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<document id="3A59A6B73C9F3BDA7EDB438A3078E921" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105362570" checkinUser="felipe" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel-Angel" docDate="2005" docId="03C487D0C451FFB5174FFD1620F1A714" docLanguage="en" docName="zt01012p037.pdf" docOrigin="Zootaxa 1012 (1012)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrodactylus montanus Arce-Perez and Moron 2000" docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="FFFDFFA8C45DFFB81647FFEA2418A64A" masterDocTitle="New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua" masterLastPageNumber="37" masterPageNumber="23" pageNumber="35" updateTime="1698980141271" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7707C27D103123C1E52E05C10336B3A7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="46B300364BC9A022738787D51ABD576E">
<mods:title id="6E04AF003D6D9D177A6036B960205496">New species and new records of species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua</mods:title>
</mods:titleInfo>
<mods:name id="78114013AC2019FE2CD32FD3D51FE5C0" type="personal">
<mods:role id="27AF9A0C566D5AC93DDE872BA365C3A9">
<mods:roleTerm id="7AC8DE48702E9C96FB04C442562E08B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F343A13E346F20C227BC97C61945044">Arce-Pérez, Roberto</mods:namePart>
</mods:name>
<mods:name id="5FEF13AF30012C648E4E343488003181" type="personal">
<mods:role id="A0B52173CB5DBE18D11625E383718231">
<mods:roleTerm id="F53ABF85C27754A30C485101959C61A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0A726EF3BC27ED29B2616714713E8C5">Morón, Miguel-Angel</mods:namePart>
</mods:name>
<mods:typeOfResource id="110BFF1727E61580A8D67AACAAFC7FD6">text</mods:typeOfResource>
<mods:relatedItem id="99DA20D0155944CA6B3CBC04F2064793" type="host">
<mods:titleInfo id="47BD4895A30F041387805DA0F49CB6AD">
<mods:title id="B73EE3A57FDE836A18778D18A18D899B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BF1C1A033D6A9517EA45E2D3EBA76696">
<mods:date id="4E51B5A87EBA6278C1F29F7FE5F523FF">2005</mods:date>
<mods:detail id="15108D5233C6CB6AF2DAB3543BAF7397" type="pubDate">
<mods:number id="2F4CA7CBA4A8A7D9171A0DFB15F7D13E">2005-06-28</mods:number>
</mods:detail>
<mods:detail id="8E86629B0837C3507B68F037CF00F831" type="volume">
<mods:number id="0FFF85654336D28EB2063ABF8735CDF0">1012</mods:number>
</mods:detail>
<mods:detail id="038151813665D378CBB1FFE0CE81A641" type="issue">
<mods:number id="665F4B82B3164001E8B9EFCF18C60009">1012</mods:number>
</mods:detail>
<mods:extent id="3D65B2B5C71E3CC92FB291C2A22D5EE0" unit="page">
<mods:start id="47CC58DC923D7495BA5B9E4ABCD9B19F">23</mods:start>
<mods:end id="BC78D29D404DA1EDB4B539703F21CA4E">37</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E20AC77314E5E06EBD673D94EE6AC831">journal article</mods:classification>
<mods:identifier id="19927E1EAE36BFA791DC421D5036F8D9" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="62EAA0CDC64FB98E33CE8B65400BA176" type="ZooBank">E84D2D81-6B26-4311-B56F-6D3C4E672C78</mods:identifier>
</mods:mods>
<treatment id="03C487D0C451FFB5174FFD1620F1A714" ID-DOI="http://doi.org/10.5281/zenodo.5049204" ID-Zenodo-Dep="5049204" LSID="urn:lsid:plazi:treatment:03C487D0C451FFB5174FFD1620F1A714" httpUri="http://treatment.plazi.org/id/03C487D0C451FFB5174FFD1620F1A714" lastPageId="13" lastPageNumber="35" pageId="12" pageNumber="35">
<subSubSection id="C377654DC451FFB4174FFD1627B2A55C" box="[264,938,764,790]" pageId="12" pageNumber="35" type="nomenclature">
<paragraph id="8BD236C6C451FFB4174FFD1627B2A55C" blockId="12.[264,938,764,790]" box="[264,938,764,790]" pageId="12" pageNumber="35">
