pageId="8" pageNumber="595">
<emphasis id="B932B06AFFEEFF9244D5FB9AFEA4FBEF" box="[151,329,1034,1056]" italics="true" pageId="8" pageNumber="595">
<taxonomicName id="4C4617FBFFEEFF9244D5FB9AFEA8FBEF" box="[151,325,1034,1056]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="8" pageNumber="595" phylum="Annelida" rank="species" species="setosus">Thelepus setosus</taxonomicName>
.
</emphasis>
(Non Quatrefages) 
<bibRefCitation id="EFD71189FFEEFF924650FB9AFD35FBEF" author="Blankensteyn" box="[530,728,1034,1056]" pageId="8" pageNumber="595" refString="Blankensteyn, A. (1988) Terebellidae e Trichobranchidae (Annelida: Polychaeta) da Costa Sudeste do Brasil (24 º - 27 ºS). M. Sc. Dissertation, Instituto de Ciencias Biologicas, Universidade Federal do Parana, Curitiba, 178 pp." type="book" year="1988">Blankensteyn 1988</bibRefCitation>
: p. 74–77, fig. 18.
</paragraph>
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<subSubSection id="C35C3FF3FFEEFF9244D5FBDDFE95FAF1" pageId="8" pageNumber="595" type="materials_examined">
<paragraph id="8BF96C78FFEEFF9244D5FBDDFE95FAF1" blockId="8.[151,1437,1101,2026]" pageId="8" pageNumber="595">
<materialsCitation id="3B2E6625FFEEFF9244D5FBDDFD39FB61" ID-GBIF-Occurrence="1455758453" collectingDate="1997-12-16" collectionCode="BIOTA, MZUSP" country="Brazil" county="Type" elevation="50" latitude="-25.198166" location="Benthic" longLatPrecision="12" longitude="-47.134834" municipality="Atlantic Ocean" pageId="8" pageNumber="595" specimenCode="MZUSP 3020, MZUSP 3045" specimenCount="3" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="B932B06AFFEEFF9244D5FBDDFECBFBA9" bold="true" box="[151,294,1101,1126]" pageId="8" pageNumber="595">
<typeStatus id="54FDD2DAFFEEFF9244D5FBDDFF3FFBA9" box="[151,210,1101,1126]" pageId="8" pageNumber="595">
<collectingCounty id="629814F4FFEEFF9244D5FBDDFF3FFBA9" box="[151,210,1101,1126]" pageId="8" pageNumber="595">Type</collectingCounty>
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series.
</emphasis>
<collectingMunicipality id="6B9DF602FFEEFF924573FBDDFE0CFBA9" box="[305,481,1101,1126]" pageId="8" pageNumber="595">Atlantic Ocean</collectingMunicipality>
, southeastern 
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, state of 
<collectingRegion id="4982A29AFFEEFF92477DFBDEFC5AFBA9" box="[831,951,1101,1126]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="595">São Paulo</collectingRegion>
, project ‘ 
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/FAPESP/ 
<location id="8E993AA3FFEEFF9240AFFBDDFAAEFBA9" LSID="urn:lsid:plazi:treatment:03EFDD6EFFEEFF9144D5FC35FEDCFCDA:8E993AA3FFEEFF9240AFFBDDFAAEFBA9" box="[1261,1347,1101,1126]" country="Brazil" county="Type" latitude="-25.198166" longLatPrecision="12" longitude="-47.134834" municipality="Atlantic Ocean" name="Benthic" pageId="8" pageNumber="595" stateProvince="Sao Paulo">Benthic</location>
marine biodiversity in the state of 
<location id="8E993AA3FFEEFF924598FBE2FDB0FB45" LSID="urn:lsid:plazi:treatment:03EFDD6EFFEEFF9144D5FC35FEDCFCDA:8E993AA3FFEEFF924598FBE2FDB0FB45" box="[474,605,1137,1162]" country="Brazil" county="Type" latitude="-25.198166" longLatPrecision="12" longitude="-47.134834" municipality="Atlantic Ocean" name="Sao Paulo'" pageId="8" pageNumber="595" stateProvince="Sao Paulo">São Paulo’</location>
: 
<typeStatus id="54FDD2DAFFEEFF92462FFBE1FD38FB45" box="[621,725,1137,1162]" pageId="8" pageNumber="595" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBE0C403FFEEFF9246ABFBE2FC60FB45" box="[745,909,1138,1162]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="595">
<collectionCode id="ED57F4BDFFEEFF9246ABFBE2FCA5FB45" box="[745,840,1138,1162]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="595">MZUSP</collectionCode>
3020
</specimenCode>
) and 
<typeStatus id="54FDD2DAFFEEFF924797FBE2FBD8FB45" box="[981,1077,1138,1162]" pageId="8" pageNumber="595" type="paratype">paratype</typeStatus>
7 (
<specimenCode id="DBE0C403FFEEFF924021FBE2FAE7FB45" box="[1123,1290,1137,1162]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="595">
<collectionCode id="ED57F4BDFFEEFF924021FBE2FB2FFB45" box="[1123,1218,1138,1162]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="595">MZUSP</collectionCode>
3045
</specimenCode>
): 
<geoCoordinate id="EE720ABFFFEEFF924160FBE1FA71FB45" box="[1314,1436,1137,1162]" direction="south" orientation="latitude" pageId="8" pageNumber="595" precision="9" value="-25.198166">25°11.89'S</geoCoordinate>
<geoCoordinate id="EE720ABFFFEEFF9244D5FB06FEF0FB61" box="[151,285,1174,1198]" direction="west" orientation="longitude" pageId="8" pageNumber="595" precision="9" value="-47.134834">47°08.09'W</geoCoordinate>
, 
<quantity id="4CBEC19DFFEEFF924565FB05FE8DFB62" box="[295,352,1173,1198]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="595" unit="m" value="50.0">
<elevation id="006B8B4BFFEEFF924565FB05FE8DFB62" box="[295,352,1173,1198]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="595" unit="m" value="50.0">50 m</elevation>
</quantity>
deep, in sand, coll. 
