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<document id="43792F46C3FB2685477599DE74CA978A" ID-CLB-Dataset="26191" ID-DOI="10.5852/ejt.2019.535" ID-GBIF-Dataset="dccc66ae-685c-4944-81ed-d0be51d7f702" ID-Zenodo-Dep="3338044" ID-ZooBank="27320418-4683-4C38-A0BC-31CC9D408428" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563280090768" checkinUser="plazi" docAuthor="Kundrata, Robin, Németh, Tamás &amp; Jarzabek-Müller, Andrea" docDate="2019" docId="C41887B51C4BA31BFD5EFEA9FE09FB39" docLanguage="en" docName="ejt-535_kundrata_nemeth_jarzabek-mueller.pdf.imf" docOrigin="European Journal of Taxonomy 535" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lacon mertliki Kundrata &amp; Németh &amp; Jarzabek-Müller 2019, sp. nov." docType="treatment" docUuid="168C9BE1-BC15-411C-9614-15303E8CB8C4" docUuidSource="ZooBank" docVersion="10" lastPageNumber="8" masterDocId="3821FFCD1C49A31CFFCEFFF1FF84FFF3" masterDocTitle="Description of Lacon mertliki sp. nov. (Coleoptera: Elateridae: Agrypninae) from the Hyrcanian forest ecoregion, with a key to the Lacon species of Iran" masterLastPageNumber="13" masterPageNumber="1" pageNumber="3" updateTime="1698745625226" updateUser="ExternalLinkService">
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<mods:title id="79951DB1F48609B1DA02184752D2E19E">Description of Lacon mertliki sp. nov. (Coleoptera: Elateridae: Agrypninae) from the Hyrcanian forest ecoregion, with a key to the Lacon species of Iran</mods:title>
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<mods:title id="B53D006520CFFDBC815EAFD28B2E97F3">European Journal of Taxonomy</mods:title>
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<heading id="174681CF1C4BA31EFD5EFEA9FC20FE81" box="[656,932,343,370]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="8BB14D201C4BA31EFD5EFEA9FCC7FE81" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[656,835,343,370]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFD5EFEA9FCC7FE81" bold="true" box="[656,835,343,370]" italics="true" pageId="2" pageNumber="3">Lacon mertliki</emphasis>
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<paragraph id="4C0E36A31C4BA31EFD9EFE6FFC60FE4B" blockId="2.[370,1217,343,440]" box="[592,996,414,440]" pageId="2" pageNumber="3">
<figureCitation id="D48A2A261C4BA31EFD9EFE6FFD10FE4B" box="[592,660,414,440]" captionStart="Fig" captionStartId="3.[189,232,1872,1898]" captionTargetBox="[220,1398,265,1822]" captionTargetId="figure@3.[189,1398,265,1830]" captionTargetPageId="3" captionText="Fig. 1. Habitus images of Lacon spp. A–C. L. mertliki sp. nov, holotype, ♂, in dorsal, ventral and lateral view, respectively. D. L. mertliki sp. nov, paratype, ♀, dorsal view. E–F. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)) in dorsal and lateral view, respectively. G–H. L. lepidopterus, ♀ from Turkey in dorsal and lateral view, respectively. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/3338046/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
A–D, 2A–B, E–H, K, 3B–G
</paragraph>
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<emphasis id="7EC5EAB11C4BA31EFF73FE2EFEB2FE0A" bold="true" box="[189,310,479,505]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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<taxonomicName id="8BB14D201C4BA31EFF73FDFEFEEAFDDB" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[189,366,526,552]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFF73FDFEFEEAFDDB" box="[189,366,526,552]" italics="true" pageId="2" pageNumber="3">Lacon mertliki</emphasis>
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is morphologically similar to 
<emphasis id="7EC5EAB11C4BA31EFC9CFDFEFB96FDDB" box="[850,1042,526,552]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="8BB14D201C4BA31EFC9CFDFEFB89FDDB" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[850,1037,526,552]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lepidopterus">L. lepidopterus</taxonomicName>
.
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Both species share the body shape and size, 
<typeStatus id="930A88011C4BA31EFEB7FDC3FE28FDBF" box="[377,428,562,588]" pageId="2" pageNumber="3">type</typeStatus>
of pubescence, short antennae, pronotal disc with tubercles and impressions, and the similar shape of genitalia. They differ in the coloration of body (reddish brown to dark brown in 
<taxonomicName id="8BB14D201C4BA31EFF73FD89FEF5FD62" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[189,369,631,657]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4BA31EFF73FD89FEF5FD62" box="[189,369,631,657]" italics="true" pageId="2" pageNumber="3">L. lepidopterus</emphasis>
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, black in 
<taxonomicName id="8BB14D201C4BA31EFE24FD89FDE3FD62" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[490,615,631,657]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFE24FD89FDE3FD62" box="[490,615,631,657]" italics="true" pageId="2" pageNumber="3">L. mertliki</emphasis>
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<taxonomicNameLabel id="65F657CA1C4BA31EFDBEFD89FD4AFD61" box="[624,718,632,658]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
) and pubescence (usually uniformly golden and black in 
<taxonomicName id="8BB14D201C4BA31EFF73FD6DFEF6FD46" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[189,370,667,693]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4BA31EFF73FD6DFEF6FD46" box="[189,370,667,693]" italics="true" pageId="2" pageNumber="3">L. lepidopterus</emphasis>
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, black and silver/yellowish with some white setae in 
<taxonomicName id="8BB14D201C4BA31EFC36FD6DFBF0FD46" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[1016,1140,667,693]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFC36FD6DFBF0FD46" box="[1016,1140,667,693]" italics="true" pageId="2" pageNumber="3">L. mertliki</emphasis>
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<taxonomicNameLabel id="65F657CA1C4BA31EFBB3FD6AFB5AFD46" box="[1149,1246,667,693]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
; 
<figureCitation id="D48A2A261C4BA31EFB25FD6AFAB4FD46" box="[1259,1328,667,693]" captionStart="Fig" captionStartId="3.[189,232,1872,1898]" captionTargetBox="[220,1398,265,1822]" captionTargetId="figure@3.[189,1398,265,1830]" captionTargetPageId="3" captionText="Fig. 1. Habitus images of Lacon spp. A–C. L. mertliki sp. nov, holotype, ♂, in dorsal, ventral and lateral view, respectively. D. L. mertliki sp. nov, paratype, ♀, dorsal view. E–F. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)) in dorsal and lateral view, respectively. G–H. L. lepidopterus, ♀ from Turkey in dorsal and lateral view, respectively. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/3338046/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
A–H, 4A), the shape of pronotum (posterior angles more divergent in 
<taxonomicName id="8BB14D201C4BA31EFC02FD4EFB06FD2B" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[972,1154,702,728]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4BA31EFC02FD4EFB06FD2B" box="[972,1154,702,728]" italics="true" pageId="2" pageNumber="3">L. lepidopterus</emphasis>
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, especially in male; 
<figureCitation id="D48A2A261C4BA31EFF73FD10FE8DFD08" box="[189,265,737,763]" captionStart="Fig" captionStartId="4.[189,232,1768,1794]" captionTargetBox="[228,1355,265,1719]" captionTargetId="figure@4.[228,1359,265,1728]" captionTargetPageId="4" captionText="Fig. 2. Details of pronotum, antenna, scutellar shield and elytral surface of Lacon spp. A–D. Pronotum, dorsal view. A. L. mertliki sp. nov., holotype, ♂. B. L. mertliki sp. nov., paratype, ♀. C. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)). D. L. lepidopterus, ♂ from Turkey. E–F. Antenna of L. mertliki sp. nov. E. Holotype, ♂. F. Paratype, ♀. G–J. Scutellar shield. G. L. mertliki sp. nov., holotype, ♂. H. L. mertliki sp. nov., paratype, ♀. I. L. lepidopterus, ♂. J. L. lepidopterus, ♀. K–L. Detail of elytral surface. K. L. mertliki sp. nov., holotype, ♂. L. L. lepidopterus, ♂ (lectotype of L. aurosquamosus). Scale bars: A–F = 1.0 mm; G–J = 0.5 mm; K–L = not to scale." httpUri="https://zenodo.org/record/3338048/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
A–D), scutellar shield (relatively narrower in 
<taxonomicName id="8BB14D201C4BA31EFC80FD13FB8CFD08" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[846,1032,737,763]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4BA31EFC80FD13FB8CFD08" box="[846,1032,737,763]" italics="true" pageId="2" pageNumber="3">L. lepidopterus</emphasis>
</taxonomicName>
; 