<heading id="D09A81AAC451FFB4174FFD1627B2A55C" bold="true" box="[264,938,764,790]" fontSize="11" level="1" pageId="12" pageNumber="35" reason="1">
<taxonomicName id="4C6D4D45C451FFB4174FFD1627B2A55C" authority="Arce-Perez and Moron, 2000" authorityName="Arce-Perez and Moron" authorityYear="2000" box="[264,938,764,790]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B919EAD4C451FFB4174FFD1627B2A55C" bold="true" box="[264,938,764,790]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4174FFD162623A55C" bold="true" box="[264,571,764,790]" italics="true" pageId="12" pageNumber="35">Macrodactylus montanus</emphasis>
Arce­Pérez and
<bibRefCitation id="EFFC4B37C451FFB4154CFD1627B2A55C" author="Moron, M. A." box="[779,938,764,790]" pageId="12" pageNumber="35" pagination="123 - 239" refId="ref6864" refString="Arce-Perez R. &amp; Moron, M. A. (2000) Taxonomia y distribucion de las especies de Macrodactylus Latreille (Coleoptera: Melolonthidae) de Mexico y America del Norte. Acta Zoologica Mexicana (n. s), 79, 123 - 239." type="journal article" year="2000">Morón, 2000</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4174FFCAC27D6A263" pageId="12" pageNumber="35" type="diagnosis">
<paragraph id="8BD236C6C451FFB4174FFCAC27D6A263" blockId="12.[264,1325,838,1905]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4174FFCAC2598A52A" bold="true" box="[264,384,838,864]" pageId="12" pageNumber="35">Diagnosis</emphasis>
. This species is distinguished by the following combination of characters: antennal club reddish­brown with the apex darkened; pronotum black with scattered setae on the borders; elytra orange­yellow with sericeous shine (
<figureCitation id="13562A43C451FFB4159CFC7D203FA5FB" box="[987,1063,919,945]" captionStart="FIGURES 7–12" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 7–12. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="12" pageNumber="35">Fig. 9</figureCitation>
); pygidium glabrous, orange­yellow; abdomen black; femora and tibiae reddish­brown; tarsi nearly black with scattered white setae; parameres with apex lanceolate and setae on the external borders; female with scattered erect setae on the pronotum (
<figureCitation id="13562A43C451FFB41519FBE527D9A263" box="[862,961,1039,1065]" captionStart="FIGURES 7–12" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 7–12. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="12" pageNumber="35">Figs. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4177FFBDC2065A353" pageId="12" pageNumber="35" type="materials_examined">
<paragraph id="8BD236C6C451FFB4177FFBDC2065A353" blockId="12.[264,1325,838,1905]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFBDC263BA21A" bold="true" box="[312,547,1078,1104]" pageId="12" pageNumber="35">Material examined</emphasis>
(
<specimenCount id="9D6BFD4FC451FFB41474FBDD2686A21B" box="[563,670,1079,1105]" pageId="12" pageNumber="35" type="male">11 males</specimenCount>
,
<specimenCount id="9D6BFD4FC451FFB414EEFBDD2736A21B" box="[681,814,1079,1105]" pageId="12" pageNumber="35" type="female">16 females</specimenCount>
).
<materialsCitation id="3B053C9BC451FFB41507FBDD27CAA2EA" collectingDate="1996-05-30" collectionCode="IEXA, NEW, COUNTRY, RECORDS" collectorName="Col. O. Gomez" country="Mexico" elevation="2250" location="Reserva Huitepec" municipality="San Cristobal de Las Casas" pageId="12" pageNumber="35" specimenCount="2" stateProvince="Chiapas" typeStatus="holotype">
<collectingCountry id="F37A7656C451FFB41507FBDD27A0A21B" box="[832,952,1079,1105]" name="Mexico" pageId="12" pageNumber="35">MEXICO</collectingCountry>
.