<date id="FFF84AB8FFEEFF92467EFB05FD39FB61" box="[572,724,1173,1198]" pageId="8" pageNumber="595" value="1997-12-16">
<collectingDate id="EFBCB350FFEEFF92467EFB05FD39FB61" box="[572,724,1173,1198]" pageId="8" pageNumber="595" value="1997-12-16">16 Dec. 1997</collectingDate>
</date>
</materialsCitation>
; paratypes 1–4 (MZUSP 3021, 3022, 3023, 3024, respectively): 23°42'541&quot;S 45°11'380&quot;W, 
<quantity id="4CBEC19DFFEEFF924594FB29FDC4FB1E" box="[470,553,1209,1234]" metricMagnitude="1" metricUnit="m" metricValue="2.53" pageId="8" pageNumber="595" unit="m" value="25.3">25.3 m</quantity>
deep, in sand, coll. 
<date id="FFF84AB8FFEEFF92475CFB2AFC59FB1D" box="[798,948,1209,1234]" pageId="8" pageNumber="595" value="2001-07-29">29 Jul. 2001</date>
; paratype 5 (MZUSP 3046): 23°25'663&quot;S 44°46'570&quot;W, 
<quantity id="4CBEC19DFFEEFF924579FB4DFE99FB3A" box="[315,372,1245,1270]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="595" unit="m" value="35.0">35 m</quantity>
deep, in sand, coll. 
<date id="FFF84AB8FFEEFF924615FB4EFD07FB39" box="[599,746,1246,1270]" pageId="8" pageNumber="595" value="2001-06-11">11 Jun. 2001</date>
; paratype 6 (MZUSP 3047): 23°45'062&quot;S 45°13'856&quot;W, 
<quantity id="4CBEC19DFFEEFF924129FB4DFF40FAD6" metricMagnitude="1" metricUnit="m" metricValue="2.53" pageId="8" pageNumber="595" unit="m" value="25.3">25.3 m</quantity>
deep, in sand, coll. 
<date id="FFF84AB8FFEEFF9245CCFA92FDCFFAD5" box="[398,546,1282,1306]" pageId="8" pageNumber="595" value="2001-10-18">18 Oct. 2001</date>
. Data on the range of variation of numerical characters within the type series is provided in 
<tableCitation id="C6C459C3FFEEFF92455CFAB5FE9FFAF1" box="[286,370,1317,1342]" captionStart="TABLE 1" captionStartId="4.[151,239,613,636]" captionTargetBox="[159,1419,664,1376]" captionTargetPageId="4" captionText="TABLE 1. Morphological variation within the type-series of Thelepus megalabiatum n. sp. and Thelepus brevitori n. sp." httpUri="http://table.plazi.org/id/DF393CF0FFE2FF9E44D5FDF5FA71FDB3" pageId="8" pageNumber="595" tableUuid="DF393CF0FFE2FF9E44D5FDF5FA71FDB3">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C3FF3FFEEFF934485FAD9FE96FED2" lastPageId="9" lastPageNumber="596" pageId="8" pageNumber="595" type="description">
<paragraph id="8BF96C78FFEEFF934485FAD9FE96FED2" blockId="8.[151,1437,1101,2026]" lastBlockId="9.[151,1437,151,573]" lastPageId="9" lastPageNumber="596" pageId="8" pageNumber="595">
<emphasis id="B932B06AFFEEFF924485FAD9FEB4FAAD" bold="true" box="[199,345,1353,1378]" pageId="8" pageNumber="595">Description.</emphasis>
Preserved body uniformly beige to yellowish, without distinct patterns of pigmentation (
<figureCitation id="137D70FDFFEEFF92417AFAD9FA9AFAAD" box="[1336,1399,1353,1378]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A– G). All specimens incomplete, longest specimens with 68 (68) segments, 26 (25) mm long and 4 (3) mm wide. Transverse prostomium attached to dorsal surface of upper lip; basal part thick, with two thin irregular rows of eyespots continuous throughout, eyespots more separated from each other mid-dorsally (
<figureCitation id="137D70FDFFEEFF92403EFA25FB50FA01" box="[1148,1213,1461,1486]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
D–E, G); distal part of prostomium low, restricted to base of upper lip (
<figureCitation id="137D70FDFFEEFF924694FA49FCF5FA3D" box="[726,792,1497,1522]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
D–E, G); buccal tentacles long and thin, reaching around segment 20 if directed posteriorly (
<figureCitation id="137D70FDFFEEFF924660FA6DFD89F9D9" box="[546,612,1533,1558]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A). Peristomium restricted to lips, upper lip short, thick, hood-like, wider than long; lower lip short, swollen, rectangular, wider than long, surrounded by mid-ventral lobe originated from segment 1 (
<figureCitation id="137D70FDFFEEFF92455AF9D5FEB7F991" box="[280,346,1605,1630]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
C, F). Anterior body highly glandular ventrally, with thick, crenulated to slightly corrugated body wall between neuropodia, progressively narrower until segment 17 or slightly more posteriorly (17) (
<figureCitation id="137D70FDFFEEFF924156F9F9FAB5F94D" box="[1300,1368,1641,1666]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A, C– F); segment 1 with ventral lobe around lower lip, marginal to mouth; segments 2–4 shorter, then segments increasing progressively in length very slightly until mid-body (