<figureCitation id="D48A2A261C4BA31EFBD0FD10FBEEFD08" box="[1054,1130,737,763]" captionStart="Fig" captionStartId="4.[189,232,1768,1794]" captionTargetBox="[228,1355,265,1719]" captionTargetId="figure@4.[228,1359,265,1728]" captionTargetPageId="4" captionText="Fig. 2. Details of pronotum, antenna, scutellar shield and elytral surface of Lacon spp. A–D. Pronotum, dorsal view. A. L. mertliki sp. nov., holotype, ♂. B. L. mertliki sp. nov., paratype, ♀. C. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)). D. L. lepidopterus, ♂ from Turkey. E–F. Antenna of L. mertliki sp. nov. E. Holotype, ♂. F. Paratype, ♀. G–J. Scutellar shield. G. L. mertliki sp. nov., holotype, ♂. H. L. mertliki sp. nov., paratype, ♀. I. L. lepidopterus, ♂. J. L. lepidopterus, ♀. K–L. Detail of elytral surface. K. L. mertliki sp. nov., holotype, ♂. L. L. lepidopterus, ♂ (lectotype of L. aurosquamosus). Scale bars: A–F = 1.0 mm; G–J = 0.5 mm; K–L = not to scale." httpUri="https://zenodo.org/record/3338048/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
G–J), male parameral apical lobes (
<figureCitation id="D48A2A261C4BA31EFEAFFCF5FE2DFCED" box="[353,425,772,798]" captionStart="Fig" captionStartId="5.[189,232,1851,1877]" captionTargetBox="[193,1395,1314,1803]" captionTargetId="figure@5.[189,1398,1310,1812]" captionTargetPageId="5" captionText="Fig. 3. Pregenital segments and genitalia of Lacon spp. A. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)), aedeagus, dorsal view. B–C. L. mertliki sp. nov., holotype, ♂. B. Aedeagus, dorsal view. C. Abdominal terminal segments. D–G. L. mertliki sp. nov., paratype, ♀. D. Abdominal tergite VIII. E. Abdominal sternite VIII. F. Ovipositor. G. Large sclerite of bursa copulatrix. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/3338050/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
A–B) and the spines of the large sclerite of bursa copulatrix (
<figureCitation id="D48A2A261C4BA31EFB6AFCF5FA87FCED" box="[1188,1283,772,798]" captionStart="Fig" captionStartId="5.[189,232,1851,1877]" captionTargetBox="[193,1395,1314,1803]" captionTargetId="figure@5.[189,1398,1310,1812]" captionTargetPageId="5" captionText="Fig. 3. Pregenital segments and genitalia of Lacon spp. A. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)), aedeagus, dorsal view. B–C. L. mertliki sp. nov., holotype, ♂. B. Aedeagus, dorsal view. C. Abdominal terminal segments. D–G. L. mertliki sp. nov., paratype, ♀. D. Abdominal tergite VIII. E. Abdominal sternite VIII. F. Ovipositor. G. Large sclerite of bursa copulatrix. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/3338050/files/figure.png" pageId="2" pageNumber="3">Fig. 3G</figureCitation>
here and fig. 
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<bibRefCitation id="28204B521C4BA31EFEF5FCD6FD8DFCB1" author="Prosvirov A. S." box="[315,521,807,834]" firstAuthor="Prosvirov" journalOrPublisher="Zootaxa" pageId="2" pageNumber="3" pagination="279 - 296" part="4168" refId="ref6264" refString="Prosvirov A. S. 2016 b. New and little-known species of the genus Lacon Laporte, 1838 (Coleoptera: Elateridae) of Afghanistan and adjacent countries. Zootaxa 4168: 279 - 296. https: // doi. org / 10.11646 / zootaxa. 4168.2.3" title="New and little-known species of the genus Lacon Laporte, 1838 (Coleoptera: Elateridae) of Afghanistan and adjacent countries" type="journal article" year="2016" yearSuffix="b">Prosvirov 2016b</bibRefCitation>
). 
<taxonomicName id="8BB14D201C4BA31EFDECFCD9FD6FFCB2" baseAuthorityName="Herbst" baseAuthorityYear="1779" box="[546,747,808,833]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="7EC5EAB11C4BA31EFDECFCD9FD6FFCB2" box="[546,747,808,833]" italics="true" pageId="2" pageNumber="3">Lacon punctatus</emphasis>
</taxonomicName>
shares with 
<taxonomicName id="8BB14D201C4BA31EFC43FCD9FB89FCB2" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[909,1037,807,833]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFC43FCD9FB89FCB2" box="[909,1037,807,833]" italics="true" pageId="2" pageNumber="3">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4BA31EFBD4FCD9FBFEFCB1" box="[1050,1146,808,834]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
the body shape and size, black coloration and scale-like body pubescence, but it differs in black and white setae, slightly longer antennae, the shape of pronotal disc (longer than wide, usually widest at posterior angles, and with posterior angles more divergent in 
<taxonomicName id="8BB14D201C4BA31EFD60FC63FCC0FC58" baseAuthorityName="Herbst" baseAuthorityYear="1779" box="[686,836,914,939]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="7EC5EAB11C4BA31EFD60FC63FCC0FC58" box="[686,836,914,939]" italics="true" pageId="2" pageNumber="3">L. punctatus</emphasis>
</taxonomicName>
; slightly wider than long, widest at half, and with posterior angles less divergent in 
<taxonomicName id="8BB14D201C4BA31EFD5DFC44FC95FC3D" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[659,785,948,974]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFD5DFC44FC95FC3D" box="[659,785,948,974]" italics="true" pageId="2" pageNumber="3">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4BA31EFCD5FC44FCF8FC3C" box="[795,892,949,975]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
), aedeagus (apical lobe of paramere more subparallel-sided and with more rounded apex in 
<taxonomicName id="8BB14D201C4BA31EFD32FC29FC08FC02" baseAuthorityName="Herbst" baseAuthorityYear="1779" box="[764,908,984,1009]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="7EC5EAB11C4BA31EFD32FC29FC08FC02" box="[764,908,984,1009]" italics="true" pageId="2" pageNumber="3">L. punctatus</emphasis>
</taxonomicName>
), and the large sclerite of bursa copulatrix (thinner sclerite in 
<taxonomicName id="8BB14D201C4BA31EFE64FC0DFDC5FBE6" baseAuthorityName="Herbst" baseAuthorityYear="1779" box="[426,577,1020,1045]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="7EC5EAB11C4BA31EFE64FC0DFDC5FBE6" box="[426,577,1020,1045]" italics="true" pageId="2" pageNumber="3">L. punctatus</emphasis>
</taxonomicName>
). Remaining 
<taxonomicName id="8BB14D201C4BA31EFD25FC0DFCB1FBE6" authorityName="Laporte" authorityYear="1838" box="[747,821,1020,1045]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7EC5EAB11C4BA31EFD25FC0DFCB1FBE6" box="[747,821,1020,1045]" italics="true" pageId="2" pageNumber="3">Lacon</emphasis>
</taxonomicName>
species from 
<collectingCountry id="34A676331C4BA31EFC29FC0AFB93FBE6" box="[999,1047,1019,1045]" name="Iran" pageId="2" pageNumber="3">Iran</collectingCountry>
have thinner, hair-like body pubescence, lack distinct tubercles and impressions on pronotal disc, and all also differ in the male and female genitalia. Additionally, 
<taxonomicName id="8BB14D201C4BA31EFDE3FBB3FD69FBA8" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[557,749,1089,1115]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="7EC5EAB11C4BA31EFDE3FBB3FD69FBA8" box="[557,749,1089,1115]" italics="true" pageId="2" pageNumber="3">Lacon modestus</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4BA31EFCE6FBB3FC2DFBA8" baseAuthorityName="Candeze" baseAuthorityYear="1889" box="[808,937,1089,1115]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mekrani">
<emphasis id="7EC5EAB11C4BA31EFCE6FBB3FC2DFBA8" box="[808,937,1089,1115]" italics="true" pageId="2" pageNumber="3">L. mekrani</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="8BB14D201C4BA31EFBF2FBB3FB3CFBA8" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[1084,1208,1089,1115]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4BA31EFBF2FBB3FB3CFBA8" box="[1084,1208,1089,1115]" italics="true" pageId="2" pageNumber="3">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4BA31EFB0FFBB3FA99FBAF" box="[1217,1309,1090,1116]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
mainly in having a shiny, lighter body with sparser punctation, distinct sublateral carina on pronotal posterior angles, and prominent oblique keels on elytra near scutellar shield covered with thin setae, 
<taxonomicName id="8BB14D201C4BA31EFB3DFB79FAF3FB52" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[1267,1399,1159,1185]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4BA31EFB3DFB79FAF3FB52" box="[1267,1399,1159,1185]" italics="true" pageId="2" pageNumber="3">L. unicolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4BA31EFF3EFB5DFEDFFB36" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[240,347,1195,1221]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4BA31EFF3EFB5DFEDFFB36" box="[240,347,1195,1221]" italics="true" pageId="2" pageNumber="3">L. nadaii</emphasis>
</taxonomicName>
in having more elongate pronotum, distinctly longer than wide, and 
<taxonomicName id="8BB14D201C4BA31EFB43FB5DFA8AFB36" baseAuthorityName="Solsky" baseAuthorityYear="1881" box="[1165,1294,1195,1221]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="7EC5EAB11C4BA31EFB43FB5DFA8AFB36" box="[1165,1294,1195,1221]" italics="true" pageId="2" pageNumber="3">L. funebris</emphasis>
</taxonomicName>
in much sparser and smaller pronotal punctation.
