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<document id="D01B56363882FA1F0E7CA1EA6150BEF2" ID-DOI="10.5281/zenodo.5430686" ID-ISSN="0253-116X" ID-Zenodo-Dep="5430686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630425214704" checkinUser="valdenar" docAuthor="Assing, V." docDate="2008" docId="03DBED0FFFEBFFD4FF43CF88FCBCFABB" docLanguage="en" docName="LinzerbiolBeitr.2008.40.2.1327-1335.pdf" docOrigin="Linzer biologische Beiträge 40 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba sultanica Assing 2008, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1334" masterDocId="FFE29577FFEEFFD3FFF2CE24FFF7F85E" masterDocTitle="New species and additional records of Geostiba from the Eastern Mediterranean region and from Middle Asia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1335" masterPageNumber="1327" pageNumber="1332" updateTime="1699264800852" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E0FBA524A5E7D5BA0C524AC3434A60DF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="CC0050412C44FAE492199DA87DDC52FD">New species and additional records of Geostiba from the Eastern Mediterranean region and from Middle Asia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="8D9154B59E4E69BD81D6D8393076E914">Assing, V.</mods:namePart>
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<mods:title id="3078D441A7850C5D47F4F4615B75C340">Linzer biologische Beiträge</mods:title>
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<mods:date id="0606F8B40A2C3CE2C9E2A020F77634DE">2008</mods:date>
<mods:detail id="BF84E9FBA351680A12EE2F84D547DB6C" type="pubDate">
<mods:number id="D8909E315932A04570F8DDE1E17F73A5">2008-12-19</mods:number>
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<mods:number id="B8398B85A7B880A1418E76687171F4B1">40</mods:number>
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<mods:number id="C82D14F891F16E695A2DB2C479817F5A">2</mods:number>
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<mods:classification id="4B63A5500FBC28D84F0552188FDB2CB1">journal article</mods:classification>
<mods:identifier id="E3B5140EF3FEE31D1160482250D7472F" type="DOI">10.5281/zenodo.5430686</mods:identifier>
<mods:identifier id="0D7483B7A3DA41D85A4099DF13DAB4B5" type="ISSN">0253-116X</mods:identifier>
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<treatment id="03DBED0FFFEBFFD4FF43CF88FCBCFABB" LSID="urn:lsid:plazi:treatment:03DBED0FFFEBFFD4FF43CF88FCBCFABB" httpUri="http://treatment.plazi.org/id/03DBED0FFFEBFFD4FF43CF88FCBCFABB" lastPageId="7" lastPageNumber="1334" pageId="5" pageNumber="1332">
<subSubSection id="C3680F92FFEBFFD6FF43CF88FDBEF99C" box="[177,585,428,450]" pageId="5" pageNumber="1332" type="nomenclature">
<paragraph id="8BCD5C19FFEBFFD6FF43CF88FDBEF99C" blockId="5.[177,585,428,450]" box="[177,585,428,450]" pageId="5" pageNumber="1332">
<heading id="D085EB75FFEBFFD6FF43CF88FDBEF99C" box="[177,585,428,450]" fontSize="8" level="2" pageId="5" pageNumber="1332" reason="2">
<emphasis id="B906800BFFEBFFD6FF43CF88FE32F99C" bold="true" box="[177,453,428,450]" pageId="5" pageNumber="1332">
<taxonomicName id="4C72279AFFEBFFD6FF43CF88FE84F99C" ID-CoL="3FSNT" authority="Assing, 2008" authorityName="Assing" authorityYear="2008" box="[177,371,428,450]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1332" phylum="Arthropoda" rank="species" species="sultanica" status="sp. nov.">
