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<document id="D7C7AC09AECD9939CA4B595154D110DC" ID-DOI="10.1080/00222933.2012.708457" ID-ISSN="1464-5262" ID-Zenodo-Dep="5202137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628964073586" checkinUser="felipe" docAuthor="Stroiński, Adam &amp; Świerczewski, Dariusz" docDate="2012" docId="03B4211AD207FFE6FE4EFC4F512E110E" docLanguage="en" docName="JNatHist.46.41-42.2577-2593.pdf" docOrigin="Journal of Natural History 46 (41 - 42)" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Urana paradoxa Melichar 1902" docType="treatment" docVersion="3" lastPageNumber="2590" masterDocId="FF8D5962D20AFFE9FFF1FF8852531560" masterDocTitle="Revision of an extraordinary Selizini genus Urana Melichar, 1902 from Madagascar (Hemiptera: Fulgoromorpha: Flatidae)" masterLastPageNumber="2593" masterPageNumber="2577" pageNumber="2588" updateTime="1699057009937" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8A41155F79DA59CA5E5DFC18FB956F32" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A807E59EF3A0F06FE6D67323FC11EC71">
<mods:title id="8CA52DD374F490704B34A569E186656E">Revision of an extraordinary Selizini genus Urana Melichar, 1902 from Madagascar (Hemiptera: Fulgoromorpha: Flatidae)</mods:title>
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<mods:name id="81ACF019781F9D29C19EF376953B1CEF" type="personal">
<mods:role id="9809F4C12A1AC39D228F380E8F6D6430">
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<mods:namePart id="A9252E75BAB1B85FF156209ED57A0141">Stroiński, Adam</mods:namePart>
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<mods:roleTerm id="9ABAE64A7F06323D0F663942B06030E3">Author</mods:roleTerm>
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<mods:namePart id="682695ABD9071DC428AE25FED1E7E052">Świerczewski, Dariusz</mods:namePart>
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<mods:typeOfResource id="41D17BCD64DDA38CBD997CCB5B3CBB74">text</mods:typeOfResource>
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<mods:title id="CE228F474F5EF1D2C2647BA5F2298CF1">Journal of Natural History</mods:title>
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<mods:part id="5751320DC337442471F34ACAC0685C04">
<mods:date id="553CFD6FA71068D154777A9FC6D22F22">2012</mods:date>
<mods:detail id="20EFD9EBFB75A4BC9B96884C69E55500" type="volume">
<mods:number id="48A86E9EF8A21D4AB1F65C554A35428A">46</mods:number>
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<mods:detail id="942C16E61B094B023F845D5578019167" type="issue">
<mods:number id="2B7A362511B4F9E723509FC38E949A58">41 - 42</mods:number>
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<mods:start id="21240A87041B1152A2A151915310B99A">2577</mods:start>
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<mods:classification id="CF84149E5815FE2D302AF501B9D3B3E9">journal article</mods:classification>
<mods:identifier id="D846909B5B9C398CA13125058423DD20" type="DOI">10.1080/00222933.2012.708457</mods:identifier>
<mods:identifier id="1CA7425B6347694589CEC6C51C4BD49E" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="43B6B7E87FA5661AF80FD34FAA356CBD" type="Zenodo-Dep">5202137</mods:identifier>
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<treatment id="03B4211AD207FFE6FE4EFC4F512E110E" LSID="urn:lsid:plazi:treatment:03B4211AD207FFE6FE4EFC4F512E110E" httpUri="http://treatment.plazi.org/id/03B4211AD207FFE6FE4EFC4F512E110E" lastPageId="15" lastPageNumber="2590" pageId="13" pageNumber="2588">
<subSubSection id="C307C387D207FFE4FE4EFC4F514C16BE" box="[447,799,966,991]" pageId="13" pageNumber="2588" type="nomenclature">
<paragraph id="8BA2900CD207FFE4FE4EFC4F514C16BE" blockId="13.[447,799,966,1022]" box="[447,799,966,991]" pageId="13" pageNumber="2588">
<heading id="D0EA2760D207FFE4FE4EFC4F514C16BE" box="[447,799,966,991]" centered="true" fontSize="36" level="2" pageId="13" pageNumber="2588" reason="7">
<taxonomicName id="4C1DEB8FD207FFE4FE4EFC4F514C16BE" ID-CoL="7DNN5" authority="Melichar, 1902" authorityName="Melichar" authorityYear="1902" box="[447,799,966,991]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B9694C1ED207FFE4FE4EFC4F514C16BE" bold="true" box="[447,799,966,991]" pageId="13" pageNumber="2588">
