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<document id="7F16D3BE1EE936EE4000961E7B1398EC" ID-DOI="10.1080/14772019.2018.1446462" ID-ISSN="1478-0941" ID-PMC="PMC6474738" ID-PubMed="31057335" ID-Zenodo-Dep="10883523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711484535305" checkinUser="felipe" docAuthor="Torres-Silva, Ana. I., Eder, Wolfgang, Hohenegger, Johann &amp; Briguglio, Antonino" docDate="2018" docId="9E6A87F1FFC32829FC75FF4993658DB4" docLanguage="en" docName="JSystPalaeontol.17.7.557-595.pdf" docOrigin="Journal of Systematic Palaeontology 17 (7)" docSource="http://dx.doi.org/10.1080/14772019.2018.1446462" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.2:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="2" docTitle="Palaeonummulites trinitatensis" docType="treatment" docVersion="3" lastPageNumber="582" masterDocId="6253FF89FFDB2833FFF3FFF491548914" masterDocTitle="Morphometric analysis of Eocene nummulitids in western and central Cuba: taxonomy, biostratigraphy and evolutionary trends" masterLastPageNumber="595" masterPageNumber="557" pageNumber="580" updateTime="1711485251774" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="209BA0A7AA4BF44FA7019DFC42FC4FB6">Morphometric analysis of Eocene nummulitids in western and central Cuba: taxonomy, biostratigraphy and evolutionary trends</mods:title>
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<mods:namePart id="27FEEEB4FB4B353257BBEE69839E8D02">Torres-Silva, Ana. I.</mods:namePart>
<mods:affiliation id="800BB106479288F8AE012981B9EC09C6">Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria;</mods:affiliation>
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<mods:namePart id="9FF9714D13F2EF9FC0F4EF0F535F6EAA">Eder, Wolfgang</mods:namePart>
<mods:affiliation id="4D27D4BAA67F098C9F7035928E040E1C">Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria;</mods:affiliation>
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<mods:namePart id="91A24599075E7E7E7100FB4D5B024975">Hohenegger, Johann</mods:namePart>
<mods:affiliation id="421B78C84471511C1F6B9376E676C634">Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria;</mods:affiliation>
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<mods:namePart id="7779B76C867299FC148F07BAE09DCAA6">Briguglio, Antonino</mods:namePart>
<mods:affiliation id="ACBBC1F0168DDCABBF554747E1BA450A">Dipartimento di Scienze della, Terra dellAmbiente e della Vita, Universita ́ degli Studi di Genova, Corso Europa, 26, I- 16132 Genova, Italy</mods:affiliation>
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<subSubSection id="5ED9656CFFC3282BFC75FF4994D689C0" box="[902,1410,189,213]" pageId="24" pageNumber="580" type="nomenclature">
<paragraph id="167C36E7FFC3282BFC75FF4994D689C0" blockId="24.[902,1410,189,244]" box="[902,1410,189,213]" pageId="24" pageNumber="580">
<heading id="4D34818BFFC3282BFC75FF4994D689C0" box="[902,1410,189,213]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="580" reason="8">
<taxonomicName id="D1C34D64FFC3282BFC75FF4994D689C0" ID-CoL="8TGFP" authority="(Nuttall, 1928)" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[902,1410,189,213]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC3282BFC75FF49958889C1" bold="true" box="[902,1244,189,213]" italics="true" pageId="24" pageNumber="580">Palaeonummulites trinitatensis</emphasis>
(
<bibRefCitation id="72524B16FFC3282BFB19FF49942C89C0" author="Nuttall, W. L. F." box="[1258,1400,189,212]" pageId="24" pageNumber="580" pagination="57 - 116" refId="ref22440" refString="Nuttall, W. L. F. 1928. Tertiary foraminifera from the Naparima region of Trinidad (British West Indies). Quarterly Journal of the Geological Society, 84, 57 - 116." type="journal article" year="1928">Nuttall, 1928</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5ED9656CFFC3282BFBB2FF2995858B47" pageId="24" pageNumber="580" type="description">
<paragraph id="167C36E7FFC3282BFBB2FF29959289E0" blockId="24.[902,1410,189,244]" box="[1089,1222,221,244]" pageId="24" pageNumber="580">
(
<figureCitation id="8EF82A62FFC3282BFBB9FF2995EA89E0" box="[1098,1214,221,244]" captionStart="Figure 17" captionStartId="25.[159,227,1642,1663]" captionTargetBox="[173,1451,185,1609]" captionTargetId="figure-68@25.[173,1453,183,1610]" captionTargetPageId="25" captionText="Figure 17. AG, Operculinoides soldadensis Vaughan &amp; Cole; A, Loma El Santo, CA-215-871; B, Loma Candelaria, 98LC-1-669; C F, Norona; C, NOR-UN 24; DF, NOR-UN 15/14; G, holotype, Trinidad. H, I, Palaeonummulites trinitatensis (Nutall); H, Loma Candelaria, 98LC-1ICT3; I, holotype of Operculinoides kugleri Vaughan &amp; Cole, Trinidad. J, Operculinoides ocalanus (Cushman), Loma Jabaco, CA-4-724. AD, GJ, A forms in equatorial section; E, A form in axial section; F, external view." figureDoi="http://doi.org/10.5281/zenodo.10883561" httpUri="https://zenodo.org/record/10883561/files/figure.png" pageId="24" pageNumber="580">Fig. 17H, I</figureCitation>
)
</paragraph>
<paragraph id="167C36E7FFC3282BFCBEFEE992DC8840" blockId="24.[845,1467,285,595]" pageId="24" pageNumber="580">
1928
<taxonomicName id="D1C34D64FFC3282BFC78FEE995B18820" authority="Nuttall" authorityName="Nuttall" authorityYear="1928" box="[907,1253,285,308]" class="Globothalamea" family="Nummulitidae" genus="Operculina" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC3282BFC78FEE995D98820" box="[907,1165,285,308]" italics="true" pageId="24" pageNumber="580">Operculina trinitatensis</emphasis>
Nuttall
</taxonomicName>
: 102, pl. 8, figs 10, 11.
</paragraph>
<paragraph id="167C36E7FFC3282BFCBEFEA994488880" blockId="24.[845,1467,285,595]" pageId="24" pageNumber="580">
1941
<taxonomicName id="D1C34D64FFC3282BFC7DFEA9947E8860" authority="(Nuttall)" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[910,1322,349,372]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC3282BFC7DFEA995E98860" box="[910,1213,349,372]" italics="true" pageId="24" pageNumber="580">Operculinoides trinitatensis</emphasis>
(Nuttall)
</taxonomicName>
; Vaughan &amp; Cole: 47, pl. 10, fig. 12, pl. 13, figs 414.
</paragraph>
<paragraph id="167C36E7FFC3282BFCBEFE69944788C7" blockId="24.[845,1467,285,595]" pageId="24" pageNumber="580">
1941
<taxonomicName id="D1C34D64FFC3282BFC78FE6995B988A0" authority="(Nuttall)" baseAuthorityName="Nuttall" box="[907,1261,413,436]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="kugleri">
<emphasis id="24B7EAF5FFC3282BFC78FE6995D788A0" box="[907,1155,413,436]" italics="true" pageId="24" pageNumber="580">Operculinoides kugleri</emphasis>
(Nuttall)
</taxonomicName>
; Vaughan &amp; Cole: 18, pl. 10, figs 35, 7, 8, pl. 13, figs 1, 2.
