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<document id="A99A2346520D4EB12DC27E48C948DE32" ID-DOI="10.11646/zootaxa.3716.4.1" ID-GBIF-Dataset="1dba1918-7c9c-4c43-acb3-53ac6c20636b" ID-ISSN="1175-5326" ID-Zenodo-Dep="223511" ID-ZooBank="579F6501-B733-4D58-9B41-6C931097A6D0" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460682526751" checkinUser="plazi" docAuthor="Pereira, Luis Alberto" docDate="2013" docId="03D0878DFFEC4510FF0E13119A3FF868" docLanguage="en" docName="zt03716p527.pdf" docOrigin="Zootaxa 3716 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ityphilus bonatoi Pereira, 2013, sp. nov." docType="treatment" docVersion="9" lastPageNumber="522" masterDocId="FFE9FFF5FFE14505FF991109993EFFB1" masterDocTitle="Further contribution to the knowledge of Ityphilus calinus Chamberlin, 1957, a poorly known ballophilid centipede from Colombia, with description of Ityphilus bonatoi, a new diminutive geophilomorph species from Brazil (Myriapoda: Chilopoda, Geophilomorpha)" masterLastPageNumber="527" masterPageNumber="501" pageNumber="514" updateTime="1698350611203" updateUser="plazi">
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<mods:title id="8C1AE2F0A9B9086A3D094BFDD4A76204">Further contribution to the knowledge of Ityphilus calinus Chamberlin, 1957, a poorly known ballophilid centipede from Colombia, with description of Ityphilus bonatoi, a new diminutive geophilomorph species from Brazil (Myriapoda: Chilopoda, Geophilomorpha)</mods:title>
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<mods:date id="9E39D7138D38C6C820BE5C9E6D0D1FE1">2013</mods:date>
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<treatment id="03D0878DFFEC4510FF0E13119A3FF868" ID-DOI="http://doi.org/10.5281/zenodo.5629982" ID-GBIF-Taxon="119580771" ID-Zenodo-Dep="5629982" LSID="urn:lsid:plazi:treatment:03D0878DFFEC4510FF0E13119A3FF868" httpUri="http://treatment.plazi.org/id/03D0878DFFEC4510FF0E13119A3FF868" lastPageId="21" lastPageNumber="522" pageId="13" pageNumber="514">
<subSubSection id="C3636510FFEC4508FF0E13119802FDE4" pageId="13" pageNumber="514" type="nomenclature">
<paragraph id="8BC6369BFFEC4508FF0E131198F7FD82" blockId="13.[151,457,536,597]" box="[151,457,536,563]" pageId="13" pageNumber="514">
<heading id="D08E81F7FFEC4508FF0E131198F7FD82" bold="true" box="[151,457,536,563]" fontSize="11" level="1" pageId="13" pageNumber="514" reason="1">
<emphasis id="B90DEA89FFEC4508FF0E131198F7FD82" bold="true" box="[151,457,536,563]" pageId="13" pageNumber="514">
<taxonomicName id="4C794D18FFEC4508FF0E1311985BFD83" box="[151,357,536,562]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFEC4508FF0E1311985BFD83" bold="true" box="[151,357,536,562]" italics="true" pageId="13" pageNumber="514">Ityphilus bonatoi</emphasis>
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<taxonomicNameLabel id="A23E57F2FFEC4508FEF5131098F7FD82" box="[364,457,537,563]" pageId="13" pageNumber="514" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BC6369BFFEC4508FF0E13359802FDE4" blockId="13.[151,457,536,597]" box="[151,316,572,597]" pageId="13" pageNumber="514">
(
<figureCitation id="13422A1EFFEC4508FF061335980DFDE4" box="[159,307,572,597]" captionStart-0="FIGURES 53 57" captionStart-1="FIGURES 58 65" captionStart-2="FIGURES 66 74" captionStart-3="FIGURES 75 82" captionStart-4="FIGURES 83 96" captionStart-5="FIGURES 97 102" captionStart-6="FIGURES 103 105" captionStartId-0="15.[151,264,1866,1889]" captionStartId-1="16.[151,264,1797,1820]" captionStartId-2="17.[151,264,1797,1820]" captionStartId-3="18.[151,264,1866,1889]" captionStartId-4="19.[151,264,1851,1874]" captionStartId-5="20.[151,264,1866,1889]" captionStartId-6="21.[151,264,1284,1307]" captionTargetBox-0="[156,1436,193,1845]" captionTargetBox-1="[151,1436,193,1773]" captionTargetBox-2="[151,1436,193,1776]" captionTargetBox-3="[151,1436,193,1845]" captionTargetBox-4="[151,1436,193,1830]" captionTargetBox-5="[151,1434,193,1845]" captionTargetBox-6="[156,1431,200,1261]" captionTargetId-0="figure@15.[151,1436,193,1845]" captionTargetId-1="figure@16.[151,1436,193,1777]" captionTargetId-2="figure@17.[151,1436,193,1777]" captionTargetId-3="figure@18.[151,1436,193,1845]" captionTargetId-4="figure@19.[151,1436,193,1830]" captionTargetId-5="figure@20.[151,1436,193,1845]" captionTargetId-6="figure@21.[151,1436,193,1264]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionTargetPageId-5="20" captionTargetPageId-6="21" captionText-0="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." captionText-1="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." captionText-2="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." captionText-3="FIGURES 75 82. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (75) Sternite 17. (76) Sternite 23. (77) Sternite 26. (78) Sternite 31. (79) Sternite 36. (80) Sternite 37. (81) Sternite 38. (82) Sternite 39. Scale bar: 0.1 mm." captionText-4="FIGURES 83 96. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (84 93); 0.1 mm (83, 94 96)." captionText-5="FIGURES 97 102. Ityphilus bona
)
</paragraph>
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<subSubSection id="C3636510FFEC4508FF0E138D982DFBC1" pageId="13" pageNumber="514" type="diagnosis">
<paragraph id="8BC6369BFFEC4508FF0E138D9B42FD54" blockId="13.[151,1436,644,2037]" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF0E138D9834FD2C" bold="true" box="[151,266,644,669]" pageId="13" pageNumber="514">Diagnosis</emphasis>
. An
<taxonomicName id="4C794D18FFEC4508FED8138D989DFD2C" box="[321,419,644,669]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA89FFEC4508FED8138D989DFD2C" box="[321,419,644,669]" italics="true" pageId="13" pageNumber="514">Ityphilus</emphasis>
</taxonomicName>
species with internal edge of forcipular tarsungulum serrate. Among the other Neotropical members of the genus having the same trait it only shares with
<taxonomicName id="4C794D18FFEC4508FCF113A09DABFD71" authority="Pereira, 2012" authorityName="Pereira" authorityYear="2012" box="[872,1173,679,704]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="donatellae">
<emphasis id="B90DEA89FFEC4508FCF113A09AC8FD71" box="[872,1014,679,704]" italics="true" pageId="13" pageNumber="514">I. donatellae</emphasis>
Pereira, 2012
</taxonomicName>
a very small body size and a low number of leg-bearing segments.
</paragraph>
<paragraph id="8BC6369BFFEC4508FF5E13E6982DFBC1" blockId="13.[151,1436,644,2037]" pageId="13" pageNumber="514">
<taxonomicName id="4C794D18FFEC4508FF5E13E698BAFCB9" box="[199,388,751,776]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFEC4508FF5E13E698BAFCB9" box="[199,388,751,776]" italics="true" pageId="13" pageNumber="514">Ityphilus bonatoi</emphasis>
</taxonomicName>
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<emphasis id="B90DEA89FFEC4508FE1213E698DAFCB9" bold="true" box="[395,484,751,776]" pageId="13" pageNumber="514">sp. nov.</emphasis>
</taxonomicNameLabel>
can be differentiated from
<taxonomicName id="4C794D18FFEC4508FC8A13F89A9EFCB9" box="[787,928,751,776]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="donatellae">
<emphasis id="B90DEA89FFEC4508FC8A13F89A9EFCB9" box="[787,928,751,776]" italics="true" pageId="13" pageNumber="514">I. donatellae</emphasis>
</taxonomicName>
by means of the following selected traits (the corresponding traits in the latter are given in parentheses): body length of the female
<quantity id="4C819B7EFFEC4508FBD3121D9DA7FC9D" box="[1098,1177,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="514" unit="mm" value="13.0">13 mm</quantity>
(female:
<quantity id="4C819B7EFFEC4508FA9A121D9C6CFC9D" box="[1283,1362,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="514" unit="mm" value="11.0">11 mm</quantity>
, male:
<quantity id="4C819B7EFFEC4508FF0E123E99CFFCE1" box="[151,241,823,848]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="514" unit="mm" value="8.5">8.5 mm</quantity>
); female with 41 leg-bearing segments (female with 43, male with 41); shape of the antennae as in
<figureCitation id="13422A1EFFEC4508FAD1123E99E2FCC4" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Figures 53, 54</figureCitation>
(as in
<figureCitation id="13422A1EFFEC4508FEBF1255989DFCC5" box="[294,419,860,885]" captionStart="FIGURES 106 118" captionStartId="22.[151,261,1755,1778]" captionTargetBox="[155,1434,210,1734]" captionTargetId="figure@22.[151,1436,193,1734]" captionTargetPageId="22" captionText="FIGURES 106 118. (106). Ityphilus calinus Chamberlin, 1957 (male holotype; COLOMBIA: Valle del Cauca Department): Anterior end of the body, dorsal. (From Chamberlin, 1957). (107). Ityphilus donatellae Pereira, 2012 (female paratype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): Left antenna dorsal. (From Pereira, 2012). (108 117). Ityphilus donatellae Pereira, 2012 (female holotype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): (108) Left a. a. XIII, dorsal (a, b, c: a, b, c, type sensilla). (109) Detail of calyx of poison gland in left forcipular telopodite, ventral. (110) Sternite 2. (111) Sternite 3. (112) Sternite 5. (113) Sternite 8. (114) Sternite 17. (115) Sternite 27. (116) Sternite 35. (117) Sternite 42. (From Pereira et al., 2000). (118). Ityphilus idanus Crabill, 1960 (female holotype; BRITISH WEST INDIES: Barbuda, Damby Cave): Right forcipular telopodite, ventral (a: detail of calyx of poison gland). (From Crabill, 1960). Scale bars: 0.05 mm (108); 0.1 mm (107, 109 117). No scales available (106, 118)." httpUri="https://zenodo.org/record/223526/files/figure.png" pageId="13" pageNumber="514">Figure 107</figureCitation>