<emphasis id="B906800BFFEBFFD6FF43CF88FE84F99C" bold="true" box="[177,371,428,450]" italics="true" pageId="5" pageNumber="1332">Geostiba sultanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2353D70FFEBFFD6FE8BCF89FE32F99C" box="[377,453,429,450]" pageId="5" pageNumber="1332" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1349409CFFEBFFD6FE20CF88FDB6F99C" box="[466,577,428,450]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Figs 10-21</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3680F92FFEBFFD4FF43CFFCFCBCFABB" lastPageId="7" lastPageNumber="1334" pageId="5" pageNumber="1332" type="materials_examined">
<paragraph id="8BCD5C19FFEBFFD6FF43CFFCFCB2FA44" blockId="5.[177,1085,472,538]" pageId="5" pageNumber="1332">
<materialsCitation id="3B1A5644FFEBFFD6FF43CFFCFE17FA5D" collectingDate="2008-04-19" collectorName="SW Sultandagi &amp; Brachat &amp; Meybohm" elevation="1730" location="Sultan Dag" longitude="38.496387" pageId="5" pageNumber="1332" specimenCount="1" stateProvince="Konya" typeStatus="holotype">
<typeStatus id="54C9E2BBFFEBFFD6FF43CFFCFEF1F9B2" box="[177,262,472,492]" pageId="5" pageNumber="1332" type="holotype">Holotype</typeStatus>
: N
<geoCoordinate id="EE463ADEFFEBFFD6FEB0CFFCFE44F9B2" box="[322,435,472,492]" degrees="38" direction="east" minutes="29" orientation="longitude" pageId="5" pageNumber="1332" precision="15" seconds="47" value="38.496387">38°29'47&quot; E</geoCoordinate>
31°11'37&quot; (23), TR
<collectingRegion id="49B692FBFFEBFFD6FD9FCFFCFD5BF9B2" box="[621,684,472,492]" country="Turkey" name="Konya" pageId="5" pageNumber="1332">Konya</collectingRegion>
,
<location id="8EAD0AC2FFEBFFD6FD4ACFFCFCD7F9B2" LSID="urn:lsid:plazi:treatment:03DBED0FFFEBFFD4FF43CF88FCBCFABB:8EAD0AC2FFEBFFD6FD4ACFFCFCD7F9B2" box="[696,800,472,492]" longitude="38.496387" name="Sultan Dag" pageId="5" pageNumber="1332" stateProvince="Konya">Sultan Dağ</location>
[ları]
<collectorName id="268739CFFFEBFFD6FCA7CFFCFC16F9B2" box="[853,993,472,492]" pageId="5" pageNumber="1332">SW Sultandağı</collectorName>
,
<quantity id="4C8AF1FCFFEBFFD6FC1DCFFCFBCEF9B2" box="[1007,1081,472,492]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="5" pageNumber="1332" unit="m" value="1730.0">
<elevation id="005FBB2AFFEBFFD6FC1DCFFCFBCEF9B2" box="[1007,1081,472,492]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="5" pageNumber="1332" unit="m" value="1730.0">1730 m</elevation>
</quantity>
,
<collectorName id="268739CFFFEBFFD6FF43CFCBFF0FFA5D" box="[177,248,495,515]" pageId="5" pageNumber="1332">Brachat</collectorName>
&amp;
<collectorName id="268739CFFFEBFFD6FEECCFCBFE8CFA5D" box="[286,379,495,515]" pageId="5" pageNumber="1332">Meybohm</collectorName>
<date id="FFCC7AD9FFEBFFD6FE77CFCBFE17FA5D" box="[389,480,495,515]" pageId="5" pageNumber="1332" value="2008-04-19">
<collectingDate id="EF888331FFEBFFD6FE77CFCBFE17FA5D" box="[389,480,495,515]" pageId="5" pageNumber="1332" value="2008-04-19">19.4.2008</collectingDate>
</date>
</materialsCitation>
/
<typeStatus id="54C9E2BBFFEBFFD6FE09CFCBFDADFA5D" box="[507,602,495,515]" pageId="5" pageNumber="1332" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C72279AFFEBFFD6FD73CFCBFCC7FA5D" authority="Assing, 2008" authorityName="Assing" authorityYear="2008" box="[641,816,495,515]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1332" phylum="Arthropoda" rank="species" species="sultanica" status="sp. nov.">
<emphasis id="B906800BFFEBFFD6FD73CFCBFCC7FA5D" box="[641,816,495,515]" italics="true" pageId="5" pageNumber="1332">Geostiba sultanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2353D70FFEBFFD6FCC8CFCBFC87FA5D" box="[826,880,495,515]" pageId="5" pageNumber="1332" rank="species">sp. n.</taxonomicNameLabel>
det.
