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<document id="9F9FA0C0B6161169EC2B2C690B89DCA0" ID-CLB-Dataset="29013" ID-DOI="10.5281/zenodo.1458814" ID-GBIF-Dataset="9993ce48-70e1-4fd4-8c3b-450408aba033" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458814" ID-ZooBank="DB533B6F-A661-46C6-9118-AAF22D7C7B96" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539289999112" checkinUser="plazi" docAuthor="Dupérré, Nadine &amp; Tapia, Elicio" docDate="2018" docId="03D98E5A193AFF90E3F2FBF679050C79" docLanguage="en" docName="zootaxa.4459.3.4.pdf" docOrigin="Zootaxa 4459 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anapis shina Dupérré &amp; Tapia 2018, new species" docType="treatment" docVersion="9" lastPageNumber="499" masterDocId="FFE0F6221934FF81E365FFE77D510D39" masterDocTitle="Further discoveries on the minuscule spiders from the Chocó region of Ecuador with the description of seven new species of Anapis (Araneae: Anapidae)" masterLastPageNumber="506" masterPageNumber="482" pageNumber="496" updateTime="1698578611331" updateUser="ExternalLinkService">
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<mods:title id="90921750ACF6F81085FA2E9016C08AE9">Further discoveries on the minuscule spiders from the Chocó region of Ecuador with the description of seven new species of Anapis (Araneae: Anapidae)</mods:title>
</mods:titleInfo>
<mods:name id="397C1512F81C0F4E38F1C74126213346" type="personal">
<mods:role id="BCC42DAC0BFA1B596033D7F397F2B6F2">
<mods:roleTerm id="1B5CA4B2385739E737A5A3FAC0005A11">Author</mods:roleTerm>
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<mods:namePart id="0B2359EA06A25E8BC1A5FF506E986247">Dupérré, Nadine</mods:namePart>
</mods:name>
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<mods:roleTerm id="905A818D07635D8B43055A0DF2C219F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA2D547B2443E8095094AB59A9587A1D">Tapia, Elicio</mods:namePart>
</mods:name>
<mods:typeOfResource id="B4679537B25DD847FCCBD6CDDB28BC76">text</mods:typeOfResource>
<mods:relatedItem id="382C6EF1180DE48055913DEEE1BA8A6E" type="host">
<mods:titleInfo id="C29422A8CBA5120E0E081D45107AC657">
<mods:title id="18DD7B1BA3AC861E51ECA858E80A4DE1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="11F4EBEE863C38B009EE579DD4A87439">
<mods:date id="2A7D8B0079CD98A6BEE9635923893BA0">2018</mods:date>
<mods:detail id="23CB4D0266C693CC78E304A9BC42BE8E" type="pubDate">
<mods:number id="916B30BC8EB2220ACE3F3C45F49668B4">2018-08-16</mods:number>
</mods:detail>
<mods:detail id="44334E98DAF26734D6918CB686479868" type="volume">
<mods:number id="B4F31463AA42BB2DDBE94C0047C60156">4459</mods:number>
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<mods:classification id="A2AB81D363D7EC6F139A35EE95B9BAD7">journal article</mods:classification>
<mods:identifier id="98F04024C07AC4DA5EC814FEF3206E37" type="CLB-Dataset">29013</mods:identifier>
<mods:identifier id="C4310154A21B1BA9ED51A2CF1EC60F5D" type="DOI">10.5281/zenodo.1458814</mods:identifier>
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<treatment id="03D98E5A193AFF90E3F2FBF679050C79" ID-DOI="http://doi.org/10.5281/zenodo.5996567" ID-GBIF-Taxon="148692830" ID-Zenodo-Dep="5996567" LSID="urn:lsid:plazi:treatment:03D98E5A193AFF90E3F2FBF679050C79" httpUri="http://treatment.plazi.org/id/03D98E5A193AFF90E3F2FBF679050C79" lastPageId="17" lastPageNumber="499" pageId="14" pageNumber="496">