); a. a. XIV conspicuously longer than wide, length/width ratio
<emphasis id="B90DEA89FFEC4508FBF912549D44FCC5" box="[1120,1146,861,884]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 1.34: 1 (about as wide as long); a.a. XIV shorter than a.a. XIXIII taken together, ratio
<emphasis id="B90DEA89FFEC4508FCC112889A4CFC29" box="[856,882,897,920]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 0.87:1 (a.a. XIV longer than a.a. XIXIII taken together, ratio
<emphasis id="B90DEA89FFEC4508FEA012AC986AFC0D" box="[313,340,933,956]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 1.25:1); shape of
<typeStatus id="54C28839FFEC4508FDBD12AC9B6AFC0C" box="[548,596,933,957]" pageId="13" pageNumber="514">type</typeStatus>
<emphasis id="B90DEA89FFEC4508FDC212AC9B59FC0D" box="[603,615,933,956]" italics="true" pageId="13" pageNumber="514">c</emphasis>
sensilla on dorsal side of a. a. IX and XIII as in
<figureCitation id="13422A1EFFEC4508FB1B12AD9DCFFC0C" box="[1154,1265,932,957]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="13" pageNumber="514">Figure 60</figureCitation>
: c (as in
<figureCitation id="13422A1EFFEC4508FACA12AD99FBFC51" captionStart="FIGURES 106 118" captionStartId="22.[151,261,1755,1778]" captionTargetBox="[155,1434,210,1734]" captionTargetId="figure@22.[151,1436,193,1734]" captionTargetPageId="22" captionText="FIGURES 106 118. (106). Ityphilus calinus Chamberlin, 1957 (male holotype; COLOMBIA: Valle del Cauca Department): Anterior end of the body, dorsal. (From Chamberlin, 1957). (107). Ityphilus donatellae Pereira, 2012 (female paratype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): Left antenna dorsal. (From Pereira, 2012). (108 117). Ityphilus donatellae Pereira, 2012 (female holotype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): (108) Left a. a. XIII, dorsal (a, b, c: a, b, c, type sensilla). (109) Detail of calyx of poison gland in left forcipular telopodite, ventral. (110) Sternite 2. (111) Sternite 3. (112) Sternite 5. (113) Sternite 8. (114) Sternite 17. (115) Sternite 27. (116) Sternite 35. (117) Sternite 42. (From Pereira et al., 2000). (118). Ityphilus idanus Crabill, 1960 (female holotype; BRITISH WEST INDIES: Barbuda, Damby Cave): Right forcipular telopodite, ventral (a: detail of calyx of poison gland). (From Crabill, 1960). Scale bars: 0.05 mm (108); 0.1 mm (107, 109 117). No scales available (106, 118)." httpUri="https://zenodo.org/record/223526/files/figure.png" pageId="13" pageNumber="514">Figure 108</figureCitation>
: c); internal edge of forcipular tarsungulum expanded inwards bearing 67 strong teeth occupying the middle and apical thirds,
<figureCitation id="13422A1EFFEC4508FEFA12E598D6FBB4" box="[355,488,1004,1029]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="13" pageNumber="514">Figs. 6870</figureCitation>
(internal edge of forcipular tarsungulum not expanded inwards bearing
<emphasis id="B90DEA89FFEC4508FA8612E49C07FBB5" box="[1311,1337,1005,1028]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 7 small teeth occupying the middle third,
<figureCitation id="13422A1EFFEC4508FDBA15069BB6FB99" box="[547,648,1039,1064]" captionStart="FIGURES 106 118" captionStartId="22.[151,261,1755,1778]" captionTargetBox="[155,1434,210,1734]" captionTargetId="figure@22.[151,1436,193,1734]" captionTargetPageId="22" captionText="FIGURES 106 118. (106). Ityphilus calinus Chamberlin, 1957 (male holotype; COLOMBIA: Valle del Cauca Department): Anterior end of the body, dorsal. (From Chamberlin, 1957). (107). Ityphilus donatellae Pereira, 2012 (female paratype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): Left antenna dorsal. (From Pereira, 2012). (108 117). Ityphilus donatellae Pereira, 2012 (female holotype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): (108) Left a. a. XIII, dorsal (a, b, c: a, b, c, type sensilla). (109) Detail of calyx of poison gland in left forcipular telopodite, ventral. (110) Sternite 2. (111) Sternite 3. (112) Sternite 5. (113) Sternite 8. (114) Sternite 17. (115) Sternite 27. (116) Sternite 35. (117) Sternite 42. (From Pereira et al., 2000). (118). Ityphilus idanus Crabill, 1960 (female holotype; BRITISH WEST INDIES: Barbuda, Damby Cave): Right forcipular telopodite, ventral (a: detail of calyx of poison gland). (From Crabill, 1960). Scale bars: 0.05 mm (108); 0.1 mm (107, 109 117). No scales available (106, 118)." httpUri="https://zenodo.org/record/223526/files/figure.png" pageId="13" pageNumber="514">Fig. 109</figureCitation>
); shape of pore-fields as in
<figureCitation id="13422A1EFFEC4508FC4B15069D48FB99" box="[978,1142,1039,1064]" captionStart-0="FIGURES 66 74" captionStart-1="FIGURES 75 82" captionStart-2="FIGURES 83 96" captionStartId-0="17.[151,264,1797,1820]" captionStartId-1="18.[151,264,1866,1889]" captionStartId-2="19.[151,264,1851,1874]" captionTargetBox-0="[151,1436,193,1776]" captionTargetBox-1="[151,1436,193,1845]" captionTargetBox-2="[151,1436,193,1830]" captionTargetId-0="figure@17.[151,1436,193,1777]" captionTargetId-1="figure@18.[151,1436,193,1845]" captionTargetId-2="figure@19.[151,1436,193,1830]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." captionText-1="FIGURES 75 82. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (75) Sternite 17. (76) Sternite 23. (77) Sternite 26. (78) Sternite 31. (79) Sternite 36. (80) Sternite 37. (81) Sternite 38. (82) Sternite 39. Scale bar: 0.1 mm." captionText-2="FIGURES 83 96. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (84 93); 0.1 mm (83, 94 96)." httpUri-0="https://zenodo.org/record/223521/files/figure.png" httpUri-1="https://zenodo.org/record/223522/files/figure.png" httpUri-2="https://zenodo.org/record/223523/files/figure.png" pageId="13" pageNumber="514">Figures 7193</figureCitation>
(as in
<figureCitation id="13422A1EFFEC4508FB5515069CAEFB99" box="[1228,1424,1039,1064]" captionStart="FIGURES 106 118" captionStartId="22.[151,261,1755,1778]" captionTargetBox="[155,1434,210,1734]" captionTargetId="figure@22.[151,1436,193,1734]" captionTargetPageId="22" captionText="FIGURES 106 118. (106). Ityphilus calinus Chamberlin, 1957 (male holotype; COLOMBIA: Valle del Cauca Department): Anterior end of the body, dorsal. (From Chamberlin, 1957). (107). Ityphilus donatellae Pereira, 2012 (female paratype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): Left antenna dorsal. (From Pereira, 2012). (108 117). Ityphilus donatellae Pereira, 2012 (female holotype; BRAZIL: Amazonas: 02 º 34´S, 60 º 06´W): (108) Left a. a. XIII, dorsal (a, b, c: a, b, c, type sensilla). (109) Detail of calyx of poison gland in left forcipular telopodite, ventral. (110) Sternite 2. (111) Sternite 3. (112) Sternite 5. (113) Sternite 8. (114) Sternite 17. (115) Sternite 27. (116) Sternite 35. (117) Sternite 42. (From Pereira et al., 2000). (118). Ityphilus idanus Crabill, 1960 (female holotype; BRITISH WEST INDIES: Barbuda, Damby Cave): Right forcipular telopodite, ventral (a: detail of calyx of poison gland). (From Crabill, 1960). Scale bars: 0.05 mm (108); 0.1 mm (107, 109 117). No scales available (106, 118)." httpUri="https://zenodo.org/record/223526/files/figure.png" pageId="13" pageNumber="514">Figures 110117</figureCitation>
); sternite 5 with
<emphasis id="B90DEA89FFEC4508FEA7153C9866FBFD" box="[318,344,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 34 pores, 8 (
<emphasis id="B90DEA89FFEC4508FE6A153C9B33FBFD" box="[499,525,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 52), 17 (
<emphasis id="B90DEA89FFEC4508FDE5153C9BA9FBFD" box="[636,663,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 72), 27 (
<emphasis id="B90DEA89FFEC4508FC9D153C9A21FBFD" box="[772,799,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 28) (sternite 5 with
<emphasis id="B90DEA89FFEC4508FB93153C9D1AFBFD" box="[1034,1060,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 21 pores, 8 (
<emphasis id="B90DEA89FFEC4508FB26153C9DE7FBFD" box="[1215,1241,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 30), 17 (
<emphasis id="B90DEA89FFEC4508FADE153C9C5FFBFD" box="[1351,1377,1077,1100]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 24), 27 (
<emphasis id="B90DEA89FFEC4508FF5A155099E3FBC1" box="[195,221,1113,1136]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 9)).
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFEC4508FF5E15759C50FB6C" pageId="13" pageNumber="514" type="discussion">
<paragraph id="8BC6369BFFEC4508FF5E15759A4AFB09" blockId="13.[151,1436,644,2037]" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF5E1575980AFB24" bold="true" box="[199,308,1148,1173]" pageId="13" pageNumber="514">Remarks</emphasis>
.