<figureCitation id="137D70FDFFEEFF924727F921FC4BF905" box="[869,934,1713,1738]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A–F). Three pairs of branchiae, on segments 2–4, segment 2 with 10–18 (15) branchial filaments, segment 3 with 5–10 (10) filaments, segment 4 with 4–8 (8) filaments, leaving wide mid-dorsal gap; origin of filaments of segment 2 dorsal to level of notopodia, not extending laterally; branchial filaments originating directly from body wall, cushion-like areas absent (
<figureCitation id="137D70FDFFEEFF9240F7F88DFB14F8F9" box="[1205,1273,1821,1846]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
G). Notopodia present until segments 
<quantity id="4CBEC19DFFEEFF9245DCF8D1FDDDF895" box="[414,560,1857,1882]" metricMagnitude="-1" metricUnit="m" metricValue="7.02183" metricValueMax="7.439659999999999" metricValueMin="6.604" pageId="8" pageNumber="595" unit="in" value="27.645" valueMax="29.29" valueMin="26.0">26–29, 29 in</quantity>
most specimens, including 
<typeStatus id="54FDD2DAFFEEFF924732F8D1FC38F895" box="[880,981,1857,1882]" pageId="8" pageNumber="595" type="holotype">holotype</typeStatus>
; relatively short and bilobed notopodia, lobes of equal size, notopodia progressively longer and more laterally placed on segments 3–6, then longitudinally aligned laterally (
<figureCitation id="137D70FDFFEEFF924522F819FE49F86D" box="[352,420,1929,1954]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A–B, D–E, G). Narrowly-winged notochaetae in both rows, poorly marked difference in length between rows, shafts expanded at bases of wings, rounded, especially in anterior row (
<figureCitation id="137D70FDFFEEFF92414CF83DFABAF809" box="[1294,1367,1965,1990]" captionStart="FIGURE 6" captionStartId="10.[151,250,1924,1946]" captionTargetBox="[155,1429,197,1895]" captionTargetId="figure@10.[151,1436,193,1902]" captionTargetPageId="10" captionText="FIGURE 6. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A – C). Notochaetae, segments 8 and 26 (2), respectively. (D – G), uncini, segments 5, 18, 33 (2), and 26, respectively. b = base of uncinus; c = crest with secondary rows of teeth; db = dorsal button; h = heel; mf = main fang; p = prow. Scale bars: A – C = 60 µm; D – F = 30 µm; F = 10 µm; G – H = 20 µm." httpUri="https://zenodo.org/record/495878/files/figure.png" pageId="8" pageNumber="595">Fig. 6</figureCitation>
A–C). Neuropodia as low, fleshy ridges on anterior segments, narrow tori of uniform width, laterally aligned (
<figureCitation id="137D70FDFFEEFF924154F841FABAF825" box="[1302,1367,2001,2026]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="8" pageNumber="595">Fig. 5</figureCitation>
A, D– E), as raised cushion-like pinnules after notopodia terminate (
<figureCitation id="137D70FDFFEFFF93472CFF07FC5EFF7F" box="[878,947,151,176]" captionStart="FIGURE 5" captionStartId="9.[151,250,1886,1908]" captionTargetBox="[157,1432,629,1857]" captionTargetId="figure@9.[151,1436,625,1865]" captionTargetPageId="9" captionText="FIGURE 5. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B – E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F – G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l 1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B – G = 0.3 mm." httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="9" pageNumber="596">Fig. 5</figureCitation>
A–B); uncini with terminal dorsal button, remarkably short prow, as protruding knob, base relatively large and convex, and crest with 2 rows of secondary teeth, each with 2 teeth, medial indentation absent (
<figureCitation id="137D70FDFFEFFF934744FF4FFCA3FF37" box="[774,846,223,248]" captionStart="FIGURE 6" captionStartId="10.[151,250,1924,1946]" captionTargetBox="[155,1429,197,1895]" captionTargetId="figure@10.[151,1436,193,1902]" captionTargetPageId="10" captionText="FIGURE 6. Thelepus brevitori n. sp. (Holotype, MZUSP 3020), (A – C). Notochaetae, segments 8 and 26 (2), respectively. (D – G), uncini, segments 5, 18, 33 (2), and 26, respectively. b = base of uncinus; c = crest with secondary rows of teeth; db = dorsal button; h = heel; mf = main fang; p = prow. Scale bars: A – C = 60 µm; D – F = 30 µm; F = 10 µm; G – H = 20 µm." httpUri="https://zenodo.org/record/495878/files/figure.png" pageId="9" pageNumber="596">Fig. 6</figureCitation>
D–H). Nephridial and genital papillae not visible. Pygidium unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35C3FF3FFEFFF914485FEB7FED2FD4A" lastPageId="11" lastPageNumber="598" pageId="9" pageNumber="596" type="discussion">
<paragraph id="8BF96C78FFEFFF934485FEB7FBBFFDF3" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="596">
<emphasis id="B932B06AFFEFFF934485FEB7FED6FE8F" bold="true" box="[199,315,295,320]" pageId="9" pageNumber="596">Remarks.</emphasis>
