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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. & Jansa, Sharon A." docDate="2022" docId="03EFDD5DF6F268E3DABAFC29182BFD41" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.8:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="8" docTitle="Yalkaparidontia Archer 1988" docType="treatment" docVersion="9" lastPageNumber="201" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="201" updateTime="1659987007554" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
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<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. & School of Biological, Earth & Environmental Sciences University of New South Wales, Australia & Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Voss, Robert S.</mods:namePart>
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<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jansa, Sharon A.</mods:namePart>
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<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
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<mods:number>2022</mods:number>
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<mods:detail type="issue">
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<mods:number>457</mods:number>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974209" ID-Zenodo-Dep="6974209" LSID="urn:lsid:plazi:treatment:03EFDD5DF6F268E3DABAFC29182BFD41" httpUri="http://treatment.plazi.org/id/03EFDD5DF6F268E3DABAFC29182BFD41" lastPageId="202" lastPageNumber="201" pageId="201" pageNumber="201">
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<subSubSection box="[177,570,903,928]" pageId="201" pageNumber="201" type="nomenclature">
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<paragraph blockId="201.[177,570,903,928]" box="[177,570,903,928]" pageId="201" pageNumber="201">
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<heading box="[177,570,903,928]" centered="true" fontSize="10" level="2" pageId="201" pageNumber="201" reason="4">
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<taxonomicName authority="Archer et al., 1988" authorityName="Archer" authorityYear="1988" box="[177,570,903,928]" genus="Yalkaparidontia" pageId="201" pageNumber="201" rank="genus">
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†Yalkaparidontia
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<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[369,570,903,928]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,435,953,977]" pageId="201" pageNumber="201" type="synonymic_list">
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<paragraph blockId="201.[108,638,953,1738]" box="[140,435,953,977]" pageId="201" pageNumber="201">
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CONTENTS: †
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<emphasis box="[285,435,953,977]" italics="true" pageId="201" pageNumber="201">
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<taxonomicName box="[285,432,953,977]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">Yalkaparidon</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="201" pageNumber="201" type="description">
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<paragraph blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">
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<collectionCode box="[140,199,986,1010]" pageId="201" pageNumber="201">STEM</collectionCode>
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AGE: 39.9 Mya (95%
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<collectionCode box="[456,515,987,1010]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13184" lsid="urn:lsid:biocol.org:col:13184" name="Hampstead Scientific Society" pageId="201" pageNumber="201" type="Herbarium">HPD</collectionCode>
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: 30.7–49.7 Mya).
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</paragraph>
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<paragraph blockId="201.[108,638,953,1738]" box="[140,457,1052,1076]" pageId="201" pageNumber="201">
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<collectionCode box="[140,224,1052,1076]" pageId="201" pageNumber="201">CROWN</collectionCode>
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AGE: Not applicable.
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</paragraph>
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<paragraph blockId="201.[108,638,953,1738]" pageId="201" pageNumber="201">
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UNAMBIGUOUS CRANIODENTAL
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<collectionCode box="[108,171,1119,1143]" pageId="201" pageNumber="201">AUTAPOMORPHIES</collectionCode>
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: Postglenoid process greatly reduced or absent (char. 75: 0→1; ci = 0.200); mandible usually with one mental foramen (char. 98: 1→0; ci = 0.063); second and third upper incisors entirely lacking enamel (char. 105: 0→1; ci = 1.000); first upper premolar absent (char. 114: 0→1; ci = 0.200); protocone absent (char. 141: 0→1; ci = 1.000); talonid greatly reduced or absent (char. 166: 0→1; ci = 0.500).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201" type="discussion">
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<paragraph blockId="201.[108,638,953,1738]" lastBlockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
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COMMENTS: The peculiar †
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<taxonomicName box="[458,607,1416,1440]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[458,607,1416,1440]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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is known only from two described species (†
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<taxonomicName class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