<taxonomicName id="4C794D18FFEC4508FEDB1574988EFB24" box="[322,432,1148,1173]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="bonatoi">
<emphasis id="B90DEA89FFEC4508FEDB1574988EFB24" box="[322,432,1148,1173]" italics="true" pageId="13" pageNumber="514">I. bonatoi</emphasis>
</taxonomicName>
can be separated from all the other Neotropical species of
<taxonomicName id="4C794D18FFEC4508FBD115759D94FB24" box="[1096,1194,1148,1173]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA89FFEC4508FBD115759D94FB24" box="[1096,1194,1148,1173]" italics="true" pageId="13" pageNumber="514">Ityphilus</emphasis>
</taxonomicName>
with internal edge of forcipular tarsungulum serrate, by using the key presented below.
</paragraph>
<paragraph id="8BC6369BFFEC4508FF5E15CC9C50FB6C" blockId="13.[151,1436,644,2037]" box="[199,1390,1220,1245]" pageId="13" pageNumber="514">
For characters differentiating
<taxonomicName id="4C794D18FFEC4508FD8D15CC9BBEFB6C" box="[532,640,1220,1245]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="bonatoi">
<emphasis id="B90DEA89FFEC4508FD8D15CC9BBEFB6C" box="[532,640,1220,1245]" italics="true" pageId="13" pageNumber="514">I. bonatoi</emphasis>
</taxonomicName>
from other Neotropical species of
<taxonomicName id="4C794D18FFEC4508FB9E15CD9D57FB6C" box="[1031,1129,1220,1245]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEA89FFEC4508FB9E15CD9D57FB6C" box="[1031,1129,1220,1245]" italics="true" pageId="13" pageNumber="514">Ityphilus</emphasis>
</taxonomicName>
, see Discussion below.
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFEC4508FF5E15EE9B38FAF9" pageId="13" pageNumber="514" type="materials_examined">
<paragraph id="8BC6369BFFEC4508FF5E15EE9AC4FA95" blockId="13.[151,1436,644,2037]" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF5E15EE98CAFAB1" bold="true" box="[199,500,1255,1280]" pageId="13" pageNumber="514">
<typeStatus id="54C28839FFEC4508FF5E15EE983CFAB1" box="[199,258,1255,1280]" pageId="13" pageNumber="514">Type</typeStatus>
material examined
</emphasis>
:
<collectingCountry id="F36E760BFFEC4508FD9015E19B4DFAB1" box="[521,627,1256,1280]" name="Brazil" pageId="13" pageNumber="514">BRAZIL</collectingCountry>
: RJ:
<collectingCountry id="F36E760BFFEC4508FD2F15EE9BDCFAB1" box="[694,738,1255,1280]" name="Iceland" pageId="13" pageNumber="514">Ilha</collectingCountry>
Grande, Enseada das Palmas,
<date id="FFC7105BFFEC4508FBC015E19C75FAB1" box="[1113,1355,1255,1280]" pageId="13" pageNumber="514" value="1997-02-02" valueMax="1997-02-12" valueMin="1997-02-02">212 February 1997</date>
, M. J. Ramírez leg.:
<typeStatus id="54C28839FFEC4508FEAD140598A8FA94" box="[308,406,1292,1317]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
female, 41 leg-bearing segments, body length
<quantity id="4C819B7EFFEC4508FC3D14059ACBFA95" box="[932,1013,1292,1316]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="514" unit="mm" value="13.0">13 mm</quantity>
.
</paragraph>
<paragraph id="8BC6369BFFEC4508FF5E14389B38FAF9" blockId="13.[151,1436,644,2037]" box="[199,518,1328,1352]" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF5E143898ABFAF9" box="[199,405,1328,1352]" italics="true" pageId="13" pageNumber="514">
Depository of
<typeStatus id="54C28839FFEC4508FEFE143898ABFAF9" box="[359,405,1329,1352]" pageId="13" pageNumber="514">type</typeStatus>
</emphasis>
: MZUSP.
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFEC4510FF5E145D9AA6F96C" lastPageId="21" lastPageNumber="522" pageId="13" pageNumber="514" type="description">
<paragraph id="8BC6369BFFEC4508FF5E145D9BE3FA4C" blockId="13.[151,1436,644,2037]" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF5E145D9866FADC" bold="true" box="[199,344,1364,1389]" pageId="13" pageNumber="514">Description.</emphasis>
Female
<typeStatus id="54C28839FFEC4508FE5C145D9B19FADC" box="[453,551,1364,1389]" pageId="13" pageNumber="514" type="holotype">holotype</typeStatus>
. Forty-one leg-bearing segments, body length
<quantity id="4C819B7EFFEC4508FBC0145D9D8CFADD" box="[1113,1202,1364,1388]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="514" unit="mm" value="13.0">13 mm</quantity>
. Trunk attenuate in anterior and posterior regions. Width of selected leg-bearing segments as follows: 1 (
<quantity id="4C819B7EFFEC4508FBF314719DE9FA21" box="[1130,1239,1400,1424]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="514" unit="mm" value="0.3">0.30 mm</quantity>
); 2 (
<quantity id="4C819B7EFFEC4508FA9514719C46FA21" box="[1292,1400,1400,1424]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="514" unit="mm" value="0.3">0.30 mm</quantity>
); 3 (
<quantity id="4C819B7EFFEC4508FF0714959836FA05" box="[158,264,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="514" unit="mm" value="0.28">0.28 mm</quantity>
); 6 (
<quantity id="4C819B7EFFEC4508FEA11495989CFA05" box="[312,418,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="514" unit="mm" value="0.32">0.32 mm</quantity>
); 12 (
<quantity id="4C819B7EFFEC4508FE7914959B77FA05" box="[480,585,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="514" unit="mm" value="0.4">0.40 mm</quantity>
); 18 (
<quantity id="4C819B7EFFEC4508FD1E14959BCFFA05" box="[647,753,1436,1461]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="514" unit="mm" value="0.45">0.45 mm</quantity>
); 22 (
<quantity id="4C819B7EFFEC4508FCB614959AA6FA05" box="[815,920,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="514" unit="mm" value="0.49">0.49 mm</quantity>
); 27 (
<quantity id="4C819B7EFFEC4508FC4E14959D7EFA05" box="[983,1088,1436,1461]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="514" unit="mm" value="0.51">0.51 mm</quantity>
); 32 (
<quantity id="4C819B7EFFEC4508FBE714959DD6FA05" box="[1150,1256,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="13" pageNumber="514" unit="mm" value="0.48">0.48 mm</quantity>
); 38 (
<quantity id="4C819B7EFFEC4508FABF14959CB1FA05" box="[1318,1423,1436,1460]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="13" pageNumber="514" unit="mm" value="0.39">0.39 mm</quantity>
); 40 (
<quantity id="4C819B7EFFEC4508FF5B14C99815FA69" box="[194,299,1472,1496]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="13" pageNumber="514" unit="mm" value="0.33">0.33 mm</quantity>
); 41 (
<quantity id="4C819B7EFFEC4508FEF314B698EAFA69" box="[362,468,1471,1496]" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="13" pageNumber="514" unit="mm" value="0.36">0.36 mm</quantity>
). Width of cephalic plate
<quantity id="4C819B7EFFEC4508FD6A14C99A64FA69" box="[755,858,1472,1496]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="514" unit="mm" value="0.32">0.32 mm</quantity>
. Width of forcipular coxosternite
<quantity id="4C819B7EFFEC4508FB4D14C99C02FA69" box="[1236,1340,1472,1496]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="514" unit="mm" value="0.32">0.32 mm</quantity>
. Ground color (of preserved specimen in alcohol) pale ocher.