</paragraph>
</subSubSection>
<subSubSection id="04AB65281C4BA31EFF73FAE1FAD4FAAA" box="[189,1360,1296,1369]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="4C0E36A31C4BA31EFF73FAE1FEC0FAD9" blockId="2.[189,324,1296,1322]" box="[189,324,1296,1322]" pageId="2" pageNumber="3">
<heading id="174681CF1C4BA31EFF73FAE1FEC0FAD9" bold="true" box="[189,324,1296,1322]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="7EC5EAB11C4BA31EFF73FAE1FEC0FAD9" bold="true" box="[189,324,1296,1322]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF73FACFFAD4FAAA" blockId="2.[189,1360,1342,1369]" box="[189,1360,1342,1369]" pageId="2" pageNumber="3">
The species name is a patronym in honor of Josef Mertlík (Opatovice nad Labem, 
<collectingCountry id="34A676331C4BA31EFB4AFACFFAC0FAAA" box="[1156,1348,1342,1369]" name="Czech Republic" pageId="2" pageNumber="3">Czech Republic</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="04AB65281C4BA31EFF73FA8EFAE6F92E" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="4C0E36A31C4BA31EFF73FA8EFEE8FA6A" blockId="2.[189,364,1407,1433]" box="[189,364,1407,1433]" pageId="2" pageNumber="3">
<heading id="174681CF1C4BA31EFF73FA8EFEE8FA6A" bold="true" box="[189,364,1407,1433]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="7EC5EAB11C4BA31EFF73FA8EFEE8FA6A" bold="true" box="[189,364,1407,1433]" pageId="2" pageNumber="3">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF24FA5FFED8FA3B" blockId="2.[189,1399,1454,1516]" box="[234,348,1454,1480]" pageId="2" pageNumber="3">
<emphasis id="7EC5EAB11C4BA31EFF24FA5FFED8FA3B" bold="true" box="[234,348,1454,1480]" pageId="2" pageNumber="3">
<typeStatus id="930A88011C4BA31EFF24FA5FFED8FA3B" box="[234,348,1454,1480]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF73FA20FAF3FA1F" blockId="2.[189,1399,1454,1516]" box="[189,1399,1488,1516]" pageId="2" pageNumber="3">
<materialsCitation id="FCD93CFE1C4BA31EFF73FA20FAF3FA1F" ID-GBIF-Occurrence="2283905919" box="[189,1399,1488,1516]" collectionCode="NHMW" collectorName="Col. Hauser" country="Iran" elevation="1160" location="Golestan" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Golestan" typeStatus="holotype">
<collectingCountry id="34A676331C4BA31EFF73FA20FE81FA18" box="[189,261,1489,1515]" name="Iran" pageId="2" pageNumber="3">IRAN</collectingCountry>
• 
<specimenCount id="5AB7FD2A1C4BA31EFED0FA23FEB3FA1F" box="[286,311,1490,1516]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
; “Pers. Kopet-Dagh., Siaret [= Iran, 
<collectingRegion id="8E75F8411C4BA31EFD2DFA20FCD4FA18" box="[739,848,1489,1515]" country="Iran" name="Golestan" pageId="2" pageNumber="3">Golestan</collectingRegion>
: Ziarat], 
<quantity id="8B499B461C4BA31EFC77FA20FB92FA1F" box="[953,1046,1489,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="3" unit="m" value="1160.0">
<elevation id="C79CD1901C4BA31EFC77FA20FB92FA1F" box="[953,1046,1489,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="3" unit="m" value="1160.0">1160 m</elevation>
</quantity>
., 5.99. 
<collectorName id="E14453751C4BA31EFBA7FA20FB7FFA18" box="[1129,1275,1489,1515]" pageId="2" pageNumber="3">Col. Hauser</collectorName>
”; 
<collectionCode id="2AA0AE661C4BA31EFADCFA23FAF7FA1F" box="[1298,1395,1490,1516]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3" type="Museum">NHMW</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF24F9E2FEE3F9DE" blockId="2.[189,1399,1555,1757]" box="[234,359,1555,1581]" pageId="2" pageNumber="3">
<emphasis id="7EC5EAB11C4BA31EFF24F9E2FEE3F9DE" bold="true" box="[234,359,1555,1581]" pageId="2" pageNumber="3">
<typeStatus id="930A88011C4BA31EFF24F9E2FEE3F9DE" box="[234,359,1555,1581]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF73F9C7FAE6F92E" blockId="2.[189,1399,1555,1757]" pageId="2" pageNumber="3">
<materialsCitation id="FCD93CFE1C4BA31EFF73F9C7FC8EF9A3" ID-GBIF-Occurrence="2283905916" box="[189,778,1590,1616]" collectionCode="NHMW" collectorName="Col. Hauser" country="Iran" elevation="1160" location="Golestan" pageId="2" pageNumber="3" specimenCount="2" specimenCount-female="2" stateProvince="Golestan" typeStatus="paratype">
<collectingCountry id="34A676331C4BA31EFF73F9C7FE80F9A3" box="[189,260,1590,1616]" name="Iran" pageId="2" pageNumber="3">IRAN</collectingCountry>
• 
<specimenCount id="5AB7FD2A1C4BA31EFED0F9C7FEE5F9A3" box="[286,353,1590,1616]" pageId="2" pageNumber="3" type="female">2 ♀♀</specimenCount>
; same data as for holotype; 
<collectionCode id="2AA0AE661C4BA31EFD65F9C7FC8EF9A3" box="[683,778,1590,1616]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3" type="Museum">NHMW</collectionCode>
</materialsCitation>
• 
<materialsCitation id="FCD93CFE1C4BA31EFCEDF9C7FD93F980" ID-GBIF-Occurrence="2283905915" collectionCode="NHMW" collectorName="Coll. Hauser" country="Iran" county="Persia" location="Iran" municipality="Astrabad." pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Golestan" typeStatus="paratype">
<specimenCount id="5AB7FD2A1C4BA31EFCEDF9C7FCD5F9A3" box="[803,849,1590,1616]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
; “ 
<collectingCounty id="A56F4E2F1C4BA31EFCA4F9C7FC32F9A3" box="[874,950,1590,1616]" pageId="2" pageNumber="3">Persia</collectingCounty>
, 
<collectingMunicipality id="AC6AACD91C4BA31EFC0FF9C7FBB5F9A3" box="[961,1073,1590,1616]" pageId="2" pageNumber="3">Astrabad.</collectingMunicipality>
[= 
<location id="496E60781C4BA31EFB94F9C7FB0FF9A3" LSID="urn:lsid:plazi:treatment:C41887B51C4BA31BFD5EFEA9FE09FB39:496E60781C4BA31EFB94F9C7FB0FF9A3" box="[1114,1163,1590,1616]" country="Iran" county="Persia" municipality="Astrabad." name="Iran" pageId="2" pageNumber="3" stateProvince="Golestan">Iran</location>
, 
<collectingRegion id="8E75F8411C4BA31EFB56F9C7FA80F9A3" box="[1176,1284,1590,1616]" country="Iran" name="Golestan" pageId="2" pageNumber="3">Golestan</collectingRegion>
: Gorgan], 5.99. 