<collectionCode id="ED63C4DCFFEBFFD6FC57CFCBFC4FFA5D" box="[933,952,495,515]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="1332" type="Museum">V</collectionCode>
. Assing 2008 (cAss).
<typeStatus id="54C9E2BBFFEBFFD6FF0ACC22FEA3FA44" box="[248,340,518,538]" pageId="5" pageNumber="1332" type="paratype">Paratypes</typeStatus>
: 10, 26: same data as
<typeStatus id="54C9E2BBFFEBFFD6FD97CC22FD42FA44" box="[613,693,518,538]" pageId="5" pageNumber="1332" type="holotype">holotype</typeStatus>
(cAss,
<collectionCode id="ED63C4DCFFEBFFD6FD08CC22FCCCFA44" box="[762,827,518,538]" pageId="5" pageNumber="1332">OÖLL</collectionCode>
).
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CC0DFC47FA05" blockId="5.[177,1085,552,603]" pageId="5" pageNumber="1332">
D e s c r i p t i o n: Body length 2.0-
<quantity id="4C8AF1FCFFEBFFD6FDB4CC0CFD6FFA60" box="[582,664,552,574]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="1332" unit="mm" value="2.5">2.5 mm</quantity>
. Habitus as in
<figureCitation id="1349409CFFEBFFD6FCC7CC0CFC75FA60" box="[821,898,552,574]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 10</figureCitation>
. Coloration: body uniformly yellowish; abdominal segment
<collectionCode id="ED63C4DCFFEBFFD6FDAACC62FD84FA05" box="[600,627,582,603]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="1332">VI</collectionCode>
occasionally weakly infuscate.
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CC4FFCDFFA87" blockId="5.[177,1085,619,729]" pageId="5" pageNumber="1332">
Head approximately as wide as long or weakly oblong (
<figureCitation id="1349409CFFEBFFD6FD0DCC4FFCBAFADF" box="[767,845,619,641]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 11</figureCitation>
); punctation extremely fine, barely noticeable; surface with shallow microreticulation. Eyes reduced to minute rudiments, without ommatidia and pigmentation (
<figureCitation id="1349409CFFEBFFD6FD59CC81FD0FFAE5" box="[683,760,677,699]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 12</figureCitation>
). Antennae distinctly incrassate apically; antennomere X approximately twice as wide as long.
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CCCCFDF9FB66" blockId="5.[177,1085,744,824]" pageId="5" pageNumber="1332">
Pronotum approximately 1.15 times as wide as head and 1.1 times as wide as long (
<figureCitation id="1349409CFFEBFFD6FBE1CCCCFF3BFB45" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 11</figureCitation>
); punctation extremely fine, barely noticeable; microreticulation similar to that of head or slightly more pronounced.
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CD6CFEBCFBC6" blockId="5.[177,1085,840,920]" pageId="5" pageNumber="1332">
Elytra with pronounced sexual dimorphism, 0.55-0.65 times as long as pronotum (
<figureCitation id="1349409CFFEBFFD6FBE1CD6CFF3BFB25" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 11</figureCitation>
); microsculpture very shallow, less pronounced than that of head and pronotum. Hind wings reduced.