<emphasis id="B9694C1ED207FFE4FE4EFC4F502116BF" bold="true" box="[447,626,966,991]" italics="true" pageId="13" pageNumber="2588">Urana paradoxa</emphasis>
<bibRefCitation id="EF8CEDFDD207FFE4FD88FC4E514C16BE" author="Melichar L" box="[633,799,966,990]" pageId="13" pageNumber="2588" pagination="1 - 256" refId="ref4777" refString="Melichar L. 1902. Monographie der Acanaloniiden und Flatiden (Homoptera). Ann k. k Naturhist Hofmus Wien. 17: 1 - 256." type="journal article" year="1902">Melichar, 1902</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C307C387D207FFE4FDFBFC6E5086169E" box="[522,725,998,1022]" pageId="13" pageNumber="2588" type="description">
<paragraph id="8BA2900CD207FFE4FDFBFC6E5086169E" blockId="13.[447,799,966,1022]" box="[522,725,998,1022]" pageId="13" pageNumber="2588">
(
<figureCitation id="13268C89D207FFE4FDE0FC6E50CB169E" box="[529,664,998,1022]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="5.[143,210,1728,1750]" captionStartId-1="6.[211,278,1648,1670]" captionStartId-2="7.[143,210,1648,1670]" captionStartId-3="9.[143,210,1645,1667]" captionStartId-4="10.[211,278,1358,1380]" captionTargetBox-0="[155,1092,162,1694]" captionTargetBox-1="[213,1170,162,1615]" captionTargetBox-2="[145,1102,162,1613]" captionTargetBox-3="[145,1102,161,1612]" captionTargetBox-4="[213,1170,169,1325]" captionTargetId-0="figure-9@5.[155,1092,161,1694]" captionTargetId-1="figure-5@6.[213,1170,162,1615]" captionTargetId-2="figure-9@7.[145,1102,162,1614]" captionTargetId-3="figure-9@9.[145,1102,161,1612]" captionTargetId-4="figure-5@10.[213,1170,169,1326]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Figure 1. Urana paradoxa Melichar, 1902, female: (A) habitus, lateral view; (B) same, dorsal view; (C) same, ventral view; (D) anterior part of body, dorsal view; (E) frons, frontal view." captionText-1="Figure 2. Urana paradoxa Melichar, 1902, female: (A) anterior part of body, dorso-lateral view; (B) same, dorsal view; (C) thorax, dorso-lateral view; (D) mesonotum and base of tegmen, dorsal view; (E) head and thorax, dorsal view; (F) frons, dorsal view; (G, H) anterior part of body, frontal view." captionText-2="Figure 3. Urana paradoxa Melichar, 1902, female: (A) head and thorax, lateral view; (B, C) antenna; (D) base of tegmen and bulla; (E) base of tegmen and clavus; (F) posterior part of tegmen; (G, H) sensory and wax gland-plates on tegmen." captionText-3="Figure 4. Urana paradoxa Melichar, 1902, female: (A) hind leg, lateral view; (B) hind tibia, ventral view; (C) hind tarsomere, ventral view; (D) genital capsule, lateral view; (E) anal tube, dorsal view; (F) end of gonoplac, lateral view; (G) pregenital sternite, ventral view; (H) same, lateral view." captionText-4="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5202139" figureDoi-1="http://doi.org/10.5281/zenodo.5202141" figureDoi-2="http://doi.org/10.5281/zenodo.5202143" figureDoi-3="http://doi.org/10.5281/zenodo.5202145" figureDoi-4="http://doi.org/10.5281/zenodo.5202147" httpUri-0="https://zenodo.org/record/5202139/files/figure.png" httpUri-1="https://zenodo.org/record/5202141/files/figure.png" httpUri-2="https://zenodo.org/record/5202143/files/figure.png" httpUri-3="https://zenodo.org/record/5202145/files/figure.png" httpUri-4="https://zenodo.org/record/5202147/files/figure.png" pageId="13" pageNumber="2588">Figures 1–5</figureCitation>
;
<figureCitation id="13268C89D207FFE4FD55FC6F50E0169E" box="[676,691,999,1022]" captionStart="Figure 7" captionStartId="12.[211,278,1616,1638]" captionTargetBox="[213,1162,161,1583]" captionTargetId="figure-5@12.[213,1170,161,1583]" captionTargetPageId="12" captionText="Figure 7. Urana paradoxa Melichar, 1902, female: (A) pregenital sternite, ventral view; (B) anal tube, dorsal view; (C) gonoplac, lateral view; (D) end of gonoplac, internal view; (E) gonapophysis VIII, lateral view; (F) gonapophysis IX and gonospiculum bridge, dorsal view; (G) same, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5202151" httpUri="https://zenodo.org/record/5202151/files/figure.png" pageId="13" pageNumber="2588">7</figureCitation>
;