<collectingRegion id="49A9F824C451FFB41585FBDD2047A21B" box="[962,1119,1079,1105]" country="Mexico" name="Chiapas" pageId="12" pageNumber="35">Chiapas state</collectingRegion>
,
<collectingMunicipality id="6BB6ACBCC451FFB4122CFBDD2598A233" pageId="12" pageNumber="35">San Cristobal de Las Casas</collectingMunicipality>
municipality,
<location id="8EB2601DC451FFB4146AFBB5271BA233" LSID="urn:lsid:plazi:treatment:03C487D0C451FFB5174FFD1620F1A714:8EB2601DC451FFB4146AFBB5271BA233" box="[557,771,1119,1145]" country="Mexico" municipality="San Cristobal de Las Casas" name="Reserva Huitepec" pageId="12" pageNumber="35" stateProvince="Chiapas">Reserva Huitepec</location>
,
<quantity id="4C959B23C451FFB41556FBB5277DA233" box="[785,869,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="12" pageNumber="35" unit="m" value="2250.0">
<elevation id="0040D1F5C451FFB41556FBB5277DA233" box="[785,869,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="12" pageNumber="35" unit="m" value="2250.0">2250m</elevation>
</quantity>
,
<date id="FFD31006C451FFB41534FBB5200EA233" box="[883,1046,1119,1145]" pageId="12" pageNumber="35" value="1996-05-30">
<collectingDate id="EF97E9EEC451FFB41534FBB5200EA233" box="[883,1046,1119,1145]" pageId="12" pageNumber="35" value="1996-05-30">30 May 1996</collectingDate>
</date>
,
<collectorName id="26985310C451FFB41262FBB520C4A233" box="[1061,1244,1119,1145]" pageId="12" pageNumber="35">Col. O. Gómez</collectorName>
(
<typeStatus id="54D68864C451FFB412AAFBB52522A2EB" pageId="12" pageNumber="35" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D68864C451FFB41731FB6D25CEA2EB" box="[374,470,1159,1185]" pageId="12" pageNumber="35" type="allotype">allotype</typeStatus>
) (
<collectionCode id="ED7CAE03C451FFB417A8FB6D2623A2EB" box="[495,571,1159,1185]" pageId="12" pageNumber="35">IEXA</collectionCode>
).
<emphasis id="B919EAD4C451FFB4141EFB6C27CAA2EA" bold="true" box="[601,978,1158,1184]" pageId="12" pageNumber="35">
<collectionCode id="ED7CAE03C451FFB4141EFB6C2687A2EA" box="[601,671,1158,1184]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="12" pageNumber="35">NEW</collectionCode>
<collectionCode id="ED7CAE03C451FFB414EFFB6C27CAA2EA" box="[680,978,1158,1184]" pageId="12" pageNumber="35">COUNTRY RECORDS</collectionCode>
</emphasis>
</materialsCitation>
:
<materialsCitation id="3B053C9BC451FFB415ABFB6D269AA283" collectorName="Guisayote" country="Honduras" location="Ocotepeque department" pageId="12" pageNumber="35" specimenCount="1" stateProvince="Ocotepeque" typeStatus="holotype">
<collectingCountry id="F37A7656C451FFB415ABFB6D208BA2EB" box="[1004,1171,1159,1185]" name="Honduras" pageId="12" pageNumber="35">HONDURAS</collectingCountry>
.
<collectingRegion id="49A9F824C451FFB412D8FB6D2597A283" country="Honduras" name="Ocotepeque" pageId="12" pageNumber="35">Ocotepeque department</collectingRegion>
,
<collectorName id="26985310C451FFB417DCFB45260DA283" box="[411,533,1199,1225]" pageId="12" pageNumber="35">Guisayote</collectorName>
,
<quantity id="4C959B23C451FFB41465FB45269AA283" box="[546,642,1199,1225]" metricMagnitude="4" metricUnit="m" metricValue="2.05" pageId="12" pageNumber="35" unit="km" value="20.5">20.5 km</quantity>
</materialsCitation>
<materialsCitation id="3B053C9BC451FFB414CCFB452096A283" box="[651,1166,1199,1225]" collectingDate="1994-06-16" collectionCode="R" collectorName="Col." country="Honduras" elevation="2170" location="Ocotepeque" pageId="12" pageNumber="35" specimenCount="1" stateProvince="Ocotepeque" typeStatus="holotype">
<collectingRegion id="49A9F824C451FFB414CCFB452702A283" box="[651,794,1199,1225]" country="Honduras" name="Ocotepeque" pageId="12" pageNumber="35">Ocotepeque</collectingRegion>
,
<quantity id="4C959B23C451FFB41561FB45279BA283" box="[806,899,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="12" pageNumber="35" unit="m" value="2170.0">