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<emphasis italics="true" pageId="201" pageNumber="201">Y. coheni</emphasis>
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</taxonomicName>
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and †
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<taxonomicName box="[256,351,1483,1506]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
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<emphasis box="[256,351,1483,1506]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
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</taxonomicName>
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) from late Oligocene to middle Miocene sites at Riversleigh World Heritage Area in northern
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<collectingCountry box="[370,471,1548,1572]" name="Australia" pageId="201" pageNumber="201">Australia</collectingCountry>
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(
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<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[176,360,1581,1605]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
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). †
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<taxonomicName box="[389,612,1581,1605]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
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<emphasis box="[389,612,1581,1605]" italics="true" pageId="201" pageNumber="201">Yalkaparidon coheni</emphasis>
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</taxonomicName>
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is by far the better known of the two species, being represented by partial cranial material, whereas †
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<taxonomicName box="[221,314,1681,1704]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
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<emphasis box="[221,314,1681,1704]" italics="true" pageId="201" pageNumber="201">Y. jonesi</emphasis>
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</taxonomicName>
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is known only from a fragmentary right mandible (
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<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[406,631,1714,1738]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,897,226,250]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
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). †
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<taxonomicName box="[937,1088,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[937,1088,226,250]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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combines some relatively plesiomorphic features of the skull (particularly of the basicranium) with a highly derived dentition that includes an enlarged “gliriform” anteriormost lower incisor and zalambdodont molars (
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<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[996,1209,391,415]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al., 1988</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D." box="[686,818,424,448]" pageId="201" pageNumber="201" pagination="1 - 17" refId="ref192600" refString="Beck, R. M. D. 2009. Was the Oligo-Miocene Australian metatherian Yalkaparidon a ' mammalian woodpecker'? Biological Journal of the Linnean Society 97: 1 - 17." type="journal article" year="2009">Beck, 2009</bibRefCitation>
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;
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[834,1039,424,448]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
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). Of the other unambiguous craniodental apomorphies identified here, extreme reduction of the postglenoid process likely reflects extensive anteroposterior movement of the lower jaw (a trait also seen, for example, in members of the placental clade Glires;
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<bibRefCitation author="Cope, E. D." box="[738,877,623,647]" pageId="201" pageNumber="201" pagination="3 - 11" refId="ref198605" refString="Cope, E. D. 1888. The mechanical causes of the origin of the dentition of the Rodentia. American Naturalist 22: 3 - 11." type="journal article" year="1888">Cope, 1888</bibRefCitation>
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;
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<bibRefCitation author="Druzinsky, R. E." box="[895,1093,622,646]" pageId="201" pageNumber="201" pagination="323 - 349" refId="ref200607" refString="Druzinsky, R. E. 2015. The oral apparatus of rodents: variations on the theme of a gnawing machine. In P. G. Cox and L. Hautier (editors), Evolution of the rodents: 323 - 349. Cambridge: Cambridge University Press." type="book chapter" year="2015">Druzinsky, 2015</bibRefCitation>
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), whereas absence of enamel from I2 and I3 is an autapomorphy that we have not seen in any other metatherian.
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</paragraph>
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<paragraph blockId="201.[684,1214,226,1738]" pageId="201" pageNumber="201">
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Based on its unusual combination of features, we agree with
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<bibRefCitation author="Archer, M. & S. Hand & H. Godthelp" box="[929,1159,788,812]" pageId="201" pageNumber="201" pagination="1528 - 1531" refId="ref190593" refString="Archer, M., S. Hand, and H. Godthelp. 1988. A new order of Tertiary zalambdodont marsupials. Science 239: 1528 - 1531." type="journal article" year="1988">Archer et al. (1988)</bibRefCitation>
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and
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[684,884,821,845]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