<figureCitation id="13268C89D207FFE4FD4EFC6E509F169E" box="[703,716,998,1022]" captionStart="Figure 8" captionStartId="14.[211,278,1537,1559]" captionTargetBox="[213,1170,168,1504]" captionTargetId="figure-5@14.[213,1170,168,1504]" captionTargetPageId="14" captionText="Figure 8. Distribution map of genus Urana Melichar, 1901." figureDoi="http://doi.org/10.5281/zenodo.5202153" httpUri="https://zenodo.org/record/5202153/files/figure.png" pageId="13" pageNumber="2588">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C307C387D207FFE4FF7EFBAF519C111E" pageId="13" pageNumber="2588" type="reference_group">
<paragraph id="8BA2900CD207FFE4FF7EFBAF53AB115E" blockId="13.[143,504,1062,1086]" box="[143,504,1062,1086]" pageId="13" pageNumber="2588">
<taxonomicName id="4C1DEB8FD207FFE4FF7EFBAF53A7115E" ID-CoL="7DNN5" authority="Melichar, 1902" authorityName="Melichar" authorityYear="1902" box="[143,500,1062,1086]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B9694C1ED207FFE4FF7EFBAF5310115E" box="[143,323,1062,1086]" italics="true" pageId="13" pageNumber="2588">Urana paradoxa</emphasis>
<bibRefCitation id="EF8CEDFDD207FFE4FEB8FBAE53A7115E" author="Melichar L" box="[329,500,1062,1086]" pageId="13" pageNumber="2588" pagination="1 - 256" refId="ref4777" refString="Melichar L. 1902. Monographie der Acanaloniiden und Flatiden (Homoptera). Ann k. k Naturhist Hofmus Wien. 17: 1 - 256." type="journal article" year="1902">Melichar, 1902</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA2900CD207FFE4FF7EFBEF519C111E" blockId="13.[143,975,1126,1150]" box="[143,975,1126,1150]" pageId="13" pageNumber="2588">
<taxonomicName id="4C1DEB8FD207FFE4FF7EFBEF53A6111E" ID-CoL="7DNN5" authority=": Melichar 1923" authorityName="Melichar" authorityYear="1923" box="[143,501,1126,1150]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B9694C1ED207FFE4FF7EFBEF5310111E" box="[143,323,1126,1150]" italics="true" pageId="13" pageNumber="2588">Urana paradoxa</emphasis>
:
<bibRefCitation id="EF8CEDFDD207FFE4FEA0FBEE53A6111E" author="Melichar L" box="[337,501,1126,1150]" pageId="13" pageNumber="2588" pagination="1 - 185" refId="ref4805" refString="Melichar L. 1923. Homoptera. Fam. Acanaloniidae, Flatidae et Ricaniidae. Genera Insectorum. 182: 1 - 185." type="journal article" year="1923">Melichar 1923</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EF8CEDFDD207FFE4FDF1FBEE50DE111E" author="Synave H" box="[512,653,1126,1150]" pageId="13" pageNumber="2588" pagination="197 - 217" refId="ref5176" refString="Synave H. 1956. Les Flatidae de Madagascar (Hemiptera-Homoptera). Mem Ins Sci Madagascar (Ser. E). 7: 197 - 217." type="journal article" year="1956">Synave 1956</bibRefCitation>
,
<bibRefCitation id="EF8CEDFDD207FFE4FD69FBEE517C111E" author="Metcalf ZP" box="[664,815,1126,1150]" pageId="13" pageNumber="2588" refId="ref4829" refString="Metcalf ZP. 1957. Flatidae and Hypochthonellidae. Raleigh (NC): North Carolina State College. Part 13. General Catalogue of the Homoptera. Fascicule IV., 565 pp." type="book" year="1957">Metcalf 1957</bibRefCitation>
,
<bibRefCitation id="EF8CEDFDD207FFE4FCCBFBEE5198111E" author="Medler J" box="[826,971,1126,1150]" pageId="13" pageNumber="2588" pagination="105 - 118" refId="ref4624" refString="Medler J. 1990. Types of Flatidae XIII, lectotype designations and taxonomic notes on African species in the Zoological Museum of the Humboldt-University Berlin (Homoptera, Fulgoroidea). Dtsch ent Z N. F. 37 (1 - 3): 105 - 118." type="journal article" year="1990">Medler 1990</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307C387D207FFE6FF7EFB42512E110E" lastPageId="15" lastPageNumber="2590" pageId="13" pageNumber="2588" type="diagnosis">
<paragraph id="8BA2900CD207FFE4FF7EFB4252A81181" blockId="13.[143,251,1226,1249]" box="[143,251,1226,1249]" pageId="13" pageNumber="2588">
<emphasis id="B9694C1ED207FFE4FF7EFB4252A81181" box="[143,251,1226,1249]" italics="true" pageId="13" pageNumber="2588">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA2900CD207FFE4FF7EFB79509210E9" blockId="13.[143,1103,1265,1417]" pageId="13" pageNumber="2588">
Closely similar to
<taxonomicName id="4C1DEB8FD207FFE4FEA9FB7A53E41069" authority="Stroiński &amp; Świerczewski, 2012" authorityName="Stroiński &amp; Świerczewski" authorityYear="2012" box="[344,439,1266,1289]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="unica" status="sp. nov.">
<emphasis id="B9694C1ED207FFE4FEA9FB7A533F1069" box="[344,364,1266,1289]" italics="true" pageId="13" pageNumber="2588">U</emphasis>
.