<subSubSection id="C36A6CC7193AFF8FE3F2FBF67CF70974" pageId="14" pageNumber="496" type="nomenclature">
<paragraph id="8BCF3F4C193AFF8FE3F2FBF67C9C0912" blockId="14.[151,461,1041,1101]" box="[151,461,1041,1067]" pageId="14" pageNumber="496">
<heading id="D0878820193AFF8FE3F2FBF67C9C0912" bold="true" box="[151,461,1041,1067]" fontSize="11" level="1" pageId="14" pageNumber="496" reason="1">
<emphasis id="B904E35E193AFF8FE3F2FBF67C9C0912" bold="true" box="[151,461,1041,1067]" pageId="14" pageNumber="496">
<taxonomicName id="4C7044CF193AFF8FE3F2FBF67C670912" ID-CoL="DFR8" authority="Dupérré &amp; Tapia, 2018" authorityName="Dupérré &amp; Tapia" authorityYear="2018" box="[151,310,1041,1067]" class="Arachnida" family="Anapidae" genus="Anapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="shina" status="sp. nov.">
<emphasis id="B904E35E193AFF8FE3F2FBF67C670912" bold="true" box="[151,310,1041,1067]" italics="true" pageId="14" pageNumber="496">Anapis shina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375E25193AFF8FE258FBF67C9C0912" box="[317,461,1041,1067]" pageId="14" pageNumber="496" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF3F4C193AFF8FE3F2FBD37CF70974" blockId="14.[151,461,1041,1101]" box="[151,422,1076,1101]" pageId="14" pageNumber="496">
<figureCitation id="134B23C9193AFF8FE3F2FBD37C460975" box="[151,279,1076,1101]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Figs 38–43</figureCitation>
,
<figureCitation id="134B23C9193AFF8FE247FBD37C3B0974" box="[290,362,1076,1101]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="14" pageNumber="496">56, 57</figureCitation>
,
<figureCitation id="134B23C9193AFF8FE213FBD37CF30974" box="[374,418,1076,1101]" captionStart="FIGURES 62–62" captionStartId="21.[151,264,1956,1979]" captionTargetBox="[202,1375,208,1917]" captionTargetId="figure@21.[185,1408,193,1930]" captionTargetPageId="21" captionText="FIGURES 62–62a. Distribution maps of Anapidae in Ecuador. 62, Distribution of previously described species and formerly known localities. 62a, Distribution of new species and new localities for previously described species." httpUri="https://zenodo.org/record/1458839/files/figure.png" pageId="14" pageNumber="496">62a</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6CC7193AFF8FE3F2FB9B79930839" pageId="14" pageNumber="496" type="materials_examined">
<paragraph id="8BCF3F4C193AFF8FE3F2FB9B79930839" blockId="14.[151,1437,1148,2036]" pageId="14" pageNumber="496">
<materialsCitation id="3B183511193AFF8FE3F2FB9B7DA809E5" ID-GBIF-Occurrence="1917413223" collectingDate="2014-11-13" collectingDateMax="2014-11-15" collectingDateMin="2014-11-13" collectionCode="QCAZ" collectorName="N. Duperre &amp; E. Tapia &amp; C. Tapia" country="Ecuador" location="BioloGical" municipality="Material" pageId="14" pageNumber="496" specimenCount="2" stateProvince="Cotopaxi ProviNcE" typeStatus="holotype">
<emphasis id="B904E35E193AFF8FE3F2FB9B7CD709AC" bold="true" box="[151,390,1148,1173]" pageId="14" pageNumber="496">
<collectingMunicipality id="6BABA536193AFF8FE3F2FB9B7DAE09AC" box="[151,255,1148,1173]" pageId="14" pageNumber="496">Material</collectingMunicipality>
examined.