</paragraph>
<paragraph id="8BC6369BFFEC450BFF5E170198C3FEF1" blockId="13.[151,1436,644,2037]" lastBlockId="14.[151,1436,151,320]" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
<emphasis id="B90DEA89FFEC4508FF5E17019806F9AE" box="[199,312,1544,1567]" italics="true" pageId="13" pageNumber="514">Antennae.</emphasis>
About 2.6 times as long as the cephalic plate, conspicuously geniculate, distally clavate (
<figureCitation id="13422A1EFFEC4508FAA1170E9989F9F4" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Figs. 53, 54</figureCitation>
). Basal a.a. slightly overlapping medially (
<figureCitation id="13422A1EFFEC4508FD3C17259BC2F9F4" box="[677,764,1580,1605]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="13" pageNumber="514">Fig. 62</figureCitation>
). Ratio of width of a.a. X (= widest a.a. of distal antennal half)
<emphasis id="B90DEA89FFEC4508FF55175999EAF9D6" box="[204,212,1616,1639]" italics="true" pageId="13" pageNumber="514">/</emphasis>
width of a.a. V (= narrowest a.a. of basal antennal half) 1.66: 1. Apical club extending over a.a. IX to XIV, a.a. VIII transitional, being narrow basally and widened distally (
<figureCitation id="13422A1EFFEC4508FCD4177D9A85F93D" box="[845,955,1652,1677]" captionStart="FIGURES 1 5" captionStartId="2.[151,264,1866,1889]" captionTargetBox="[153,1428,199,1845]" captionTargetId="figure@2.[151,1436,193,1845]" captionTargetPageId="2" captionText="FIGURES 1 5. Ityphilus calinus Chamberlin, 1957 (male holotype; COLOMBIA: Valle del Cauca Department: 13 miles W of Santiago de Cali): (1) Right antenna, ventral. (2) Right antenna, dorsal. (3) Apical portion of right a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (4) Right a. a. II, ventral (b: b type sensilla). (5) Right a. a. V, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (3 5); 0.2 mm (1, 2)." httpUri="https://zenodo.org/record/223512/files/figure.png" pageId="13" pageNumber="514">Figs. 1, 2</figureCitation>
). A.a. XIV apically blunt, shorter than the three previous a.a. taken together (ratio of length of a.a. XIV
<emphasis id="B90DEA89FFEC4508FCA517919A7AF91E" box="[828,836,1688,1711]" italics="true" pageId="13" pageNumber="514">/</emphasis>
length of a.a. XIXIII,
<emphasis id="B90DEA89FFEC4508FBDC17909D5EF901" box="[1093,1120,1689,1712]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 0.87:1). Length
<emphasis id="B90DEA89FFEC4508FA8317919C1CF91E" box="[1306,1314,1688,1711]" italics="true" pageId="13" pageNumber="514">/</emphasis>
width ratio of left a.a. IXIV (in dorsoventral position) as follows: I (0.69: 1); II (0.77: 1); III (0.80: 1); IV (0.88: 1); V (1.04: 1); VI (0.86: 1); VII (0.56: 1); VIII (0.35: 1); IX (0.28: 1); X (0.29: 1); XI (0.36: 1); XII (0.37: 1); XIII (0.40: 1); XIV (1.35: 1). Ventral chaetotaxy: setae on a.a. I to VIII of various lengths, and relatively few in number, those of a.a. IX to XIV much shorter and very numerous (
<figureCitation id="13422A1EFFEC4508FD24162E9A2EF8F1" box="[701,784,1831,1856]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Fig. 53</figureCitation>
). Dorsal chaetotaxy: setae on a.a. I to VIII similar to those on ventral side, setae on a.a. IX to XIV much longer and less numerous than those on ventral side (
<figureCitation id="13422A1EFFEC4508FA9A16459C67F8D4" box="[1283,1369,1868,1893]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Fig. 54</figureCitation>
). A.a. XIV with
<emphasis id="B90DEA89FFEC4508FE9316789815F839" box="[266,299,1905,1928]" italics="true" pageId="13" pageNumber="514">ca.</emphasis>
11 claviform sensilla on the external border, and
<emphasis id="B90DEA89FFEC4508FCF916789A44F839" box="[864,890,1905,1928]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 6 on the internal border (
<figureCitation id="13422A1EFFEC4508FB3C16669DC7F839" box="[1189,1273,1903,1928]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Fig. 55</figureCitation>
); distal end of this a.a. with
<emphasis id="B90DEA89FFEC4508FEB2169C987BF81D" box="[299,325,1941,1964]" italics="true" pageId="13" pageNumber="514">ca</emphasis>
. 11 very small hyaline specialized sensilla apparently not split apically (
<figureCitation id="13422A1EFFEC4508FBF2169D9D80F81C" box="[1131,1214,1940,1965]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Fig. 55</figureCitation>
). Ventral and dorsal surface of a.a. II, V, IX, and XIII with very small specialized sensilla. Ventrally, specialized sensilla distributed apically (
<figureCitation id="13422A1EFFEC4508FE9B16D598AFF844" box="[258,401,2012,2037]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="13" pageNumber="514">Figs. 56, 57</figureCitation>
), and represented by two different
<typeStatus id="54C28839FFEC4508FCB316D49A57F844" box="[810,873,2013,2037]" pageId="13" pageNumber="514">types</typeStatus>
:
<emphasis id="B90DEA89FFEC4508FCEE16D49ABBF845" box="[887,901,2013,2036]" italics="true" pageId="13" pageNumber="514">a</emphasis>
and
<emphasis id="B90DEA89FFEC4508FC5B16D59AEEF844" box="[962,976,2012,2037]" italics="true" pageId="13" pageNumber="514">b</emphasis>
.
<typeStatus id="54C28839FFEC4508FC7816D49D27F844" box="[993,1049,2013,2037]" pageId="13" pageNumber="514">Type</typeStatus>
<emphasis id="B90DEA89FFEC4508FBBA16D49D0FF845" box="[1059,1073,2013,2036]" italics="true" pageId="13" pageNumber="514">a</emphasis>
sensilla very thin and not split apically (
<figureCitation id="13422A1EFFEF450BFF67119E986CFF01" box="[254,338,151,176]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="14" pageNumber="515">Fig. 56</figureCitation>
: a);
<typeStatus id="54C28839FFEF450BFE191191988EFF01" box="[384,432,152,176]" pageId="14" pageNumber="515">type</typeStatus>
<emphasis id="B90DEA89FFEF450BFE2F119E98FAFF01" box="[438,452,151,176]" italics="true" pageId="14" pageNumber="515">b</emphasis>
sensilla (
<figureCitation id="13422A1EFFEF450BFDB4119E9BBEFF01" box="[557,640,151,176]" captionStart="FIGURES 53 57" captionStartId="15.[151,264,1866,1889]" captionTargetBox="[156,1436,193,1845]" captionTargetId="figure@15.[151,1436,193,1845]" captionTargetPageId="15" captionText="FIGURES 53 57. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a. a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a. a. V, ventral (a, b: a, b type sensilla). (57) Left a. a. IX, ventral (a, b: a, b type sensilla). Scale bars: 0.05 mm (55 57); 0.2 mm (53, 54)." httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="14" pageNumber="515">Fig. 56</figureCitation>
: b) very similar to those on the apex of a.a. XIV. Specialized sensilla on dorsal side distributed apically (
<figureCitation id="13422A1EFFEF450BFE6711B59BBBFF64" box="[510,645,188,213]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Figs. 5860</figureCitation>
) and represented by three different
<typeStatus id="54C28839FFEF450BFB8C11B49D6AFF64" box="[1045,1108,189,213]" pageId="14" pageNumber="515">types</typeStatus>
:
<emphasis id="B90DEA89FFEF450BFBC611B49D53FF65" box="[1119,1133,189,212]" italics="true" pageId="14" pageNumber="515">a</emphasis>
and
<emphasis id="B90DEA89FFEF450BFB3D11B59D8CFF64" box="[1188,1202,188,213]" italics="true" pageId="14" pageNumber="515">b</emphasis>
similar to
<emphasis id="B90DEA89FFEF450BFAB211B49C07FF65" box="[1323,1337,189,212]" italics="true" pageId="14" pageNumber="515">a</emphasis>
and
<emphasis id="B90DEA89FFEF450BFAF611B59C43FF64" box="[1391,1405,188,213]" italics="true" pageId="14" pageNumber="515">b</emphasis>
of ventral side (
<figureCitation id="13422A1EFFEF450BFEB411D698BAFF49" box="[301,388,223,248]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 60</figureCitation>
: a, b), and
<typeStatus id="54C28839FFEF450BFD9111E99B06FF49" box="[520,568,224,248]" pageId="14" pageNumber="515">type</typeStatus>
<emphasis id="B90DEA89FFEF450BFDDB11E89B70FF49" box="[578,590,225,248]" italics="true" pageId="14" pageNumber="515">c</emphasis>
sensilla “spine-like, larger, not divided apically, and slightly darker (pale brownishochreous) (
<figureCitation id="13422A1EFFEF450BFE12100D98E1FEAC" box="[395,479,260,285]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 60</figureCitation>
: c). Number and distribution of specialized sensilla on ventral and dorsal sides of a.a. II, V, IX, and XIII as in
<tableCitation id="C6FB0320FFEF450BFE3A102E98C6FEF1" box="[419,504,295,320]" captionStart="TABLE 3" captionStartId="14.[151,238,364,387]" captionTargetBox="[159,1337,446,675]" captionTargetPageId="14" captionText="TABLE 3. Number of type a, b and c sensilla on antennal articles II, V, IX and XIII in the female holotype of Ityphilus bonatoi sp. nov." httpUri="http://table.plazi.org/id/DF066613FFEF450BFF0E10659801FE12" pageId="14" pageNumber="515" tableUuid="DF066613FFEF450BFF0E10659801FE12">Table 3</tableCitation>
.