Members of 
<taxonomicName id="4C4617FBFFEFFF9345A4FEB9FD87FE8F" box="[486,618,295,320]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEFFF9345A4FEB9FD87FE8F" box="[486,618,295,320]" italics="true" pageId="9" pageNumber="596">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEFFF93463BFEB7FD52FE8F" bold="true" box="[633,703,295,320]" pageId="9" pageNumber="596">
<taxonomicNameLabel id="A2010D11FFEFFF93463BFEB7FD52FE8F" box="[633,703,295,320]" pageId="9" pageNumber="596" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differ from both species described above, 
<taxonomicName id="4C4617FBFFEFFF93408DFEB9FAAEFE8F" box="[1231,1347,297,320]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="setosus">
<emphasis id="B932B06AFFEFFF93408DFEB9FAAEFE8F" box="[1231,1347,297,320]" italics="true" pageId="9" pageNumber="596">T. setosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4617FBFFEFFF9341C5FEB9FEDAFEAA" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="megalabiatum">
<emphasis id="B932B06AFFEFFF9341C5FEB9FEDAFEAA" italics="true" pageId="9" pageNumber="596">T. megalabiatum</emphasis>
</taxonomicName>
, in the morphology of branchiae, with few and relatively short and thick filaments, leaving a wide mid-dorsal gap, and in only having up to 27 pairs of notopodia, extending until segment 29. In contrast, in members of both other species the branchiae have more filaments and these are longer and thinner, originating from cushionlike areas in 
<taxonomicName id="4C4617FBFFEFFF934566FE29FE32FE1F" box="[292,479,439,464]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="megalabiatum">
<emphasis id="B932B06AFFEFFF934566FE29FE32FE1F" box="[292,479,439,464]" italics="true" pageId="9" pageNumber="596">T. megalabiatum</emphasis>
</taxonomicName>
, and notopodia extending until segment 
<quantity id="4CBEC19DFFEFFF9347E5FE27FC32FE1F" box="[935,991,439,464]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="9" pageNumber="596" unit="in" value="56.0">56 in</quantity>
the 
<typeStatus id="54FDD2DAFFEFFF93404DFE27FB9CFE1F" box="[1039,1137,439,464]" pageId="9" pageNumber="596" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C4617FBFFEFFF9340D7FE29FAEFFE1F" box="[1173,1282,441,464]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="setosus">
<emphasis id="B932B06AFFEFFF9340D7FE29FAEFFE1F" box="[1173,1282,441,464]" italics="true" pageId="9" pageNumber="596">T. setosus</emphasis>
</taxonomicName>
, and segment 44–61+ in the case of members of 
<taxonomicName id="4C4617FBFFEFFF93466DFE4DFD00FE3A" box="[559,749,476,501]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="megalabiatum">
<emphasis id="B932B06AFFEFFF93466DFE4DFD00FE3A" box="[559,749,476,501]" italics="true" pageId="9" pageNumber="596">T. megalabiatum</emphasis>
</taxonomicName>
(
<tableCitation id="C6C459C3FFEFFF9346BDFE4CFCB4FE3B" box="[767,857,476,501]" captionStart="TABLE 2" captionStartId="7.[151,239,620,643]" captionTargetBox="[159,1431,743,1126]" captionTargetPageId="7" captionText="TABLE 2. Comparison between the most important numerical characters of the species of Thelepus most similar to T. megalabiatum n. sp. and T. brevitori n. sp. Sources: Hutchings &amp; Glasby (1987); Londoño-Mesa (2009); Capa &amp; Hutchings (2006); characters of T. setosus examined directly from type-material." httpUri="http://table.plazi.org/id/DF393CF0FFE1FF9D44D5FDFCFC19FD0C" pageId="9" pageNumber="596" tableUuid="DF393CF0FFE1FF9D44D5FDFCFC19FD0C">Table 2</tableCitation>
). In addition, 
<taxonomicName id="4C4617FBFFEFFF9347BCFE4CFB23FE3A" box="[1022,1230,476,501]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEFFF9347BCFE4CFB23FE3A" box="[1022,1230,476,501]" italics="true" pageId="9" pageNumber="596">Thelepus brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEFFF93409AFE4CFAF7FE3A" bold="true" box="[1240,1306,476,501]" pageId="9" pageNumber="596">
<taxonomicNameLabel id="A2010D11FFEFFF93409AFE4CFAF7FE3A" box="[1240,1306,476,501]" pageId="9" pageNumber="596" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, is a much smaller species, as the 
<typeStatus id="54FDD2DAFFEFFF9345DAFE6FFE17FDD7" box="[408,506,511,536]" pageId="9" pageNumber="596" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C4617FBFFEFFF934662FD91FD60FDD7" box="[544,653,513,536]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="setosus">
<emphasis id="B932B06AFFEFFF934662FD91FD60FDD7" box="[544,653,513,536]" italics="true" pageId="9" pageNumber="596">T. setosus</emphasis>
</taxonomicName>
has ~93 segments and it is ~ 