<collectorName id="E14453751C4BA31EFECCF9A8FE1AF980" box="[258,414,1625,1651]" pageId="2" pageNumber="3">Coll. Hauser</collectorName>
”; 
<collectionCode id="2AA0AE661C4BA31EFE76F9A8FD93F980" box="[440,535,1625,1651]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="3" type="Museum">NHMW</collectionCode>
</materialsCitation>
• 
<materialsCitation id="FCD93CFE1C4BA31EFDF6F9A8FD67F965" ID-GBIF-Occurrence="2283905918" collectingDate="2016-10-05" collectionCode="PCAJM" collectorName="J. Muller" country="Iran" elevation="494" latitude="37.39472" location="Tangrah" longLatPrecision="1" longitude="55.79771" municipality="Iran" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Golestan" typeStatus="paratype">
<specimenCount id="5AB7FD2A1C4BA31EFDF6F9A8FDECF980" box="[568,616,1625,1651]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
; “ 
<collectingMunicipality id="AC6AACD91C4BA31EFD4BF9A8FD3DF980" box="[645,697,1625,1651]" pageId="2" pageNumber="3">Iran</collectingMunicipality>
, 
<collectingRegion id="8E75F8411C4BA31EFD06F9A8FCFEF980" box="[712,890,1625,1651]" country="Iran" name="Golestan" pageId="2" pageNumber="3">Golestan Prov.</collectingRegion>
, near 
<location id="496E60781C4BA31EFC0BF9A8FBA3F980" LSID="urn:lsid:plazi:treatment:C41887B51C4BA31BFD5EFEA9FE09FB39:496E60781C4BA31EFC0BF9A8FBA3F980" box="[965,1063,1625,1651]" country="Iran" latitude="37.39472" longLatPrecision="1" longitude="55.79771" municipality="Iran" name="Tangrah" pageId="2" pageNumber="3" stateProvince="Golestan">Tangrah</location>
, 
<elevation id="C79CD1901C4BA31EFBF8F9A8FB42F980" box="[1078,1222,1625,1651]" metricMagnitude="2" metricUnit="m" metricValue="4.9399999999999995" pageId="2" pageNumber="3" unit="m" value="494.0">
<quantity id="8B499B461C4BA31EFBF8F9A8FB00F980" box="[1078,1156,1625,1651]" metricMagnitude="2" metricUnit="m" metricValue="4.9399999999999995" pageId="2" pageNumber="3" unit="m" value="494.0">494 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="298550641C4BA31EFB19F9A8FAF6F980" box="[1239,1394,1625,1651]" degrees="37.39472" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="1" value="37.39472">37.39472° N</geoCoordinate>
, 
<geoCoordinate id="298550641C4BA31EFF73F98DFECBF964" box="[189,335,1660,1687]" degrees="55.79771" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="1" value="55.79771">55.79771° E</geoCoordinate>
, 
<date id="380F10631C4BA31EFE94F98DFE4DF964" box="[346,457,1660,1687]" pageId="2" pageNumber="3" value="2016-10-05">
<collectingDate id="284BE98B1C4BA31EFE94F98DFE4DF964" box="[346,457,1660,1687]" pageId="2" pageNumber="3" value="2016-10-05">5.X.2016</collectingDate>
</date>
. leg. 
<collectorName id="E14453751C4BA31EFDCDF98CFDF5F965" box="[515,625,1660,1687]" pageId="2" pageNumber="3">J. Müller</collectorName>
”; 
<collectionCode id="2AA0AE661C4BA31EFD4BF98DFD67F965" box="[645,739,1660,1686]" pageId="2" pageNumber="3">PCAJM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="FCD93CFE1C4BA31EFD37F98DFCA7F94A" ID-GBIF-Occurrence="2283905920" collectingDate="2016-10-06" collectionCode="PCFL" collectorName="F. Lange" country="Iran" elevation="1200" location="Iran" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Mazandaran" typeStatus="paratype">
<specimenCount id="5AB7FD2A1C4BA31EFD37F98DFCA0F964" box="[761,804,1660,1687]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
; “ 
<location id="496E60781C4BA31EFCF5F98DFCEBF965" LSID="urn:lsid:plazi:treatment:C41887B51C4BA31BFD5EFEA9FE09FB39:496E60781C4BA31EFCF5F98DFCEBF965" box="[827,879,1660,1686]" country="Iran" name="Iran" pageId="2" pageNumber="3" stateProvince="Mazandaran">Iran</location>
, 
<collectingRegion id="8E75F8411C4BA31EFCB7F98DFBCAF964" box="[889,1102,1660,1687]" country="Iran" name="Mazandaran" pageId="2" pageNumber="3">Mazandaran Prov.</collectingRegion>
, S Galugah, 
<elevation id="C79CD1901C4BA31EFB13F98DFAF5F965" box="[1245,1393,1660,1687]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="1200.0">
<quantity id="8B499B461C4BA31EFB13F98DFAB0F964" box="[1245,1332,1660,1687]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="3" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
, in brown rotting log, 
<date id="380F10631C4BA31EFE7DF96EFDA7F94A" box="[435,547,1695,1721]" pageId="2" pageNumber="3" value="2016-10-06">
<collectingDate id="284BE98B1C4BA31EFE7DF96EFDA7F94A" box="[435,547,1695,1721]" pageId="2" pageNumber="3" value="2016-10-06">6.X.2016</collectingDate>
</date>
. leg. 
<collectorName id="E14453751C4BA31EFD91F951FD4DF94A" box="[607,713,1695,1722]" pageId="2" pageNumber="3">F. Lange</collectorName>
”; 
<collectionCode id="2AA0AE661C4BA31EFD11F96EFCA7F94A" box="[735,803,1695,1721]" pageId="2" pageNumber="3">PCFL</collectionCode>
</materialsCitation>
• 
<materialsCitation id="FCD93CFE1C4BA31EFCF2F96EFADBF92E" ID-GBIF-Occurrence="2283905917" collectingDate="2018-07" collectionCode="PCAJM" collectorName="F. Kazerani" country="Iran" county="Iran" elevation="1138" latitude="36.652" location="Natural forest" longLatPrecision="70" longitude="53.841" municipality="Shafarood forest" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Gilan" typeStatus="paratype">
<specimenCount id="5AB7FD2A1C4BA31EFCF2F96EFCECF949" box="[828,872,1695,1722]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
; “ 
<collectingCounty id="A56F4E2F1C4BA31EFC4FF96EFC31F94A" box="[897,949,1695,1721]" pageId="2" pageNumber="3">Iran</collectingCounty>
, 
<collectingRegion id="8E75F8411C4BA31EFC0EF96EFBC2F949" box="[960,1094,1695,1722]" country="Iran" name="Gilan" pageId="2" pageNumber="3">Gilan Prov.</collectingRegion>
, 
<collectingMunicipality id="AC6AACD91C4BA31EFB9FF96EFA91F94A" box="[1105,1301,1695,1721]" pageId="2" pageNumber="3">Shafarood forest</collectingMunicipality>
, 
<elevation id="C79CD1901C4BA31EFAEEF96EFF70F92F" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="2" pageNumber="3" unit="m" value="1138.0">
<quantity id="8B499B461C4BA31EFAEEF96EFAF3F949" box="[1312,1399,1695,1722]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="2" pageNumber="3" unit="m" value="1138.0">1138 m</quantity>
a.s.l.
</elevation>
, 
<geoCoordinate id="298550641C4BA31EFECFF933FEF8F92E" box="[257,380,1730,1757]" degrees="36.652" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="55" value="36.652">36.652° N</geoCoordinate>
, 
<geoCoordinate id="298550641C4BA31EFE49F933FE7AF92E" box="[391,510,1730,1757]" degrees="53.841" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="55" value="53.841">53.841° E</geoCoordinate>
, 
<location id="496E60781C4BA31EFDC4F933FD29F92F" LSID="urn:lsid:plazi:treatment:C41887B51C4BA31BFD5EFEA9FE09FB39:496E60781C4BA31EFDC4F933FD29F92F" box="[522,685,1730,1756]" country="Iran" county="Iran" latitude="36.652" longLatPrecision="70" longitude="53.841" municipality="Shafarood forest" name="Natural forest" pageId="2" pageNumber="3" stateProvince="Gilan">Natural forest</location>
, 
<collectingMethod id="95F04EB41C4BA31EFD76F933FCC9F92E" box="[696,845,1730,1757]" pageId="2" pageNumber="3">window trap</collectingMethod>
, month 
<date id="380F10631C4BA31EFC64F933FB9AF92E" box="[938,1054,1730,1757]" pageId="2" pageNumber="3" value="2018-07">
<collectingDate id="284BE98B1C4BA31EFC64F933FB9AF92E" box="[938,1054,1730,1757]" pageId="2" pageNumber="3" value="2018-07">July 2018</collectingDate>
</date>
, leg. 