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that †
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<taxonomicName box="[960,1106,821,845]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[960,1106,821,845]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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warrants classification within its own family and order. †
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<taxonomicName box="[698,844,887,911]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[698,844,887,911]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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is the only definitive member of this order currently known, although the enigmatic †
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<taxonomicName box="[767,1056,953,977]" class="Mammalia" family="Yingabalanaridae" genus="Yingabalanara" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="species">
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<emphasis box="[767,1056,953,977]" italics="true" pageId="201" pageNumber="201">Yingabalanara richardsoni</emphasis>
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</taxonomicName>
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(known from two lower molars from the early Miocene of Riversleigh;
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<bibRefCitation author="Archer, M. & R. G. Every & H. Godthelp & S. Hand & K. Scally" box="[823,1033,1019,1043]" pageId="201" pageNumber="201" pagination="193 - 202" refId="ref190672" refString="Archer, M., R. G. Every, H. Godthelp, S. Hand, and K. Scally. 1990. Yingabalanaridae, a new family of enigmatic mammals from Tertiary deposits of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum 28: 193 - 202." type="journal article" year="1990">Archer et al., 1990</bibRefCitation>
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) may be a dentally plesiomorphic yalkaparidontian (see
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014: 155</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="201.[684,1214,226,1738]" lastBlockId="202.[108,638,226,646]" lastPageId="202" lastPageNumber="202" pageId="201" pageNumber="201">
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Our undated (fig. 32) and dated (
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<figureCitation box="[1127,1205,1118,1143]" captionStart="FIG" captionStartId="181.[108,150,938,959]" captionTargetId="figure-12@181.[241,1216,224,1485]" captionTargetPageId="181" captionText="FIG. 33. (opposite page and at right) Fifty-per- cent majority rule consensus of post-burn-in trees that results from dated Bayesian analysis (using combined tip-and-node dating and separate Independent Gamma Rates [IGR] clock models for the molecular and morpho- logical partitions) of our total evidence data- set. Black dots at nodes indicate ≥0.95 Bayesian posterior probability (“strong support”); dark gray dots indicate 0.75–0.94 Bayesian poste- rior probability (“moderate support”); light gray dots indicate 0.50–0.74 Bayesian poste- rior probability (“weak support”). Nodes with- out dots were constrained a priori so that their ages could be calibrated (see tables 6, 13). Orange bars represent 95% Highest Posterior Density (HPD) intervals on the ages of nodes. For clarity, 95% HPD intervals are not shown for the ages of fossil terminals." pageId="201" pageNumber="201">fig. 33</figureCitation>
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) total-evidence analyses both recover a clade that unites †
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<taxonomicName box="[776,923,1184,1208]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[776,923,1184,1208]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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with paucituberculatans. However, isolated tarsals tentatively referred to †
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<taxonomicName box="[697,841,1251,1275]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
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<emphasis box="[697,841,1251,1275]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
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</taxonomicName>
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by
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<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[879,1067,1251,1275]" pageId="201" pageNumber="201" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al. (2014)</bibRefCitation>
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show greater derived similarities to australidelphians than to paucituberculatans and Beck et al.’s (2014) accompanying phylogenetic analyses also supported australidelphian affinities for this taxon (see also
|
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<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[795,995,1416,1440]" pageId="201" pageNumber="201" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">Beck et al., 2016</bibRefCitation>
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). Nevertheless, the subsequent phylogenetic analyses of
|
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<bibRefCitation author="Beck, R. M. D." pageId="201" pageNumber="201" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck (2017a)</bibRefCitation>
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and
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<bibRefCitation author="Zimicz and Goin" box="[837,1127,1482,1506]" firstAuthor="Zimicz" pageId="201" pageNumber="201" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin (2020)</bibRefCitation>
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, which included †
|
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|
|
<taxonomicName box="[813,963,1515,1539]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="201" pageNumber="201" phylum="Chordata" rank="genus">
|