</paragraph>
<paragraph id="167C36E7FFC3282BFCBEFE28956B8B07" blockId="24.[845,1467,285,595]" pageId="24" pageNumber="580">
1975
<taxonomicName id="D1C34D64FFC3282BFC7AFE2895B888E7" authority="(Nuttall)" baseAuthorityName="Nuttall" box="[905,1260,476,499]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="spiralis">
<emphasis id="24B7EAF5FFC3282BFC7AFE2895D188E7" box="[905,1157,476,499]" italics="true" pageId="24" pageNumber="580">Operculinoides spiralis</emphasis>
(Nuttall)
</taxonomicName>
; Caudri: 542, pl. 1, fig. 20, pl. 8, fig. 13.
</paragraph>
<paragraph id="167C36E7FFC3282BFCBEFDE895858B47" blockId="24.[845,1467,285,595]" pageId="24" pageNumber="580">
1975
<taxonomicName id="D1C34D64FFC3282BFC7FFDE894708B27" authority="(Nuttall)" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[908,1316,540,563]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC3282BFC7FFDE895EE8B27" box="[908,1210,540,563]" italics="true" pageId="24" pageNumber="580">Operculinoides trinitatensis</emphasis>
(Nuttall)
</taxonomicName>
; Caudri: 541, pl. 1, figs 10, 16, pl. 8, figs 14, 15.
</paragraph>
</subSubSection>
<subSubSection id="5ED9656CFFC3282BFCBEFD7494508BA2" pageId="24" pageNumber="580" type="materials_examined">
<paragraph id="167C36E7FFC3282BFCBEFD7494508BA2" blockId="24.[845,1466,640,694]" pageId="24" pageNumber="580">
<emphasis id="24B7EAF5FFC3282BFCBEFD7492E38B83" bold="true" box="[845,951,640,663]" pageId="24" pageNumber="580">Material.</emphasis>
Four megalospheric specimens in equatorial section from Loma Candelaria (98LC-1).
</paragraph>
</subSubSection>
<subSubSection id="5ED9656CFFC3282AFCBEFD1790B48E43" lastPageId="25" lastPageNumber="581" pageId="24" pageNumber="580" type="description">
<paragraph id="167C36E7FFC3282BFCBEFD17928C8BEE" blockId="24.[845,984,739,762]" box="[845,984,739,762]" pageId="24" pageNumber="580">
<emphasis id="24B7EAF5FFC3282BFCBEFD17928C8BEE" bold="true" box="[845,984,739,762]" pageId="24" pageNumber="580">Description.</emphasis>
</paragraph>
<paragraph id="167C36E7FFC3282BFC86FCF8952E8A97" blockId="24.[845,1466,780,899]" pageId="24" pageNumber="580">
<emphasis id="24B7EAF5FFC3282BFC86FCF895138A37" bold="true" box="[885,1095,780,803]" pageId="24" pageNumber="580">External features.</emphasis>
The test is planispiral, involute, laterally slightly compressed. No trace of septal sutures and ornamentation is visible due to poor preservation of the individuals studied here.
</paragraph>
<paragraph id="167C36E7FFC3282BFC86FC4492968DB2" blockId="24.[845,1467,944,1349]" pageId="24" pageNumber="580">
<emphasis id="24B7EAF5FFC3282BFC86FC44951C8AD3" bold="true" box="[885,1096,944,967]" pageId="24" pageNumber="580">Internal features.</emphasis>
Megalospheric generation with spherical proloculus (mean diameter = 0.1 mm) followed by a reniform deuteroloculus and a moderately tightly coiled spiral with commonly three whorls. Rapid increase in height of the last spiral, with chamber height roughly 3 times greater than chamber length. Operculine primary septa with strong backbend angle gently tapered towards inner ends.
</paragraph>
<paragraph id="167C36E7FFC3282BFC94FB5B92948C51" blockId="24.[845,1467,944,1349]" pageId="24" pageNumber="580">
Characters and attributes for
<taxonomicName id="D1C34D64FFC3282BFB52FB5B92DE8DF2" baseAuthorityName="Nuttall" baseAuthorityYear="1928" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC3282BFB52FB5B92DE8DF2" italics="true" pageId="24" pageNumber="580">Palaeonummulites trinitatensis</emphasis>
</taxonomicName>
and comparisons with
<taxonomicName id="D1C34D64FFC3282BFB7CFB3B92388C12" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="striatoreticulatus">
<emphasis id="24B7EAF5FFC3282BFB7CFB3B92388C12" italics="true" pageId="24" pageNumber="580">Nummulites striatoreticulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="D1C34D64FFC3282BFC78FB1B95E38C12" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[907,1207,1263,1286]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC3282BFC78FB1B95E38C12" box="[907,1207,1263,1286]" italics="true" pageId="24" pageNumber="580">Operculinoides floridensis</emphasis>
</taxonomicName>
(tightly coiled),
<taxonomicName id="D1C34D64FFC3282BFA52FB1B92E88C32" baseAuthorityName="Heilprin" baseAuthorityYear="1885" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC3282BFA52FB1B92E88C32" italics="true" pageId="24" pageNumber="580">O. floridensis</emphasis>
</taxonomicName>
(loosely coiled) and
<taxonomicName id="D1C34D64FFC3282BFB59FAFB941E8C32" authorityName="Vaughan &amp; Cole" authorityYear="1941" box="[1194,1354,1295,1318]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="species" species="soldadensis">
<emphasis id="24B7EAF5FFC3282BFB59FAFB941E8C32" box="[1194,1354,1295,1318]" italics="true" pageId="24" pageNumber="580">O. soldadensis</emphasis>
</taxonomicName>
are given in
<tableCitation id="5B41035CFFC3282BFC9BFADA92EC8C51" box="[872,952,1326,1349]" captionStart="Table 5" captionStartId="26.[159,217,186,207]" captionText="Table 5. Characters and attributes (means and standard deviations, SD, in mm) for Palaeonummulites trinitatensis and comparisons with Nummulites striatoreticulatus, Operculinoides floridensis (tightly coiled), O. floridensis (loosely coiled) and O. soldadensis. Symbol key: ++, strong positive differences with &lt;1% error probability; +, differences with &lt;5% error probability; 0, no significant differences; -, negative differences with &lt;5% error probability; ――, strong negative differences with &lt;1% error probability." pageId="24" pageNumber="580">Table 5</tableCitation>
.
</paragraph>
<caption id="42BC666FFFC3282BFF6CFA7F90E68CC1" pageId="24" pageNumber="580" startId="24.[159,217,1419,1440]" targetType="table">
<paragraph id="167C36E7FFC3282BFF6CFA7F90E68CC1" blockId="24.[159,1466,1419,1493]" pageId="24" pageNumber="580">
<emphasis id="24B7EAF5FFC3282BFF6CFA7F91A68CB4" bold="true" box="[159,242,1419,1440]" pageId="24" pageNumber="580">Table 4.</emphasis>
Characters and attributes (means and standard deviations, SD, in mm) for
<emphasis id="24B7EAF5FFC3282BFC37FA7F953E8CB4" box="[964,1130,1419,1440]" italics="true" pageId="24" pageNumber="580">Palaeoummulites</emphasis>
and comparisons with
<taxonomicName id="D1C34D64FFC3282BFABBFA7F94EE8CB4" box="[1352,1466,1419,1440]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC3282BFABBFA7F94EE8CB4" box="[1352,1466,1419,1440]" italics="true" pageId="24" pageNumber="580">Nummulites</emphasis>
</taxonomicName>
and
<emphasis id="24B7EAF5FFC3282BFF3BFA5190368CAE" box="[200,354,1445,1466]" italics="true" pageId="24" pageNumber="580">
<taxonomicName id="D1C34D64FFC3282BFF3BFA51900B8CAE" authorityName="Hanzawa" authorityYear="1935" box="[200,351,1445,1466]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="genus">Operculinoides</taxonomicName>
.