<collectorName id="E14453751C4BA31EFB92F932FB63F92F" box="[1116,1255,1730,1757]" pageId="2" pageNumber="3">F. Kazerani</collectorName>
”; 
<collectionCode id="2AA0AE661C4BA31EFB33F932FADBF92E" box="[1277,1375,1731,1757]" pageId="2" pageNumber="3">PCAJM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="04AB65281C4BA31AFF73F8F5FCF0F8DA" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
<paragraph id="4C0E36A31C4BA31EFF73F8F5FECAF8ED" blockId="2.[189,334,1796,1822]" box="[189,334,1796,1822]" pageId="2" pageNumber="3">
<heading id="174681CF1C4BA31EFF73F8F5FECAF8ED" bold="true" box="[189,334,1796,1822]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="7EC5EAB11C4BA31EFF73F8F5FECAF8ED" bold="true" box="[189,334,1796,1822]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF24F8C2FCA1F8BE" blockId="2.[189,805,1842,1904]" box="[234,805,1842,1869]" pageId="2" pageNumber="3">
<emphasis id="7EC5EAB11C4BA31EFF24F8C2FE19F8BE" bold="true" box="[234,413,1843,1869]" pageId="2" pageNumber="3">
Male 
<typeStatus id="930A88011C4BA31EFEFFF8C2FE19F8BE" box="[305,413,1843,1869]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="D48A2A261C4BA31EFE62F8C3FE76F8BE" box="[428,498,1842,1869]" captionStart="Fig" captionStartId="3.[189,232,1872,1898]" captionTargetBox="[220,1398,265,1822]" captionTargetId="figure@3.[189,1398,265,1830]" captionTargetPageId="3" captionText="Fig. 1. Habitus images of Lacon spp. A–C. L. mertliki sp. nov, holotype, ♂, in dorsal, ventral and lateral view, respectively. D. L. mertliki sp. nov, paratype, ♀, dorsal view. E–F. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)) in dorsal and lateral view, respectively. G–H. L. lepidopterus, ♀ from Turkey in dorsal and lateral view, respectively. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/3338046/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
A–C, 2A, E, G, K, 3B–C)
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF73F8A7FDD4F89C" blockId="2.[189,805,1842,1904]" box="[189,592,1877,1904]" pageId="2" pageNumber="3">
SIZE. 
<quantity id="8B499B461C4BA31EFF32F8A4FEE1F883" box="[252,357,1877,1904]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="2" pageNumber="3" unit="mm" value="15.3">15.3 mm</quantity>
long, 
<quantity id="8B499B461C4BA31EFE60F8A7FD8CF883" box="[430,520,1878,1904]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="2" pageNumber="3" unit="mm" value="4.8">4.8 mm</quantity>
wide.
</paragraph>
<paragraph id="4C0E36A31C4BA31EFF73F869FAF3F827" blockId="2.[189,1399,1943,2005]" pageId="2" pageNumber="3">BODY AND COLOR. Elongate, very slightly convex, matt, blackish; articulations of legs and tarsi reddish brown to dark brown. Body densely covered with combination of black (majority) and silver/yellowish</paragraph>
<caption id="18CE662B1C4AA31FFF73F8A1FACDF827" httpUri="https://zenodo.org/record/3338046/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1872,1898]" targetBox="[220,1398,265,1822]" targetPageId="3">
<paragraph id="4C0E36A31C4AA31FFF73F8A1FACDF827" blockId="3.[189,1399,1872,2004]" pageId="3" pageNumber="4">
<emphasis id="7EC5EAB11C4AA31FFF73F8A1FE8AF899" bold="true" box="[189,270,1872,1898]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
Habitus images of 
<taxonomicName id="8BB14D201C4AA31FFDCCF8A0FDCFF899" authorityName="Laporte" authorityYear="1838" box="[514,587,1873,1898]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7EC5EAB11C4AA31FFDCCF8A0FDCFF899" box="[514,587,1873,1898]" italics="true" pageId="3" pageNumber="4">Lacon</emphasis>
</taxonomicName>
spp. 
<emphasis id="7EC5EAB11C4AA31FFD5CF8A1FD4FF899" bold="true" box="[658,715,1872,1898]" pageId="3" pageNumber="4">A–C</emphasis>
. 
<taxonomicName id="8BB14D201C4AA31FFD13F8A0FCDFF899" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[733,859,1872,1898]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4AA31FFD13F8A0FCDFF899" box="[733,859,1872,1898]" italics="true" pageId="3" pageNumber="4">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4AA31FFCA9F8A1FC44F899" box="[871,960,1872,1898]" pageId="3" pageNumber="4" rank="species">sp. nov</taxonomicNameLabel>
, holotype, ♂, in dorsal, ventral and lateral view, respectively. 
<emphasis id="7EC5EAB11C4AA31FFE22F882FD85F87E" bold="true" box="[492,513,1907,1933]" pageId="3" pageNumber="4">D</emphasis>
. 
<taxonomicName id="8BB14D201C4AA31FFDDEF885FD0DF87E" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[528,649,1907,1933]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4AA31FFDDEF885FD0DF87E" box="[528,649,1907,1933]" italics="true" pageId="3" pageNumber="4">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4AA31FFD5EF882FD62F87E" box="[656,742,1907,1933]" pageId="3" pageNumber="4" rank="species">sp. nov</taxonomicNameLabel>
, paratype, ♀, dorsal view. 
<emphasis id="7EC5EAB11C4AA31FFBD2F882FBD4F87E" bold="true" box="[1052,1104,1907,1933]" pageId="3" pageNumber="4">E–F</emphasis>
. 
<taxonomicName id="8BB14D201C4AA31FFB91F885FE80F843" authority="(Panzer, 1801)" authorityName="Panzer" authorityYear="1801" baseAuthorityName="Panzer" baseAuthorityYear="1801" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4AA31FFB91F885FA94F87E" box="[1119,1296,1907,1933]" italics="true" pageId="3" pageNumber="4">L. lepidopterus</emphasis>
(Panzer, 1801)
</taxonomicName>
, ♂ (lectotype of 
<taxonomicName id="8BB14D201C4AA31FFE1CF866FC15F843" authority="(Jagemann, 1944))" authorityName="" authorityYear="1944" baseAuthorityName="Jagemann" baseAuthorityYear="1944" box="[466,913,1942,1968]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aurosquamosus">
<emphasis id="7EC5EAB11C4AA31FFE1CF866FD2EF843" box="[466,682,1943,1968]" italics="true" pageId="3" pageNumber="4">L. aurosquamosus</emphasis>
(Jagemann, 1944))
</taxonomicName>
in dorsal and lateral view, respectively. 
<emphasis id="7EC5EAB11C4AA31FFF73F848FF7DF820" bold="true" box="[189,249,1977,2003]" pageId="3" pageNumber="4">G–H</emphasis>
. 
<taxonomicName id="8BB14D201C4AA31FFEC6F84BFE3EF820" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[264,442,1977,2003]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4AA31FFEC6F84BFE3EF820" box="[264,442,1977,2003]" italics="true" pageId="3" pageNumber="4">L. lepidopterus</emphasis>
</taxonomicName>
, ♀ from Turkey in dorsal and lateral view, respectively. Scale bars = 2.0 mm.
</paragraph>
</caption>
<caption id="18CE662B1C4DA318FF73F919FB02F826" httpUri="https://zenodo.org/record/3338048/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1768,1794]" targetBox="[228,1355,265,1719]" targetPageId="4">
<paragraph id="4C0E36A31C4DA318FF73F919FB02F826" blockId="4.[189,1399,1768,2006]" pageId="4" pageNumber="5">
<emphasis id="7EC5EAB11C4DA318FF73F919FE8EF8F1" bold="true" box="[189,266,1768,1794]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
Details of pronotum, antenna, scutellar shield and elytral surface of 
<taxonomicName id="8BB14D201C4DA318FBE2F918FBF1F8F1" authorityName="Laporte" authorityYear="1838" box="[1068,1141,1769,1794]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="7EC5EAB11C4DA318FBE2F918FBF1F8F1" box="[1068,1141,1769,1794]" italics="true" pageId="4" pageNumber="5">Lacon</emphasis>
</taxonomicName>
spp. 
<emphasis id="7EC5EAB11C4DA318FB7AF919FB69F8F1" bold="true" box="[1204,1261,1768,1794]" pageId="4" pageNumber="5">A–D</emphasis>
. Pronotum, dorsal view. 
<emphasis id="7EC5EAB11C4DA318FE9CF8FAFEE3F8D6" bold="true" box="[338,359,1803,1829]" pageId="4" pageNumber="5">A</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FEB8F8FDFE74F8D6" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[374,496,1803,1829]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FEB8F8FDFE74F8D6" box="[374,496,1803,1829]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FE36F8FDFDD1F8D5" box="[504,597,1804,1830]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂. 
<emphasis id="7EC5EAB11C4DA318FD35F8FAFC8BF8D6" bold="true" box="[763,783,1803,1829]" pageId="4" pageNumber="5">B</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FCD3F8FDFC13F8D6" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[797,919,1803,1829]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FCD3F8FDFC13F8D6" box="[797,919,1803,1829]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FC51F8FDFC78F8D5" box="[927,1020,1804,1830]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀. 
<emphasis id="7EC5EAB11C4DA318FB6FF8FAFB32F8D6" bold="true" box="[1185,1206,1803,1829]" pageId="4" pageNumber="5">C</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FB0BF8FDFEEDF8BB" authority="(Panzer, 1801)" authorityName="Panzer" authorityYear="1801" baseAuthorityName="Panzer" baseAuthorityYear="1801" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4DA318FB0BF8FDFAF3F8D6" box="[1221,1399,1803,1829]" italics="true" pageId="4" pageNumber="5">L. lepidopterus</emphasis>
(Panzer, 1801)
</taxonomicName>
, ♂ (lectotype of 
<taxonomicName id="8BB14D201C4DA318FDE9F8DEFC5AF8BB" authority="(Jagemann, 1944))" authorityName="" authorityYear="1944" baseAuthorityName="Jagemann" baseAuthorityYear="1944" box="[551,990,1838,1864]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aurosquamosus">
<emphasis id="7EC5EAB11C4DA318FDE9F8DEFD7EF8BB" box="[551,762,1839,1864]" italics="true" pageId="4" pageNumber="5">L. aurosquamosus</emphasis>
(Jagemann, 1944))
</taxonomicName>
. 