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<emphasis box="[813,963,1515,1539]" italics="true" pageId="201" pageNumber="201">Yalkaparidon</emphasis>
|
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|
</taxonomicName>
|
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and used character scores from these referred tarsals also placed this taxon in a clade with paucituberculatans (including argyrolagids, a fossil group not included in our study; but see
|
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|
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<bibRefCitation author="Abello, M. A. & A. M. Candela" pageId="201" pageNumber="201" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
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|
). Although this clade is worthy of further investigation, we suspect that placement of †
|
|||
|
|
<taxonomicName box="[149,292,226,250]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
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|
|
<emphasis box="[149,292,226,250]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
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|
|
</taxonomicName>
|
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|
with paucituberculatans largely reflects convergent acquisitions of a gliriform lower incisor. Current evidence (including the analyses presented here) indicates that diprotodontians evolved a similar lower incisor independently of paucituberculatans, so it is plausible that †
|
|||
|
|
<taxonomicName box="[174,319,424,448]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
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|
|
<emphasis box="[174,319,424,448]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
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|
|
</taxonomicName>
|
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|
|
represents a third origin of this tooth
|
|||
|
|
<typeStatus box="[226,276,458,482]" pageId="202" pageNumber="202">type</typeStatus>
|
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|
|
. Based on our results and those of other recent studies (
|
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|
|
<bibRefCitation author="Beck, R. M. D. & K. J. Travouillon & K. P. Aplin & H. Godthelp & M. Archer" box="[375,564,490,514]" pageId="202" pageNumber="202" pagination="127 - 172" refId="ref192966" refString="Beck, R. M. D., K. J. Travouillon, K. P. Aplin, H. Godthelp, and M. Archer. 2014. The osteology and systematics of the enigmatic Australian Oligo-Miocene metatherian Yalkaparidon (Yalkaparidontidae; Yalkaparidontia;? Australidelphia; Marsupialia). Journal of Mammalian Evolution 21 (2): 127 - 172." type="journal article" year="2014">Beck et al., 2014</bibRefCitation>
|
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|
|
,
|
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|
|
<bibRefCitation author="Beck, R. M. D. & N. M. Warburton & M. Archer & S. J. Hand & K. P. Aplin" box="[577,632,490,514]" pageId="202" pageNumber="202" pagination="151 - 171" refId="ref193033" refString="Beck, R. M. D., N. M. Warburton, M. Archer, S. J. Hand, and K. P. Aplin. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151 - 171." type="journal article" year="2016">2016</bibRefCitation>
|
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|
|
;
|
|||
|
|
<bibRefCitation author="Beck, R. M. D." box="[108,244,523,547]" pageId="202" pageNumber="202" pagination="373 - 414" refId="ref192706" refString="Beck, R. M. D. 2017 a. The skull of Epidolops ameghinoi from the early Eocene Itaborai fauna, southeastern Brazil, and the affinities of the extinct marsupialiform order Polydolopimorphia. Journal of Mammalian Evolution 24 (4): 373 - 414." type="journal article" year="2017">Beck, 2017a</bibRefCitation>
|
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|
|
;
|
|||
|
|
<bibRefCitation author="Abello, M. A. & A. M. Candela" box="[256,546,523,547]" pageId="202" pageNumber="202" pagination="419 - 444" refId="ref188032" refString="Abello, M. A., and A. M. Candela. 2019. Paleobiology of Argyrolagus (Marsupialia, Argyrolagidae): an astonishing case of bipedalism among South American mammals. Journal of Mammalian Evolution 27: 419 - 444." type="journal article" year="2019">Abello and Candela, 2019</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation author="Zimicz and Goin" firstAuthor="Zimicz" pageId="202" pageNumber="202" pagination="975 - 992" refId="ref237724" refString="Zimicz, A. N., and F. J. Goin. 2020. A reassessment of the genus Groeberia Patterson, 1952 (Mammalia, Metatheria): functional and phylogenetic implications. Journal of Systematic Palaeontology 18 (12): 975 - 992." type="journal article" year="2020">Zimicz and Goin, 2020</bibRefCitation>
|
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|
|
), we consider the higher-level affinities of †
|
|||
|
|
<taxonomicName box="[273,423,589,613]" class="Mammalia" family="Yalkaparidontidae" genus="Yalkaparidon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="202" pageNumber="202" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis box="[273,423,589,613]" italics="true" pageId="202" pageNumber="202">Yalkaparidon</emphasis>
|
|||
|
|
</taxonomicName>
|
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|
|
to be uncertain, beyond its being a member of
|
|||
|
|
<taxonomicName authorityName=", Illiger" authorityYear="1811" box="[448,581,622,646]" class="Mammalia" kingdom="Animalia" order="Marsupialia" pageId="202" pageNumber="202" phylum="Chordata" rank="order">Marsupialia</taxonomicName>
|
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|
|
.
|
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|
|
</paragraph>
|
|||
|
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</subSubSection>
|
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|
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</treatment>
|
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|
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</document>
|