<quantity id="4CBEC19DFFEFFF934791FD90FBDDFDD8" box="[979,1072,512,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="596" unit="mm" value="110.0">110 mm</quantity>
long and 
<quantity id="4CBEC19DFFEFFF9340E2FD90FB1BFDD8" box="[1184,1270,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="596" unit="mm" value="3.9">3.9 mm</quantity>
wide, and that of 
<taxonomicName id="4C4617FBFFEFFF9344F7FDB5FE9CFDF2" box="[181,369,548,573]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="megalabiatum">
<emphasis id="B932B06AFFEFFF9344F7FDB5FE9CFDF2" box="[181,369,548,573]" italics="true" pageId="9" pageNumber="596">T. megalabiatum</emphasis>
</taxonomicName>
has 78 segments and it is 
<quantity id="4CBEC19DFFEFFF9346D4FDB4FD1FFDF3" box="[662,754,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="9" pageNumber="596" unit="mm" value="113.0">113 mm</quantity>
long and 
<quantity id="4CBEC19DFFEFFF934723FDB5FC4FFDF3" box="[865,930,549,572]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="596" unit="mm" value="7.0">7 mm</quantity>
wide (
<tableCitation id="C6C459C3FFEFFF9347AFFDB4FBA8FDF3" box="[1005,1093,548,573]" captionStart="TABLE 2" captionStartId="7.[151,239,620,643]" captionTargetBox="[159,1431,743,1126]" captionTargetPageId="7" captionText="TABLE 2. Comparison between the most important numerical characters of the species of Thelepus most similar to T. megalabiatum n. sp. and T. brevitori n. sp. Sources: Hutchings &amp; Glasby (1987); Londoño-Mesa (2009); Capa &amp; Hutchings (2006); characters of T. setosus examined directly from type-material." httpUri="http://table.plazi.org/id/DF393CF0FFE1FF9D44D5FDFCFC19FD0C" pageId="9" pageNumber="596" tableUuid="DF393CF0FFE1FF9D44D5FDFCFC19FD0C">Table 2</tableCitation>
).
</paragraph>
<caption id="DF393CF0FFEFFF9344D5F8CEFD57F820" httpUri="https://zenodo.org/record/495877/files/figure.png" pageId="9" pageNumber="596" targetBox="[157,1432,629,1857]" targetPageId="9">
<paragraph id="8BF96C78FFEFFF9344D5F8CEFD57F820" blockId="9.[151,1436,1886,2031]" pageId="9" pageNumber="596">
<emphasis id="B932B06AFFEFFF9344D5F8CEFEF8F8BB" bold="true" box="[151,277,1886,1908]" pageId="9" pageNumber="596">FIGURE 5.</emphasis>
<taxonomicName id="4C4617FBFFEFFF93455DF8CEFE31F8BB" box="[287,476,1886,1908]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="9" pageNumber="596" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEFFF93455DF8CEFE31F8BB" box="[287,476,1886,1908]" italics="true" pageId="9" pageNumber="596">Thelepus brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEFFF9345A7F8CEFDCCF8BB" bold="true" box="[485,545,1886,1908]" pageId="9" pageNumber="596">
<taxonomicNameLabel id="A2010D11FFEFFF9345A7F8CEFDCCF8BB" box="[485,545,1886,1908]" pageId="9" pageNumber="596" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Holotype, MZUSP 3020), (A). Incomplete worm, left lateral view; (B–E). Anterior body, dorsal, ventral, left and right lateral views, respectively; (F–G). Close ups of the anterior end, ventral and dorsal views, respectively. Numbers refer to the segments; e = eyespots; g = glandular ventral areas; ll = lower lip; l1 = lobe on segment 1; no = notopodia; n = neuropodium; P = basal part of prostomium; po = pharyngeal organ; ul = upper lip; * distal part of prostomium. Scale bars: A = 0.5 mm; B–G = 0.3 mm.
</paragraph>
</caption>
<caption id="DF393CF0FFECFF9044D5F814FAE5F818" httpUri="https://zenodo.org/record/495878/files/figure.png" pageId="10" pageNumber="597" targetBox="[155,1429,197,1895]" targetPageId="10">
<paragraph id="8BF96C78FFECFF9044D5F814FAE5F818" blockId="10.[151,1436,1924,2007]" pageId="10" pageNumber="597">
<emphasis id="B932B06AFFECFF9044D5F814FEFFF855" bold="true" box="[151,274,1924,1946]" pageId="10" pageNumber="597">FIGURE 6.</emphasis>
<taxonomicName id="4C4617FBFFECFF90455BF814FE3FF855" box="[281,466,1924,1946]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="10" pageNumber="597" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFECFF90455BF814FE3FF855" box="[281,466,1924,1946]" italics="true" pageId="10" pageNumber="597">Thelepus brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFECFF90459AF814FDFDF855" bold="true" box="[472,528,1924,1946]" pageId="10" pageNumber="597">
<taxonomicNameLabel id="A2010D11FFECFF90459AF814FDFDF855" box="[472,528,1924,1946]" pageId="10" pageNumber="597" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Holotype, MZUSP 3020), (A–C). Notochaetae, segments 8 and 26 (2), respectively. (D– G), uncini, segments 5, 18, 33 (2), and 26, respectively. b = base of uncinus; c = crest with secondary rows of teeth; db = dorsal button; h = heel; mf = main fang; p = prow. Scale bars: A–C = 60 µm; D–F = 30 µm; F = 10 µm; G–H = 20 µm.