</emphasis>
Symbol key: +, differences with &lt;5% error probability; 0, no significant differences; , strong negative differences with &lt;1% error probability.
</paragraph>
</caption>
<paragraph id="167C36E7FFC3282BFF6CFA0694D38E63" pageId="24" pageNumber="580">
<table id="64C3C447FFC3D7CCFF6CFA0694EE8E63" box="[159,1466,1522,1911]" gridcols="5" gridrows="12" pageId="24" pageNumber="580">
<tr id="A8F334A5FFC3D7CCFF6CFA0694EE8F13" box="[159,1466,1522,1543]" gridrow="0" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CFA06909C8F13" box="[159,456,1522,1543]" gridcol="0" gridrow="0" pageId="24" pageNumber="580">
<taxonomicName id="D1C34D64FFC3282BFF6CFA0690058F13" authority="Mean SD" authorityName="Mean SD" box="[159,337,1522,1543]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC3282BFF6CFA0690058F13" box="[159,337,1522,1543]" italics="true" pageId="24" pageNumber="580">Palaeonummulites</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC3D7CCFDAFFA0693FE8F13" box="[604,682,1522,1543]" gridcol="1" gridrow="0" pageId="24" pageNumber="580">Mean</th>
<th id="EB225DD9FFC3D7CCFCCDFA0692D88F13" box="[830,908,1522,1543]" gridcol="2" gridrow="0" pageId="24" pageNumber="580">SD</th>
<th id="EB225DD9FFC3D7CCFBD3FA0695E08F13" box="[1056,1204,1522,1543]" gridcol="3" gridrow="0" pageId="24" pageNumber="580">
<taxonomicName id="D1C34D64FFC3282BFBD3FA0695E08F13" authorityName="Hanzawa" authorityYear="1935" box="[1056,1204,1522,1543]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC3282BFBD3FA0695E08F13" box="[1056,1204,1522,1543]" italics="true" pageId="24" pageNumber="580">Operculinoides</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC3D7CCFABBFA0694EE8F13" box="[1352,1466,1522,1543]" gridcol="4" gridrow="0" pageId="24" pageNumber="580">
<taxonomicName id="D1C34D64FFC3282BFABBFA0694EE8F13" authority="First" authorityName="First" box="[1352,1466,1522,1543]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="24" pageNumber="580" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC3282BFABBFA0694EE8F13" box="[1352,1466,1522,1543]" italics="true" pageId="24" pageNumber="580">Nummulites</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF9D794EE8F2D" box="[159,1466,1571,1593]" gridrow="1" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF9D7909C8F2D" box="[159,456,1571,1593]" gridcol="0" gridrow="1" pageId="24" pageNumber="580">First chamber length</th>
<td id="EB225DD9FFC3D7CCFDAFF9D793FE8F2D" box="[604,682,1571,1593]" gridcol="1" gridrow="1" pageId="24" pageNumber="580">243.3</td>
<td id="EB225DD9FFC3D7CCFCCDF9D792D88F2D" box="[830,908,1571,1593]" gridcol="2" gridrow="1" pageId="24" pageNumber="580">38.59</td>
<td id="EB225DD9FFC3D7CCFBD3F9D795E08F2D" box="[1056,1204,1571,1593]" gridcol="3" gridrow="1" pageId="24" pageNumber="580">――</td>
<td id="EB225DD9FFC3D7CCFABBF9D794EE8F2D" box="[1352,1466,1571,1593]" gridcol="4" gridrow="1" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF9B794EE8F4D" box="[159,1466,1603,1625]" gridrow="2" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF9B7909C8F4D" box="[159,456,1603,1625]" gridcol="0" gridrow="2" pageId="24" pageNumber="580">Proloculus nominal diameter</th>
<td id="EB225DD9FFC3D7CCFDAFF9B793FE8F4D" box="[604,682,1603,1625]" gridcol="1" gridrow="2" pageId="24" pageNumber="580">113.0</td>
<td id="EB225DD9FFC3D7CCFCCDF9B792D88F4D" box="[830,908,1603,1625]" gridcol="2" gridrow="2" pageId="24" pageNumber="580">25.84</td>
<td id="EB225DD9FFC3D7CCFBD3F9B795E08F4D" box="[1056,1204,1603,1625]" gridcol="3" gridrow="2" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF9B794EE8F4D" box="[1352,1466,1603,1625]" gridcol="4" gridrow="2" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF99794EE8F6D" box="[159,1466,1635,1657]" gridrow="3" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF997909C8F6D" box="[159,456,1635,1657]" gridcol="0" gridrow="3" pageId="24" pageNumber="580">Deuteroloculus ratio</th>
<td id="EB225DD9FFC3D7CCFDAFF99793FE8F6D" box="[604,682,1635,1657]" gridcol="1" gridrow="3" pageId="24" pageNumber="580">0.888</td>
<td id="EB225DD9FFC3D7CCFCCDF99792D88F6D" box="[830,908,1635,1657]" gridcol="2" gridrow="3" pageId="24" pageNumber="580">0.1258</td>
<td id="EB225DD9FFC3D7CCFBD3F99795E08F6D" box="[1056,1204,1635,1657]" gridcol="3" gridrow="3" pageId="24" pageNumber="580">――</td>
<td id="EB225DD9FFC3D7CCFABBF99794EE8F6D" box="[1352,1466,1635,1657]" gridcol="4" gridrow="3" pageId="24" pageNumber="580">0</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF97694EE8F8C" box="[159,1466,1666,1688]" gridrow="4" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF976909C8F8C" box="[159,456,1666,1688]" gridcol="0" gridrow="4" pageId="24" pageNumber="580">Initial marginal radius</th>
<td id="EB225DD9FFC3D7CCFDAFF97693FE8F8C" box="[604,682,1666,1688]" gridcol="1" gridrow="4" pageId="24" pageNumber="580">140.1</td>
<td id="EB225DD9FFC3D7CCFCCDF97692D88F8C" box="[830,908,1666,1688]" gridcol="2" gridrow="4" pageId="24" pageNumber="580">28.76</td>
<td id="EB225DD9FFC3D7CCFBD3F97695E08F8C" box="[1056,1204,1666,1688]" gridcol="3" gridrow="4" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF97694EE8F8C" box="[1352,1466,1666,1688]" gridcol="4" gridrow="4" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF95794EE8FAC" box="[159,1466,1699,1720]" gridrow="5" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF957909C8FAC" box="[159,456,1699,1720]" gridcol="0" gridrow="5" pageId="24" pageNumber="580">Marginal radius increase</th>
<td id="EB225DD9FFC3D7CCFDAFF95793FE8FAC" box="[604,682,1699,1720]" gridcol="1" gridrow="5" pageId="24" pageNumber="580">0.094</td>
<td id="EB225DD9FFC3D7CCFCCDF95792D88FAC" box="[830,908,1699,1720]" gridcol="2" gridrow="5" pageId="24" pageNumber="580">0.0195</td>
<td id="EB225DD9FFC3D7CCFBD3F95795E08FAC" box="[1056,1204,1699,1720]" gridcol="3" gridrow="5" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF95794EE8FAC" box="[1352,1466,1699,1720]" gridcol="4" gridrow="5" pageId="24" pageNumber="580">0</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF93694EE8FCC" box="[159,1466,1730,1752]" gridrow="6" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF936909C8FCC" box="[159,456,1730,1752]" gridcol="0" gridrow="6" pageId="24" pageNumber="580">Spiral chamber height increase</th>