<elevation id="0040D1F5C451FFB41561FB45279BA283" box="[806,899,1199,1225]" metricMagnitude="3" metricUnit="m" metricValue="2.17" pageId="12" pageNumber="35" unit="m" value="2170.0">2170 m</elevation>
</quantity>
,
<date id="FFD31006C451FFB415C8FB452028A283" box="[911,1072,1199,1225]" pageId="12" pageNumber="35" value="1994-06-16">
<collectingDate id="EF97E9EEC451FFB415C8FB452028A283" box="[911,1072,1199,1225]" pageId="12" pageNumber="35" value="1994-06-16">16 June 1994</collectingDate>
</date>
,
<collectorName id="26985310C451FFB4127AFB452077A283" box="[1085,1135,1199,1225]" pageId="12" pageNumber="35">Col.</collectorName>
<collectionCode id="ED7CAE03C451FFB41230FB452096A283" box="[1143,1166,1199,1225]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="35">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B053C9BC451FFB412DDFB4526A5A2BB" collectionCode="CMNC, MXAL" country="Honduras" location="Anderson" pageId="12" pageNumber="35" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Ocotepeque" typeStatus="holotype">
<location id="8EB2601DC451FFB412DDFB452117A283" LSID="urn:lsid:plazi:treatment:03C487D0C451FFB5174FFD1620F1A714:8EB2601DC451FFB412DDFB452117A283" box="[1178,1295,1199,1225]" country="Honduras" name="Anderson" pageId="12" pageNumber="35" stateProvince="Ocotepeque">Anderson</location>
,
<specimenCount id="9D6BFD4FC451FFB4135BFB452557A2BB" pageId="12" pageNumber="35" type="male">7 males</specimenCount>
,
<specimenCount id="9D6BFD4FC451FFB41726FB3D25CFA2BB" box="[353,471,1239,1265]" pageId="12" pageNumber="35" type="female">6 females</specimenCount>
(
<collectionCode id="ED7CAE03C451FFB417AAFB3D2650A2BB" box="[493,584,1239,1265]" pageId="12" pageNumber="35">CMNC</collectionCode>
,
<collectionCode id="ED7CAE03C451FFB4141EFB3D26ADA2BB" box="[601,693,1239,1265]" pageId="12" pageNumber="35">MXAL</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B053C9BC451FFB41489FB3D2061A353" collectingDate="1999-07-17" collectionCode="UVGC, IEXA" collectorName="Col. J. Monzon" country="Honduras" elevation="1800" location="Cerro de Celaque" municipality="Corquin" pageId="12" pageNumber="35" specimenCount="12" specimenCount-female="9" specimenCount-male="3" stateProvince="Lempira" typeStatus="holotype">
<collectingRegion id="49A9F824C451FFB41489FB3D27DDA2BB" box="[718,965,1239,1265]" country="Honduras" name="Lempira" pageId="12" pageNumber="35">Lempira department</collectingRegion>
,
<collectingMunicipality id="6BB6ACBCC451FFB41591FB3D2021A2BB" box="[982,1081,1239,1265]" pageId="12" pageNumber="35">Corquin</collectingMunicipality>
,
<location id="8EB2601DC451FFB4120CFB3D213EA2BB" LSID="urn:lsid:plazi:treatment:03C487D0C451FFB5174FFD1620F1A714:8EB2601DC451FFB4120CFB3D213EA2BB" box="[1099,1318,1239,1265]" country="Honduras" municipality="Corquin" name="Cerro de Celaque" pageId="12" pageNumber="35" stateProvince="Lempira">Cerro de Celaque</location>
,
<quantity id="4C959B23C451FFB4174FFB15257BA353" box="[264,355,1279,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="35" unit="m" value="1800.0">
<elevation id="0040D1F5C451FFB4174FFB15257BA353" box="[264,355,1279,1305]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="35" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFD31006C451FFB41728FB152611A353" box="[367,521,1279,1305]" pageId="12" pageNumber="35" value="1999-07-17">
<collectingDate id="EF97E9EEC451FFB41728FB152611A353" box="[367,521,1279,1305]" pageId="12" pageNumber="35" value="1999-07-17">17 July 1999</collectingDate>
</date>
,
<collectorName id="26985310C451FFB41452FB1526D5A353" box="[533,717,1279,1305]" pageId="12" pageNumber="35">Col. J. Monzón</collectorName>
,
<specimenCount id="9D6BFD4FC451FFB4149EFB15272EA353" box="[729,822,1279,1305]" pageId="12" pageNumber="35" type="male">3 males</specimenCount>
,
<specimenCount id="9D6BFD4FC451FFB41505FB1527ABA353" box="[834,947,1279,1305]" pageId="12" pageNumber="35" type="female">9 females</specimenCount>