<emphasis id="7EC5EAB11C4DA318FC28F8DFFC7FF8BB" bold="true" box="[998,1019,1838,1864]" pageId="4" pageNumber="5">D</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FBC6F8DEFB3CF8BB" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[1032,1208,1838,1864]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4DA318FBC6F8DEFB3CF8BB" box="[1032,1208,1838,1864]" italics="true" pageId="4" pageNumber="5">L. lepidopterus</emphasis>
</taxonomicName>
, ♂ from Turkey. 
<emphasis id="7EC5EAB11C4DA318FF73F8A3FF75F89F" bold="true" box="[189,241,1874,1900]" pageId="4" pageNumber="5">E–F</emphasis>
. Antenna of 
<taxonomicName id="8BB14D201C4DA318FE4AF8A3FE79F898" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[388,509,1873,1899]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FE4AF8A3FE79F898" box="[388,509,1873,1899]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FDCDF8A3FDD8F89F" box="[515,604,1874,1900]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EC5EAB11C4DA318FDACF8A3FDF2F89F" bold="true" box="[610,630,1874,1900]" pageId="4" pageNumber="5">E</emphasis>
. Holotype, ♂. 
<emphasis id="7EC5EAB11C4DA318FCEEF8A3FCB6F89F" bold="true" box="[800,818,1874,1900]" pageId="4" pageNumber="5">F</emphasis>
. Paratype, ♀. 
<emphasis id="7EC5EAB11C4DA318FC19F8A0FB8FF898" bold="true" box="[983,1035,1873,1899]" pageId="4" pageNumber="5">G–J</emphasis>
. Scutellar shield. 
<emphasis id="7EC5EAB11C4DA318FB14F8A0FB75F898" bold="true" box="[1242,1265,1873,1899]" pageId="4" pageNumber="5">G</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FB30F8A3FAF2F898" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[1278,1398,1873,1899]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FB30F8A3FAF2F898" box="[1278,1398,1873,1899]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FF73F884FE9FF87C" box="[189,283,1909,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂. 
<emphasis id="7EC5EAB11C4DA318FE0AF884FE5FF87C" bold="true" box="[452,475,1909,1935]" pageId="4" pageNumber="5">H</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FE25F884FDE2F87C" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[491,614,1909,1935]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FE25F884FDE2F87C" box="[491,614,1909,1935]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FDA1F884FD49F87C" box="[623,717,1909,1935]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀. 
<emphasis id="7EC5EAB11C4DA318FCBAF884FCFBF87C" bold="true" box="[884,895,1909,1935]" pageId="4" pageNumber="5">I</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FC5EF884FBC7F87C" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[912,1091,1909,1935]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4DA318FC5EF884FBC7F87C" box="[912,1091,1909,1935]" italics="true" pageId="4" pageNumber="5">L. lepidopterus</emphasis>
</taxonomicName>
, ♂. 
<emphasis id="7EC5EAB11C4DA318FBB6F884FB03F87C" bold="true" box="[1144,1159,1909,1935]" pageId="4" pageNumber="5">J</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FB58F884FACEF87C" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[1174,1354,1909,1935]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4DA318FB58F884FACEF87C" box="[1174,1354,1909,1935]" italics="true" pageId="4" pageNumber="5">L. lepidopterus</emphasis>
</taxonomicName>
, ♀. 
<emphasis id="7EC5EAB11C4DA318FF73F869FF72F841" bold="true" box="[189,246,1944,1970]" pageId="4" pageNumber="5">K–L</emphasis>
. Detail of elytral surface. 
<emphasis id="7EC5EAB11C4DA318FDE3F869FDC0F841" bold="true" box="[557,580,1944,1970]" pageId="4" pageNumber="5">K</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FD9AF868FD4AF841" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[596,718,1944,1970]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4DA318FD9AF868FD4AF841" box="[596,718,1944,1970]" italics="true" pageId="4" pageNumber="5">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4DA318FD19F869FCB1F841" box="[727,821,1944,1970]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂. 
<emphasis id="7EC5EAB11C4DA318FC13F869FC75F841" bold="true" box="[989,1009,1944,1970]" pageId="4" pageNumber="5">L</emphasis>
. 
<taxonomicName id="8BB14D201C4DA318FBCEF868FB37F841" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[1024,1203,1944,1970]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4DA318FBCEF868FB37F841" box="[1024,1203,1944,1970]" italics="true" pageId="4" pageNumber="5">L. lepidopterus</emphasis>
</taxonomicName>
, ♂ (lectotype of 
<taxonomicName id="8BB14D201C4DA318FF73F84DFE17F826" authorityName="" baseAuthorityName="Jagemann" baseAuthorityYear="1944" box="[189,403,1980,2005]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aurosquamosus">
<emphasis id="7EC5EAB11C4DA318FF73F84DFE17F826" box="[189,403,1980,2005]" italics="true" pageId="4" pageNumber="5">L. aurosquamosus</emphasis>
</taxonomicName>
). Scale bars: A–F = 1.0 mm; G–J = 0.5 mm; K–L = not to scale.
</paragraph>
</caption>
<paragraph id="4C0E36A31C4CA319FF73FEFAFB10FEBA" blockId="5.[189,1398,267,329]" pageId="5" pageNumber="6">scale-like setae, with some additional white setae. Antennomeres II–XI covered with combination of blackish and yellowish hair-like; mouthparts covered with yellowish hair-like setae.</paragraph>
<paragraph id="4C0E36A31C4CA319FF73FE81FCBBFD19" blockId="5.[189,1399,368,746]" pageId="5" pageNumber="6">HEAD. Wider than long; frons with moderately deep median depression. Punctures coarse and dense, almost contiguous; surface between punctures smooth. Labrum transverse, convex, finely punctate, anteriorly rounded, with long, semi-erect hair-like pubescence. Mandible robust, relatively short, bidentate, basally with large punctures and long hair-like setae. Maxillary palp relatively short, apical palpomere securiform, more than 1.5 times as long as wide, apically obliquely widened, flattened and truncated. Antennae reaching near middle of pronotum, distinctly serrate from antennomere III. Scape long, robust, notably thicker than others; length ratio of antennomeres II–V as 1.0:2.0:1.8:1.8; antennomere II minute, short, slightly wider than long; antennomere III elongate; antennomeres IV–X about equal in shape, about as long as wide to slightly wider; last antennomere almost obovate. Antennae covered with moderately dense, decumbent, blackish hair-like setae, inner sides of antennomeres II–XI covered with additional erect yellowish hair-like setae.</paragraph>
<paragraph id="4C0E36A31C4CA31AFF73FCE3FE4EFC30" blockId="5.[189,1399,785,1199]" lastBlockId="6.[189,1399,267,963]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
PRONOTUM. Slightly wider than long (length 
<quantity id="8B499B461C4CA319FD17FCE3FCBEFCDF" box="[729,826,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="6" unit="mm" value="4.2">4.2 mm</quantity>
; width 
<quantity id="8B499B461C4CA319FC5BFCE3FC73FCDF" box="[917,1015,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="6" unit="mm" value="4.3">4.3 mm</quantity>
), widest medially; sides widely rounded, slightly sinuate before posterior angles, from about half distinctly narrowed toward anterior margin; anterior angles short, sharp, reaching about half length of each eye; posterior angles rather short, flattened, very slightly divergent, almost rectangular. Pronotal disc with two pairs of tubercles in centre of disc laterally from median impression; anterior pair somewhat larger than posterior pair; with two deep distinct impressions before half, laterally from centre of disc, and two pairs of smaller and moderately deep impressions in the anterior and posterior third, respectively. Median depression on pronotal disc somewhat shortly divided by central part with tubercles; posterior part of depression very wide and deep; anterior part of depression shorter, rather narrow and shallow. Punctation of pronotum similar to punctation on head. Pronotosternal sutures deeply grooved almost over entire length. Prosternum elongate, about 2.5 times as long as wide, punctate similarly as pronotum; prosternal lobe short, with equally rounded anterior margin; surface rough, wrinkled. Hypomeron with slightly denser and smaller punctures than punctures on prosternum; basally with smooth depression. Prosternal process slightly more than 3 times as long as diameter of procoxal cavity, with weak prominence near apex, narrowly rounded apically. Mesoventrite transverse; mesoventral cavity with well-defined walls; mesoventral process wide, apically truncate. Metaventrite robust, with large punctures; discrimen long, almost reaching anterior margin. Metacoxal plate narrowed at about half; wider part with posterior margin bisinuate. Scutellar shield about 1.35 times as long as wide, distinctly declined, flat, tongue-shaped, anterior margin weakly emarginate medially, sides subparallel, posteriorly weakly rounded, with large deep punctures, pubescence same as on elytra. Elytra slightly wider than pronotum, widest at about ⅓, together about 2.2 times as long as wide (length 
<quantity id="8B499B461C4FA31AFACDFDD4FAF6FDCC" box="[1283,1394,549,576]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="6" pageNumber="7" unit="mm" value="10.5">10.5 mm</quantity>
; width 
<quantity id="8B499B461C4FA31AFEC8FDB9FEE0FD90" box="[262,356,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="7" unit="mm" value="4.8">4.8 mm</quantity>
), about 2.5 times as long as pronotum; subparallel-sided from humeri to posterior ⅓, then slightly tapering to apex. Disc of elytra slightly convex near humeri, abruptly declined near scutellar shield, almost flattened after humeri, without striae but with elongate longitudinal keel running from humeri to about ⅔ of elytral length, moderately densely covered with punctures subequal in size and shape, usually separated by about diameter of puncture. Epipleura complete, distinctly narrowed after metacoxae; its broader part covered with scale-like setae, narrower part covered with hair-like setae. Hind wings completely developed, reaching apex of elytra. Leg moderately long; tarsomere I slightly longer than combined lengths of tarsomeres II and III, but shorter than combined lengths of tarsomeres II–IV; tarsomeres II–IV longer than wide but relatively short, each one slightly shorter than preceding one; apical tarsomere longest, slender; pretarsal claws simple, slightly curved, each basally with long seta.