<emphasis id="B9694C1ED207FFE4FE8DFB7A53E41069" box="[380,439,1266,1289]" italics="true" pageId="13" pageNumber="2588">unica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25AF165D207FFE4FE4DFB7A505C1069" box="[444,527,1266,1289]" pageId="13" pageNumber="2588" rank="species">sp. nov.</taxonomicNameLabel>
but differs by the male genital structures: basal part of periandrium with three dorsal teeth (
<figureCitation id="13268C89D207FFE4FDB3FA99509D1049" box="[578,718,1297,1321]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="13" pageNumber="2588">Figure 5A,B</figureCitation>
) – more in
<taxonomicName id="4C1DEB8FD207FFE4FCB9FA9A51F51049" authorityName="Stroiński &amp; Świerczewski" authorityYear="2012" box="[840,934,1298,1321]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="unica">
<emphasis id="B9694C1ED207FFE4FCB9FA9A510F1049" box="[840,860,1298,1321]" italics="true" pageId="13" pageNumber="2588">U</emphasis>
.
<emphasis id="B9694C1ED207FFE4FC9AFA9A51F51049" box="[875,934,1298,1321]" italics="true" pageId="13" pageNumber="2588">unica</emphasis>
</taxonomicName>
(
<figureCitation id="13268C89D207FFE4FC43FA9956111049" box="[946,1090,1297,1321]" captionStart="Figure 6" captionStartId="11.[143,210,1642,1664]" captionTargetBox="[146,1102,164,1605]" captionTargetId="figure-9@11.[145,1102,163,1610]" captionTargetPageId="11" captionText="Figure 6. Urana unica sp. nov., male: (A) anal tube, dorsal view; (B) same, lateral view; (C) stylus, lateral view; (D, E) phallic complex, lateral view; (F) apical part of ventral periandrium; (G) aedeagus, lateral view; (H) same, dorsal view; (I) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202149" httpUri="https://zenodo.org/record/5202149/files/figure.png" pageId="13" pageNumber="2588">Figure 6D,E</figureCitation>
); ventral part with lower keel-shaped median lobe –
<figureCitation id="13268C89D207FFE4FD4DFAB9517E1029" box="[700,813,1329,1353]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="13" pageNumber="2588">Figure 5A</figureCitation>
(
<taxonomicName id="4C1DEB8FD207FFE4FCCDFABA51C91029" authorityName="Stroiński &amp; Świerczewski" authorityYear="2012" box="[828,922,1330,1353]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="unica">
<emphasis id="B9694C1ED207FFE4FCCDFABA51031029" box="[828,848,1330,1353]" italics="true" pageId="13" pageNumber="2588">U</emphasis>
.
<emphasis id="B9694C1ED207FFE4FCAEFABA51C91029" box="[863,922,1330,1353]" italics="true" pageId="13" pageNumber="2588">unica</emphasis>
</taxonomicName>
– without lobe); upper ventral lobes elongated and narrow with deep median split (
<figureCitation id="13268C89D207FFE4FC74FAD956471009" box="[901,1044,1361,1385]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="13" pageNumber="2588">Figure 5A,B</figureCitation>
) – in
<taxonomicName id="4C1DEB8FD207FFE4FF7EFAFA52BC10E9" authorityName="Stroiński &amp; Świerczewski" authorityYear="2012" box="[143,239,1393,1417]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="2588" phylum="Arthropoda" rank="species" species="unica">
<emphasis id="B9694C1ED207FFE4FF7EFAFA52F010E9" box="[143,163,1394,1417]" italics="true" pageId="13" pageNumber="2588">U</emphasis>
.
<emphasis id="B9694C1ED207FFE4FF45FAFA52BC10E9" box="[180,239,1394,1417]" italics="true" pageId="13" pageNumber="2588">unica</emphasis>
</taxonomicName>
with three upper short lobes (
<figureCitation id="13268C89D207FFE4FDCEFAF950E710E9" box="[575,692,1393,1417]" captionStart="Figure 6" captionStartId="11.[143,210,1642,1664]" captionTargetBox="[146,1102,164,1605]" captionTargetId="figure-9@11.[145,1102,163,1610]" captionTargetPageId="11" captionText="Figure 6. Urana unica sp. nov., male: (A) anal tube, dorsal view; (B) same, lateral view; (C) stylus, lateral view; (D, E) phallic complex, lateral view; (F) apical part of ventral periandrium; (G) aedeagus, lateral view; (H) same, dorsal view; (I) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202149" httpUri="https://zenodo.org/record/5202149/files/figure.png" pageId="13" pageNumber="2588">Figure 6F</figureCitation>
).
</paragraph>
<paragraph id="8BA2900CD207FFE4FF7EFA5D535D108C" blockId="13.[143,270,1493,1516]" box="[143,270,1493,1516]" pageId="13" pageNumber="2588">
<emphasis id="B9694C1ED207FFE4FF7EFA5D535D108C" box="[143,270,1493,1516]" italics="true" pageId="13" pageNumber="2588">Description</emphasis>
</paragraph>
<paragraph id="8BA2900CD207FFE4FF7EFA7453EA1374" blockId="13.[143,441,1532,1556]" box="[143,441,1532,1556]" pageId="13" pageNumber="2588">
Total length
<quantity id="4CE53DE9D207FFE4FEEFFA7453E61374" box="[286,437,1532,1556]" metricMagnitude="-2" metricUnit="m" metricValue="1.005" metricValueMax="1.1" metricValueMin="0.91" pageId="13" pageNumber="2588" unit="cm" value="1.005" valueMax="1.1" valueMin="0.91">0.91–1.10 cm</quantity>
.