<td id="EB225DD9FFC3D7CCFDAFF93693FE8FCC" box="[604,682,1730,1752]" gridcol="1" gridrow="6" pageId="24" pageNumber="580">1.7</td>
<td id="EB225DD9FFC3D7CCFCCDF93692D88FCC" box="[830,908,1730,1752]" gridcol="2" gridrow="6" pageId="24" pageNumber="580">0.07</td>
<td id="EB225DD9FFC3D7CCFBD3F93695E08FCC" box="[1056,1204,1730,1752]" gridcol="3" gridrow="6" pageId="24" pageNumber="580">――</td>
<td id="EB225DD9FFC3D7CCFABBF93694EE8FCC" box="[1352,1466,1730,1752]" gridcol="4" gridrow="6" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF91694EE8FEC" box="[159,1466,1762,1784]" gridrow="7" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF916909C8FEC" box="[159,456,1762,1784]" gridcol="0" gridrow="7" pageId="24" pageNumber="580">Initial spiral chamber height</th>
<td id="EB225DD9FFC3D7CCFDAFF91693FE8FEC" box="[604,682,1762,1784]" gridcol="1" gridrow="7" pageId="24" pageNumber="580">46.5</td>
<td id="EB225DD9FFC3D7CCFCCDF91692D88FEC" box="[830,908,1762,1784]" gridcol="2" gridrow="7" pageId="24" pageNumber="580">10.02</td>
<td id="EB225DD9FFC3D7CCFBD3F91695E08FEC" box="[1056,1204,1762,1784]" gridcol="3" gridrow="7" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF91694EE8FEC" box="[1352,1466,1762,1784]" gridcol="4" gridrow="7" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF8F694EE8E0C" box="[159,1466,1794,1816]" gridrow="8" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF8F6909C8E0C" box="[159,456,1794,1816]" gridcol="0" gridrow="8" pageId="24" pageNumber="580">Backbend angle</th>
<td id="EB225DD9FFC3D7CCFDAFF8F693FE8E0C" box="[604,682,1794,1816]" gridcol="1" gridrow="8" pageId="24" pageNumber="580">0.495</td>
<td id="EB225DD9FFC3D7CCFCCDF8F692D88E0C" box="[830,908,1794,1816]" gridcol="2" gridrow="8" pageId="24" pageNumber="580">0.1320</td>
<td id="EB225DD9FFC3D7CCFBD3F8F695E08E0C" box="[1056,1204,1794,1816]" gridcol="3" gridrow="8" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF8F694EE8E0C" box="[1352,1466,1794,1816]" gridcol="4" gridrow="8" pageId="24" pageNumber="580">+</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF8D694EE8E2C" box="[159,1466,1826,1848]" gridrow="9" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF8D6909C8E2C" box="[159,456,1826,1848]" gridcol="0" gridrow="9" pageId="24" pageNumber="580">Initial chamber length</th>
<td id="EB225DD9FFC3D7CCFDAFF8D693FE8E2C" box="[604,682,1826,1848]" gridcol="1" gridrow="9" pageId="24" pageNumber="580">139.0</td>
<td id="EB225DD9FFC3D7CCFCCDF8D692D88E2C" box="[830,908,1826,1848]" gridcol="2" gridrow="9" pageId="24" pageNumber="580">17.17</td>
<td id="EB225DD9FFC3D7CCFBD3F8D695E08E2C" box="[1056,1204,1826,1848]" gridcol="3" gridrow="9" pageId="24" pageNumber="580">――</td>
<td id="EB225DD9FFC3D7CCFABBF8D694EE8E2C" box="[1352,1466,1826,1848]" gridcol="4" gridrow="9" pageId="24" pageNumber="580">――</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF8B694EE8E43" box="[159,1466,1858,1879]" gridrow="10" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF8B6909C8E43" box="[159,456,1858,1879]" gridcol="0" gridrow="10" pageId="24" pageNumber="580">Chamber length increase</th>
<td id="EB225DD9FFC3D7CCFDAFF8B693FE8E43" box="[604,682,1858,1879]" gridcol="1" gridrow="10" pageId="24" pageNumber="580">0.028</td>
<td id="EB225DD9FFC3D7CCFCCDF8B692D88E43" box="[830,908,1858,1879]" gridcol="2" gridrow="10" pageId="24" pageNumber="580">0.0175</td>
<td id="EB225DD9FFC3D7CCFBD3F8B695E08E43" box="[1056,1204,1858,1879]" gridcol="3" gridrow="10" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF8B694EE8E43" box="[1352,1466,1858,1879]" gridcol="4" gridrow="10" pageId="24" pageNumber="580">0</td>
</tr>
<tr id="A8F334A5FFC3D7CCFF6CF89694EE8E63" box="[159,1466,1890,1911]" gridrow="11" pageId="24" pageNumber="580">
<th id="EB225DD9FFC3D7CCFF6CF896909C8E63" box="[159,456,1890,1911]" gridcol="0" gridrow="11" pageId="24" pageNumber="580">Perimeter ratio</th>
<td id="EB225DD9FFC3D7CCFDAFF89693FE8E63" box="[604,682,1890,1911]" gridcol="1" gridrow="11" pageId="24" pageNumber="580">1.277</td>
<td id="EB225DD9FFC3D7CCFCCDF89692D88E63" box="[830,908,1890,1911]" gridcol="2" gridrow="11" pageId="24" pageNumber="580">0.1076</td>
<td id="EB225DD9FFC3D7CCFBD3F89695E08E63" box="[1056,1204,1890,1911]" gridcol="3" gridrow="11" pageId="24" pageNumber="580">0</td>
<td id="EB225DD9FFC3D7CCFABBF89694EE8E63" box="[1352,1466,1890,1911]" gridcol="4" gridrow="11" pageId="24" pageNumber="580">0</td>
</tr>
</table>
</paragraph>
<caption id="42BC666FFFC2282AFF6CF99E95C08FDB" ID-DOI="http://doi.org/10.5281/zenodo.10883561" ID-Zenodo-Dep="10883561" httpUri="https://zenodo.org/record/10883561/files/figure.png" pageId="25" pageNumber="581" startId="25.[159,227,1642,1663]" targetBox="[173,1451,185,1609]" targetPageId="25" targetType="figure">
<paragraph id="167C36E7FFC2282AFF6CF99E95C08FDB" blockId="25.[159,1467,1642,1743]" pageId="25" pageNumber="581">
<emphasis id="24B7EAF5FFC2282AFF6CF99E901D8F6B" bold="true" box="[159,329,1642,1663]" pageId="25" pageNumber="581">Figure 17. AG,</emphasis>
<taxonomicName id="D1C34D64FFC2282AFEBCF99E92578F6B" authority="Vaughan &amp; Cole" authorityName="Vaughan &amp; Cole" authorityYear="1941" box="[335,771,1642,1663]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="soldadensis">
<emphasis id="24B7EAF5FFC2282AFEBCF99E930C8F6B" box="[335,600,1642,1663]" italics="true" pageId="25" pageNumber="581">Operculinoides soldadensis</emphasis>
Vaughan &amp; Cole
</taxonomicName>
;
<emphasis id="24B7EAF5FFC2282AFCFEF99E92708F6B" bold="true" box="[781,804,1642,1663]" pageId="25" pageNumber="581">A,</emphasis>
Loma El Santo, CA-215-871;
<emphasis id="24B7EAF5FFC2282AFBBFF99E95368F6B" bold="true" box="[1100,1122,1642,1663]" pageId="25" pageNumber="581">B,</emphasis>
Loma Candelaria, 98LC-1-669;
<emphasis id="24B7EAF5FFC2282AFA6EF99E91E08F8E" bold="true" pageId="25" pageNumber="581">C F,</emphasis>
Norona;
<emphasis id="24B7EAF5FFC2282AFEFCF97190728F8E" bold="true" box="[271,294,1669,1690]" pageId="25" pageNumber="581">C,</emphasis>
NOR-UN 24;
<emphasis id="24B7EAF5FFC2282AFE47F97190B28F8E" bold="true" box="[436,486,1669,1690]" pageId="25" pageNumber="581">DF,</emphasis>
NOR-UN 15/14;
<emphasis id="24B7EAF5FFC2282AFD61F97193FF8F8E" bold="true" box="[658,683,1669,1690]" pageId="25" pageNumber="581">G,</emphasis>
holotype, Trinidad.