</paragraph>
<caption id="DF066613FFEF450BFF0E10659801FE12" ID-Table-UUID="DF066613FFEF450BFF0E10659801FE12" httpUri="http://table.plazi.org/id/DF066613FFEF450BFF0E10659801FE12" pageId="14" pageNumber="515" targetBox="[159,1337,446,675]" targetIsTable="true" targetPageId="14">
<paragraph id="8BC6369BFFEF450BFF0E10659801FE12" blockId="14.[151,1436,363,419]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF0E1065983CFE32" bold="true" box="[151,258,364,387]" pageId="14" pageNumber="515">TABLE 3</emphasis>
. Number of type
<emphasis id="B90DEA89FFEF450BFE5D106598EFFE32" box="[452,465,364,387]" italics="true" pageId="14" pageNumber="515">a</emphasis>
,
<emphasis id="B90DEA89FFEF450BFE79106298D3FE32" box="[480,493,363,387]" italics="true" pageId="14" pageNumber="515">b</emphasis>
and
<emphasis id="B90DEA89FFEF450BFDBD10659B0EFE32" box="[548,560,364,387]" italics="true" pageId="14" pageNumber="515">c</emphasis>
sensilla on antennal articles II, V, IX and XIII in the female holotype of
<taxonomicName id="4C794D18FFEF450BFADB106299D8FE12" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFEF450BFADB106299D8FE12" italics="true" pageId="14" pageNumber="515">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFEF450BFF7510859801FE12" box="[236,319,396,419]" pageId="14" pageNumber="515" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BC6369BFFEF450BFECF10B79C39FD12" pageId="14" pageNumber="515">
<table id="F979C43BFFEFBAFAFF0610B79C07FD12" box="[159,1337,446,675]" gridcols="7" gridrows="6" pageId="14" pageNumber="515">
<tr id="354934D9FFEFBAFAFF0610B79C07FE65" box="[159,1337,446,468]" gridrow="0" pageId="14" pageNumber="515" rowspan-0="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<th id="76985DA5FFEFBAFAFECF10B798A1FE65" box="[342,415,446,468]" gridcol="1" gridrow="0" pageId="14" pageNumber="515">Ventral</th>
<th id="76985DA5FFEFBAFAFD5F10B79A34FE65" box="[710,778,446,468]" gridcol="3" gridrow="0" pageId="14" pageNumber="515">Dorsal</th>
<th id="76985DA5FFEFBAFAFB7410B79C07FE65" box="[1261,1337,446,468]" gridcol="6" gridrow="0" pageId="14" pageNumber="515">Figures</th>
</tr>
<tr id="354934D9FFEFBAFAFF0610E19C07FE4F" box="[159,1337,488,510]" gridrow="1" pageId="14" pageNumber="515" rowspan-0="1" rowspan-6="1">
<td id="76985DA5FFEFBAFAFECF10E198A1FE4F" box="[342,415,488,510]" gridcol="1" gridrow="1" pageId="14" pageNumber="515">a</td>
<td id="76985DA5FFEFBAFAFD9710E19B25FE4F" box="[526,539,488,510]" gridcol="2" gridrow="1" pageId="14" pageNumber="515">b</td>
<td id="76985DA5FFEFBAFAFD5F10E19A34FE4F" box="[710,778,488,510]" gridcol="3" gridrow="1" pageId="14" pageNumber="515">a</td>
<td id="76985DA5FFEFBAFAFCE710E19AB5FE4F" box="[894,907,488,510]" gridcol="4" gridrow="1" pageId="14" pageNumber="515">b</td>
<td id="76985DA5FFEFBAFAFBAF10E19D7DFE4F" box="[1078,1091,488,510]" gridcol="5" gridrow="1" pageId="14" pageNumber="515">c</td>
</tr>
<tr id="354934D9FFEFBAFAFF0613189C07FD96" box="[159,1337,529,551]" gridrow="2" pageId="14" pageNumber="515" rowspan-6="1">
<th id="76985DA5FFEFBAFAFF06131899F4FD96" box="[159,202,529,551]" gridcol="0" gridrow="2" pageId="14" pageNumber="515">II</th>
<td id="76985DA5FFEFBAFAFECF131898A1FD96" box="[342,415,529,551]" gridcol="1" gridrow="2" pageId="14" pageNumber="515">-</td>
<td id="76985DA5FFEFBAFAFD9713189B25FD96" box="[526,539,529,551]" gridcol="2" gridrow="2" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD5F13189A34FD96" box="[710,778,529,551]" gridcol="3" gridrow="2" pageId="14" pageNumber="515">-</td>
<td id="76985DA5FFEFBAFAFCE713189AB5FD96" box="[894,907,529,551]" gridcol="4" gridrow="2" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFBAF13189D7DFD96" box="[1078,1091,529,551]" gridcol="5" gridrow="2" pageId="14" pageNumber="515">-</td>
</tr>
<tr id="354934D9FFEFBAFAFF0613339C07FDE1" box="[159,1337,570,592]" gridrow="3" pageId="14" pageNumber="515">
<th id="76985DA5FFEFBAFAFF06133399F4FDE1" box="[159,202,570,592]" gridcol="0" gridrow="3" pageId="14" pageNumber="515">V</th>
<td id="76985DA5FFEFBAFAFECF133398A1FDE1" box="[342,415,570,592]" gridcol="1" gridrow="3" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD9713339B25FDE1" box="[526,539,570,592]" gridcol="2" gridrow="3" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD5F13339A34FDE1" box="[710,778,570,592]" gridcol="3" gridrow="3" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFCE713339AB5FDE1" box="[894,907,570,592]" gridcol="4" gridrow="3" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFBAF13339D7DFDE1" box="[1078,1091,570,592]" gridcol="5" gridrow="3" pageId="14" pageNumber="515">-</td>
<td id="76985DA5FFEFBAFAFB7413339C07FDE1" box="[1261,1337,570,592]" gridcol="6" gridrow="3" pageId="14" pageNumber="515">56, 58</td>
</tr>
<tr id="354934D9FFEFBAFAFF06136D9C07FDCB" box="[159,1337,612,634]" gridrow="4" pageId="14" pageNumber="515">
<th id="76985DA5FFEFBAFAFF06136D99F4FDCB" box="[159,202,612,634]" gridcol="0" gridrow="4" pageId="14" pageNumber="515">IX</th>
<td id="76985DA5FFEFBAFAFECF136D98A1FDCB" box="[342,415,612,634]" gridcol="1" gridrow="4" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD97136D9B25FDCB" box="[526,539,612,634]" gridcol="2" gridrow="4" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD5F136D9A34FDCB" box="[710,778,612,634]" gridcol="3" gridrow="4" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFCE7136D9AB5FDCB" box="[894,907,612,634]" gridcol="4" gridrow="4" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFBAF136D9D7DFDCB" box="[1078,1091,612,634]" gridcol="5" gridrow="4" pageId="14" pageNumber="515">2</td>
<td id="76985DA5FFEFBAFAFB74136D9C07FDCB" box="[1261,1337,612,634]" gridcol="6" gridrow="4" pageId="14" pageNumber="515">57, 59</td>
</tr>
<tr id="354934D9FFEFBAFAFF0613849C07FD12" box="[159,1337,653,675]" gridrow="5" pageId="14" pageNumber="515">
<th id="76985DA5FFEFBAFAFF06138499F4FD12" box="[159,202,653,675]" gridcol="0" gridrow="5" pageId="14" pageNumber="515">XIII</th>
<td id="76985DA5FFEFBAFAFECF138498A1FD12" box="[342,415,653,675]" gridcol="1" gridrow="5" pageId="14" pageNumber="515">-</td>
<td id="76985DA5FFEFBAFAFD9713849B25FD12" box="[526,539,653,675]" gridcol="2" gridrow="5" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFD5F13849A34FD12" box="[710,778,653,675]" gridcol="3" gridrow="5" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFCE713849AB5FD12" box="[894,907,653,675]" gridcol="4" gridrow="5" pageId="14" pageNumber="515">1</td>
<td id="76985DA5FFEFBAFAFBAF13849D7DFD12" box="[1078,1091,653,675]" gridcol="5" gridrow="5" pageId="14" pageNumber="515">2</td>
<td id="76985DA5FFEFBAFAFB7413849C07FD12" box="[1261,1337,653,675]" gridcol="6" gridrow="5" pageId="14" pageNumber="515">60</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E13ED9B02FC91" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E13ED984AFD4C" box="[199,372,740,765]" italics="true" pageId="14" pageNumber="515">Cephalic plate.</emphasis>
About as wide as the forcipular tergite; slightly wider than long (width
<emphasis id="B90DEA89FFEF450BFB3313EC9D8CFD4D" box="[1194,1202,741,764]" italics="true" pageId="14" pageNumber="515">/</emphasis>
length ratio, 1.13:1); shape and chaetotaxy as in
<figureCitation id="13422A1EFFEF450BFE51120E9B09FC91" box="[456,567,775,800]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Figure 61</figureCitation>
.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E12259A32FCF4" blockId="14.[151,1437,740,2025]" box="[199,780,812,837]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E1225981FFCF4" box="[199,289,812,837]" italics="true" pageId="14" pageNumber="515">Clypeus</emphasis>
. With 7+5 setae distributed as in
<figureCitation id="13422A1EFFEF450BFD0112259A36FCF4" box="[664,776,812,837]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Figure 62</figureCitation>
.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E12469B21FC3C" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E1246981EFCD9" box="[199,288,847,872]" italics="true" pageId="14" pageNumber="515">Labrum</emphasis>
. Poorly pigmented; mid-piece membranous, smooth, slightly concave; sidepieces with 2+2 diminutive sharply pointed denticles (
<figureCitation id="13422A1EFFEF450BFE27127D9B2CFC3C" box="[446,530,884,909]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 63</figureCitation>
).
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E129E9BD1FC64" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E129E9803FC01" box="[199,317,919,944]" italics="true" pageId="14" pageNumber="515">Mandibles</emphasis>
. Dentate lamella not subdivided into blocks, 9 teeth in the right mandible (
<figureCitation id="13422A1EFFEF450BFB01129E9DD3FC01" box="[1176,1261,919,944]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 64</figureCitation>
), 7 teeth in left mandible; pectinate lamella with
<emphasis id="B90DEA89FFEF450BFD9512B49B18FC65" box="[524,550,957,980]" italics="true" pageId="14" pageNumber="515">ca</emphasis>
. 17 hyaline teeth.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E12E89A44FBD4" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E12E8984EFC49" box="[199,368,991,1016]" italics="true" pageId="14" pageNumber="515">First maxillae.</emphasis>
Coxosternite apparently without lappets, telopodites bearing lappets (
<figureCitation id="13422A1EFFEF450BFB0712D69DCBFC49" box="[1182,1269,991,1016]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 65</figureCitation>
). Coxosternite devoid of setae; coxal projections subtriangular, round tipped and provided with 1+1 setae (
<figureCitation id="13422A1EFFEF450BFB36150D9C3AFBAC" box="[1199,1284,1028,1053]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 65</figureCitation>
). Telopodites without visible suture between presumptive basal and distal articles, ventral surface with 1+1 setae near the central part of the medial edge (
<figureCitation id="13422A1EFFEF450BFE31154598C5FBD4" box="[424,507,1100,1125]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 65</figureCitation>
), dorsal surface with 1+1 sensilla.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E1566984CFB61" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E15669841FB39" box="[199,383,1135,1160]" italics="true" pageId="14" pageNumber="515">Second maxillae</emphasis>
. Coxosternite without any trace of suture along the sagittal plane and provided with 4+5 setae arranged as in
<figureCitation id="13422A1EFFEF450BFED9159D988EFB1C" box="[320,432,1172,1197]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Figure 65</figureCitation>
. Apical claw of telopodites well developed, bipectinate, dorsal and ventral edges with 7 teeth (
<figureCitation id="13422A1EFFEF450BFF4515BE9858FB61" box="[220,358,1207,1232]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Figs. 66, 67</figureCitation>
).