</paragraph>
<paragraph id="8BA2900CD207FFE4FF7EF9CA538813FA" blockId="13.[143,1104,1602,1690]" pageId="13" pageNumber="2588">
<emphasis id="B9694C1ED207FFE4FF7EF9CA5280133A" box="[143,211,1602,1626]" italics="true" pageId="13" pageNumber="2588">Head.</emphasis>
Vertex: proportion A
<emphasis id="B9694C1ED207FFE4FE21F9CA538F133A" bold="true" box="[464,476,1602,1626]" italics="true" pageId="13" pageNumber="2588">/</emphasis>
B = 6.00–9.00; anterior margin in the specimen from Anjanaharibe weakly concave in median portion. Frons: proportion C
<emphasis id="B9694C1ED207FFE4FC6FF9EA51F9131A" bold="true" box="[926,938,1634,1658]" italics="true" pageId="13" pageNumber="2588">/</emphasis>
E = 0.70–0.84; proportion D
<emphasis id="B9694C1ED207FFE4FED6F90A536013FA" bold="true" box="[295,307,1666,1690]" italics="true" pageId="13" pageNumber="2588">/</emphasis>
E = 0.97–1.02.
</paragraph>
<caption id="DF62C084D204FFE7FF22F989517E1377" ID-DOI="http://doi.org/10.5281/zenodo.5202153" ID-Zenodo-Dep="5202153" box="[211,813,1537,1559]" httpUri="https://zenodo.org/record/5202153/files/figure.png" pageId="14" pageNumber="2589" startId="14.[211,278,1537,1559]" targetBox="[213,1170,168,1504]" targetPageId="14">
<paragraph id="8BA2900CD204FFE7FF22F989517E1377" blockId="14.[211,813,1537,1559]" box="[211,813,1537,1559]" pageId="14" pageNumber="2589">
Figure 8. Distribution map of genus
<taxonomicName id="4C1DEB8FD204FFE7FDBAF98A517A1377" authority="Melichar, 1901" authorityName="Melichar" authorityYear="1901" box="[587,809,1537,1559]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="2589" phylum="Arthropoda" rank="genus">
<emphasis id="B9694C1ED204FFE7FDBAF98A50D91377" box="[587,650,1538,1559]" italics="true" pageId="14" pageNumber="2589">Urana</emphasis>
<bibRefCitation id="EF8CEDFDD204FFE7FD61F989517A1377" author="Melichar L" box="[656,809,1537,1559]" pageId="14" pageNumber="2589" pagination="178 - 258" refId="ref4749" refString="Melichar L. 1901. Monographie der Acanaloniiden und Flatiden (Homoptera). Ann k. k Naturhist Hofmus Wien. 16: 178 - 258." type="journal article" year="1901">Melichar, 1901</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA2900CD204FFE7FF22F9CD505E13FC" blockId="14.[211,1172,1604,1692]" pageId="14" pageNumber="2589">
<emphasis id="B9694C1ED204FFE7FF22F9CD537E133D" box="[211,301,1605,1629]" italics="true" pageId="14" pageNumber="2589">Thorax.</emphasis>
Pronotum: proportion F
<emphasis id="B9694C1ED204FFE7FDA2F9CD500C133D" bold="true" box="[595,607,1605,1629]" italics="true" pageId="14" pageNumber="2589">/</emphasis>
B = 3.00–6.00. Mesonotum: proportion G
<emphasis id="B9694C1ED204FFE7FBA4F9CD5632133D" bold="true" box="[1109,1121,1605,1629]" italics="true" pageId="14" pageNumber="2589">/</emphasis>
F = 3.33–3.78, proportion G
<emphasis id="B9694C1ED204FFE7FE19F9EC53A7131C" bold="true" box="[488,500,1636,1660]" italics="true" pageId="14" pageNumber="2589">/</emphasis>
B+F = 1.82–2.94, proportion G
<emphasis id="B9694C1ED204FFE7FC96F9EC5120131C" bold="true" box="[871,883,1636,1660]" italics="true" pageId="14" pageNumber="2589">/</emphasis>
H = 0.80–0.94. Tegmina: proportion I
<emphasis id="B9694C1ED204FFE7FE91F90C533F13FC" bold="true" box="[352,364,1668,1692]" italics="true" pageId="14" pageNumber="2589">/</emphasis>
J = 2.62–2.75.
