<documentid="6DD52B840D6A243E2F068312D27A3D3D"ID-DOI="10.11646/zootaxa.4718.4.6"ID-GBIF-Dataset="df425f1f-fbf0-4996-b51b-07cb9cb136c9"ID-ISSN="1175-5326"ID-Zenodo-Dep="3602714"ID-ZooBank="E315343C-3418-4F43-BB85-61C0E8AACC81"IM.bibliography_approvedBy="julia"IM.materialsCitations_approvedBy="julia"IM.metadata_approvedBy="julia"IM.tables_approvedBy="julia"IM.taxonomicNames_approvedBy="julia"IM.treatments_approvedBy="julia"checkinTime="1578565593362"checkinUser="plazi"docAuthor="Bahder, Brian W., Bartlett, Charles R., Helmick, Ericka E., Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Goss, Erica M. & Ascunce, Marina S."docDate="2020"docId="03FE2B29FF9FFF9BFF53B43221D0FBFE"docLanguage="en"docName="zootaxa.4718.4.6.pdf"docOrigin="Zootaxa 4718 (4)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Agoo dahliana Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce 2020, sp. n."docType="treatment"docVersion="9"lastPageNumber="531"masterDocId="FFC75351FF97FF91FFC4B3362007FC4F"masterDocTitle="Revised status of Omolicna subgenus Agoo (Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) with a new species from Costa Rica and new country records"masterLastPageNumber="535"masterPageNumber="521"pageNumber="529"updateTime="1729260785061"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="C3634C9342C200A0453E42E33278F036">Revised status of Omolicna subgenus Agoo (Hemiptera: Auchenorrhyncha: Fulgoroidea: Derbidae) with a new species from Costa Rica and new country records</mods:title>
<taxonomicNameid="4C57E1BCFF9FFF99FF53B4322231FB51"ID-CoL="5TSKK"authority="Bahder & Bartlett"authorityName="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce"authorityYear="2020"box="[151,566,1796,1823]"class="Insecta"family="Derbidae"genus="Agoo"kingdom="Animalia"order="Hemiptera"pageId="8"pageNumber="529"phylum="Arthropoda"rank="species"species="dahliana"status="sp. nov.">
<figureCitationid="136C86BAFF9FFF99FF5BB41E2133FB0D"box="[159,308,1832,1858]"captionStart-0="FIGURE 7"captionStart-1="FIGURE 8"captionStart-2="FIGURE 9"captionStart-3="FIGURE 10"captionStart-4="FIGURE 11"captionStartId-0="9.[151,250,1567,1592]"captionStartId-1="10.[151,250,972,997]"captionStartId-2="11.[151,250,914,939]"captionStartId-3="11.[151,250,2005,2030]"captionStartId-4="12.[151,250,614,639]"captionTargetBox-0="[239,1347,625,1539]"captionTargetBox-1="[155,1429,413,946]"captionTargetBox-2="[244,1344,174,889]"captionTargetBox-3="[255,1334,952,1982]"captionTargetBox-4="[154,1429,179,590]"captionTargetId-0="figure@9.[231,1357,616,1547]"captionTargetId-1="figure@10.[151,1436,409,952]"captionTargetId-2="figure@11.[239,1349,174,895]"captionTargetId-3="figure@11.[253,1334,951,1983]"captionTargetId-4="figure@12.