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CD83FE7BFC4B" blockId="5.[177,1085,935,1045]" pageId="5" pageNumber="1332">Abdomen approximately 1.3 times as wide as elytra; punctation very fine and moderately sparse; microreticulation distinct everywhere; posterior margin of tergite VII without palisade fringe; tergite VII with sexual dimorphism; posterior margin of tergite VIII convex in both sexes.</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF34CA01FD5FFD7B" blockId="5.[177,1085,1061,1317]" pageId="5" pageNumber="1332">
: elytra with weakly granulose punctation, at suture strongly elevated, forming pronounced and sharp sutural carinae extending over full length of suture, these carinae highest and broadest anteriorly, and decreasing in height and width posteriad (
<figureCitation id="1349409CFFEBFFD6FC10CA7BFBD8FC2B" box="[994,1071,1119,1141]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 11</figureCitation>
); tergite VII with pronounced long median tubercle, in large males extending approximately from anterior 1/4 of tergite to posterior margin, this tubercle smooth, glossy, and tapering in posterior 1/4 of tergite (
<figureCitation id="1349409CFFEBFFD6FDE7CA93FD70FC93" box="[533,647,1207,1229]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Figs 13-14</figureCitation>
); posterior margin of sternite VIII obtusely pointed in the middle (
<figureCitation id="1349409CFFEBFFD6FE69CAF0FE1FFCB4" box="[411,488,1236,1258]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 15</figureCitation>
); median lobe of aedeagus with long flagellum in internal sac (
<figureCitation id="1349409CFFEBFFD6FF13CAD6FEA2FD56" box="[225,341,1266,1288]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Figs 16-18</figureCitation>
); apical lobe of paramere moderately stout, with one stout long median seta and three fine short subapical setae (
<figureCitation id="1349409CFFEBFFD6FDA2CB2BFD6CFD7B" box="[592,667,1295,1317]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 19</figureCitation>
).
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF34CB11FF2DFDFC" blockId="5.[177,1085,1332,1442]" pageId="5" pageNumber="1332">
: elytra with fine, non-granulose punctation and without sutural carinae; tergite VII unmodified; sternite VIII weakly concave in the middle (
<figureCitation id="1349409CFFEBFFD6FCF4CB75FCA2FD39" box="[774,853,1361,1383]" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 20</figureCitation>
); spermatheca minute, only
<quantity id="4C8AF1FCFFEBFFD6FF15CB4BFEB3FDDA" box="[231,324,1391,1413]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="1332" unit="mm" value="0.08">0.08 mm</quantity>
long, with conspicuously short and proximally simply curved duct (
<figureCitation id="1349409CFFEBFFD6FBE6CB4BFF3AFDFC" captionStart="Figs 9-21" captionStartId="6.[177,217,1372,1394]" captionTargetBox="[178,1085,316,1353]" captionTargetId="figure-196@6.[178,1085,316,1363]" captionTargetPageId="6" captionText="Figs 9-21: Geostiba kirghisica nov.sp. (9) and G. sultanica nov.sp. (10-21): (9, 21) spermatheca; (10) male habitus; (11) male forebody; (12) head in lateral view; (13-14) male tergites VII-VIII in dorsal and in lateral view; (15) male sternite VIII; (16-18) median lobe of aedeagus in lateral and in ventral view; (19) apical lobe of paramere; (20) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="5" pageNumber="1332">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BCD5C19FFEBFFD6FF43CB96FE54FDBB" blockId="5.[177,1085,1457,1509]" pageId="5" pageNumber="1332">
E t y m o l o g y: The name (adjective) is derived from the Sultan Dağları, where the
<typeStatus id="54C9E2BBFFEBFFD6FF43CBF4FF2AFDBB" box="[177,221,1488,1509]" pageId="5" pageNumber="1332">type</typeStatus>
locality is situated.