</emphasis>
Male
<typeStatus id="54CB81EE193AFF8FE2BFFB9B7F6D09AC" box="[474,572,1148,1173]" pageId="14" pageNumber="496" type="holotype">holotype</typeStatus>
and two female
<typeStatus id="54CB81EE193AFF8FE06AFB9B7E2B09AD" box="[783,890,1148,1172]" pageId="14" pageNumber="496" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F3677FDC193AFF8FE0AEFB9B797B09AC" box="[971,1066,1148,1173]" name="Ecuador" pageId="14" pageNumber="496">Ecuador</collectingCountry>
,
<collectingRegion id="49B4F1AE193AFF8FE75EFB9B784709AC" box="[1083,1302,1148,1173]" country="Ecuador" name="Cotopaxi" pageId="14" pageNumber="496">Cotopaxi Province</collectingRegion>
, OTONGA
<location id="8EAF6997193AFF8FE3F2FB787C5D0981" LSID="urn:lsid:plazi:treatment:03D98E5A193AFF90E3F2FBF679050C79:8EAF6997193AFF8FE3F2FB787C5D0981" box="[151,268,1183,1208]" country="Ecuador" municipality="Material" name="BioloGical" pageId="14" pageNumber="496" stateProvince="Cotopaxi ProviNcE">Biological</location>
Reserve (-00.41994 -79.00623) 1,997m,
<collectingDate id="EF8AE064193AFF8FE184FB787ED40981" box="[737,901,1183,1208]" pageId="14" pageNumber="496" value="2014-11-13" valueMax="2014-11-15" valueMin="2014-11-13">
<date id="FFCE198C193AFF8FE184FB787ED40981" box="[737,901,1183,1208]" pageId="14" pageNumber="496" value="2014-11-13" valueMax="2014-11-15" valueMin="2014-11-13">13–15.xi.2014</date>
</collectingDate>
, sifting mosses,
<collectorName id="26855A9A193AFF8FE724FB4779950981" box="[1089,1220,1184,1208]" pageId="14" pageNumber="496">N. Dupérré</collectorName>
,
<collectorName id="26855A9A193AFF8FE7B5FB4778600981" box="[1232,1329,1183,1208]" pageId="14" pageNumber="496">E. Tapia</collectorName>
,
<collectorName id="26855A9A193AFF8FE658FB4778CD0981" box="[1341,1436,1183,1208]" pageId="14" pageNumber="496">C. Tapia</collectorName>
(
<collectionCode id="ED61A789193AFF8FE3FAFB237DA309E5" box="[159,242,1220,1244]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="14" pageNumber="496">QCAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B183511193AFF8FE263FB2379EF0839" ID-GBIF-Occurrence="1917413211" collectingDate="2014-09-04" collectingDateMax="2014-11-25" collectingDateMin="2014-09-04" collectionCode="ZMH" collectorName="N. Duperre &amp; E. Tapia &amp; C. Tapia" country="Ecuador" location="BioloGical" pageId="14" pageNumber="496" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Cotopaxi ProviNcE" typeStatus="paratype">
<typeStatus id="54CB81EE193AFF8FE263FB237C2909E5" box="[262,376,1220,1244]" pageId="14" pageNumber="496" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3677FDC193AFF8FE2E1FB237CB209E4" box="[388,483,1220,1245]" name="Ecuador" pageId="14" pageNumber="496">Ecuador</collectingCountry>
,