<emphasis id="24B7EAF5FFC2282AFC83F97192CA8F8E" bold="true" box="[880,926,1669,1690]" pageId="25" pageNumber="581">H, I,</emphasis>
<taxonomicName id="D1C34D64FFC2282AFC57F97194718F8E" authority="(Nutall)" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[932,1317,1669,1690]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC2282AFC57F971959B8F8E" box="[932,1231,1669,1690]" italics="true" pageId="25" pageNumber="581">Palaeonummulites trinitatensis</emphasis>
(Nutall)
</taxonomicName>
;
<emphasis id="24B7EAF5FFC2282AFADCF971941C8F8E" bold="true" box="[1327,1352,1669,1690]" pageId="25" pageNumber="581">H,</emphasis>
Loma Candelaria, 98LC-1ICT3;
<emphasis id="24B7EAF5FFC2282AFE8BF96B90D38FA0" bold="true" box="[376,391,1695,1716]" pageId="25" pageNumber="581">I,</emphasis>
holotype of
<taxonomicName id="D1C34D64FFC2282AFDF7F96B92C58FA0" authority="Vaughan &amp; Cole" authorityName="Vaughan &amp; Cole" authorityYear="1941" box="[516,913,1695,1716]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="kugleri">
<emphasis id="24B7EAF5FFC2282AFDF7F96B93B78FA0" box="[516,739,1695,1716]" italics="true" pageId="25" pageNumber="581">Operculinoides kugleri</emphasis>
Vaughan &amp; Cole
</taxonomicName>
, Trinidad.
<emphasis id="24B7EAF5FFC2282AFC09F96B95588FA0" bold="true" box="[1018,1036,1695,1716]" pageId="25" pageNumber="581">J,</emphasis>
<taxonomicName id="D1C34D64FFC2282AFBE0F96B942C8FA0" authority="(Cushman)" baseAuthorityName="Cushman" box="[1043,1400,1695,1716]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="ocalanus">
<emphasis id="24B7EAF5FFC2282AFBE0F96B94508FA0" box="[1043,1284,1695,1716]" italics="true" pageId="25" pageNumber="581">Operculinoides ocalanus</emphasis>
(Cushman)
</taxonomicName>
, Loma Jabaco, CA-4-724.
<emphasis id="24B7EAF5FFC2282AFEAAF94E90908FDB" bold="true" box="[345,452,1722,1743]" pageId="25" pageNumber="581">AD, GJ,</emphasis>
A forms in equatorial section;
<emphasis id="24B7EAF5FFC2282AFD1EF94E92568FDB" bold="true" box="[749,770,1722,1743]" pageId="25" pageNumber="581">E,</emphasis>
A form in axial section;
<emphasis id="24B7EAF5FFC2282AFC02F94E95518FDB" bold="true" box="[1009,1029,1722,1743]" pageId="25" pageNumber="581">F,</emphasis>
external view.
</paragraph>
</caption>
<paragraph id="167C36E7FFC2282AFF6CF8D490B48E43" blockId="25.[159,781,1824,1879]" pageId="25" pageNumber="581">
<emphasis id="24B7EAF5FFC2282AFF6CF8D490788E23" bold="true" box="[159,300,1824,1847]" pageId="25" pageNumber="581">Occurrence.</emphasis>
Late middle Eocene to late Eocene NP 16/ 17, Loma Candela Formation.
</paragraph>
</subSubSection>
<subSubSection id="5ED9656CFFC22829FF6CF88393658DB4" lastPageId="26" lastPageNumber="582" pageId="25" pageNumber="581" type="discussion">
<paragraph id="167C36E7FFC22829FF6CF88390418D50" blockId="25.[159,781,1911,1966]" lastBlockId="26.[159,781,846,1092]" lastPageId="26" lastPageNumber="582" pageId="25" pageNumber="581">
<emphasis id="24B7EAF5FFC2282AFF6CF883905A8E9A" bold="true" box="[159,270,1911,1934]" pageId="25" pageNumber="581">Remarks.</emphasis>
<taxonomicName id="D1C34D64FFC2282AFEEFF88393278E9A" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[284,627,1911,1934]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="tinitantensis">
<emphasis id="24B7EAF5FFC2282AFEEFF88393278E9A" box="[284,627,1911,1934]" italics="true" pageId="25" pageNumber="581">Palaeonummulites tinitantensis</emphasis>
</taxonomicName>
is not abundant in the Eocene of western and central
<collectingCountry id="6ED47677FFC2282AFD7AF86393958EBA" box="[649,705,1943,1966]" name="Cuba" pageId="25" pageNumber="581">Cuba</collectingCountry>
but is sporadically present at the Loma Candelaria locality.