</paragraph>
<caption id="18CE662B1C4CA319FF73F8CAFDBFF811" httpUri="https://zenodo.org/record/3338050/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1851,1877]" targetBox="[193,1395,1314,1803]" targetPageId="5">
<paragraph id="4C0E36A31C4CA319FF73F8CAFDBFF811" blockId="5.[189,1399,1851,2019]" pageId="5" pageNumber="6">
<emphasis id="7EC5EAB11C4CA319FF73F8CAFE8FF8A6" bold="true" box="[189,267,1851,1877]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
Pregenital segments and genitalia of 
<taxonomicName id="8BB14D201C4CA319FD0DF8CDFC88F8A6" authorityName="Laporte" authorityYear="1838" box="[707,780,1852,1877]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="7EC5EAB11C4CA319FD0DF8CDFC88F8A6" box="[707,780,1852,1877]" italics="true" pageId="5" pageNumber="6">Lacon</emphasis>
</taxonomicName>
spp. 
<emphasis id="7EC5EAB11C4CA319FC82F8CAFCE5F8A6" bold="true" box="[844,865,1851,1877]" pageId="5" pageNumber="6">A</emphasis>
. 
<taxonomicName id="8BB14D201C4CA319FCBEF8CDFB53F8A6" authority="(Panzer, 1801)" authorityName="Panzer" authorityYear="1801" baseAuthorityName="Panzer" baseAuthorityYear="1801" box="[880,1239,1851,1877]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lepidopterus">
<emphasis id="7EC5EAB11C4CA319FCBEF8CDFBA6F8A6" box="[880,1058,1851,1877]" italics="true" pageId="5" pageNumber="6">L. lepidopterus</emphasis>
(Panzer, 1801)
</taxonomicName>
, ♂ (lectotype of 
<taxonomicName id="8BB14D201C4CA319FF2EF8AEFD27F88B" authority="(Jagemann, 1944))" authorityName="" authorityYear="1944" baseAuthorityName="Jagemann" baseAuthorityYear="1944" box="[224,675,1886,1912]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aurosquamosus">
<emphasis id="7EC5EAB11C4CA319FF2EF8AEFE3CF88B" box="[224,440,1887,1912]" italics="true" pageId="5" pageNumber="6">L. aurosquamosus</emphasis>
(Jagemann, 1944))
</taxonomicName>
, aedeagus, dorsal view. 
<emphasis id="7EC5EAB11C4CA319FC04F8AFFB85F88B" bold="true" box="[970,1025,1886,1912]" pageId="5" pageNumber="6">B–C</emphasis>
. 
<taxonomicName id="8BB14D201C4CA319FBDDF8AEFB14F88B" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[1043,1168,1886,1912]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4CA319FBDDF8AEFB14F88B" box="[1043,1168,1886,1912]" italics="true" pageId="5" pageNumber="6">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4CA319FB54F8AEFB7EF88A" box="[1178,1274,1887,1913]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂. 
<emphasis id="7EC5EAB11C4CA319FF2BF873FF7DF86F" bold="true" box="[229,249,1922,1948]" pageId="5" pageNumber="6">B</emphasis>
. Aedeagus, dorsal view. 
<emphasis id="7EC5EAB11C4CA319FDE7F870FDBAF868" bold="true" box="[553,574,1921,1947]" pageId="5" pageNumber="6">C</emphasis>
. Abdominal terminal segments. 
<emphasis id="7EC5EAB11C4CA319FC09F870FB86F868" bold="true" box="[967,1026,1921,1947]" pageId="5" pageNumber="6">D–G</emphasis>
. 
<taxonomicName id="8BB14D201C4CA319FBDAF873FB15F868" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[1044,1169,1921,1947]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4CA319FBDAF873FB15F868" box="[1044,1169,1921,1947]" italics="true" pageId="5" pageNumber="6">L. mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4CA319FB55F873FB7FF86F" box="[1179,1275,1922,1948]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀. 
<emphasis id="7EC5EAB11C4CA319FF2BF854FF7EF84C" bold="true" box="[229,250,1957,1983]" pageId="5" pageNumber="6">D</emphasis>
. Abdominal tergite VIII. 
<emphasis id="7EC5EAB11C4CA319FDFDF854FDC3F84C" bold="true" box="[563,583,1957,1983]" pageId="5" pageNumber="6">E</emphasis>
. Abdominal sternite VIII. 
<emphasis id="7EC5EAB11C4CA319FC45F854FC19F84C" bold="true" box="[907,925,1957,1983]" pageId="5" pageNumber="6">F</emphasis>
. Ovipositor. 