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E15D598F9FA41" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E15D59895FB45" box="[199,427,1244,1269]" italics="true" pageId="14" pageNumber="515">Forcipular segment</emphasis>
. When closed, telopodites not reaching the anterior margin of the head (
<figureCitation id="13422A1EFFEF450BFAAA15D59CB1FB44" box="[1331,1423,1244,1269]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 61</figureCitation>
). Forcipular tergite slightly wider than the tergite of the first leg-bearing segment (ratio 1.10:1), chaetotaxy represented by two irregular transverse rows each of
<emphasis id="B90DEA89FFEF450BFD77142C9A36FA8D" box="[750,776,1317,1340]" italics="true" pageId="14" pageNumber="515">ca</emphasis>
.1113 setae (
<figureCitation id="13422A1EFFEF450BFC3D142D9AC9FA8C" box="[932,1015,1316,1341]" captionStart="FIGURES 58 65" captionStartId="16.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1773]" captionTargetId="figure@16.[151,1436,193,1777]" captionTargetPageId="16" captionText="FIGURES 58 65. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a. a. V, dorsal (a, b: a, b type sensilla). (59) Letft a. a. IX, dorsal (a, b, c: a, b, c type sensilla). (60) Left a. a. XIII, dorsal (a, b, c: a, b, c type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (58 60); 0.1 mm (63, 65); 0.2 mm (61, 62)." httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="14" pageNumber="515">Fig. 61</figureCitation>
). Coxosternite: with complete chitinlines (
<figureCitation id="13422A1EFFEF450BFF42144E9811FAD1" box="[219,303,1351,1376]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Fig. 68</figureCitation>
: a); maximum width
<emphasis id="B90DEA89FFEF450BFD8C14419B23FAEE" box="[533,541,1352,1375]" italics="true" pageId="14" pageNumber="515">/</emphasis>
length at the middle ratio, 1.83:1; central part of anterior margin as in
<figureCitation id="13422A1EFFEF450BFAB0144E9CA6FAD1" box="[1321,1432,1351,1376]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Figure 68</figureCitation>
. Telopodites: all articles without teeth; trochanteroprefemur with greatest length
<emphasis id="B90DEA89FFEF450BFB9314649D2CFA35" box="[1034,1042,1389,1412]" italics="true" pageId="14" pageNumber="515">/</emphasis>
greatest width 1.12:1; internal edge of tarsungula expanded inwards, bearing 67 strong teeth occupying the middle and apical thirds (
<figureCitation id="13422A1EFFEF450BFA9D14869CAEFA19" box="[1284,1424,1423,1448]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Figs. 6870</figureCitation>
). Calyx of poison gland with shape and relative size as in
<figureCitation id="13422A1EFFEF450BFC8F14BD9A85FA7D" box="[790,955,1460,1485]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Figures 69, 70</figureCitation>
. Shape and chaetotaxy of coxosternite and telopodites as in
<figureCitation id="13422A1EFFEF450BFECD14DE98FDFA41" box="[340,451,1495,1520]" captionStart="FIGURES 66 74" captionStartId="17.[151,264,1797,1820]" captionTargetBox="[151,1436,193,1776]" captionTargetId="figure@17.[151,1436,193,1777]" captionTargetPageId="17" captionText="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="14" pageNumber="515">Figure 68</figureCitation>
.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E14F49D3AF9A4" blockId="14.[151,1437,740,2025]" box="[199,1028,1532,1557]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E14F49819F9A5" box="[199,295,1533,1556]" italics="true" pageId="14" pageNumber="515">Tergites.</emphasis>
Surface of pretergites and metatergites smooth, sulci not evident.
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E17299AC6F979" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E17299BC2F989" box="[199,764,1567,1592]" italics="true" pageId="14" pageNumber="515">Sternites of leg-bearing segments 1 to penultimate</emphasis>
. Pore-fields present in an uninterrupted series from sternite 2 to penultimate (40) inclusive. All pore-fields undivided and located on raised prominences. Form and relative size of pore-fields changing along the trunk as in
<figureCitation id="13422A1EFFEF450BFD5D176E9A56F931" box="[708,872,1639,1664]" captionStart-0="FIGURES 66 74" captionStart-1="FIGURES 75 82" captionStart-2="FIGURES 83 96" captionStartId-0="17.[151,264,1797,1820]" captionStartId-1="18.[151,264,1866,1889]" captionStartId-2="19.[151,264,1851,1874]" captionTargetBox-0="[151,1436,193,1776]" captionTargetBox-1="[151,1436,193,1845]" captionTargetBox-2="[151,1436,193,1830]" captionTargetId-0="figure@17.[151,1436,193,1777]" captionTargetId-1="figure@18.[151,1436,193,1845]" captionTargetId-2="figure@19.[151,1436,193,1830]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionText-0="FIGURES 66 74. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (69 74); 0.2 mm (68)." captionText-1="FIGURES 75 82. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (75) Sternite 17. (76) Sternite 23. (77) Sternite 26. (78) Sternite 31. (79) Sternite 36. (80) Sternite 37. (81) Sternite 38. (82) Sternite 39. Scale bar: 0.1 mm." captionText-2="FIGURES 83 96. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (84 93); 0.1 mm (83, 94 96)." httpUri-0="https://zenodo.org/record/223521/files/figure.png" httpUri-1="https://zenodo.org/record/223522/files/figure.png" httpUri-2="https://zenodo.org/record/223523/files/figure.png" pageId="14" pageNumber="515">Figures 7193</figureCitation>
. Number of pores on selected sternites as follows: sternite 2 (7); 4 (26); 5 (34); 8 (52); 9 (64); 12 (72); 13 (76); 14 (66); 15 (71); 16 (70); 17 (72); 18 (64); 20 (57); 23 (45); 26 (29); 27 (28); 31 (16); 35 (13); 36 (12); 37 (6); 38 (4); 39 (3); 40 (2).
</paragraph>
<paragraph id="8BC6369BFFEF450BFF5E17DC98B8F8E9" blockId="14.[151,1437,740,2025]" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E17DC9B3CF95C" box="[199,514,1748,1773]" italics="true" pageId="14" pageNumber="515">Legs (pair 1 to penultimate)</emphasis>
. First pair slightly shorter than second pair (ratio 0.90: 1), aspect and relative size as in
<figureCitation id="13422A1EFFEF450BFF4D17FE9844F8A1" box="[212,378,1783,1808]" captionStart="FIGURES 83 96" captionStartId="19.[151,264,1851,1874]" captionTargetBox="[151,1436,193,1830]" captionTargetId="figure@19.[151,1436,193,1830]" captionTargetPageId="19" captionText="FIGURES 83 96. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (84 93); 0.1 mm (83, 94 96)." httpUri="https://zenodo.org/record/223523/files/figure.png" pageId="14" pageNumber="515">Figures 94, 95</figureCitation>
. Chaetotaxy of legs similar throughout the entire body. Distribution, number and relative size of setae as in
<figureCitation id="13422A1EFFEF450BFEAC161598E5F885" box="[309,475,1820,1845]" captionStart-0="FIGURES 83 96" captionStart-1="FIGURES 97 102" captionStartId-0="19.[151,264,1851,1874]" captionStartId-1="20.[151,264,1866,1889]" captionTargetBox-0="[151,1436,193,1830]" captionTargetBox-1="[151,1434,193,1845]" captionTargetId-0="figure@19.[151,1436,193,1830]" captionTargetId-1="figure@20.[151,1436,193,1845]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 83 96. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (84 93); 0.1 mm (83, 94 96)." captionText-1="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri-0="https://zenodo.org/record/223523/files/figure.png" httpUri-1="https://zenodo.org/record/223524/files/figure.png" pageId="14" pageNumber="515">Figures 9499</figureCitation>
. Claws with three thin pale accessory spines ventrobasally, arrangement and relative size as in
<figureCitation id="13422A1EFFEF450BFE9D163698BFF8E9" box="[260,385,1855,1880]" captionStart="FIGURES 97 102" captionStartId="20.[151,264,1866,1889]" captionTargetBox="[151,1434,193,1845]" captionTargetId="figure@20.[151,1436,193,1845]" captionTargetPageId="20" captionText="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="14" pageNumber="515">Figure 100</figureCitation>
.
</paragraph>
<paragraph id="8BC6369BFFEF4510FF5E166D9C79F9C1" blockId="14.[151,1437,740,2025]" lastBlockId="21.[151,1437,1408,2009]" lastPageId="21" lastPageNumber="522" pageId="14" pageNumber="515">
<emphasis id="B90DEA89FFEF450BFF5E166D9B25F8CD" box="[199,539,1892,1917]" italics="true" pageId="14" pageNumber="515">Ultimate leg-bearing segment</emphasis>
. About 1.10 times as wide as the penultimate; intercalary pleurites present at both sides of ultimate pretergite; ultimate presternite divided along the sagittal plane; length
<emphasis id="B90DEA89FFEF450BFB3316819D8CF82E" box="[1194,1202,1928,1951]" italics="true" pageId="14" pageNumber="515">/</emphasis>
width ratio of tergite 0.72: 1; length
<emphasis id="B90DEA89FFEF450BFEA416A4987BF875" box="[317,325,1965,1988]" italics="true" pageId="14" pageNumber="515">/</emphasis>
width ratio of sternite 1.0: 1. Shape and chaetotaxy of tergite and sternite as in
<figureCitation id="13422A1EFFEF450BFB4D16A59C63F875" box="[1236,1373,1964,1989]" captionStart="FIGURES 97 102" captionStartId="20.[151,264,1866,1889]" captionTargetBox="[151,1434,193,1845]" captionTargetId="figure@20.[151,1436,193,1845]" captionTargetPageId="20" captionText="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="14" pageNumber="515">Figures 102</figureCitation>
,
<figureCitation id="13422A1EFFEF450BFAF216A59CA6F875" box="[1387,1432,1964,1988]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="14" pageNumber="515">103</figureCitation>
. Coxopleura very slightly protruding at distal-internal ventral ends, setae distributed on almost the whole ventral and lateral surfaces, dorsal side with setae on the external half only (
<figureCitation id="13422A1EFFF44510FC0014899D3BFA29" box="[921,1029,1408,1433]" captionStart="FIGURES 97 102" captionStartId="20.[151,264,1866,1889]" captionTargetBox="[151,1434,193,1845]" captionTargetId="figure@20.[151,1436,193,1845]" captionTargetPageId="20" captionText="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="21" pageNumber="522">Figs. 102</figureCitation>
,
<figureCitation id="13422A1EFFF44510FB8914899D00FA29" box="[1040,1086,1408,1432]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">103</figureCitation>
). Two single (“homogeneous”) coxal organs in each coxopleuron, the anterior being smaller than the posterior, shape as in
<figureCitation id="13422A1EFFF44510FB1614AD9C35FA0C" box="[1167,1291,1444,1469]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Figure 105</figureCitation>
. Coxal pores opening on the membrane between coxopleuron and sternite, partially covered by the latter (
<figureCitation id="13422A1EFFF44510FB5314C19C11FA50" box="[1226,1327,1480,1505]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Fig. 105</figureCitation>
), internal cuticular structure of coxal organs as shown in
<figureCitation id="13422A1EFFF44510FD2514E59A07F9B4" box="[700,825,1516,1541]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Figure 105</figureCitation>
(a: mucous layer). Ultimate legs with seven articles. Articles strongly thickened, subconically narrowing from base to distal end, width of trochanter
<emphasis id="B90DEA89FFF44510FB4C17189DE3F999" box="[1237,1245,1553,1576]" italics="true" pageId="21" pageNumber="522">/</emphasis>
width of tarsus 2
<emphasis id="B90DEA89FFF44510FF0E173C998CF9FD" box="[151,178,1589,1612]" italics="true" pageId="21" pageNumber="522">ca</emphasis>
. 2.66: 1. Length of telopodites of ultimate legs
<emphasis id="B90DEA89FFF44510FD5A173C9BF5F9FD" box="[707,715,1589,1612]" italics="true" pageId="21" pageNumber="522">/</emphasis>
length of sternite 1.71: 1. Length of telopodites of ultimate legs
<emphasis id="B90DEA89FFF44510FA0D173C9CA2F9FD" box="[1428,1436,1589,1612]" italics="true" pageId="21" pageNumber="522">/</emphasis>
length of legs of the penultimate pair 0.72: 1. Shape and chaetotaxy of ultimate legs as in
<figureCitation id="13422A1EFFF44510FB1A17519C34F9C1" box="[1155,1290,1624,1649]" captionStart="FIGURES 97 102" captionStartId="20.[151,264,1866,1889]" captionTargetBox="[151,1434,193,1845]" captionTargetId="figure@20.[151,1436,193,1845]" captionTargetPageId="20" captionText="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="21" pageNumber="522">Figures 102</figureCitation>
,
<figureCitation id="13422A1EFFF44510FA8F17519C7DF9C1" box="[1302,1347,1624,1648]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">103</figureCitation>
.