[151,1436,174,595]"captionTargetPageId-0="9"captionTargetPageId-1="10"captionTargetPageId-2="11"captionTargetPageId-3="11"captionTargetPageId-4="12"captionText-0="FIGURE 7. Adult male habitus Agoo dahliana sp. n.; A. body lateral view and B. body dorsal view (B), scale = 1 mm."captionText-1="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."captionText-2="FIGURE 9. Agoo dahliana sp. n., forewing venation."captionText-3="FIGURE 10. Male terminalia of Agoo dahliana sp. n.; A. lateral view, B. ventral view, and C. dorsal view."captionText-4="FIGURE 11. Aedeagus of adult male Agoo dahliana sp. n.; A. right lateral view, B. left lateral view, and C. dorsal view."figureDoi-0="http://doi.org/10.5281/zenodo.3602728"figureDoi-1="http://doi.org/10.5281/zenodo.3602730"figureDoi-2="http://doi.org/10.5281/zenodo.3602732"figureDoi-3="http://doi.org/10.5281/zenodo.3602734"figureDoi-4="http://doi.org/10.5281/zenodo.3602736"httpUri-0="https://zenodo.org/record/3602728/files/figure.png"httpUri-1="https://zenodo.org/record/3602730/files/figure.png"httpUri-2="https://zenodo.org/record/3602732/files/figure.png"httpUri-3="https://zenodo.org/record/3602734/files/figure.png"httpUri-4="https://zenodo.org/record/3602736/files/figure.png"pageId="8"pageNumber="529">Figures 7–11</figureCitation>
Body overall testaceous with darker areas. Orange patches on dorsal portion of clypeus and genae. Dark patch on venter of abdomen and fuscous bands on pronotal foveae. Forewings with many dark spots in cells. Male pygofer with median ventral process wider than long; broad near base, attenuating distally to broadly rounded apex (lateral teeth lacking). Parameres, in lateral view, possess a single, large ventral lobe; dorsal surface bearing large lobe with two weakly sclerotized processes; basal process angled posteriorly; in ventral view parameres possess large, hooked lobes with acute sclerotized apices; paramere apices rounded with distinct, anterior facing, sclerotized spine. Aedeagal apex with two elongate processes on each side, one pair at apex, one pair subapical, in addition to complex endosoma bearing additional processes. Segment X (= “anal tube”) in lateral view robust, caudal portion stout, and ventrally sinuate, apex acute and moderately downcurved; in dorsal view deeply notched in caudal aspect.
<figureCitationid="136C86BAFF9EFF98FC9EB2B92399FDE5"box="[858,926,399,426]"captionStart="FIGURE 7"captionStartId="9.[151,250,1567,1592]"captionTargetBox="[239,1347,625,1539]"captionTargetId="figure@9.[231,1357,616,1547]"captionTargetPageId="9"captionText="FIGURE 7. Adult male habitus Agoo dahliana sp. n.; A. body lateral view and B. body dorsal view (B), scale = 1 mm."figureDoi="http://doi.org/10.5281/zenodo.3602728"httpUri="https://zenodo.org/record/3602728/files/figure.png"pageId="9"pageNumber="530">Fig. 7</figureCitation>
); face deep orange on dorsal third of clypeus, frons above frontoclypeal suture, and genae below antennae (
<figureCitationid="136C86BAFF9EFF98FC81B2842399FD82"box="[837,926,434,461]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="9"pageNumber="530">Fig. 8A</figureCitation>
); lateral carinae of front and vertex darkened. Paranotal folia with dark spots on inner surface. Mesonotum diffusely infuscate with orange, medially pale (giving appearance of weakly contrasted median vitta); lateral-most portion of mesonotum and tegulae pale. Forewings faintly yellowish, veins white (distally) to clouded (basally); distal cells with diffuse dark spots. Basal cells also possessing orange shading. Dorsum of abdomen orangish; aedeagus appearing dark brown within paler terminalia.