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFF2053CD149E" blockId="15.[143,1104,167,510]" pageId="15" pageNumber="2590">
<emphasis id="B9694C1ED205FFE6FF7EFF20536915A0" box="[143,314,168,192]" italics="true" pageId="15" pageNumber="2590">Male genitalia.</emphasis>
Anal tube (in lateral view,
<figureCitation id="13268C89D205FFE6FD8DFF2F50A715DF" box="[636,756,167,191]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="15" pageNumber="2590">Figure 5C</figureCitation>
): lower margin convex; apical part distinctly narrower than median. Anal tube (in dorsal view,
<figureCitation id="13268C89D205FFE6FC88FF4F51A015BF" box="[889,1011,199,223]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="15" pageNumber="2590">Figure 5D</figureCitation>
): lateral margins in median portion convex, apical portion tapering apicad. Genital styles: posterior margin not concave in median portion; lower part of genital styles destroyed. Periandrium (
<figureCitation id="13268C89D205FFE6FEDCFEAF5393145F" box="[301,448,295,319]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="15" pageNumber="2590">Figure 5A,B</figureCitation>
): basal part with three dorsal teeth (median straight, lateral aurochs-horn-like); ventral part with three lobes: lower one laterally flattened, keel-shaped, surpassing half of upper lobes length; upper lobes extremely elongated and narrow with deep median split; lateral part of periandrium apically with huge single-armed appendage with basal one or two processes and short teeth. Aedeagus s.s.: lateral lobes smaller than those of
<taxonomicName id="4C1DEB8FD205FFE6FDBBFE4F50F814BE" authorityName="Stroiński &amp; Świerczewski" authorityYear="2012" box="[586,683,455,478]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="2590" phylum="Arthropoda" rank="species" species="unica">
<emphasis id="B9694C1ED205FFE6FDBBFE4F500D14BE" box="[586,606,455,478]" italics="true" pageId="15" pageNumber="2590">U</emphasis>
.
<emphasis id="B9694C1ED205FFE6FD81FE4F50F814BE" box="[624,683,455,478]" italics="true" pageId="15" pageNumber="2590">unica</emphasis>
</taxonomicName>
and placed subapically, dorso-apical lobes as in
<figureCitation id="13268C89D205FFE6FEFAFE6E53C9149E" box="[267,410,486,510]" captionStart="Figure 5" captionStartId="10.[211,278,1358,1380]" captionTargetBox="[213,1170,169,1325]" captionTargetId="figure-5@10.[213,1170,169,1326]" captionTargetPageId="10" captionText="Figure 5. Urana paradoxa Melichar, 1902, male: (A) periandrium, lateral view; (B) same, dorsal view; (C) anal tube, lateral view; (D) same, dorsal view; (E) stylus, lateral view; (F) aedeagus, lateral view; (G) same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5202147" httpUri="https://zenodo.org/record/5202147/files/figure.png" pageId="15" pageNumber="2590">Figure 5F,G</figureCitation>
.
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFDB2537B1732" blockId="15.[143,296,570,594]" box="[143,296,570,594]" pageId="15" pageNumber="2590">
<emphasis id="B9694C1ED205FFE6FF7EFDB2537B1732" box="[143,296,570,594]" italics="true" pageId="15" pageNumber="2590">Type material</emphasis>
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFDE9533017D9" blockId="15.[143,1103,609,761]" pageId="15" pageNumber="2590">
<materialsCitation id="3B759A51D205FFE6FF7EFDE9533017D9" collectionCode="ZMB" collectorName="Sikora J. F." country="Madagascar" location="Antananarivo" pageId="15" pageNumber="2590" specimenCount="1" specimenCount-male="1" stateProvince="Antananarivo" typeStatus="holotype">
<typeStatus id="54A62EAED205FFE6FF7EFDE952AF1719" box="[143,252,609,633]" pageId="15" pageNumber="2590" type="holotype">Holotype</typeStatus>
,
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: [
<collectingCountry id="F30AD09CD205FFE6FECCFDE9539A1719" box="[317,457,609,633]" name="Madagascar" pageId="15" pageNumber="2590">Madagascar</collectingCountry>
,
<collectingRegion id="49D95EEED205FFE6FE2AFDE950281719" box="[475,635,609,633]" country="Madagascar" name="Antananarivo" pageId="15" pageNumber="2590">Antananarivo</collectingRegion>
,
<collectorName id="26E8F5DAD205FFE6FD7DFDE951591719" box="[652,778,609,633]" pageId="15" pageNumber="2590">Sikora J.F.</collectorName>
], [Hymenopt mimicry Fall] [
<taxonomicName id="4C1DEB8FD205FFE6FF67FD0A539C17F9" authority="Melichar" authorityName="Melichar" authorityYear="1902" box="[150,463,641,665]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="2590" phylum="Arthropoda" rank="species" species="paradoxa">Urana paradoxa Melichar</taxonomicName>
det.], [Typus], [Zool. Mus. Berlin]. The
<typeStatus id="54A62EAED205FFE6FC3AFD09567C17F9" box="[971,1071,641,665]" pageId="15" pageNumber="2590" type="holotype">holotype</typeStatus>
is deposited in
<collectionCode id="ED0C08C9D205FFE6FEEEFD2A530C17D9" box="[287,351,674,697]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="15" pageNumber="2590" type="Museum">ZMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFD4A53BA1799" blockId="15.[143,1103,609,761]" pageId="15" pageNumber="2590">
Note: In original description (
<bibRefCitation id="EF8CEDFDD205FFE6FE15FD4950DA17B9" author="Melichar L" box="[484,649,705,729]" pageId="15" pageNumber="2590" pagination="1 - 256" refId="ref4777" refString="Melichar L. 1902. Monographie der Acanaloniiden und Flatiden (Homoptera). Ann k. k Naturhist Hofmus Wien. 17: 1 - 256." type="journal article" year="1902">Melichar 1902</bibRefCitation>
) and in
<bibRefCitation id="EF8CEDFDD205FFE6FD19FD4951DA17B9" author="Medler J" box="[744,905,705,729]" pageId="15" pageNumber="2590" pagination="105 - 118" refId="ref4624" refString="Medler J. 1990. Types of Flatidae XIII, lectotype designations and taxonomic notes on African species in the Zoological Museum of the Humboldt-University Berlin (Homoptera, Fulgoroidea). Dtsch ent Z N. F. 37 (1 - 3): 105 - 118." type="journal article" year="1990">Medler (1990)</bibRefCitation>
the name of
<typeStatus id="54A62EAED205FFE6FBEEFD4A561C17BA" box="[1055,1103,706,730]" pageId="15" pageNumber="2590">type</typeStatus>
locality is given as Antanarivo.