</paragraph>
<paragraph id="8BCD5C19FFEBFFD5FF43CBD0FDABF975" blockId="5.[177,1085,1524,1605]" lastBlockId="6.[177,1085,219,299]" lastPageId="6" lastPageNumber="1333" pageId="5" pageNumber="1332">
C o m p a r a t i v e n o t e s:
<taxonomicName id="4C72279AFFEBFFD6FDFFCBD0FD25FE54" authorityName="Assing" authorityYear="2008" box="[525,722,1524,1546]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1332" phylum="Arthropoda" rank="species" species="sultanica">
<emphasis id="B906800BFFEBFFD6FDFFCBD0FD25FE54" box="[525,722,1524,1546]" italics="true" pageId="5" pageNumber="1332">Geostiba sultanica</emphasis>
</taxonomicName>
is readily distinguished from all other known congeners recorded from
<collectingCountry id="F3651C89FFEBFFD6FD91C835FD5BFE79" box="[611,684,1553,1575]" name="Turkey" pageId="5" pageNumber="1332">Turkey</collectingCountry>
based on external characters alone, particularly by the presence of a long median tubercle on the male abdominal tergite VII. In addition, it is separated from its congeners by the morphology of the aedeagus, especially the presence and shape of the flagellum in the internal sac, as well as by the shape and minute size of the spermatheca.
</paragraph>
<caption id="DF0D0C91FFE8FFD5FF43CB78FD2CFD92" ID-DOI="http://doi.org/10.5281/zenodo.5430690" ID-Zenodo-Dep="5430690" httpUri="https://zenodo.org/record/5430690/files/figure.png" pageId="6" pageNumber="1333" startId="6.[177,217,1372,1394]" targetBox="[178,1085,316,1353]" targetPageId="6">
<paragraph id="8BCD5C19FFE8FFD5FF43CB78FD2CFD92" blockId="6.[177,1085,1372,1484]" pageId="6" pageNumber="1333">
<emphasis id="B906800BFFE8FFD5FF43CB78FEFCFD2F" bold="true" box="[177,267,1372,1394]" pageId="6" pageNumber="1333">Figs 9-21</emphasis>
:
<taxonomicName id="4C72279AFFE8FFD5FEEBCB79FE39FD2F" authority="Assing, 2008" authorityName="Assing" authorityYear="2008" box="[281,462,1373,1393]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="species" species="kirghisica" status="sp. nov.">
<emphasis id="B906800BFFE8FFD5FEEBCB79FE39FD2F" box="[281,462,1373,1393]" italics="true" pageId="6" pageNumber="1333">Geostiba kirghisica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2353D70FFE8FFD5FE27CB79FDEFFD2F" box="[469,536,1373,1393]" pageId="6" pageNumber="1333" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B906800BFFE8FFD5FDD4CB78FDC6FD2F" bold="true" box="[550,561,1372,1393]" pageId="6" pageNumber="1333">9</emphasis>
) and
<taxonomicName id="4C72279AFFE8FFD5FD98CB79FD2CFD2F" authority="Assing, 2008" authorityName="Assing" authorityYear="2008" box="[618,731,1373,1393]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="species" species="sultanica" status="sp. nov.">
<emphasis id="B906800BFFE8FFD5FD98CB79FD2CFD2F" box="[618,731,1373,1393]" italics="true" pageId="6" pageNumber="1333">G. sultanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2353D70FFE8FFD5FD11CB79FCD1FD2F" box="[739,806,1373,1393]" pageId="6" pageNumber="1333" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B906800BFFE8FFD5FCC6CB78FC9EFD2F" bold="true" box="[820,873,1372,1393]" pageId="6" pageNumber="1333">10-21</emphasis>
): (
<emphasis id="B906800BFFE8FFD5FC74CB78FC41FD2F" bold="true" box="[902,950,1372,1393]" pageId="6" pageNumber="1333">9, 21</emphasis>