<collectingRegion id="49B4F1AE193AFF8FE294FB237F9909E4" box="[497,712,1220,1245]" country="Ecuador" name="Cotopaxi" pageId="14" pageNumber="496">Cotopaxi Province</collectingRegion>
, OTONGA
<location id="8EAF6997193AFF8FE031FB237E9809E4" LSID="urn:lsid:plazi:treatment:03D98E5A193AFF90E3F2FBF679050C79:8EAF6997193AFF8FE031FB237E9809E4" box="[852,969,1220,1245]" country="Ecuador" name="BioloGical" pageId="14" pageNumber="496" stateProvince="Cotopaxi ProviNcE">Biological</location>
Reserve (-00.41994 -79.00623) 1,997m,
<collectingDate id="EF8AE064193AFF8FE3F2FB007C4E0839" box="[151,287,1255,1280]" pageId="14" pageNumber="496" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">
<date id="FFCE198C193AFF8FE3F2FB007C4E0839" box="[151,287,1255,1280]" pageId="14" pageNumber="496" value="2014-09-04" valueMax="2014-09-07" valueMin="2014-09-04">4–7.ix.2014</date>
</collectingDate>
,
<specimenCount id="9D76F4C5193AFF8FE24EFB0F7C040839" box="[299,341,1256,1280]" pageId="14" pageNumber="496" type="male">2♂</specimenCount>
<specimenCount id="9D76F4C5193AFF8FE233FB0F7C230839" box="[342,370,1256,1280]" pageId="14" pageNumber="496" type="female">1♀</specimenCount>
,
<collectingDate id="EF8AE064193AFF8FE21BFB007F720839" box="[382,547,1255,1280]" pageId="14" pageNumber="496" value="2014-11-13" valueMax="2014-11-25" valueMin="2014-11-13">
<date id="FFCE198C193AFF8FE21BFB007F720839" box="[382,547,1255,1280]" pageId="14" pageNumber="496" value="2014-11-13" valueMax="2014-11-25" valueMin="2014-11-13">13–25.xi.2014</date>
</collectingDate>
,
<specimenCount id="9D76F4C5193AFF8FE14AFB0F7F050839" box="[559,596,1256,1280]" pageId="14" pageNumber="496" type="male">2♂</specimenCount>
, sifting mosses,
<collectorName id="26855A9A193AFF8FE068FB0F7EDF0839" box="[781,910,1256,1280]" pageId="14" pageNumber="496">N. Dupérré</collectorName>
,
<collectorName id="26855A9A193AFF8FE0FCFB0F7EA80839" box="[921,1017,1255,1280]" pageId="14" pageNumber="496">E. Tapia</collectorName>
,
<collectorName id="26855A9A193AFF8FE761FB0F79330839" box="[1028,1122,1255,1280]" pageId="14" pageNumber="496">C. Tapia</collectorName>
(
<collectionCode id="ED61A789193AFF8FE714FB0F79E40839" box="[1137,1205,1256,1280]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="14" pageNumber="496">ZMH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6CC7193AFF8FE3A2FAEB7826081C" box="[199,1399,1292,1317]" pageId="14" pageNumber="496" type="etymology">
<paragraph id="8BCF3F4C193AFF8FE3A2FAEB7826081C" blockId="14.[151,1437,1148,2036]" box="[199,1399,1292,1317]" pageId="14" pageNumber="496">
<emphasis id="B904E35E193AFF8FE3A2FAEB7C1F081C" bold="true" box="[199,334,1292,1317]" pageId="14" pageNumber="496">Etymology.</emphasis>
The specific name is a noun in apposition taken from the Kichwa language meaning “similar”.