</paragraph>
</caption>
<paragraph id="8BF96C78FFEDFF914485FF08FA71FEAB" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="598">
In regards to the other species previously identified as 
<taxonomicName id="4C4617FBFFEDFF914767FF09FC7CFF7F" box="[805,913,153,176]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="setosus">
<emphasis id="B932B06AFFEDFF914767FF09FC7CFF7F" box="[805,913,153,176]" italics="true" pageId="11" pageNumber="598">T. setosus</emphasis>
</taxonomicName>
, i.e., 
<taxonomicName id="4C4617FBFFEDFF91478FFF09FBBEFF7F" box="[973,1107,151,176]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="haitiensis">
<emphasis id="B932B06AFFEDFF91478FFF09FBBEFF7F" box="[973,1107,151,176]" italics="true" pageId="11" pageNumber="598">T. haitiensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4617FBFFEDFF9140CBFF09FADDFF7F" box="[1161,1328,151,176]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="fraggleorum">
<emphasis id="B932B06AFFEDFF9140CBFF09FADDFF7F" box="[1161,1328,151,176]" italics="true" pageId="11" pageNumber="598">T. fraggleorum</emphasis>
</taxonomicName>
, from the Caribbean, and 
<taxonomicName id="4C4617FBFFEDFF914509FF2DFE2BFF1B" box="[331,454,189,212]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="extensus">
<emphasis id="B932B06AFFEDFF914509FF2DFE2BFF1B" box="[331,454,189,212]" italics="true" pageId="11" pageNumber="598">T. extensus</emphasis>
</taxonomicName>
, from 
<collectingCountry id="F3512CE8FFEDFF914655FF2CFD6CFF1A" box="[535,641,188,213]" name="Australia" pageId="11" pageNumber="598">Australia</collectingCountry>
, members of 
<taxonomicName id="4C4617FBFFEDFF91475CFF2DFC70FF1A" box="[798,925,188,213]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEDFF91475CFF2DFC70FF1A" box="[798,925,188,213]" italics="true" pageId="11" pageNumber="598">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEDFF9147E7FF2CFC0BFF1A" bold="true" box="[933,998,188,213]" pageId="11" pageNumber="598">
<taxonomicNameLabel id="A2010D11FFEDFF9147E7FF2CFC0BFF1A" box="[933,998,188,213]" pageId="11" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
also differ mostly in numbers of both branchial filaments and pairs of notopodia. Specimens of 
<taxonomicName id="4C4617FBFFEDFF914774FF71FC2DFF37" box="[822,960,223,248]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="haitiensis">
<emphasis id="B932B06AFFEDFF914774FF71FC2DFF37" box="[822,960,223,248]" italics="true" pageId="11" pageNumber="598">T. haitiensis</emphasis>
</taxonomicName>
have many more filaments, arising from cushion-like bases, and 41–46 pairs of notopodia, and members of 
<taxonomicName id="4C4617FBFFEDFF91473CFE95FBC8FED2" box="[894,1061,260,285]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="fraggleorum">
<emphasis id="B932B06AFFEDFF91473CFE95FBC8FED2" box="[894,1061,260,285]" italics="true" pageId="11" pageNumber="598">T. fraggleorum</emphasis>
</taxonomicName>
have similar number of branchial filaments to that observed in 
<taxonomicName id="4C4617FBFFEDFF91459CFEB9FDB6FE8F" box="[478,603,295,320]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEDFF91459CFEB9FDB6FE8F" box="[478,603,295,320]" italics="true" pageId="11" pageNumber="598">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEDFF914620FEB7FD4FFE8F" bold="true" box="[610,674,295,320]" pageId="11" pageNumber="598">
<taxonomicNameLabel id="A2010D11FFEDFF914620FEB7FD4FFE8F" box="[610,674,295,320]" pageId="11" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(see 
<tableCitation id="C6C459C3FFEDFF91469FFEB7FCDFFE8F" box="[733,818,295,320]" captionStart="TABLE 2" captionStartId="7.[151,239,620,643]" captionTargetBox="[159,1431,743,1126]" captionTargetPageId="7" captionText="TABLE 2. Comparison between the most important numerical characters of the species of Thelepus most similar to T. megalabiatum n. sp. and T. brevitori n. sp. Sources: Hutchings &amp; Glasby (1987); Londoño-Mesa (2009); Capa &amp; Hutchings (2006); characters of T. setosus examined directly from type-material." httpUri="http://table.plazi.org/id/DF393CF0FFE1FF9D44D5FDFCFC19FD0C" pageId="11" pageNumber="598" tableUuid="DF393CF0FFE1FF9D44D5FDFCFC19FD0C">Table 2</tableCitation>
), but they have notopodia extending up to segment 72. Members of 
<taxonomicName id="4C4617FBFFEDFF914567FEDDFE72FEAB" box="[293,415,333,356]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="extensus">
<emphasis id="B932B06AFFEDFF914567FEDDFE72FEAB" box="[293,415,333,356]" italics="true" pageId="11" pageNumber="598">T. extensus</emphasis>
</taxonomicName>
also have many more branchial filaments and notopodia extend until segment 37 (