) spermatheca; (
<emphasis id="B906800BFFE8FFD5FF4BCB57FF27FDD6" bold="true" box="[185,208,1395,1416]" pageId="6" pageNumber="1333">10</emphasis>
) male habitus; (
<emphasis id="B906800BFFE8FFD5FE95CB57FE89FDD6" bold="true" box="[359,382,1395,1416]" pageId="6" pageNumber="1333">11</emphasis>
) male forebody; (
<emphasis id="B906800BFFE8FFD5FDD7CB57FDCBFDD6" bold="true" box="[549,572,1395,1416]" pageId="6" pageNumber="1333">12</emphasis>
) head in lateral view; (
<emphasis id="B906800BFFE8FFD5FCE4CB57FCBDFDD6" bold="true" box="[790,842,1395,1416]" pageId="6" pageNumber="1333">13-14</emphasis>
) male tergites VII-VIII in dorsal and in lateral view; (
<emphasis id="B906800BFFE8FFD5FE59CBADFE35FDC0" bold="true" box="[427,450,1417,1438]" pageId="6" pageNumber="1333">15</emphasis>
) male sternite VIII; (
<emphasis id="B906800BFFE8FFD5FD76CBADFD4EFDC0" bold="true" box="[644,697,1417,1438]" pageId="6" pageNumber="1333">16-18</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="B906800BFFE8FFD5FECBCB84FEA7FDEB" bold="true" box="[313,336,1440,1461]" pageId="6" pageNumber="1333">19</emphasis>
) apical lobe of paramere; (
<emphasis id="B906800BFFE8FFD5FDBDCB84FD91FDEB" bold="true" box="[591,614,1440,1461]" pageId="6" pageNumber="1333">20</emphasis>
) female sternite VIII. Scale bars: 10: 1.0 mm; 11: 0.5 mm; 12-15, 20: 0.2 mm; 9, 16-19: 0.1 mm; 21: 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BCD5C19FFE8FFD4FF43CBFFFC5EF938" blockId="6.[177,1085,1499,1609]" lastBlockId="7.[177,1085,219,358]" lastPageId="7" lastPageNumber="1334" pageId="6" pageNumber="1333">
The subgeneric affiliations of the species are unclear. Based on the modifications of the male abdominal tergite VII,
<taxonomicName id="4C72279AFFE8FFD5FE10CBDCFD94FE50" authorityName="Assing" authorityYear="2008" box="[482,611,1528,1550]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="species" species="sultanica">
<emphasis id="B906800BFFE8FFD5FE10CBDCFD94FE50" box="[482,611,1528,1550]" italics="true" pageId="6" pageNumber="1333">G. sultanica</emphasis>
</taxonomicName>
would have to be assigned to the subgenus
<taxonomicName id="4C72279AFFE8FFD5FF43C831FE5BFE75" authority="CASEY 1906" authorityName="CASEY" authorityYear="1906" box="[177,428,1557,1579]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="subGenus" subGenus="Typhlusida">
<emphasis id="B906800BFFE8FFD5FF43C831FED6FE75" box="[177,289,1557,1579]" italics="true" pageId="6" pageNumber="1333">Typhlusida</emphasis>
CASEY 1906
</taxonomicName>
. The geographically closest representatives of this subgenus are
<taxonomicName id="4C72279AFFE8FFD5FF43C817FE0CFE17" authority="(RAMBOUSEK 1924)" baseAuthorityName="RAMBOUSEK" baseAuthorityYear="1924" box="[177,507,1587,1609]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="species" species="rhilensis">
<emphasis id="B906800BFFE8FFD5FF43C817FEDFFE17" box="[177,296,1587,1609]" italics="true" pageId="6" pageNumber="1333">G. rhilensis</emphasis>
(RAMBOUSEK 1924)
</taxonomicName>
from
<collectingCountry id="F3651C89FFE8FFD5FDC8C817FD66FE17" box="[570,657,1587,1609]" name="Bulgaria" pageId="6" pageNumber="1333">Bulgaria</collectingCountry>
and