</paragraph>
<caption id="DF066613FFEE450AFF0E16439CB9F82F" httpUri="https://zenodo.org/record/223519/files/figure.png" pageId="15" pageNumber="516" targetBox="[156,1436,193,1845]" targetPageId="15">
<paragraph id="8BC6369BFFEE450AFF0E16439CB9F82F" blockId="15.[151,1436,1866,1950]" pageId="15" pageNumber="516">
<emphasis id="B90DEA89FFEE450AFF0E16439819F8D1" bold="true" box="[151,295,1866,1889]" pageId="15" pageNumber="516">FIGURES 53</emphasis>
<emphasis id="B90DEA89FFEE450AFEAD1642986DF8D1" bold="true" box="[308,339,1867,1888]" pageId="15" pageNumber="516">57.</emphasis>
<taxonomicName id="4C794D18FFEE450AFEC316429B38F8D0" box="[346,518,1867,1889]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFEE450AFEC316429B38F8D0" box="[346,518,1867,1889]" italics="true" pageId="15" pageNumber="516">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFEE450AFD9516429B62F8D1" box="[524,604,1867,1888]" pageId="15" pageNumber="516" rank="species">
<emphasis id="B90DEA89FFEE450AFD9516429B62F8D1" bold="true" box="[524,604,1867,1888]" pageId="15" pageNumber="516">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (53) Left antenna, ventral. (54) Left antenna, dorsal. (55) Apical portion of left a.a. XIV, dorsal (a: claviform sensilla, b: apical specialized sensilla). (56) Left a.a. V, ventral (a, b:
<emphasis id="B90DEA89FFEE450AFE8016809803F82F" box="[281,317,1928,1950]" italics="true" pageId="15" pageNumber="516">a, b</emphasis>
type sensilla). (57) Left a.a. IX, ventral (a, b:
<emphasis id="B90DEA89FFEE450AFD6616809A1AF82F" box="[767,804,1928,1950]" italics="true" pageId="15" pageNumber="516">a, b</emphasis>
type sensilla). Scale bars: 0.05 mm (5557); 0.2 mm (53, 54).
</paragraph>
</caption>
<caption id="DF066613FFF14515FF0E160C9DF4F826" httpUri="https://zenodo.org/record/223520/files/figure.png" pageId="16" pageNumber="517" targetBox="[151,1436,193,1773]" targetPageId="16">
<paragraph id="8BC6369BFFF14515FF0E160C9DF4F826" blockId="16.[151,1436,1797,1943]" pageId="16" pageNumber="517">
<emphasis id="B90DEA89FFF14515FF0E160C9816F8AA" bold="true" box="[151,296,1797,1820]" pageId="16" pageNumber="517">FIGURES 58</emphasis>
<emphasis id="B90DEA89FFF14515FEAC160F986BF8AA" bold="true" box="[309,341,1798,1819]" pageId="16" pageNumber="517">65.</emphasis>
<taxonomicName id="4C794D18FFF14515FEC4160F9B34F8AD" box="[349,522,1798,1820]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF14515FEC4160F9B34F8AD" box="[349,522,1798,1820]" italics="true" pageId="16" pageNumber="517">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF14515FD8A160E9B5BF8AD" box="[531,613,1799,1820]" pageId="16" pageNumber="517" rank="species">
<emphasis id="B90DEA89FFF14515FD8A160E9B5BF8AD" bold="true" box="[531,613,1799,1820]" pageId="16" pageNumber="517">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (58) Left a.a. V, dorsal (a, b:
<emphasis id="B90DEA89FFF14515FAEC160E9CA2F8AD" box="[1397,1436,1798,1820]" italics="true" pageId="16" pageNumber="517">a, b</emphasis>
type sensilla). (59) Letft a.a. IX, dorsal (a, b, c:
<emphasis id="B90DEA89FFF14515FDEA162F9B8FF88A" box="[627,689,1829,1851]" italics="true" pageId="16" pageNumber="517">a, b, c</emphasis>
type sensilla). (60) Left a.a. XIII, dorsal (a, b, c:
<emphasis id="B90DEA89FFF14515FB07162F9DE2F88A" box="[1182,1244,1829,1851]" italics="true" pageId="16" pageNumber="517">a, b, c</emphasis>
type sensilla). (61) Dorsal view of anterior region of the body, showing cephalic plate, bases of antennae, forcipular segment, and anterior portion of leg-bearing segment 1. (62) Clypeus and bases of antennae. (63) Labrum. (64) Dentate lamella of right mandible, ventral. (65) First and second maxillae, ventral. Scale bars: 0.02 mm (64); 0.05 mm (5860); 0.1 mm (63, 65); 0.2 mm (61, 62).
</paragraph>
</caption>
<caption id="DF066613FFF04514FF0E160C9A5DF826" httpUri="https://zenodo.org/record/223521/files/figure.png" pageId="17" pageNumber="518" targetBox="[151,1436,193,1776]" targetPageId="17">
<paragraph id="8BC6369BFFF04514FF0E160C9A5DF826" blockId="17.[151,1437,1797,1943]" pageId="17" pageNumber="518">
<emphasis id="B90DEA89FFF04514FF0E160C9814F8AA" bold="true" box="[151,298,1797,1820]" pageId="17" pageNumber="518">FIGURES 66</emphasis>
<emphasis id="B90DEA89FFF04514FEAE160F9869F8AA" bold="true" box="[311,343,1798,1819]" pageId="17" pageNumber="518">74.</emphasis>
<taxonomicName id="4C794D18FFF04514FEF9160F9B2EF8AD" box="[352,528,1798,1820]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF04514FEF9160F9B2EF8AD" box="[352,528,1798,1820]" italics="true" pageId="17" pageNumber="518">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF04514FD83160E9B50F8AD" box="[538,622,1799,1820]" pageId="17" pageNumber="518" rank="species">
<emphasis id="B90DEA89FFF04514FD83160E9B50F8AD" bold="true" box="[538,622,1799,1820]" pageId="17" pageNumber="518">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (66) Claw of left telopodite of second maxillae, ventral. (67) Claw of left telopodite of second maxillae, dorsal. (68) Forcipular segment, ventral (a: chitin-lines). (69) Detail of duct (a), calyx (b), and poison gland (c), of venom apparatus in left forcipular telopodite, ventral. (70) Detail of duct (a) and calyx (b) of venom apparatus in right forcipular telopodite, ventral. (71) Sternite 2. (72) Sternite 4. (73) Sternite 9. (74) Sternite 13. Scale bars: 0.02 mm (66, 67); 0.1 mm (6974); 0.2 mm (68).
</paragraph>
</caption>
<caption id="DF066613FFF34517FF0E16439C0EF8CE" httpUri="https://zenodo.org/record/223522/files/figure.png" pageId="18" pageNumber="519" targetBox="[151,1436,193,1845]" targetPageId="18">
<paragraph id="8BC6369BFFF34517FF0E16439C0EF8CE" blockId="18.[151,1436,1866,1919]" pageId="18" pageNumber="519">
<emphasis id="B90DEA89FFF34517FF0E16439816F8D1" bold="true" box="[151,296,1866,1889]" pageId="18" pageNumber="519">FIGURES 75</emphasis>
<emphasis id="B90DEA89FFF34517FEAC1642986BF8D1" bold="true" box="[309,341,1867,1888]" pageId="18" pageNumber="519">82.</emphasis>
<taxonomicName id="4C794D18FFF34517FEC416429B34F8D0" box="[349,522,1867,1889]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF34517FEC416429B34F8D0" box="[349,522,1867,1889]" italics="true" pageId="18" pageNumber="519">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF34517FD8A16429B5BF8D1" box="[531,613,1867,1888]" pageId="18" pageNumber="519" rank="species">
<emphasis id="B90DEA89FFF34517FD8A16429B5BF8D1" bold="true" box="[531,613,1867,1888]" pageId="18" pageNumber="519">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (75) Sternite 17. (76) Sternite 23. (77) Sternite 26. (78) Sternite 31. (79) Sternite 36. (80) Sternite 37. (81) Sternite 38. (82) Sternite 39. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="DF066613FFF24516FF0E16329BE5F87D" httpUri="https://zenodo.org/record/223523/files/figure.png" pageId="19" pageNumber="520" targetBox="[151,1436,193,1830]" targetPageId="19">
<paragraph id="8BC6369BFFF24516FF0E16329BE5F87D" blockId="19.[151,1436,1851,1996]" pageId="19" pageNumber="520">
<emphasis id="B90DEA89FFF24516FF0E16329819F8E0" bold="true" box="[151,295,1851,1874]" pageId="19" pageNumber="520">FIGURES 83</emphasis>
<emphasis id="B90DEA89FFF24516FEAD1635986AF8E0" bold="true" box="[308,340,1852,1873]" pageId="19" pageNumber="520">96.</emphasis>
<taxonomicName id="4C794D18FFF24516FEC216359B36F8E3" box="[347,520,1852,1874]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF24516FEC216359B36F8E3" box="[347,520,1852,1874]" italics="true" pageId="19" pageNumber="520">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF24516FD9616359B5EF8E0" box="[527,608,1852,1873]" pageId="19" pageNumber="520" rank="species">
<emphasis id="B90DEA89FFF24516FD9616359B5EF8E0" bold="true" box="[527,608,1852,1873]" pageId="19" pageNumber="520">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (83) Sternite 40. (84) Pore-field on sternite 5. (85) Pore-field on sternite 8. (86) Pore-field on sternite 12. (87) Pore-field on sternite 14. (88) Pore-field on sternite 15. (89) Pore-field on sternite 16. (90) Pore-field on sternite 18. (91) Pore-field on sternite 20. (92) Pore-field on sternite 27. (93) Porefield on sternite 35. (94) Left leg (pair 1), posteroventral view. (95) Left leg (pair 2), posteroventral view. (96) Left leg (pair 14), ventral. Scale bars: 0.03 mm (8493); 0.1 mm (83, 9496).