</paragraph>
</subSubSection>
<subSubSection id="C36A6CC7193AFF8FE3A2FAC8790708C4" pageId="14" pageNumber="496" type="diagnosis">
<paragraph id="8BCF3F4C193AFF8FE3A2FAC8790708C4" blockId="14.[151,1437,1148,2036]" pageId="14" pageNumber="496">
<emphasis id="B904E35E193AFF8FE3A2FAC87C100871" bold="true" box="[199,321,1327,1352]" pageId="14" pageNumber="496">Diagnosis.</emphasis>
Males closely resemble
<taxonomicName id="4C7044CF193AFF8FE13BFAD77E720871" authority="Dupérré &amp; Tapia, 2018" authorityName="Dupérré &amp; Tapia" authorityYear="2018" box="[606,803,1328,1352]" class="Arachnida" family="Anapidae" genus="Anapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="mariebertheae" status="sp. nov.">
<emphasis id="B904E35E193AFF8FE13BFAD77E720871" box="[606,803,1328,1352]" italics="true" pageId="14" pageNumber="496">A. mariebertheae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375E25193AFF8FE049FAC87E3C0871" box="[812,877,1327,1352]" pageId="14" pageNumber="496" rank="species">
<emphasis id="B904E35E193AFF8FE049FAC87E3C0871" bold="true" box="[812,877,1327,1352]" pageId="14" pageNumber="496">n. sp.</emphasis>
</taxonomicNameLabel>
but can be distinguished from the latter by their higher clypeus, deeper cephalic groove (
<figureCitation id="134B23C9193AFF8FE110FAB37F810854" box="[629,720,1364,1389]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="14" pageNumber="496">Fig. 56</figureCitation>
); from all other species the male is distinguished by a dark elongated conductor and cymbial apical extension (
<figureCitation id="134B23C9193AFF8FE18DFA907E1108A9" box="[744,832,1399,1424]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Fig. 38</figureCitation>
), females most resembles
<taxonomicName id="4C7044CF193AFF8FE716FA9F7C7B088D" authority="PlatNick &amp; SHadab 1978" authorityName="PlatNick &amp; SHadab" authorityYear="1978" class="Arachnida" family="Anapidae" genus="Anapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="monteverde">
<emphasis id="B904E35E193AFF8FE716FA9F784708A9" box="[1139,1302,1400,1424]" italics="true" pageId="14" pageNumber="496">A. monteverde</emphasis>
<bibRefCitation id="EFE142BD193AFF8FE645FA907C7B088D" pageId="14" pageNumber="496" refId="ref12803" refString="Platnick, N. I. &amp; Shadab, M. U. (1978) A review of the spider genus Anapis (Araneae, Anapidae), with a dual cladistic analysis. American Museum Novitates, 2663, 1 - 23." type="journal article">Platnick &amp; Shadab 1978</bibRefCitation>
</taxonomicName>
but can be distinguished by their non-pointed abodomen (
<figureCitation id="134B23C9193AFF8FE0A0FA7B794A088C" box="[965,1051,1436,1461]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="14" pageNumber="496">Fig. 57</figureCitation>
), pointed in the latter (
<bibRefCitation id="EFE142BD193AFF8FE645FA7B7C7C08E1" pageId="14" pageNumber="496" refId="ref12803" refString="Platnick, N. I. &amp; Shadab, M. U. (1978) A review of the spider genus Anapis (Araneae, Anapidae), with a dual cladistic analysis. American Museum Novitates, 2663, 1 - 23." type="journal article">Platnick &amp; Shadab 1978</bibRefCitation>
); from all other species by their short copulatory ducts and long, sinuous fertilization ducts (
<figureCitation id="134B23C9193AFF8FE65BFA587DE608C5" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Figs 42, 43</figureCitation>
); long copulatory ducts and short fertilization ducts in
<taxonomicName id="4C7044CF193AFF8FE079FA037EE708C5" authority="Dupérré &amp; Tapia, 2018" authorityName="Dupérré &amp; Tapia" authorityYear="2018" box="[796,950,1508,1532]" class="Arachnida" family="Anapidae" genus="Anapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="496" phylum="Arthropoda" rank="species" species="anabelleae" status="sp. nov.">
<emphasis id="B904E35E193AFF8FE079FA037EE708C5" box="[796,950,1508,1532]" italics="true" pageId="14" pageNumber="496">A. anabelleae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375E25193AFF8FE0D9FA037EAA08C4" box="[956,1019,1508,1533]" pageId="14" pageNumber="496" rank="species">