<taxonomicName id="4C72279AFFE8FFD5FD37C817FC4AFE17" authority="(KRAATZ 1856)" baseAuthorityName="KRAATZ" baseAuthorityYear="1856" box="[709,957,1587,1609]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1333" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B906800BFFE8FFD5FD37C817FCEFFE17" box="[709,792,1587,1609]" italics="true" pageId="6" pageNumber="1333">G. flava</emphasis>
(KRAATZ 1856)
</taxonomicName>
from south- eastern
<collectingCountry id="F3651C89FFE9FFD4FEF2CEFFFEBDF8AF" box="[256,330,219,241]" name="Austria" pageId="7" pageNumber="1334">Austria</collectingCountry>
and
<collectingCountry id="F3651C89FFE9FFD4FE8FCEFFFE2EF8AF" box="[381,473,219,241]" name="Slovenia" pageId="7" pageNumber="1334">Slovenia</collectingCountry>
. The new species is readily separated from both of them by the modifications of the male elytra, the shape of the tubercle on the male tergite VII, and particularly by the morphology of the genitalia. For illustrations of the male secondary sexual characters and the genitalia of
<taxonomicName id="4C72279AFFE9FFD4FDCDCF16FD63F916" baseAuthorityName="KRAATZ" baseAuthorityYear="1856" box="[575,660,306,328]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="B906800BFFE9FFD4FDCDCF16FD63F916" box="[575,660,306,328]" italics="true" pageId="7" pageNumber="1334">G. flava</emphasis>
</taxonomicName>
and
<taxonomicName id="4C72279AFFE9FFD4FD38CF16FCB4F916" baseAuthorityName="RAMBOUSEK" baseAuthorityYear="1924" box="[714,835,306,328]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="rhilensis">
<emphasis id="B906800BFFE9FFD4FD38CF16FCB4F916" box="[714,835,306,328]" italics="true" pageId="7" pageNumber="1334">G. rhilensis</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFE321E8FFE9FFD4FC84CF16FBF8F916" author="ASSING V" box="[886,1039,306,328]" pageId="7" pageNumber="1334" pagination="79 - 85" refId="ref4000" refString="ASSING V. (2000): On some species of Geostiba THOMSON, 1858 from the eastern Alps (Coleoptera: Staphylinidae, Aleocharinae). - Koleopterologische Rundschau 70: 79 - 85." type="journal article" year="2000">ASSING (2000)</bibRefCitation>
and
<bibRefCitation id="EFE321E8FFE9FFD4FF43CF74FECFF938" author="PACE R" box="[177,312,336,358]" pageId="7" pageNumber="1334" pagination="3 - 46" refId="ref4286" refString="PACE R. (1983 b): Specie del genere Geostiba THOMSON raccolte dal Dr. C. Besuchet e collaboratori in Marocco, nella Penisola Iberica e Balcanica, e nel Medio Oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 90: 3 - 46." type="journal article" year="1983">PACE (1983b</bibRefCitation>
,
<bibRefCitation id="EFE321E8FFE9FFD4FEB0CF74FE81F938" author="PACE R" box="[322,374,336,358]" pageId="7" pageNumber="1334" pagination="211 - 219" refId="ref4338" refString="PACE R. (1984): Nuove Aleocharinae microftalme mediterranee e dell'Iran, del Museum d'Histoire naturelle di Ginevra (Coleoptera Staphylinidae). - Archives Scientifiques Geneve 37: 211 - 219." type="journal article" year="1984">1984</bibRefCitation>
) (under the synonym
<taxonomicName id="4C72279AFFE9FFD4FDA6CF74FCE5F93A" authority="PACE" authorityName="PACE" box="[596,786,336,358]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="bulgarica">
<emphasis id="B906800BFFE9FFD4FDA6CF74FD22F938" box="[596,725,336,358]" italics="true" pageId="7" pageNumber="1334">G. bulgarica</emphasis>
PACE
</taxonomicName>
), respectively.