<bibRefCitation id="72524B16FFC2282AFA7BF8D492C08E43" author="Cole, W. S." pageId="25" pageNumber="581" pagination="28 - 39" refId="ref20256" refString="Cole, W. S. 1961. An analysis of certain taxonomic problems in the larger Foraminifera. Bulletins of American Paleontology, 43, 28 - 39." type="journal article" year="1961">Cole (1961)</bibRefCitation>
admitted that it is difficult to distinguish between
<taxonomicName id="D1C34D64FFC2282AFCBEF89492A58E63" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[845,1009,1888,1911]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC2282AFCBEF89492A58E63" box="[845,1009,1888,1911]" italics="true" pageId="25" pageNumber="581">P. trinitatensis</emphasis>
</taxonomicName>
and
<taxonomicName id="D1C34D64FFC2282AFBC7F89495FA8E63" baseAuthorityName="Cole" baseAuthorityYear="1941" box="[1076,1198,1888,1911]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="25" pageNumber="581" phylum="Foraminifera" rank="species" species="willcoxi">
<emphasis id="24B7EAF5FFC2282AFBC7F89495FA8E63" box="[1076,1198,1888,1911]" italics="true" pageId="25" pageNumber="581">P. willcoxi</emphasis>
</taxonomicName>
; the latter is the most widely recognized nummulitid in the Caribbean province and is absent at the studied localities. The lack of an easily recognizable
<typeStatus id="C9788845FFC12829FEC2FC9990DA8A90" box="[305,398,877,900]" pageId="26" pageNumber="582" type="holotype">holotype</typeStatus>
has led to many different morphotypes being described as
<taxonomicName id="D1C34D64FFC12829FE4EFC7993688AB0" box="[445,572,909,932]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="genus">
<emphasis id="24B7EAF5FFC12829FE4EFC7993688AB0" box="[445,572,909,932]" italics="true" pageId="26" pageNumber="582">Nummulites</emphasis>
</taxonomicName>
or
<taxonomicName id="D1C34D64FFC12829FD9AFC7991A08AD0" baseAuthorityName="Cole" baseAuthorityYear="1941" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="willcoxi">
<emphasis id="24B7EAF5FFC12829FD9AFC7991A08AD0" italics="true" pageId="26" pageNumber="582">Operculinoides willcoxi</emphasis>
</taxonomicName>
. We regard the specimen illustrated by
<bibRefCitation id="72524B16FFC12829FD37FC5991B28AF0" author="Barker, R. W." pageId="26" pageNumber="582" pagination="305 - 330" refId="ref18981" refString="Barker, R. W. 1939. Species of the foraminiferal family Camerinidae in the Tertiary and Cretaceous of Mexico. Proceedings of the United States National Museum, 86, 305 - 330." type="journal article" year="1939">Barker (1939)</bibRefCitation>
as the most similar to the original description, whereas specimens illustrated in
<bibRefCitation id="72524B16FFC12829FDEEFC1993F18D10" author="Cole, W. S." box="[541,677,1005,1028]" pageId="26" pageNumber="582" pagination="1 - 53" refId="ref20077" refString="Cole, W. S. 1941. Stratigraphic and paleontologic studies of wells in Florida. Florida Geological Survey Bulletin, 19, 1 - 53." type="journal article" year="1941">Cole (1941)</bibRefCitation>
conform more closely to the moderately tightly coiled
<taxonomicName id="D1C34D64FFC12829FD00FBF9905A8D50" baseAuthorityName="Heilprin" baseAuthorityYear="1885" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC12829FD00FBF9905A8D50" italics="true" pageId="26" pageNumber="582">O. floridensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="42BC666FFFC12829FF6CFF4E9469880B" pageId="26" pageNumber="582" startId="26.[159,217,186,207]" targetType="table">
<paragraph id="167C36E7FFC12829FF6CFF4E9469880B" blockId="26.[159,1467,186,287]" pageId="26" pageNumber="582">
<emphasis id="24B7EAF5FFC12829FF6CFF4E91A089DB" bold="true" box="[159,244,186,207]" pageId="26" pageNumber="582">Table 5.</emphasis>
Characters and attributes (means and standard deviations, SD, in mm) for
<taxonomicName id="D1C34D64FFC12829FC12FF4E945A89DB" baseAuthorityName="Nuttall" baseAuthorityYear="1928" box="[993,1294,186,207]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC12829FC12FF4E945A89DB" box="[993,1294,186,207]" italics="true" pageId="26" pageNumber="582">Palaeonummulites trinitatensis</emphasis>
</taxonomicName>
and comparisons with
<emphasis id="24B7EAF5FFC12829FF22FF2093AE89FD" box="[209,762,212,233]" italics="true" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FF22FF2090A589FD" box="[209,497,212,233]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="striatoreticulatus">Nummulites striatoreticulatus</taxonomicName>
,
<taxonomicName id="D1C34D64FFC12829FE0FFF2093AE89FD" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[508,762,212,233]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="floridensis">Operculinoides floridensis</taxonomicName>
</emphasis>
(tightly coiled),
<taxonomicName id="D1C34D64FFC12829FC6EFF20954B89FD" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[925,1055,212,233]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC12829FC6EFF20954B89FD" box="[925,1055,212,233]" italics="true" pageId="26" pageNumber="582">O. floridensis</emphasis>
</taxonomicName>
(loosely coiled) and
<emphasis id="24B7EAF5FFC12829FB1EFF2094D589FD" box="[1261,1409,212,233]" italics="true" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FB1EFF20942989FD" authorityName="Vaughan &amp; Cole" authorityYear="1941" box="[1261,1405,212,233]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="soldadensis">O. soldadensis</taxonomicName>
.
</emphasis>
Symbol key: ++, strong positive differences with &lt;1% error probability; +, differences with &lt;5% error probability; 0, no significant differences; -, negative differences with &lt;5% error probability; ――, strong negative differences with &lt;1% error probability.
