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103 lines
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<document id="F8444A1746683EE851DE3B2FF0CBBE98" ID-DOI="10.26879/855" ID-ISSN="1094-8074" ID-Zenodo-Dep="11188160" ID-ZooBank="0383B140-3EC0-4E45-904E-6ECBF9DFF202" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1714012836433" checkinUser="felipe" docAuthor="Palencia, Lorena, Peñalver, Enrique, Prieto, Carlos E. & Poyato-Ariza, Francisco José" docDate="2019" docId="044D87C3FFC5FFEE2D4ACC0519EBFD9B" docLanguage="en" docName="PalaeontolElectron.22.1.5A.pdf" docOrigin="Palaeontologia Electronica (5 A) 22 (1)" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Opiliones Sundevall 1833" docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="F874FFBBFFC7FFED2C51CB331813FFD9" masterDocTitle="First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1715639263252" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="5F263651E278DF6FF937D1DFDF953AA0">First fossil harvestmen (Arachnida: Opiliones) from Spain and notes on the fossil record of Opiliones</mods:title>
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<mods:namePart id="C9D86EC2D9553BB2E8D4678164C84D9B">Palencia, Lorena</mods:namePart>
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</mods:role>
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<mods:namePart id="5613A98A056277C64B5D2583CDCA7976">Peñalver, Enrique</mods:namePart>
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</mods:name>
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<mods:namePart id="CCE48A9B7BCD0380170078B562D307F0">Prieto, Carlos E.</mods:namePart>
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</mods:name>
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<mods:name id="90D9FE5C6E9C1BA9C350DE7F5796AD22" type="personal">
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<mods:namePart id="D728969CAA141AB7080625053E9B886A">Poyato-Ariza, Francisco José</mods:namePart>
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<mods:title id="8222047DBD2C6E2A950E5E29BE1EE4E8">Palaeontologia Electronica</mods:title>
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<mods:part id="0E870A9CA6629D45F493439F7EC69FCC">
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<mods:date id="28FB461C4A5C0D50031A75F5C8EF734F">2019</mods:date>
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<mods:title id="1917C007E5ACE5618844215B9D6E2016">5 A</mods:title>
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</mods:detail>
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<mods:number id="7B540C0EA05320643137CE80BA325AF5">2019-02-28</mods:number>
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<mods:detail id="E403E5A07D992049FF9267EC26FB7E28" type="volume">
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<mods:number id="D3EA1D72F8FAF98BBDA1500241753C48">22</mods:number>
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<mods:detail id="B7C64A1DFD9705D40F738CDB5A08A4F9" type="issue">
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<mods:number id="9C84B5291CE159618F310164AAD2558E">1</mods:number>
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<mods:start id="DAB2BDF2EF6E2967861E0B6AAEC3CB7C">1</mods:start>
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<mods:end id="6D350A68348CFB381092434E1C3FD7D7">18</mods:end>
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<mods:classification id="6FA809275D39000F5849D19140CD6ABF">journal article</mods:classification>
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<mods:identifier id="11DC1EA9317C34C9790A197D8093BC93" type="DOI">10.26879/855</mods:identifier>
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<mods:identifier id="1E58254EBF84061F2FF3D173AD5AA85D" type="ISSN">1094-8074</mods:identifier>
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<mods:identifier id="96929925310DBFB33B90126A8EEEC91A" type="Zenodo-Dep">11188160</mods:identifier>
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<mods:identifier id="84DA477D552836759EFD7D433056BEBA" type="ZooBank">0383B140-3EC0-4E45-904E-6ECBF9DFF202</mods:identifier>
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<treatment id="044D87C3FFC5FFEE2D4ACC0519EBFD9B" ID-DOI="http://doi.org/10.5281/zenodo.11188181" ID-Zenodo-Dep="11188181" LSID="urn:lsid:plazi:treatment:044D87C3FFC5FFEE2D4ACC0519EBFD9B" httpUri="http://treatment.plazi.org/id/044D87C3FFC5FFEE2D4ACC0519EBFD9B" lastPageId="3" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="C4FE655EFFC5FFEF2D4ACC051AAEF896" box="[283,701,1846,1872]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8C5B36D5FFC5FFEF2D4ACC051AAEF896" blockId="2.[283,701,1814,1872]" box="[283,701,1846,1872]" pageId="2" pageNumber="3">
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Order
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<taxonomicName id="4BE44D56FFC5FFEF2D36CC051AAEF896" ID-CoL="3Q2" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[359,701,1846,1872]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="order">OPILIONES Sundevall, 1833</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4FE655EFFC5FFEF2848CB951CC3FF66" box="[1049,1232,166,192]" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="8C5B36D5FFC5FFEF2848CB951CC3FF66" blockId="2.[1049,1232,166,223]" box="[1049,1232,166,192]" pageId="2" pageNumber="3">
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<heading id="D71381B9FFC5FFEF2848CB951CC3FF66" box="[1049,1232,166,192]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="3">Suborder indet.</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4FE655EFFC5FFEF2812CBF51B9BFD53" pageId="2" pageNumber="3" type="description">
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<paragraph id="8C5B36D5FFC5FFEF2812CBF51CB6FF06" blockId="2.[1049,1232,166,223]" box="[1091,1189,198,223]" pageId="2" pageNumber="3">