</paragraph>
<paragraph id="8BCD5C19FFE9FFD4FF43CF59FC27FA1D" blockId="7.[177,1085,381,579]" pageId="7" pageNumber="1334">
The only Turkish representative of the genus with a long flagellum in the internal sac is
<taxonomicName id="4C72279AFFE9FFD4FF43CFBEFE70F9EE" authority="PACE 1983" authorityName="PACE" authorityYear="1983" box="[177,391,410,432]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="uhligi">
<emphasis id="B906800BFFE9FFD4FF43CFBEFEFAF9EE" box="[177,269,410,432]" italics="true" pageId="7" pageNumber="1334">G. uhligi</emphasis>
PACE 1983
</taxonomicName>
, which is currently attributed to the subgenus
<taxonomicName id="4C72279AFFE9FFD4FC9BCFBEFBCFF9EE" authority="CASEY 1906" authorityName="CASEY" authorityYear="1906" box="[873,1080,410,432]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="subGenus" subGenus="Sibiota">
<emphasis id="B906800BFFE9FFD4FC9BCFBEFC46F9EE" box="[873,945,410,432]" italics="true" pageId="7" pageNumber="1334">Sibiota</emphasis>
CASEY 1906
</taxonomicName>
, but which, based on the synapomorphically derived morphology of the internal structures of the aedeagus, may be the closest relative of
<taxonomicName id="4C72279AFFE9FFD4FD56CFF1FCD4F9B5" authorityName="Assing" authorityYear="2008" box="[676,803,469,491]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="sultanica">
<emphasis id="B906800BFFE9FFD4FD56CFF1FCD4F9B5" box="[676,803,469,491]" italics="true" pageId="7" pageNumber="1334">G. sultanica</emphasis>
</taxonomicName>
. It is readily distinguished from the new species by the larger eyes with ommatidia and pigmentation, by the presence of a pair of carinae on the male tergite VII, as well as by the different morphology of the genitalia. For illustrations of the genitalia of
<taxonomicName id="4C72279AFFE9FFD4FD45CC09FCE6FA1D" authorityName="PACE" authorityYear="1983" box="[695,785,557,579]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1334" phylum="Arthropoda" rank="species" species="uhligi">
<emphasis id="B906800BFFE9FFD4FD45CC09FCE6FA1D" box="[695,785,557,579]" italics="true" pageId="7" pageNumber="1334">G. uhligi</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFE321E8FFE9FFD4FCCCCC09FC3AFA1D" author="PACE R" box="[830,973,557,579]" pageId="7" pageNumber="1334" pagination="129 - 139" refId="ref4253" refString="PACE R. (1983 a): Nuove specie europee ed asiatiche del genere Geostiba THOMSON (Coleoptera Staphylinidae). - Giornale italiano di Entomologia 1: 129 - 139." type="journal article" year="1983">PACE (1983a)</bibRefCitation>
.
</paragraph>
<paragraph id="8BCD5C19FFE9FFD4FF43CC7FFCBCFABB" blockId="7.[177,1085,602,741]" pageId="7" pageNumber="1334">
D i s t r i b u t i o n a n d b i o n o m i c s: As can be inferred from the adaptive reductions of the eyes, pigmentation, and wings, as well as from the absence of records from other localities, the species is probably endemic to the Sultan Dağları in southwestern
<collectingCountry id="F3651C89FFE9FFD4FF29CC96FEC2FA96" box="[219,309,690,712]" name="Turkey" pageId="7" pageNumber="1334">Anatolia</collectingCountry>
. The
<typeStatus id="54C9E2BBFFE9FFD4FE80CC96FE69FA99" box="[370,414,690,711]" pageId="7" pageNumber="1334">type</typeStatus>
specimens were collected in an oak forest by sifting moist leaf litter and moss at an altitude of
<quantity id="4C8AF1FCFFE9FFD4FE03CCEBFDCAFABB" box="[497,573,719,741]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="7" pageNumber="1334" unit="m" value="1730.0">1730 m</quantity>
(MEYBOHM pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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