</paragraph>
</caption>
<caption id="DF066613FFF54511FF0E16439A07F86A" httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="20" pageNumber="521" targetBox="[151,1434,193,1845]" targetPageId="20">
<paragraph id="8BC6369BFFF54511FF0E16439A07F86A" blockId="20.[151,1436,1866,2011]" pageId="20" pageNumber="521">
<emphasis id="B90DEA89FFF54511FF0E16439810F8D1" bold="true" box="[151,302,1866,1889]" pageId="20" pageNumber="521">FIGURES 97</emphasis>
<emphasis id="B90DEA89FFF54511FEA216429856F8D1" bold="true" box="[315,360,1867,1888]" pageId="20" pageNumber="521">102.</emphasis>
<taxonomicName id="4C794D18FFF54511FEEC16429B16F8D0" box="[373,552,1867,1889]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="20" pageNumber="521" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF54511FEEC16429B16F8D0" box="[373,552,1867,1889]" italics="true" pageId="20" pageNumber="521">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF54511FDAE16429BB1F8D1" box="[567,655,1867,1888]" pageId="20" pageNumber="521" rank="species">
<emphasis id="B90DEA89FFF54511FDAE16429BB1F8D1" bold="true" box="[567,655,1867,1888]" pageId="20" pageNumber="521">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (9799); 0.2 mm (101); 0.3 mm (102).
</paragraph>
</caption>
<caption id="DF066613FFF44510FF0E140D9A67FAE8" httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522" targetBox="[156,1431,200,1261]" targetPageId="21">
<paragraph id="8BC6369BFFF44510FF0E140D9A67FAE8" blockId="21.[151,1436,1284,1369]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF0E140D9806FAAB" bold="true" box="[151,312,1284,1307]" pageId="21" pageNumber="522">FIGURES 103</emphasis>
<emphasis id="B90DEA89FFF44510FEDC140C984FFAAB" bold="true" box="[325,369,1285,1306]" pageId="21" pageNumber="522">105.</emphasis>
<taxonomicName id="4C794D18FFF44510FEE5140C9B12FAAA" box="[380,556,1285,1307]" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" pageId="21" pageNumber="522" phylum="Arthropoda" rank="species" species="bonatoi" status="sp. nov.">
<emphasis id="B90DEA89FFF44510FEE5140C9B12FAAA" box="[380,556,1285,1307]" italics="true" pageId="21" pageNumber="522">Ityphilus bonatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23E57F2FFF44510FDA1140F9BB3FAAA" box="[568,653,1286,1307]" pageId="21" pageNumber="522" rank="species">
<emphasis id="B90DEA89FFF44510FDA1140F9BB3FAAA" bold="true" box="[568,653,1286,1307]" pageId="21" pageNumber="522">sp. nov.</emphasis>
</taxonomicNameLabel>
(female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103).
</paragraph>
</caption>
<paragraph id="8BC6369BFFF44510FF5E17759AA6F96C" blockId="21.[151,1437,1408,2009]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF5E17759898F925" box="[199,422,1660,1685]" italics="true" pageId="21" pageNumber="522">Postpedal segments</emphasis>
. Intermediate tergite with posterior margin convex (
<figureCitation id="13422A1EFFF44510FC6617759D5DF924" box="[1023,1123,1660,1685]" captionStart="FIGURES 97 102" captionStartId="20.[151,264,1866,1889]" captionTargetBox="[151,1434,193,1845]" captionTargetId="figure@20.[151,1436,193,1845]" captionTargetPageId="20" captionText="FIGURES 97 102. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (97) Left leg (pair 22), anteroventral view. (98) Left leg (pair 32), ventral. (99) Left leg (pair 40), anteroventral view. (100) Claw of the left leg (pair 13), anteroventral view. (101) Anterior and posterior spermathecae at level of leg-bearing segment 38, dorsal (a: spermatozoa, b: contour of spermatheca, c: pretergite, d: metatergite). (102) Ultimate leg-bearing segment and postpedal segments, dorsal. Scale bars: 0.03 mm (100); 0.1 mm (97 99); 0.2 mm (101); 0.3 mm (102)." httpUri="https://zenodo.org/record/223524/files/figure.png" pageId="21" pageNumber="522">Fig. 102</figureCitation>
); intermediate sternite with posterior margin straight to very slightly convex, posterior margin of first genital sternite slightly concave in midline (
<figureCitation id="13422A1EFFF44510FF6517CD9860F96C" box="[252,350,1732,1757]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Fig. 103</figureCitation>
). Gonopods uniarticulate, without setae (
<figureCitation id="13422A1EFFF44510FCB017CD9AB5F96C" box="[809,907,1732,1757]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFF44510FF5E17E198F6F894" pageId="21" pageNumber="522" type="discussion">
<paragraph id="8BC6369BFFF44510FF5E17E198F6F894" blockId="21.[151,1437,1408,2009]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF5E17E1980AF8B0" bold="true" box="[199,308,1768,1793]" pageId="21" pageNumber="522">Remarks</emphasis>
. The adult condition of the female
<typeStatus id="54C28839FFF44510FD4F17E19A06F8B0" box="[726,824,1768,1793]" pageId="21" pageNumber="522" type="holotype">holotype</typeStatus>
is indicated by the presence of spermatozoa in both spermathecae (
<figureCitation id="13422A1EFFF44510FEA71605989FF894" box="[318,417,1804,1829]" captionStart="FIGURES 103 105" captionStartId="21.[151,264,1284,1307]" captionTargetBox="[156,1431,200,1261]" captionTargetId="figure@21.[151,1436,193,1264]" captionTargetPageId="21" captionText="FIGURES 103 105. Ityphilus bonatoi sp. nov. (female holotype; BRAZIL: RJ: Ilha Grande): (103) Ultimate leg-bearing segment and postpedal segments, ventral. (104) Detail of distal end of right ultimate leg, ventral. (105) Left coxal organs, ventral (a: mucous layer). Scale bars 0.03 mm (104); 0.1 mm (105); 0.3 mm (103)." httpUri="https://zenodo.org/record/223525/files/figure.png" pageId="21" pageNumber="522">Fig. 103</figureCitation>
: a).
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFF44510FF5E16399DD2F820" pageId="21" pageNumber="522" type="etymology">
<paragraph id="8BC6369BFFF44510FF5E16399DD2F820" blockId="21.[151,1437,1408,2009]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF5E16399870F8F8" bold="true" box="[199,334,1840,1865]" pageId="21" pageNumber="522">Etymology.</emphasis>
The species is dedicated to my distinguished colleague Lucio Bonato (Dipartimento di Biologia, University of Padova,
<collectingCountry id="F36E760BFFF44510FE07165D98EDF8DC" box="[414,467,1876,1901]" name="Italy" pageId="21" pageNumber="522">Italy</collectingCountry>
) in recognition of his extraordinary skill, dedication, and hard work as a leader in analysing and establishing a common terminology for the external anatomy of all centipede orders.
</paragraph>
</subSubSection>
<subSubSection id="C3636510FFF44510FF5E16959A3FF868" pageId="21" pageNumber="522" type="materials_examined">
<paragraph id="8BC6369BFFF44510FF5E16959AB6F804" blockId="21.[151,1437,1408,2009]" box="[199,904,1948,1973]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF5E1695985BF804" bold="true" box="[199,357,1948,1973]" pageId="21" pageNumber="522">
<typeStatus id="54C28839FFF44510FF5E1695983CF804" box="[199,258,1948,1973]" pageId="21" pageNumber="522">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F36E760BFFF44510FEF5169498E8F804" box="[364,470,1949,1973]" name="Brazil" pageId="21" pageNumber="522">BRAZIL</collectingCountry>
: RJ:
<collectingCountry id="F36E760BFFF44510FD9516959B06F804" box="[524,568,1948,1973]" name="Iceland" pageId="21" pageNumber="522">Ilha</collectingCountry>
Grande, Enseada das Palmas.
</paragraph>
<paragraph id="8BC6369BFFF44510FF5E16C99A3FF868" blockId="21.[151,1437,1408,2009]" box="[199,769,1984,2009]" pageId="21" pageNumber="522">
<emphasis id="B90DEA89FFF44510FF5E16C9984EF868" bold="true" box="[199,368,1984,2009]" pageId="21" pageNumber="522">Known range.</emphasis>
Only known from the
<typeStatus id="54C28839FFF44510FDE916C89B9EF868" box="[624,672,1985,2009]" pageId="21" pageNumber="522">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>