<emphasis id="B904E35E193AFF8FE0D9FA037EAA08C4" bold="true" box="[956,1019,1508,1533]" pageId="14" pageNumber="496">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="134B23C9193AFF8FE76CFA03791808C4" box="[1033,1097,1508,1533]" captionStart="FIGURES 1–6" captionStartId="4.[151,264,1932,1955]" captionTargetBox="[203,1380,209,1897]" captionTargetId="figure@4.[201,1387,195,1909]" captionTargetPageId="4" captionText="FIGURES 1–6. Anapis anabelleae n. sp. 1, Male palp prolateral view. 2, Male palp retrolateral view. 2a, Male palp conductor and embolus, retrolateral view. 3, Male left metatarsus and tarsus I, prolateral view. 4, Male, left metatarsus and tarsus II, prolateral view. 5, Female epigastric region, ventral view. 6, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458816/files/figure.png" pageId="14" pageNumber="496">Fig 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6CC7193AFF90E3A2F9E07C9B0DC1" lastPageId="17" lastPageNumber="499" pageId="14" pageNumber="496" type="description">
<paragraph id="8BCF3F4C193AFF8FE3A2F9E07D810ACD" blockId="14.[151,1437,1148,2036]" pageId="14" pageNumber="496">
<emphasis id="B904E35E193AFF8FE3A2F9E07C080B19" bold="true" box="[199,345,1543,1568]" pageId="14" pageNumber="496">Description.</emphasis>
<emphasis id="B904E35E193AFF8FE200F9EF7F4E0B19" box="[357,543,1544,1568]" italics="true" pageId="14" pageNumber="496">
Male (
<typeStatus id="54CB81EE193AFF8FE2D1F9EF7F470B19" box="[436,534,1544,1568]" pageId="14" pageNumber="496" type="holotype">holotype</typeStatus>
)
</emphasis>
: Total length: 1.14; carapace length: 0.42; carapace width: 0.42; abdomen length: 0.72; abdomen width: 0.75; clypeus height: 0.1.Cephalothorax: Carapace orange; pars cephalica puntated, Y-shaped punctation reaching LE; cephalic groove deep, punctated; pars thoracica dorsally rugose, laterally punctated (
<figureCitation id="134B23C9193AFF8FE270F9937C3F0BB4" box="[277,366,1652,1677]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="14" pageNumber="496">Fig. 56</figureCitation>
). Sternum orange, suffused with dark gray, punctated, longer than wide, covered with setae. Labral spur present. Clypeus orange, punctated. Chelicerae orange, excavated medially with characteristic denticulate plate apically. Eyes: 6 eyes, rounded; AME absent, ALE separated by their diameter, ALE-PLE contiguous, LE-PME separated by their radius, PME contiguous. Abdomen: rounded, with dorsal orange scutum; soft portion whitish covered with small setose sclerites, laterally with rows of small setose sclerites slightly suffused with dark gray apically (
<figureCitation id="134B23C9193AFF8FE17BF8C07F270A79" box="[542,630,1831,1856]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="14" pageNumber="496">Fig. 56</figureCitation>
), complete spinneret scutum. Legs: Orange; metatarsus I and tarsus I without cusps (
<figureCitation id="134B23C9193AFF8FE22FF8AB7CF50A5D" box="[330,420,1868,1893]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Fig. 40</figureCitation>
); metatarsus II slightly enlarged with one ventral; tarsus II with four cusps (
<figureCitation id="134B23C9193AFF8FE650F8AB78DE0A5D" box="[1333,1423,1868,1893]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Fig. 41</figureCitation>
). Genitalia: Palpal patella with small, ventrally curved retrolateral apophysis; palpal tibia with one retrolateral trichobothrium; large scoop-shaped retrolateral apophysis (
<figureCitation id="134B23C9193AFF8FE05CF8737EC30A95" box="[825,914,1940,1965]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Fig. 39</figureCitation>
). Cymbium cup-shaped, with extension (
<figureCitation id="134B23C9193AFF8FE60CF8737DB30AE9" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Figs 38, 39</figureCitation>
). Embolus not reaching tip of conductor; conductor dark, strongly ridged, long, rounded apically (
<figureCitation id="134B23C9193AFF8FE65BF85F7D950ACD" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="14" pageNumber="496">Figs 39, 39a</figureCitation>
).