<tableCitation id="C6C459C3FFEDFF914175FEDCFA63FEAB" box="[1335,1422,332,357]" captionStart="TABLE 2" captionStartId="7.[151,239,620,643]" captionTargetBox="[159,1431,743,1126]" captionTargetPageId="7" captionText="TABLE 2. Comparison between the most important numerical characters of the species of Thelepus most similar to T. megalabiatum n. sp. and T. brevitori n. sp. Sources: Hutchings &amp; Glasby (1987); Londoño-Mesa (2009); Capa &amp; Hutchings (2006); characters of T. setosus examined directly from type-material." httpUri="http://table.plazi.org/id/DF393CF0FFE1FF9D44D5FDFCFC19FD0C" pageId="11" pageNumber="598" tableUuid="DF393CF0FFE1FF9D44D5FDFCFC19FD0C">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BF96C78FFEDFF914485FEE0FBF0FDD7" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="598">
Two other species share with 
<taxonomicName id="4C4617FBFFEDFF91465EFEE1FD77FE47" box="[540,666,367,392]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEDFF91465EFEE1FD77FE47" box="[540,666,367,392]" italics="true" pageId="11" pageNumber="598">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEDFF9146E1FEFFFD09FE47" bold="true" box="[675,740,367,392]" pageId="11" pageNumber="598">
<taxonomicNameLabel id="A2010D11FFEDFF9146E1FEFFFD09FE47" box="[675,740,367,392]" pageId="11" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
the presence of ~30 pairs of notopodia. These species are 
<taxonomicName id="4C4617FBFFEDFF9141C5FEE1FE59FE63" authority="Fauchald, 1977" authorityName="Fauchald" authorityYear="1977" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="pascua">
<emphasis id="B932B06AFFEDFF9141C5FEE1FF0BFE63" italics="true" pageId="11" pageNumber="598">T. pascua</emphasis>
(
<bibRefCitation id="EFD71189FFEDFF9144B5FE04FE46FE63" author="Fauchald" box="[247,427,404,429]" pageId="11" pageNumber="598" refString="Fauchald, K. (1977) Polychaetes from intertidal areas in Panama, with a review of previous shallow-water records. Smithsonian Contributions to Zoology, 221, 1 - 81." type="journal article" year="1977">Fauchald, 1977</bibRefCitation>
)
</taxonomicName>
, from 
<collectingCountry id="F3512CE8FFEDFF914640FE05FDB0FE63" box="[514,605,405,428]" name="Panama" pageId="11" pageNumber="598">
<collectingRegion id="4982A29AFFEDFF914640FE05FDB0FE63" box="[514,605,405,428]" country="Panama" name="Panama" pageId="11" pageNumber="598">Panama</collectingRegion>
</collectingCountry>
, and 
<taxonomicName id="4C4617FBFFEDFF9146DFFE05FBA5FE63" authority="Treadwell, 1901" authorityName="Treadwell" authorityYear="1901" box="[669,1096,404,429]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="crassibranchiatus">
<emphasis id="B932B06AFFEDFF9146DFFE05FC6EFE62" box="[669,899,404,429]" italics="true" pageId="11" pageNumber="598">T. crassibranchiatus</emphasis>
<bibRefCitation id="EFD71189FFEDFF9147CFFE04FBA5FE63" author="Treadwell" box="[909,1096,404,429]" pageId="11" pageNumber="598" refString="Treadwell, A. L. (1901) The polychaetous annelids of Porto Rico. Bulletin of the United States Fish Commission, 20 (2), 181 - 210." type="journal article" year="1901">Treadwell, 1901</bibRefCitation>
</taxonomicName>
, from 
<collectingCountry id="F3512CE8FFEDFF9140D5FE05FACFFE62" box="[1175,1314,404,429]" name="Puerto Rico" pageId="11" pageNumber="598">Puerto Rico</collectingCountry>
. However, both these species have only two pairs of branchiae, instead of three pairs as in members of 
<taxonomicName id="4C4617FBFFEDFF9140E0FE29FAF3FE1F" box="[1186,1310,439,464]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEDFF9140E0FE29FAF3FE1F" box="[1186,1310,439,464]" italics="true" pageId="11" pageNumber="598">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEDFF914164FE27FA88FE1F" bold="true" box="[1318,1381,439,464]" pageId="11" pageNumber="598">
<taxonomicNameLabel id="A2010D11FFEDFF914164FE27FA88FE1F" box="[1318,1381,439,464]" pageId="11" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and these branchiae have only 4 filaments in the first pair and 
<quantity id="4CBEC19DFFEDFF914761FE4CFCA3FE3A" box="[803,846,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="598" unit="in" value="2.0">2 in</quantity>
the second pair in members of 
<taxonomicName id="4C4617FBFFEDFF9140F2FE4DFA78FE3A" box="[1200,1429,476,501]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="crassibranchiatus">
<emphasis id="B932B06AFFEDFF9140F2FE4DFA78FE3A" box="[1200,1429,476,501]" italics="true" pageId="11" pageNumber="598">T. crassibranchiatus</emphasis>
</taxonomicName>
, and a single and long filament on each side, in both pairs, in 
<taxonomicName id="4C4617FBFFEDFF91477DFD91FC44FDD7" box="[831,937,513,536]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="pascua">