without wings. Head. In lateral view, anterior margin of head smoothly rounded (
<figureCitationid="136C86BAFF9EFF98FEC5B593215EFAF0"box="[257,345,1701,1727]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="9"pageNumber="530">Fig. 8C</figureCitation>
<figureCitationid="136C86BAFF9EFF98FC74B5932400FAF0"box="[944,1031,1701,1727]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="9"pageNumber="530">Fig. 8B</figureCitation>
), broadest near base, tapering distally; lateral margins decidedly keeled, bearing two (somewhat irregular) rows of sensorial pits, disc depressed, without distinct carinae. Transverse apical carina separating vertex from frons absent. Vertex length males:
. Frons with lateral carinae strongly keeled, narrowest between compound eyes, diverging slightly ventrad until reaching the frontoclypeal suture (
<figureCitationid="136C86BAFF9EFF98FAC5B463255DFB20"box="[1281,1370,1877,1903]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="9"pageNumber="530">Fig. 8A</figureCitation>
); sensorial pits next to each carina for entire length, carinae absent on disc of frons. Frons length males:
<taxonomicNameid="4C57E1BCFF9EFF98FE24B5162271FA77"authority="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce, 2020"authorityName="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce"authorityYear="2020"box="[480,630,1568,1592]"class="Insecta"family="Derbidae"genus="Agoo"kingdom="Animalia"order="Hemiptera"pageId="9"pageNumber="530"phylum="Arthropoda"rank="species"species="dahliana"status="sp. nov.">
Thorax. Pronotum short, anterior margin following contours of posterior margin of head (
<figureCitationid="136C86BAFF9DFF9BFB16B3AF252BFCFC"box="[1234,1324,153,179]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="10"pageNumber="531">Fig. 8B</figureCitation>
<figureCitationid="136C86BAFF9DFF9BFB87B3D72495FCB4"box="[1091,1170,225,251]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="10"pageNumber="531">Figs. 8</figureCitation>
. Mesonotum appearing slightly elevated in lateral view (
<figureCitationid="136C86BAFF9DFF9BFF5BB21F2132FD0C"box="[159,309,297,323]"captionStart="FIGURE 8"captionStartId="10.[151,250,972,997]"captionTargetBox="[155,1429,413,946]"captionTargetId="figure@10.[151,1436,409,952]"captionTargetPageId="10"captionText="FIGURE 8. Adult male Agoo dahliana sp. n.; A. head frontal view, B. head, pronotum and mesonotum dorsal view, C. head, pronotum and mesonotum lateral view, scale=1mm."figureDoi="http://doi.org/10.5281/zenodo.3602730"httpUri="https://zenodo.org/record/3602730/files/figure.png"pageId="10"pageNumber="531">Figs. 8B, 8C</figureCitation>
); in dorsal view, with three subparallel longitudinal carinae, indistinctly reaching posterior margin. Mesonotum length at midline males:
<taxonomicNameid="4C57E1BCFF9DFF9BFE50B0FB222CFFAA"authority="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce, 2020"authorityName="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce"authorityYear="2020"box="[404,555,973,997]"class="Insecta"family="Derbidae"genus="Agoo"kingdom="Animalia"order="Hemiptera"pageId="10"pageNumber="531"phylum="Arthropoda"rank="species"species="dahliana"status="sp. nov.">
of ScP+R (nearly to fork of ScP) and third row along basal half of postcubitus. Forks of R and CuA veins approximately at the same level, both well proximad of claval apex. Claval apex just beyond midpoint of wing, fork of MP just distad of claval apex; RA 1-branched, RP 1-branched, MP 4-branched and CuA 3-branched. Junction of Pcu+A
Terminalia. Pygofer, in lateral view, narrow, distal and proximal margins sinuate (
<figureCitationid="136C86BAFF9DFF9BFB96B7DD24BEF94A"box="[1106,1209,1259,1285]"captionStart="FIGURE 10"captionStartId="11.[151,250,2005,2030]"captionTargetBox="[255,1334,952,1982]"captionTargetId="figure@11.[253,1334,951,1983]"captionTargetPageId="11"captionText="FIGURE 10. Male terminalia of Agoo dahliana sp. n.; A. lateral view, B. ventral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602734"httpUri="https://zenodo.org/record/3602734/files/figure.png"pageId="10"pageNumber="531">Fig. 10A</figureCitation>
), broadest ventrally; in ventral view, ventral opening of pygofer bearing rounded lobe (