</paragraph>
<caption id="DF0F6FC4193BFF8EE3F2F89179C80AF3" httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="15" pageNumber="497" startId="15.[151,264,1910,1933]" targetBox="[234,1366,211,1878]" targetPageId="15">
<paragraph id="8BCF3F4C193BFF8EE3F2F89179C80AF3" blockId="15.[151,1435,1910,1994]" pageId="15" pageNumber="497">
<emphasis id="B904E35E193BFF8EE3F2F8917C050AB5" bold="true" box="[151,340,1910,1933]" pageId="15" pageNumber="497">FIGURES 38–43.</emphasis>
<taxonomicName id="4C7044CF193BFF8EE23CF89F7C8C0AB4" authority="Dupérré &amp; Tapia, 2018" authorityName="Dupérré &amp; Tapia" authorityYear="2018" box="[345,477,1911,1933]" class="Arachnida" family="Anapidae" genus="Anapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="497" phylum="Arthropoda" rank="species" species="shina" status="sp. nov.">
<emphasis id="B904E35E193BFF8EE23CF89F7C8C0AB4" box="[345,477,1911,1933]" italics="true" pageId="15" pageNumber="497">Anapis shina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375E25193BFF8EE286F8907F4A0AB4" box="[483,539,1911,1933]" pageId="15" pageNumber="497" rank="species">
<emphasis id="B904E35E193BFF8EE286F8907F440AB5" bold="true" box="[483,533,1911,1932]" pageId="15" pageNumber="497">n. sp</emphasis>
.
</taxonomicNameLabel>
38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view.
</paragraph>
</caption>
<caption id="DF0F6FC41924FF91E3F2F94F78C50B86" box="[151,1428,1704,1727]" httpUri="https://zenodo.org/record/1458832/files/figure.png" pageId="16" pageNumber="498" startId="16.[151,264,1704,1727]" targetBox="[179,1412,212,1653]" targetPageId="16">
<paragraph id="8BCF3F4C1924FF91E3F2F94F78C50B86" blockId="16.[151,1428,1704,1727]" box="[151,1428,1704,1727]" pageId="16" pageNumber="498">
<emphasis id="B904E35E1924FF91E3F2F94F7C020B87" bold="true" box="[151,339,1704,1727]" pageId="16" pageNumber="498">FIGURES 44–45.</emphasis>
<taxonomicName id="4C7044CF1924FF91E23FF94E7E590B86" authority="Platnick &amp; Shadab, 1979" authorityName="Platnick &amp; Shadab" authorityYear="1979" box="[346,776,1705,1727]" class="Arachnida" family="Anapidae" genus="Anapisona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="498" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B904E35E1924FF91E23FF94E7F500B86" box="[346,513,1705,1727]" italics="true" pageId="16" pageNumber="498">Anapisona pecki</emphasis>
Platnick &amp; Shadab, 1979
</taxonomicName>
. 44, Male palp retrolateral view. 45, Male palp retrolateral view.
</paragraph>
</caption>
<paragraph id="8BCF3F4C1924FF90E3A2F9017C9B0DC1" blockId="16.[151,1436,1766,2007]" lastBlockId="17.[151,1437,151,320]" lastPageId="17" lastPageNumber="499" pageId="16" pageNumber="498">
<emphasis id="B904E35E1924FF91E3A2F9017CCD0BC7" box="[199,412,1766,1790]" italics="true" pageId="16" pageNumber="498">
Female (
<typeStatus id="54CB81EE1924FF91E248F9007CC20BC7" box="[301,403,1767,1790]" pageId="16" pageNumber="498" type="paratype">paratype</typeStatus>
)
</emphasis>
: Total length: 1.22; carapace length: 0.45; carapace width: 0.41; abdomen length: 0.77; abdomen width: 0.63. Cephalothorax: Same coloration and pattern of punctation as male (
<figureCitation id="134B23C91924FF91E7EEF8ED79B00A1A" box="[1163,1249,1802,1827]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="16" pageNumber="498">Fig. 57</figureCitation>