(
<collectionCode id="ED7CAE03C451FFB41585FB152002A353" box="[962,1050,1279,1305]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="12" pageNumber="35">UVGC</collectionCode>
,
<collectionCode id="ED7CAE03C451FFB41261FB152069A353" box="[1062,1137,1279,1305]" pageId="12" pageNumber="35">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4177FFACC2706A3DB" pageId="12" pageNumber="35" type="biology_ecology">
<paragraph id="8BD236C6C451FFB4177FFACC2706A3DB" blockId="12.[264,1325,838,1905]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFACC2584A30A" bold="true" box="[312,412,1318,1344]" pageId="12" pageNumber="35">Biology.</emphasis>
This species inhabits cloud forests and pine oak forests located at altitudes from
<quantity id="4C959B23C451FFB4170FFAA5261DA323" box="[328,517,1359,1385]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.25" metricValueMin="1.8" pageId="12" pageNumber="35" unit="m" value="2025.0" valueMax="2250.0" valueMin="1800.0">1800 to 2250 m</quantity>
. Months of collection: May (2), June (13), July (12). The
<typeStatus id="54D68864C451FFB412EBFAA520C6A323" box="[1196,1246,1359,1385]" pageId="12" pageNumber="35">type</typeStatus>
series was collected on an unidentified
<taxonomicName id="4C6D4D45C451FFB414CCFA9D2702A3DB" box="[651,794,1399,1425]" class="Magnoliopsida" family="Styracaceae" kingdom="Plantae" order="Ericales" pageId="12" pageNumber="35" phylum="Tracheophyta" rank="family">Styracaceae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB4177FFA7425E9A3AB" pageId="12" pageNumber="35" type="distribution">
<paragraph id="8BD236C6C451FFB4177FFA7425E9A3AB" blockId="12.[264,1325,838,1905]" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFA7425C9A3F2" bold="true" box="[312,465,1438,1464]" pageId="12" pageNumber="35">Distribution</emphasis>
. Northeastern
<collectingRegion id="49A9F824C451FFB41438FA7526F8A3F3" box="[639,736,1439,1465]" country="Mexico" name="Chiapas" pageId="12" pageNumber="35">Chiapas</collectingRegion>
,
<collectingCountry id="F37A7656C451FFB414ABFA75275EA3F3" box="[748,838,1439,1465]" name="Mexico" pageId="12" pageNumber="35">México</collectingCountry>
to western
<collectingCountry id="F37A7656C451FFB41588FA752059A3F3" box="[975,1089,1439,1465]" name="Honduras" pageId="12" pageNumber="35">Honduras</collectingCountry>
(14° 05'–
<geoCoordinate id="EE595001C451FFB412F1FA752130A3F3" box="[1206,1320,1439,1465]" degrees="16" direction="north" minutes="44" orientation="latitude" pageId="12" pageNumber="35" precision="925" value="16.733334">16° 44' N</geoCoordinate>
, 88° 40'–
<geoCoordinate id="EE595001C451FFB4172CFA2D25FCA3AB" box="[363,484,1479,1505]" degrees="92" direction="west" minutes="38" orientation="longitude" pageId="12" pageNumber="35" precision="925" value="-92.63333">92° 38' W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C377654DC451FFB5177FFA0420F1A714" lastPageId="13" lastPageNumber="36" pageId="12" pageNumber="35" type="discussion">
<paragraph id="8BD236C6C451FFB5177FFA0420F1A714" blockId="12.[264,1325,838,1905]" lastBlockId="13.[264,1322,284,350]" lastPageId="13" lastPageNumber="36" pageId="12" pageNumber="35">
<emphasis id="B919EAD4C451FFB4177FFA0425B2A042" bold="true" box="[312,426,1518,1544]" pageId="12" pageNumber="35">Remarks</emphasis>
. This species was originally described with one pair of specimens from “Reserva Huitepec”,
<collectingRegion id="49A9F824C451FFB41446F9FD267AA07B" box="[513,610,1559,1585]" country="Mexico" name="Chiapas" pageId="12" pageNumber="35">Chiapas</collectingRegion>
,
<collectingCountry id="F37A7656C451FFB41429F9FD26D3A07B" box="[622,715,1559,1585]" name="Mexico" pageId="12" pageNumber="35">México</collectingCountry>
. The large series of specimens collected in
<collectingCountry id="F37A7656C451FFB41296F9FD2532A013" name="Honduras" pageId="12" pageNumber="35">Honduras</collectingCountry>