<figureCitationid="136C86BAFF9DFF9BFCBEB63923D8F966"box="[890,991,1295,1321]"captionStart="FIGURE 10"captionStartId="11.[151,250,2005,2030]"captionTargetBox="[255,1334,952,1982]"captionTargetId="figure@11.[253,1334,951,1983]"captionTargetPageId="11"captionText="FIGURE 10. Male terminalia of Agoo dahliana sp. n.; A. lateral view, B. ventral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602734"httpUri="https://zenodo.org/record/3602734/files/figure.png"pageId="10"pageNumber="531">Fig. 10B</figureCitation>
), widest at base and attenuating distally to rounded apex, lobe at base wider than long. Parameres, in ventral view, broadest ventrally with margins running parallel until expanding into a large, rounded tooth at midlength, strongly hooked with sclerotized apex pointing anteriorly; portion distal to hooked tooth of uniform width, narrower than basal portion until expanding slightly at terminus which possesses an anteriorly facing sclerotized spine (
<figureCitationid="136C86BAFF9DFF9BFCB2B6A923FCF9F6"box="[886,1019,1439,1465]"captionStart="FIGURE 10"captionStartId="11.[151,250,2005,2030]"captionTargetBox="[255,1334,952,1982]"captionTargetId="figure@11.[253,1334,951,1983]"captionTargetPageId="11"captionText="FIGURE 10. Male terminalia of Agoo dahliana sp. n.; A. lateral view, B. ventral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602734"httpUri="https://zenodo.org/record/3602734/files/figure.png"pageId="10"pageNumber="531">Figure 10B</figureCitation>
); in lateral view possessing a single, large ventral lobe; dorsal surface bearing large lobe with an invagination on posterior aspect, resulting in two lightly sclerotized processes, with posterior process pointing dorsally and anterior process angled posteriorly. Aedeagus with four elongate processes, one shorter pair at apex, one large pair subapical, all angled ventrad and pointing anteriorly. Complex endosoma with large retrorse median process, medium sized pair of retrorse lateral processes, and dorsal pair of posteriorly facing processes (
<figureCitationid="136C86BAFF9DFF9BFDB7B56522C4FA22"box="[627,707,1619,1645]"captionStart="FIGURE 11"captionStartId="12.[151,250,614,639]"captionTargetBox="[154,1429,179,590]"captionTargetId="figure@12.[151,1436,174,595]"captionTargetPageId="12"captionText="FIGURE 11. Aedeagus of adult male Agoo dahliana sp. n.; A. right lateral view, B. left lateral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602736"httpUri="https://zenodo.org/record/3602736/files/figure.png"pageId="10"pageNumber="531">Fig. 11</figureCitation>
). Endosoma complex, retrorse, bearing additional projections and asymmetrical 3-dimensional structures, but appearing to have two lateral processes, one medial process and a pair of dorsal processes (
<figureCitationid="136C86BAFF9DFF9BFEB9B5AD21CEFAF9"box="[381,457,1691,1718]"captionStart="FIGURE 11"captionStartId="12.[151,250,614,639]"captionTargetBox="[154,1429,179,590]"captionTargetId="figure@12.[151,1436,174,595]"captionTargetPageId="12"captionText="FIGURE 11. Aedeagus of adult male Agoo dahliana sp. n.; A. right lateral view, B. left lateral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602736"httpUri="https://zenodo.org/record/3602736/files/figure.png"pageId="10"pageNumber="531">Fig.11</figureCitation>
). Segment X (= “anal tube”), in lateral view, robust, caudal portion stout, and ventrally sinuate, apex acute and moderately downcurved; in dorsal view, deeply notched on caudal aspect (
<figureCitationid="136C86BAFF9DFF9BFB20B589253FFA96"box="[1252,1336,1727,1753]"captionStart="FIGURE 10"captionStartId="11.[151,250,2005,2030]"captionTargetBox="[255,1334,952,1982]"captionTargetId="figure@11.[253,1334,951,1983]"captionTargetPageId="11"captionText="FIGURE 10. Male terminalia of Agoo dahliana sp. n.; A. lateral view, B. ventral view, and C. dorsal view."figureDoi="http://doi.org/10.5281/zenodo.3602734"httpUri="https://zenodo.org/record/3602734/files/figure.png"pageId="10"pageNumber="531">Fig. 10</figureCitation>
<taxonomicNameid="4C57E1BCFF9DFF9BFDD0B5D22397FAB1"authority="H. F. Loomis"authorityName="H. F. Loomis"box="[532,912,1763,1790]"class="Liliopsida"family="Arecaceae"genus="Astrocaryum"kingdom="Plantae"order="Arecales"pageId="10"pageNumber="531"phylum="Tracheophyta"rank="species"species="alatum">
<taxonomicNameid="4C57E1BCFF9DFF9BFD78B4422364FBC2"authorityName="Bahder & Bartlett & Helmick & Barrantes Barrantes & Zumbado Echavarria & Goss & Ascunce"authorityYear="2020"box="[700,867,1907,1934]"class="Insecta"family="Derbidae"genus="Omolicna"kingdom="Animalia"order="Hemiptera"pageId="10"pageNumber="522"phylum="Arthropoda"rank="subGenus"subGenus="Agoo">