<emphasis id="B932B06AFFEDFF91477DFD91FC44FDD7" box="[831,937,513,536]" italics="true" pageId="11" pageNumber="598">T. pascua</emphasis>
</taxonomicName>
(
<tableCitation id="C6C459C3FFEDFF9147FAFE6FFBFDFDD7" box="[952,1040,511,536]" captionStart="TABLE 2" captionStartId="7.[151,239,620,643]" captionTargetBox="[159,1431,743,1126]" captionTargetPageId="7" captionText="TABLE 2. Comparison between the most important numerical characters of the species of Thelepus most similar to T. megalabiatum n. sp. and T. brevitori n. sp. Sources: Hutchings &amp; Glasby (1987); Londoño-Mesa (2009); Capa &amp; Hutchings (2006); characters of T. setosus examined directly from type-material." httpUri="http://table.plazi.org/id/DF393CF0FFE1FF9D44D5FDFCFC19FD0C" pageId="11" pageNumber="598" tableUuid="DF393CF0FFE1FF9D44D5FDFCFC19FD0C">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BF96C78FFEDFF914485FDB4FED2FD4A" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="598">
Finally, the notochaetae from the anterior row of 
<taxonomicName id="4C4617FBFFEDFF9146BAFDB5FC9BFDF2" box="[760,886,548,573]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori" status="sp. nov.">
<emphasis id="B932B06AFFEDFF9146BAFDB5FC9BFDF2" box="[760,886,548,573]" italics="true" pageId="11" pageNumber="598">T. brevitori</emphasis>
</taxonomicName>
<emphasis id="B932B06AFFEDFF91473CFDB4FC53FDF2" bold="true" box="[894,958,548,573]" pageId="11" pageNumber="598">
<taxonomicNameLabel id="A2010D11FFEDFF91473CFDB4FC53FDF2" box="[894,958,548,573]" pageId="11" pageNumber="598" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, are unique, not found in any of the other species in this genus, with expanded and laterally rounded shaft at the base of the wing, and narrow wing of uniform width.
</paragraph>
</subSubSection>
<subSubSection id="C35C3FF3FFEDFF914485FD1FFE13FD02" pageId="11" pageNumber="598" type="etymology">
<paragraph id="8BF96C78FFEDFF914485FD1FFE13FD02" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="598">
<emphasis id="B932B06AFFEDFF914485FD1FFEA3FD67" bold="true" box="[199,334,655,680]" pageId="11" pageNumber="598">Etymology.</emphasis>
We attribute the epithet “ 
<taxonomicName id="4C4617FBFFEDFF9146D1FD1FFD19FD67" box="[659,756,655,680]" class="Polychaeta" family="Terebellidae" genus="Thelepus" kingdom="Animalia" order="Terebellida" pageId="11" pageNumber="598" phylum="Annelida" rank="species" species="brevitori">
<emphasis id="B932B06AFFEDFF9146D1FD1FFD19FD67" box="[659,756,655,680]" italics="true" pageId="11" pageNumber="598">brevitori</emphasis>
</taxonomicName>
” to this taxon in reference to the narrow width of the neuropodial tori along the body.
</paragraph>
</subSubSection>
<subSubSection id="C35C3FF3FFEDFF914485FD47FEDCFCDA" pageId="11" pageNumber="598" type="materials_examined">
<paragraph id="8BF96C78FFEDFF914485FD47FEDCFCDA" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="598">
<materialsCitation id="3B2E6625FFEDFF914485FD47FEDCFCDA" ID-GBIF-Occurrence="1455758449" country="Brazil" elevation="50" latitude="-47.134834" location="Atlantic Ocean" pageId="11" pageNumber="598" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="B932B06AFFEDFF914485FD47FE85FD3F" bold="true" box="[199,360,727,752]" pageId="11" pageNumber="598">
<typeStatus id="54FDD2DAFFEDFF914485FD47FEEFFD3F" box="[199,258,727,752]" pageId="11" pageNumber="598">Type</typeStatus>
locality.
</emphasis>
<location id="8E993AA3FFEDFF914533FD47FDCDFD3F" LSID="urn:lsid:plazi:treatment:03EFDD6EFFEEFF9144D5FC35FEDCFCDA:8E993AA3FFEDFF914533FD47FDCDFD3F" box="[369,544,727,752]" country="Brazil" latitude="-47.134834" name="Atlantic Ocean" pageId="11" pageNumber="598" stateProvince="Rio de Janeiro">Atlantic Ocean</location>
, Southeastern 
<collectingCountry id="F3512CE8FFEDFF914685FD47FCE3FD3F" box="[711,782,727,752]" name="Brazil" pageId="11" pageNumber="598">Brazil</collectingCountry>
, state of 
<collectingRegion id="4982A29AFFEDFF91473BFD47FBC9FD3F" box="[889,1060,727,752]" country="Brazil" name="Rio de Janeiro" pageId="11" pageNumber="598">Rio de Janeiro</collectingRegion>
: 25°11.89'‟ 
<geoCoordinate id="EE720ABFFFEDFF9140EBFD47FAC0FD3F" box="[1193,1325,727,752]" direction="south" orientation="latitude" pageId="11" pageNumber="598" precision="9" value="-47.134834">S 47°08.09'</geoCoordinate>
‟W, 
<quantity id="4CBEC19DFFEDFF914123FD47FA71FD20" box="[1377,1436,727,752]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="598" unit="m" value="50.0">
<elevation id="006B8B4BFFEDFF914123FD47FA71FD20" box="[1377,1436,727,752]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="598" unit="m" value="50.0">50 m</elevation>
</quantity>
deep, in sand.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>