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFCBC5323162C" blockId="15.[143,368,820,844]" box="[143,368,820,844]" pageId="15" pageNumber="2590">
<emphasis id="B9694C1ED205FFE6FF7EFCBC5323162C" box="[143,368,820,844]" italics="true" pageId="15" pageNumber="2590">Additional materials</emphasis>
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFCD5531D1693" blockId="15.[143,1103,860,1011]" pageId="15" pageNumber="2590">
<materialsCitation id="3B759A51D205FFE6FF7EFCD553191693" collectingDate="1971-01-15" collectionCode="R, IRSNB, MNHN" collectorName="Two &amp; Madagascar-East &amp; Andapa Anjanaharibe &amp; Soga &amp; Institut Scientifique &amp; I. Sc. &amp; Andringitra East Anjavidilava &amp; December" country="Madagascar" county="Centre" determinerName="H. Synave" elevation="1600" location="East Anjavidilava" municipality="Museum Paris" pageId="15" pageNumber="2590" specimenCode="FDHM 2" specimenCount="61" specimenCount-female="1">
<collectorName id="26E8F5DAD205FFE6FF7EFCD552EC1614" box="[143,191,861,884]" pageId="15" pageNumber="2590">Two</collectorName>
<specimenCount id="9D1B5B85D205FFE6FF34FCD452B11614" box="[197,226,860,884]" pageId="15" pageNumber="2590" type="female">♀♀</specimenCount>
: [
<collectorName id="26E8F5DAD205FFE6FF07FCD453EF1614" box="[246,444,860,884]" pageId="15" pageNumber="2590">Madagascar-East</collectorName>
dct.
<collectorName id="26E8F5DAD205FFE6FE03FCD450BD1614" box="[498,750,860,884]" pageId="15" pageNumber="2590">Andapa Anjanaharibe</collectorName>
<quantity id="4CE53DE9D205FFE6FD05FCD451161614" box="[756,837,860,884]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="2590" unit="m" value="1600.0">
<elevation id="0030773FD205FFE6FD05FCD451161614" box="[756,837,860,884]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="2590" unit="m" value="1600.0">1600 m</elevation>
</quantity>
IV-
<specimenCount id="9D1B5B85D205FFE6FC9EFCD451F71614" box="[879,932,860,884]" pageId="15" pageNumber="2590" type="generic">60 P.</specimenCount>
<collectorName id="26E8F5DAD205FFE6FC58FCD451B71614" box="[937,996,860,884]" pageId="15" pageNumber="2590">Soga</collectorName>
], [
<collectorName id="26E8F5DAD205FFE6FC0AFCD4534416F4" pageId="15" pageNumber="2590">Institut Scientifique</collectorName>
], [
<collectionCode id="ED0C08C9D205FFE6FEC1FCF4531416F4" box="[304,327,892,916]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="2590">R</collectionCode>
.