</paragraph>
</caption>
<paragraph id="167C36E7FFC12829FF6CFEA294308BC8" pageId="26" pageNumber="582">
<table id="64C3C447FFC1D7CCFF6CFEC894EE8BC8" box="[159,1466,316,732]" gridcols="7" gridrows="12" pageId="26" pageNumber="582" rowsContinueIn="26.[159,1466,1521,1911]">
<tr id="A8F334A5FFC1D7CCFF6CFEC894EE887F" box="[159,1466,316,363]" gridrow="0" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFEC8909C887F" box="[159,456,316,363]" gridcol="0" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FF6CFEA29078887F" authority="Mean SD" authorityName="Mean SD" box="[159,300,342,363]" class="Globothalamea" family="Nummulitidae" genus="Palaeonummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="trinitatensis">
<emphasis id="24B7EAF5FFC12829FF6CFEA29078887F" box="[159,300,342,363]" italics="true" pageId="26" pageNumber="582">P. trinitatensis</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC1D7CCFE1DFEC8936F887F" box="[494,571,316,363]" gridcol="1" gridrow="0" pageId="26" pageNumber="582">Mean</th>
<th id="EB225DD9FFC1D7CCFD91FEC893FB887F" box="[610,687,316,363]" gridcol="2" gridrow="0" pageId="26" pageNumber="582">SD</th>
<th id="EB225DD9FFC1D7CCFD25FEC89230887F" box="[726,868,316,363]" gridcol="3" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FD2EFEC8920A8845" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[733,862,316,337]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC12829FD2EFEC8920A8845" box="[733,862,316,337]" italics="true" pageId="26" pageNumber="582">O. floridensis</emphasis>
</taxonomicName>
(tightly coiled)
</th>
<th id="EB225DD9FFC1D7CCFC78FEC89574887F" box="[907,1056,316,363]" gridcol="4" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FC65FEC895438845" baseAuthorityName="Heilprin" baseAuthorityYear="1885" box="[918,1047,316,337]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="floridensis">
<emphasis id="24B7EAF5FFC12829FC65FEC895438845" box="[918,1047,316,337]" italics="true" pageId="26" pageNumber="582">O. floridensis</emphasis>
</taxonomicName>
(loosely coiled)
</th>
<th id="EB225DD9FFC1D7CCFBB4FEC89580887F" box="[1095,1236,316,363]" gridcol="5" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FBB4FEA29580887F" authorityName="Vaughan &amp; Cole" authorityYear="1941" box="[1095,1236,342,363]" class="Globothalamea" family="Nummulitidae" genus="Operculinoides" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="soldadensis">
<emphasis id="24B7EAF5FFC12829FBB4FEA29580887F" box="[1095,1236,342,363]" italics="true" pageId="26" pageNumber="582">O. soldadensis</emphasis>
</taxonomicName>
</th>
<th id="EB225DD9FFC1D7CCFB09FEC894EE887F" box="[1274,1466,316,363]" gridcol="6" gridrow="0" pageId="26" pageNumber="582">
<taxonomicName id="D1C34D64FFC12829FB09FEA294EE887F" authority="First" authorityName="First" box="[1274,1466,342,363]" class="Globothalamea" family="Nummulitidae" genus="Nummulites" kingdom="Chromista" order="Rotaliida" pageId="26" pageNumber="582" phylum="Foraminifera" rank="species" species="striatoreticulatus">
<emphasis id="24B7EAF5FFC12829FB09FEA294EE887F" box="[1274,1466,342,363]" italics="true" pageId="26" pageNumber="582">N. striatoreticulatus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFE7394EE8889" box="[159,1466,391,413]" gridrow="1" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFE73909C8889" box="[159,456,391,413]" gridcol="0" gridrow="1" pageId="26" pageNumber="582">First chamber length</th>
<td id="EB225DD9FFC1D7CCFE1DFE73936F8889" box="[494,571,391,413]" gridcol="1" gridrow="1" pageId="26" pageNumber="582">218.2</td>
<td id="EB225DD9FFC1D7CCFD91FE7393FB8889" box="[610,687,391,413]" gridcol="2" gridrow="1" pageId="26" pageNumber="582">53.81</td>
<td id="EB225DD9FFC1D7CCFD25FE7392308889" box="[726,868,391,413]" gridcol="3" gridrow="1" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FE7395748889" box="[907,1056,391,413]" gridcol="4" gridrow="1" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FE7395808889" box="[1095,1236,391,413]" gridcol="5" gridrow="1" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FE7394EE8889" box="[1274,1466,391,413]" gridcol="6" gridrow="1" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFE5394EE88A9" box="[159,1466,423,445]" gridrow="2" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFE53909C88A9" box="[159,456,423,445]" gridcol="0" gridrow="2" pageId="26" pageNumber="582">Proloculus nominal diameter</th>
<td id="EB225DD9FFC1D7CCFE1DFE53936F88A9" box="[494,571,423,445]" gridcol="1" gridrow="2" pageId="26" pageNumber="582">99.0</td>
<td id="EB225DD9FFC1D7CCFD91FE5393FB88A9" box="[610,687,423,445]" gridcol="2" gridrow="2" pageId="26" pageNumber="582">28.91</td>
<td id="EB225DD9FFC1D7CCFD25FE53923088A9" box="[726,868,423,445]" gridcol="3" gridrow="2" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FE53957488A9" box="[907,1056,423,445]" gridcol="4" gridrow="2" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FE53958088A9" box="[1095,1236,423,445]" gridcol="5" gridrow="2" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FE5394EE88A9" box="[1274,1466,423,445]" gridcol="6" gridrow="2" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFE3394EE88C9" box="[159,1466,455,477]" gridrow="3" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFE33909C88C9" box="[159,456,455,477]" gridcol="0" gridrow="3" pageId="26" pageNumber="582">Deuteroloculus ratio</th>
<td id="EB225DD9FFC1D7CCFE1DFE33936F88C9" box="[494,571,455,477]" gridcol="1" gridrow="3" pageId="26" pageNumber="582">0.950</td>
<td id="EB225DD9FFC1D7CCFD91FE3393FB88C9" box="[610,687,455,477]" gridcol="2" gridrow="3" pageId="26" pageNumber="582">0.1376</td>
<td id="EB225DD9FFC1D7CCFD25FE33923088C9" box="[726,868,455,477]" gridcol="3" gridrow="3" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFC78FE33957488C9" box="[907,1056,455,477]" gridcol="4" gridrow="3" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FE33958088C9" box="[1095,1236,455,477]" gridcol="5" gridrow="3" pageId="26" pageNumber="582">-</td>
<td id="EB225DD9FFC1D7CCFB09FE3394EE88C9" box="[1274,1466,455,477]" gridcol="6" gridrow="3" pageId="26" pageNumber="582">0</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFE1394EE88E9" box="[159,1466,487,509]" gridrow="4" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFE13909C88E9" box="[159,456,487,509]" gridcol="0" gridrow="4" pageId="26" pageNumber="582">Initial marginal radius</th>
<td id="EB225DD9FFC1D7CCFE1DFE13936F88E9" box="[494,571,487,509]" gridcol="1" gridrow="4" pageId="26" pageNumber="582">126.8</td>
<td id="EB225DD9FFC1D7CCFD91FE1393FB88E9" box="[610,687,487,509]" gridcol="2" gridrow="4" pageId="26" pageNumber="582">32.48</td>
<td id="EB225DD9FFC1D7CCFD25FE13923088E9" box="[726,868,487,509]" gridcol="3" gridrow="4" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FE13957488E9" box="[907,1056,487,509]" gridcol="4" gridrow="4" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FE13958088E9" box="[1095,1236,487,509]" gridcol="5" gridrow="4" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FE1394EE88E9" box="[1274,1466,487,509]" gridcol="6" gridrow="4" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFDF394EE8B09" box="[159,1466,519,541]" gridrow="5" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFDF3909C8B09" box="[159,456,519,541]" gridcol="0" gridrow="5" pageId="26" pageNumber="582">Marginal radius increase</th>