have some differences from the types as follows: body length
<quantity id="4C959B23C451FFB41271F9D520A3A013" box="[1078,1211,1599,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="12" pageNumber="35" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
, humeral width 3.5–4.0 mm; clypeus, antennal funicle, elytra, pygidium, and legs reddish­brown (
<figureCitation id="13562A43C451FFB41756F9652571A0E3" box="[273,361,1679,1705]" captionStart="FIGURES 7–12" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 7–12. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="12" pageNumber="35">Fig. 11</figureCitation>
); abdominal sternites 2–5 each with 6–8 spine­like, erect, reddish­brown, long setae at the sides of midline; apex of meso­ and metatibia and tarsal segments darkened; parameres slightly larger with the crest on the inner border more developed (
<figureCitation id="13562A43C451FFB412C9F9352106A0B3" box="[1166,1310,1759,1785]" captionStart="FIGURES 13–21" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 13–21. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="12" pageNumber="35">Figs. 19–20</figureCitation>
). Female pronotum with scattered mixture of short and long erect setae (
<figureCitation id="13562A43C451FFB41220F8ED20D9A16B" box="[1127,1217,1799,1825]" captionStart="FIGURES 7–12" captionStartId="9.[264,383,1444,1468]" captionTargetBox="[372,1215,284,1437]" captionTargetId="figure-108@9.[372,1216,281,1438]" captionTargetPageId="9" captionText="FIGURES 7–12. 7­­­ Macrodactylus tibialis Arce­Pérez and Morón, sp. nov. male; 8­­­ M. hondurensis Arce­Pérez and Morón, sp. nov. male; 9­­­ M. montanus Arce­Pérez and Morón, male; 10­­­ M. tibialis, female; 11­­­M. hondurensis, female; 12­­­ M. montanus, female." pageId="12" pageNumber="35">Fig. 12</figureCitation>
); genital plates (
<figureCitation id="13562A43C451FFB4171BF8C525A9A103" box="[348,433,1839,1865]" captionStart="FIGURES 13–21" captionStartId="11.[264,383,1430,1454]" captionTargetBox="[319,1270,326,1406]" captionTargetId="figure-135@11.[318,1270,326,1406]" captionTargetPageId="11" captionText="FIGURES 13–21. Diagnostic structures of species of Macrodactylus: M. tibialis, 13­­­Parameres, distal view; 14­­­Parameres, lateral view; 15­­­Female genital plates, ventral view; M. hondurensis, 16­­­Parameres, distal view; 17­­­Parameres, lateral view; 18­­­Female genital plates, ventral view; M. montanus, 19­­­Parameres, distal view; 20­­­Parameres, lateral view; 21­­­Female genital plates, ventral view." pageId="12" pageNumber="35">Fig. 21</figureCitation>
) slightly longer and narrower than in the
<typeStatus id="54D68864C451FFB415DEF8C527E3A103" box="[921,1019,1839,1865]" pageId="12" pageNumber="35" type="allotype">allotype</typeStatus>
. The new records in
<collectingCountry id="F37A7656C451FFB412A9F8C5255FA13B" name="Honduras" pageId="12" pageNumber="35">Honduras</collectingCountry>
extend the distribution
<quantity id="4C959B23C451FFB4142AF8BD26D1A13B" box="[621,713,1879,1905]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="35" unit="km" value="250.0">250 km</quantity>
to southeast of San Cristobal Las Casas, Chiapas across the high mountain ranges in Chiapas,
<collectingCountry id="F37A7656C450FFB51555FEF6278CA77C" box="[786,916,284,310]" name="Guatemala" pageId="13" pageNumber="36">Guatemala</collectingCountry>
, and
<collectingCountry id="F37A7656C450FFB51597FEF6205EA77C" box="[976,1094,284,310]" name="Honduras" pageId="13" pageNumber="36">Honduras</collectingCountry>
. It is likely that this species will be found in Guatemalan humid forests located above
<quantity id="4C959B23C450FFB5124CFEAE207CA714" box="[1035,1124,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="36" unit="m" value="2000.0">2000 m</quantity>
in altitude.
</paragraph>
</subSubSection>
</treatment>
</document>
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