). Chelicerae and sternum as male. Labral spur present. Eyes: 6 eyes, AME absent, ALE separated by their diameter, ALE-PLE contiguous, LE-PME separated by half their diameter, PME contiguous. Abdomen: Light gray, rounded without dorsal scutum; dorsal soft portion with small, rounded setose sclerites; eight large non setose sclerites; dorsolaterally with six-seven large non setose sclerites; laterally with numerous small, rounded setose sclerites, slightly suffused with gray apically (
<figureCitation id="134B23C91924FF91E2BCF8597F7E0AEE" box="[473,559,1982,2007]" captionStart="FIGURES 54–61" captionStartId="20.[151,264,1897,1920]" captionTargetBox="[230,1363,233,1851]" captionTargetId="figure@20.[202,1387,193,1868]" captionTargetPageId="20" captionText="FIGURES 54–61. Habitus, lateral view. 54, Male Anapis nawchi n. sp. 55, Female Anapis nawchi n. sp. 56, Male Anapis shina n. sp. 57, Female Anapis shina n. sp. 58, Male Anapis chiriboga Platnick &amp; Shadab, 1978. 59, Male Anapisona pecki Platnick &amp; Shadab, 1979. 60, Male Pseudanapis domingo Platnick &amp; Shadab, 1979. 61, Female Pseudanapis domingo Platnick &amp; Shadab, 1979. Scale bars: 0.5mm" httpUri="https://zenodo.org/record/1458837/files/figure.png" pageId="16" pageNumber="498">Fig. 57</figureCitation>
); spinneret scutum complete. Legs: Orange; without cusps. Genitalia: Ventral scutum bright orange, visible through the scutum: a pair of large rounded spermathecae, straight copulatory ducts (
<figureCitation id="134B23C91925FF90E3FBFF5B7DA70DED" box="[158,246,188,213]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="17" pageNumber="499">Fig. 42</figureCitation>
). Internal genitalia with large rounded spermathecae; copulatory ducts short; fertilization ducts sinuous, directed medially (
<figureCitation id="134B23C91925FF90E20DFF387CEF0DC1" box="[360,446,223,248]" captionStart="FIGURES 38–43" captionStartId="15.[151,264,1910,1933]" captionTargetBox="[234,1366,211,1878]" captionTargetId="figure@15.[219,1368,193,1889]" captionTargetPageId="15" captionText="FIGURES 38–43. Anapis shina n. sp. 38, Male palp prolateral view. 39, Male palp retrolateral view. 39a. Male palp conductor and embolus, retrolateral view. 40, Male left metatarsus and tarsus I, prolateral view. 41, Male, left metatarsus and tarsus II, prolateral view. 42, Female epigastric region, ventral view. 43, Female internal genitalia, dorsal view." httpUri="https://zenodo.org/record/1458830/files/figure.png" pageId="17" pageNumber="499">Fig. 43</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6CC71925FF90E3A2FEE379050C79" pageId="17" pageNumber="499" type="distribution">
<paragraph id="8BCF3F4C1925FF90E3A2FEE37E0C0C24" blockId="17.[151,1437,151,320]" box="[199,861,260,285]" pageId="17" pageNumber="499">
<emphasis id="B904E35E1925FF90E3A2FEE37C310C24" bold="true" box="[199,352,260,285]" pageId="17" pageNumber="499">Distribution.</emphasis>
<collectingCountry id="F3677FDC1925FF90E202FEE37C980C24" box="[359,457,260,285]" name="Ecuador" pageId="17" pageNumber="499">Ecuador</collectingCountry>
: Known only from the
<typeStatus id="54CB81EE1925FF90E1A9FEE27FAD0C24" box="[716,764,261,285]" pageId="17" pageNumber="499">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BCF3F4C1925FF90E3A2FEC079050C79" blockId="17.[151,1437,151,320]" box="[199,1108,295,320]" pageId="17" pageNumber="499">
<emphasis id="B904E35E1925FF90E3A2FEC07CDA0C79" bold="true" box="[199,395,295,320]" pageId="17" pageNumber="499">Natural History.</emphasis>
All specimens were taken at an elevation of 1,997m in mosses.
</paragraph>
</subSubSection>
</treatment>
</document>
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