<collectorName id="26E8F5DAD205FFE6FEBAFCF4532E16F4" box="[331,381,892,916]" pageId="15" pageNumber="2590">I.Sc.</collectorName>
N.B. I.G. 22.889], [
<determinerName id="7E7E7CC8D205FFE6FDA5FCF5509416F4" box="[596,711,892,916]" pageId="15" pageNumber="2590">H. Synave</determinerName>
det., 1963
<taxonomicName id="4C1DEB8FD205FFE6FCB5FCF5566616F4" authority="Mel" authorityName="Melichar" authorityYear="1902" box="[836,1077,892,916]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="2590" phylum="Arthropoda" rank="species" species="paradoxa">Urana paradoxa Mel</taxonomicName>
] – (
<collectionCode id="ED0C08C9D205FFE6FF66FC1452A316D4" box="[151,240,924,948]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="15" pageNumber="2590">IRSNB</collectionCode>
); [
<collectorName id="26E8F5DAD205FFE6FEE0FC14502216D4" box="[273,625,924,948]" pageId="15" pageNumber="2590">
Andringitra
<location id="8EC2C6D7D205FFE6FE57FC14502216D4" LSID="urn:lsid:plazi:treatment:03B4211AD207FFE6FE4EFC4F512E110E:8EC2C6D7D205FFE6FE57FC14502216D4" box="[422,625,924,948]" country="Madagascar" county="Centre" municipality="Museum Paris" name="East Anjavidilava" pageId="15" pageNumber="2590">East Anjavidilava</location>
</collectorName>
<quantity id="4CE53DE9D205FFE6FD8EFC14514916D3" box="[639,794,924,948]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="15" pageNumber="2590" unit="m" value="1900.0" valueMax="1950.0" valueMin="1850.0">
<elevation id="0030773FD205FFE6FD8EFC14514916D3" box="[639,794,924,948]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="15" pageNumber="2590" unit="m" value="1900.0" valueMax="1950.0" valueMin="1850.0">1850–1950 m</elevation>
</quantity>
18
<collectorName id="26E8F5DAD205FFE6FCBEFC14519116D4" box="[847,962,924,948]" pageId="15" pageNumber="2590">December</collectorName>
<emphasis id="B9694C1ED205FFE6FC33FC14519D16D4" bold="true" box="[962,974,924,948]" italics="true" pageId="15" pageNumber="2590">/</emphasis>
<date id="FFA3B6CCD205FFE6FC3EFC14529416B3" pageId="15" pageNumber="2590" value="1971-01-15">
<collectingDate id="EFE74F24D205FFE6FC3EFC14529416B3" pageId="15" pageNumber="2590" value="1971-01-15">15 January 1971</collectingDate>
</date>
], [
<specimenCode id="DBBB3877D205FFE6FF15FC34530516B3" box="[228,342,955,979]" collectionCode="FDHM" pageId="15" pageNumber="2590">FDHM 2</specimenCode>
], [
<collectingMunicipality id="6BC60A76D205FFE6FE85FC34504F16B3" box="[372,540,955,979]" pageId="15" pageNumber="2590">Museum Paris</collectingMunicipality>
<collectingCountry id="F30AD09CD205FFE6FDD7FC3350E216B3" box="[550,689,955,979]" name="Madagascar" pageId="15" pageNumber="2590">Madagascar</collectingCountry>
<collectingCounty id="62C3E880D205FFE6FD4DFC33515B16B3" box="[700,776,955,979]" pageId="15" pageNumber="2590">Centre</collectingCounty>
mission C.N.
<collectionCode id="ED0C08C9D205FFE6FC58FC33519316B3" box="[937,960,955,979]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="2590">R</collectionCode>
.S.
<collectionCode id="ED0C08C9D205FFE6FC12FC3351AA16B3" box="[995,1017,955,979]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="15" pageNumber="2590">R</collectionCode>
.C.P. n
<superScript id="7C683D44D205FFE6FBB4FC30561D16A9" attach="left" box="[1093,1102,952,969]" fontSize="7" pageId="15" pageNumber="2590">◦</superScript>
225] – (
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)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFBA753471127" blockId="15.[143,276,1071,1095]" box="[143,276,1071,1095]" pageId="15" pageNumber="2590">
<emphasis id="B9694C1ED205FFE6FF7EFBA753471127" box="[143,276,1071,1095]" italics="true" pageId="15" pageNumber="2590">Distribution</emphasis>
</paragraph>
<paragraph id="8BA2900CD205FFE6FF7EFBDE512E110E" blockId="15.[143,893,1110,1134]" box="[143,893,1110,1134]" pageId="15" pageNumber="2590">
<materialsCitation id="3B759A51D205FFE6FF7EFBDE512A110E" box="[143,889,1110,1134]" country="Madagascar" location="Antsiranana" pageId="15" pageNumber="2590" specimenCount="1" stateProvince="Antsiranana">
<collectingCountry id="F30AD09CD205FFE6FF7EFBDE534D110E" box="[143,286,1110,1134]" name="Madagascar" pageId="15" pageNumber="2590">Madagascar</collectingCountry>
:
<collectingRegion id="49D95EEED205FFE6FED9FBDE53EB110E" box="[296,440,1110,1134]" country="Madagascar" name="Antsiranana" pageId="15" pageNumber="2590">Antsiranana</collectingRegion>
,
<collectingRegion id="49D95EEED205FFE6FE33FBDE5031110E" box="[450,610,1110,1134]" country="Madagascar" name="Antananarivo" pageId="15" pageNumber="2590">Antananarivo</collectingRegion>
,
<collectingRegion id="49D95EEED205FFE6FD9CFBDE5157110E" box="[621,772,1110,1134]" country="Madagascar" name="Fianarantsoa" pageId="15" pageNumber="2590">Fianarantsoa</collectingRegion>
Provinces
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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