<td id="EB225DD9FFC1D7CCFE1DFDF3936F8B09" box="[494,571,519,541]" gridcol="1" gridrow="5" pageId="26" pageNumber="582">0.099</td>
<td id="EB225DD9FFC1D7CCFD91FDF393FB8B09" box="[610,687,519,541]" gridcol="2" gridrow="5" pageId="26" pageNumber="582">0.0172</td>
<td id="EB225DD9FFC1D7CCFD25FDF392308B09" box="[726,868,519,541]" gridcol="3" gridrow="5" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFC78FDF395748B09" box="[907,1056,519,541]" gridcol="4" gridrow="5" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FDF395808B09" box="[1095,1236,519,541]" gridcol="5" gridrow="5" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFB09FDF394EE8B09" box="[1274,1466,519,541]" gridcol="6" gridrow="5" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFDD394EE8B29" box="[159,1466,551,573]" gridrow="6" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFDD3909C8B29" box="[159,456,551,573]" gridcol="0" gridrow="6" pageId="26" pageNumber="582">Spiral chamber height increase</th>
<td id="EB225DD9FFC1D7CCFE1DFDD3936F8B29" box="[494,571,551,573]" gridcol="1" gridrow="6" pageId="26" pageNumber="582">1.7</td>
<td id="EB225DD9FFC1D7CCFD91FDD393FB8B29" box="[610,687,551,573]" gridcol="2" gridrow="6" pageId="26" pageNumber="582">0.30</td>
<td id="EB225DD9FFC1D7CCFD25FDD392308B29" box="[726,868,551,573]" gridcol="3" gridrow="6" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FDD395748B29" box="[907,1056,551,573]" gridcol="4" gridrow="6" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFBB4FDD395808B29" box="[1095,1236,551,573]" gridcol="5" gridrow="6" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FDD394EE8B29" box="[1274,1466,551,573]" gridcol="6" gridrow="6" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFDB294EE8B48" box="[159,1466,582,604]" gridrow="7" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFDB2909C8B48" box="[159,456,582,604]" gridcol="0" gridrow="7" pageId="26" pageNumber="582">Initial spiral chamber height</th>
<td id="EB225DD9FFC1D7CCFE1DFDB2936F8B48" box="[494,571,582,604]" gridcol="1" gridrow="7" pageId="26" pageNumber="582">35.5</td>
<td id="EB225DD9FFC1D7CCFD91FDB293FB8B48" box="[610,687,582,604]" gridcol="2" gridrow="7" pageId="26" pageNumber="582">17.71</td>
<td id="EB225DD9FFC1D7CCFD25FDB292308B48" box="[726,868,582,604]" gridcol="3" gridrow="7" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FDB295748B48" box="[907,1056,582,604]" gridcol="4" gridrow="7" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FDB295808B48" box="[1095,1236,582,604]" gridcol="5" gridrow="7" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FDB294EE8B48" box="[1274,1466,582,604]" gridcol="6" gridrow="7" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFD9294EE8B68" box="[159,1466,614,636]" gridrow="8" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFD92909C8B68" box="[159,456,614,636]" gridcol="0" gridrow="8" pageId="26" pageNumber="582">Backbend angle</th>
<td id="EB225DD9FFC1D7CCFE1DFD92936F8B68" box="[494,571,614,636]" gridcol="1" gridrow="8" pageId="26" pageNumber="582">0.445</td>
<td id="EB225DD9FFC1D7CCFD91FD9293FB8B68" box="[610,687,614,636]" gridcol="2" gridrow="8" pageId="26" pageNumber="582">0.1165</td>
<td id="EB225DD9FFC1D7CCFD25FD9292308B68" box="[726,868,614,636]" gridcol="3" gridrow="8" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFC78FD9295748B68" box="[907,1056,614,636]" gridcol="4" gridrow="8" pageId="26" pageNumber="582">-</td>
<td id="EB225DD9FFC1D7CCFBB4FD9295808B68" box="[1095,1236,614,636]" gridcol="5" gridrow="8" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFB09FD9294EE8B68" box="[1274,1466,614,636]" gridcol="6" gridrow="8" pageId="26" pageNumber="582">++</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFD7294EE8B88" box="[159,1466,646,668]" gridrow="9" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFD72909C8B88" box="[159,456,646,668]" gridcol="0" gridrow="9" pageId="26" pageNumber="582">Initial chamber length</th>
<td id="EB225DD9FFC1D7CCFE1DFD72936F8B88" box="[494,571,646,668]" gridcol="1" gridrow="9" pageId="26" pageNumber="582">151.9</td>
<td id="EB225DD9FFC1D7CCFD91FD7293FB8B88" box="[610,687,646,668]" gridcol="2" gridrow="9" pageId="26" pageNumber="582">26.76</td>
<td id="EB225DD9FFC1D7CCFD25FD7292308B88" box="[726,868,646,668]" gridcol="3" gridrow="9" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFC78FD7295748B88" box="[907,1056,646,668]" gridcol="4" gridrow="9" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFBB4FD7295808B88" box="[1095,1236,646,668]" gridcol="5" gridrow="9" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFB09FD7294EE8B88" box="[1274,1466,646,668]" gridcol="6" gridrow="9" pageId="26" pageNumber="582">――</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFD5294EE8BA8" box="[159,1466,678,700]" gridrow="10" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFD52909C8BA8" box="[159,456,678,700]" gridcol="0" gridrow="10" pageId="26" pageNumber="582">Chamber length increase</th>
<td id="EB225DD9FFC1D7CCFE1DFD52936F8BA8" box="[494,571,678,700]" gridcol="1" gridrow="10" pageId="26" pageNumber="582">0.025</td>
<td id="EB225DD9FFC1D7CCFD91FD5293FB8BA8" box="[610,687,678,700]" gridcol="2" gridrow="10" pageId="26" pageNumber="582">0.0149</td>
<td id="EB225DD9FFC1D7CCFD25FD5292308BA8" box="[726,868,678,700]" gridcol="3" gridrow="10" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFC78FD5295748BA8" box="[907,1056,678,700]" gridcol="4" gridrow="10" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFBB4FD5295808BA8" box="[1095,1236,678,700]" gridcol="5" gridrow="10" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFB09FD5294EE8BA8" box="[1274,1466,678,700]" gridcol="6" gridrow="10" pageId="26" pageNumber="582">0</td>
</tr>
<tr id="A8F334A5FFC1D7CCFF6CFD3294EE8BC8" box="[159,1466,710,732]" gridrow="11" pageId="26" pageNumber="582">
<th id="EB225DD9FFC1D7CCFF6CFD32909C8BC8" box="[159,456,710,732]" gridcol="0" gridrow="11" pageId="26" pageNumber="582">Perimeter ratio</th>
<td id="EB225DD9FFC1D7CCFE1DFD32936F8BC8" box="[494,571,710,732]" gridcol="1" gridrow="11" pageId="26" pageNumber="582">1.263</td>
<td id="EB225DD9FFC1D7CCFD91FD3293FB8BC8" box="[610,687,710,732]" gridcol="2" gridrow="11" pageId="26" pageNumber="582">0.0859</td>
<td id="EB225DD9FFC1D7CCFD25FD3292308BC8" box="[726,868,710,732]" gridcol="3" gridrow="11" pageId="26" pageNumber="582">0</td>
<td id="EB225DD9FFC1D7CCFC78FD3295748BC8" box="[907,1056,710,732]" gridcol="4" gridrow="11" pageId="26" pageNumber="582">-</td>
<td id="EB225DD9FFC1D7CCFBB4FD3295808BC8" box="[1095,1236,710,732]" gridcol="5" gridrow="11" pageId="26" pageNumber="582">――</td>
<td id="EB225DD9FFC1D7CCFB09FD3294EE8BC8" box="[1274,1466,710,732]" gridcol="6" gridrow="11" pageId="26" pageNumber="582">+</td>
</tr>
</table>
</paragraph>
<paragraph id="167C36E7FFC12829FF6CFB9D93658DB4" blockId="26.[159,781,1129,1184]" pageId="26" pageNumber="582">
<emphasis id="24B7EAF5FFC12829FF6CFB9D939B8D94" bold="true" box="[159,719,1129,1152]" pageId="26" pageNumber="582">Stratigraphical and geographical distribution.</emphasis>
Late Eocene (Priabonian);
<collectingCountry id="6ED47677FFC12829FE7BFB7D90978DB4" box="[392,451,1161,1184]" name="Cuba" pageId="26" pageNumber="582">Cuba</collectingCountry>
,
<collectingCountry id="6ED47677FFC12829FE3EFB7D93798DB4" box="[461,557,1161,1184]" name="Trinidad and Tobago" pageId="26" pageNumber="582">Trinidad</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>