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<figureCitation id="14DF2A50FFC5FFEF2812CBF51CB6FF06" box="[1091,1189,198,223]" captionStart="FIGURE 1" captionStartId="2.[837,928,1803,1826]" captionTargetBox="[839,1436,923,1732]" captionTargetId="figure-500@2.[839,1436,923,1770]" captionTargetPageId="2" captionText="FIGURE 1. Cretaceous harvestman of uncertain subor- der, LH-29969, from Las Hoyas. 1, Photograph of LH- 29969. 2, Camera lucida drawing of LH-29969. Arrows indicate the detected leg articulations." figureDoi="http://doi.org/10.5281/zenodo.11188162" httpUri="https://zenodo.org/record/11188162/files/figure.png" pageId="2" pageNumber="3">Figure 1</figureCitation>
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</paragraph>
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<paragraph id="8C5B36D5FFC5FFEF2F19CBC21D82FE53" blockId="2.[840,1441,241,842]" pageId="2" pageNumber="3">
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<emphasis id="BE90EAC7FFC5FFEF2F19CBC21D12FED3" bold="true" box="[840,1281,241,266]" pageId="2" pageNumber="3">Description (specimen LH-29969).</emphasis>
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Typical harvestman habitus, compact body and slender legs. The eight legs appear radiating closely together. Leg lengths are indicated on
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<figureCitation id="14DF2A50FFC5FFEF28CCCA621D13FEB3" box="[1181,1280,337,362]" captionStart="FIGURE 1" captionStartId="2.[837,928,1803,1826]" captionTargetBox="[839,1436,923,1732]" captionTargetId="figure-500@2.[839,1436,923,1770]" captionTargetPageId="2" captionText="FIGURE 1. Cretaceous harvestman of uncertain subor- der, LH-29969, from Las Hoyas. 1, Photograph of LH- 29969. 2, Camera lucida drawing of LH-29969. Arrows indicate the detected leg articulations." figureDoi="http://doi.org/10.5281/zenodo.11188162" httpUri="https://zenodo.org/record/11188162/files/figure.png" pageId="2" pageNumber="3">Figure 1</figureCitation>
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. Leg articulations only are clear in three locations (
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<figureCitation id="14DF2A50FFC5FFEF2957CA421D95FE53" box="[1286,1414,369,394]" captionStart="FIGURE 1" captionStartId="2.[837,928,1803,1826]" captionTargetBox="[839,1436,923,1732]" captionTargetId="figure-500@2.[839,1436,923,1770]" captionTargetPageId="2" captionText="FIGURE 1. Cretaceous harvestman of uncertain subor- der, LH-29969, from Las Hoyas. 1, Photograph of LH- 29969. 2, Camera lucida drawing of LH-29969. Arrows indicate the detected leg articulations." figureDoi="http://doi.org/10.5281/zenodo.11188162" httpUri="https://zenodo.org/record/11188162/files/figure.png" pageId="2" pageNumber="3">Figure 1.2</figureCitation>
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).
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</paragraph>
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<paragraph id="8C5B36D5FFC5FFEF2F19CAA21B9BFD53" blockId="2.[840,1441,241,842]" pageId="2" pageNumber="3">
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<emphasis id="BE90EAC7FFC5FFEF2F19CAA21BF1FE73" bold="true" box="[840,994,401,426]" pageId="2" pageNumber="3">Taphonomy.</emphasis>
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The body of this specimen is poorly preserved, lacking any detail; chelicera and pedipalps are not observable. Anatomical orientation is uncertain due to the poor preservation of the body, which was collapsed in a small area during fossilization. Legs appear radiating very close, nearly together, probably due to this collapsing of the body.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4FE655EFFC5FFEE2F19C9A219EBFD9B" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="8C5B36D5FFC5FFEE2F19C9A219EBFD9B" blockId="2.[840,1441,241,842]" lastBlockId="3.[192,793,169,578]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<emphasis id="BE90EAC7FFC5FFEF2F19C9A21BD3FD73" bold="true" box="[840,960,657,682]" pageId="2" pageNumber="3">Remarks.</emphasis>
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The habitus is typical of harvestmen, especially that of the suborders Laniatores and
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<taxonomicName id="4BE44D56FFC5FFEF2F19C9E21B8CFD33" authorityName="Hansen and SOrensen" authorityYear="1904" box="[840,927,721,746]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="8" phylum="Arthropoda" rank="subOrder" subOrder="Eupnoi">Eupnoi</taxonomicName>
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. However, no distinctive morphological details such as pedipalps and tarsal claws of legs are preserved. Although most long-legged harvestmen show a typical I–IV leg length pattern (short, longest, short, long), this feature is not discernible in the specimen. A few articulations in the legs can, nonetheless, be observed. The radiation and habitus of the legs are consistent with those of harvestmen. The habitus and the thin legs are similar to those of the family
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<taxonomicName id="4BE44D56FFC4FFEE2DF0CA7A1A09FEBB" box="[417,538,329,354]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
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, Order
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<taxonomicName id="4BE44D56FFC4FFEE2E23CA7A1AC8FEBB" box="[626,731,329,354]" class="Arachnida" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="4" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
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. This family shows a body clearly divided into two tagmata, but the body of the fossil specimen, in spite of its poor preservation, seems to present fused tagmata (“compact body”). In spite of its unsatisfactory preservation, this specimen is worth reporting because of the Mesozoic fossil record of harvestmen is remarkably scarce.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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