<emphasis id="7EC5EAB11C4CA319FBF3F855FBD0F84D" bold="true" box="[1085,1108,1956,1982]" pageId="5" pageNumber="6">G</emphasis>
. Large sclerite of bursa copulatrix. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="4C0E36A31C4FA31AFF73FC1BFEBDFA90" blockId="6.[189,1399,1001,1379]" pageId="6" pageNumber="7">ABDOMEN. Ventrites moderately densely covered with small punctures; punctures larger and denser on ventrite 1 and on sides of each ventrite; apical ventrite rounded apically. Tergite IX transverse, wider than long, medially gradually emarginate. Tergite X slightly longer than wide, apically gradually rounded, finely punctate and covered with sparse setae. Sternite IX elongate, less than three times as long as wide, basally somewhat truncate, sides slightly constricted medially, apically equally rounded, covered with short setae, mainly at apex. Aedeagus elongate, about 3.2 times as long as wide. Median lobe elongate, slightly shorter than parameres, with short, robust basal struts, narrowed toward apex, apically narrowly rounded. Paramere elongate, gradually slightly narrowed toward apical lobe; apical lobe robust, elongate, rounded apically, with outer margin slightly emarginate, with short, stout, distinct lateral subapical hook. Phallobase relatively small, short, wider than long, U-shaped.</paragraph>
<paragraph id="4C0E36A31C4FA31AFF24FA7AFD02FA56" blockId="6.[189,1398,1419,1833]" box="[234,646,1419,1446]" pageId="6" pageNumber="7">
<emphasis id="7EC5EAB11C4FA31AFF24FA7AFEC1FA56" bold="true" box="[234,325,1419,1445]" pageId="6" pageNumber="7">Female</emphasis>
(
<figureCitation id="D48A2A261C4FA31AFE9BFA7AFE32FA56" box="[341,438,1419,1445]" captionStart="Fig" captionStartId="3.[189,232,1872,1898]" captionTargetBox="[220,1398,265,1822]" captionTargetId="figure@3.[189,1398,265,1830]" captionTargetPageId="3" captionText="Fig. 1. Habitus images of Lacon spp. A–C. L. mertliki sp. nov, holotype, ♂, in dorsal, ventral and lateral view, respectively. D. L. mertliki sp. nov, paratype, ♀, dorsal view. E–F. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)) in dorsal and lateral view, respectively. G–H. L. lepidopterus, ♀ from Turkey in dorsal and lateral view, respectively. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/3338046/files/figure.png" pageId="6" pageNumber="7">Figs 1D</figureCitation>
, 
<figureCitation id="D48A2A261C4FA31AFE0DFA7AFDA2FA55" box="[451,550,1419,1446]" captionStart="Fig" captionStartId="4.[189,232,1768,1794]" captionTargetBox="[228,1355,265,1719]" captionTargetId="figure@4.[228,1359,265,1728]" captionTargetPageId="4" captionText="Fig. 2. Details of pronotum, antenna, scutellar shield and elytral surface of Lacon spp. A–D. Pronotum, dorsal view. A. L. mertliki sp. nov., holotype, ♂. B. L. mertliki sp. nov., paratype, ♀. C. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)). D. L. lepidopterus, ♂ from Turkey. E–F. Antenna of L. mertliki sp. nov. E. Holotype, ♂. F. Paratype, ♀. G–J. Scutellar shield. G. L. mertliki sp. nov., holotype, ♂. H. L. mertliki sp. nov., paratype, ♀. I. L. lepidopterus, ♂. J. L. lepidopterus, ♀. K–L. Detail of elytral surface. K. L. mertliki sp. nov., holotype, ♂. L. L. lepidopterus, ♂ (lectotype of L. aurosquamosus). Scale bars: A–F = 1.0 mm; G–J = 0.5 mm; K–L = not to scale." httpUri="https://zenodo.org/record/3338048/files/figure.png" pageId="6" pageNumber="7">2B, F, H</figureCitation>
, 
<figureCitation id="D48A2A261C4FA31AFDFAFA7AFDC4FA56" box="[564,576,1419,1445]" captionStart="Fig" captionStartId="5.[189,232,1851,1877]" captionTargetBox="[193,1395,1314,1803]" captionTargetId="figure@5.[189,1398,1310,1812]" captionTargetPageId="5" captionText="Fig. 3. Pregenital segments and genitalia of Lacon spp. A. L. lepidopterus (Panzer, 1801), ♂ (lectotype of L. aurosquamosus (Jagemann, 1944)), aedeagus, dorsal view. B–C. L. mertliki sp. nov., holotype, ♂. B. Aedeagus, dorsal view. C. Abdominal terminal segments. D–G. L. mertliki sp. nov., paratype, ♀. D. Abdominal tergite VIII. E. Abdominal sternite VIII. F. Ovipositor. G. Large sclerite of bursa copulatrix. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/3338050/files/figure.png" pageId="6" pageNumber="7">3</figureCitation>
D–G)
</paragraph>
<paragraph id="4C0E36A31C4FA31AFF73FA5FFCF0F8DA" blockId="6.[189,1398,1419,1833]" pageId="6" pageNumber="7">
Similar in body size and shape to male (body 
<quantity id="8B499B461C4FA31AFD23FA5FFC1EFA3A" box="[749,922,1454,1481]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.75" metricValueMin="1.55" pageId="6" pageNumber="7" unit="mm" value="16.5" valueMax="17.5" valueMin="15.5">15.5–17.5 mm</quantity>
long; 
<quantity id="8B499B461C4FA31AFC24FA5FFBFEFA3A" box="[1002,1146,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.5" metricValueMin="4.6" pageId="6" pageNumber="7" unit="mm" value="5.05" valueMax="5.5" valueMin="4.6">4.6–5.5 mm</quantity>
wide); slightly more robust than in male. Antenna slightly shorter, serrated antennomeres of different shape than in male, with serrations much wider and more distinctly widely rounded; erect setae on inner sides of antennomeres less numerous. Pronotal tubercles and impressions more developed; posterior angles slightly more divergent. Scutellar shield relatively wider, 1.15–1.20 times longer than wide. Tergite VIII subtriangular, wider than long, covered with fine punctures, apico-laterally with several short setae. Sternite VIII widely triangular, apically rounded, medially less sclerotized, sparsely covered with fine punctures and short setae; spiculum ventrale about 2.75 times sternite length. Ovipositor relatively long; paraprocts about 4.5 times as long as gonocoxites; styli present, short, subapically attached to gonocoxites. 
<collectingRegion id="8E75F8411C4FA31AFAFCF939FAF2F911" box="[1330,1398,1736,1762]" country="Turkey" name="Bursa" pageId="6" pageNumber="7">Bursa</collectingRegion>
copulatrix membranous, sac-like, with large sclerite covered with short spinules and long spines, narrowed medially; distal small sclerite slender, transverse.
</paragraph>
</subSubSection>
<subSubSection id="04AB65281C4FA31BFF73F8BEFC9CFD9A" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="distribution">
<paragraph id="4C0E36A31C4FA31AFF73F8BEFED2F89A" blockId="6.[189,342,1871,1897]" box="[189,342,1871,1897]" pageId="6" pageNumber="7">
<heading id="174681CF1C4FA31AFF73F8BEFED2F89A" bold="true" box="[189,342,1871,1897]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="7EC5EAB11C4FA31AFF73F8BEFED2F89A" bold="true" box="[189,342,1871,1897]" pageId="6" pageNumber="7">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4C0E36A31C4FA31AFF73F88EFEBFF848" blockId="6.[189,1398,1918,1979]" pageId="6" pageNumber="7">
<collectingCountry id="34A676331C4FA31AFF73F88EFF68F86A" box="[189,236,1919,1945]" name="Iran" pageId="6" pageNumber="7">Iran</collectingCountry>
(
<collectingRegion id="8E75F8411C4FA31AFF30F88FFEC7F86B" box="[254,323,1918,1944]" country="Iran" name="Gilan" pageId="6" pageNumber="7">Gilan</collectingRegion>
, 
<collectingRegion id="8E75F8411C4FA31AFE9CF88FFE39F86B" box="[338,445,1918,1944]" country="Iran" name="Golestan" pageId="6" pageNumber="7">Golestan</collectingRegion>
, 
<collectingRegion id="8E75F8411C4FA31AFE05F88FFDE6F86A" box="[459,610,1918,1945]" country="Iran" name="Mazandaran" pageId="6" pageNumber="7">Mazandaran</collectingRegion>
). This species is probably distributed across the whole Hyrcanian forest belt.
</paragraph>
<paragraph id="4C0E36A31C4EA31BFF73FEFAFE70FED6" blockId="7.[189,500,267,293]" box="[189,500,267,293]" pageId="7" pageNumber="8">
<heading id="174681CF1C4EA31BFF73FEFAFE70FED6" bold="true" box="[189,500,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7EC5EAB11C4EA31BFF73FEFAFE70FED6" bold="true" box="[189,500,267,293]" pageId="7" pageNumber="8">Collecting circumstances</emphasis>
</heading>
</paragraph>
<paragraph id="4C0E36A31C4EA31BFF73FEC8FBD7FDD4" blockId="7.[189,1399,313,552]" pageId="7" pageNumber="8">
Fresh material was collected in floodplain forests of the 
<collectingRegion id="8E75F8411C4EA31BFC8CFEC8FC2EFEA0" box="[834,938,313,339]" country="Iran" name="Golestan" pageId="7" pageNumber="8">Golestan</collectingRegion>
National Park and in montane primeval beech forests in the 
<collectingRegion id="8E75F8411C4EA31BFE60FEACFDBBFE84" box="[430,575,349,375]" country="Iran" name="Mazandaran" pageId="7" pageNumber="8">Mazandaran</collectingRegion>
and 
<collectingRegion id="8E75F8411C4EA31BFDB2FEACFD39FE84" box="[636,701,349,375]" country="Iran" name="Gilan" pageId="7" pageNumber="8">Gilan</collectingRegion>
provinces (
<figureCitation id="D48A2A261C4EA31BFC84FEACFC2EFE84" box="[842,938,348,375]" captionStart="Fig" captionStartId="10.[189,232,1948,1974]" captionTargetBox="[189,1398,302,1867]" captionTargetId="figure@10.[189,1398,302,1867]" captionTargetPageId="10" captionText="Fig. 5. Habitat of Lacon mertliki sp. nov. in Iran, Mazandaran province. A. Old-growth forest. B. Advanced brown rotten log." httpUri="https://zenodo.org/record/3338054/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
). 
<taxonomicName id="8BB14D201C4EA31BFC0EFEACFBE9FE84" authority="Kundrata &amp; Németh &amp; Jarzabek-Müller, 2019" authorityName="Kundrata &amp; Németh &amp; Jarzabek-Müller" authorityYear="2019" box="[960,1133,349,375]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mertliki" status="sp. nov.">
<emphasis id="7EC5EAB11C4EA31BFC0EFEACFBE9FE84" box="[960,1133,349,375]" italics="true" pageId="7" pageNumber="8">Lacon mertliki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65F657CA1C4EA31BFBB9FEACFB57FE84" box="[1143,1235,349,375]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
was found in large logs of brown rotten wood (
<figureCitation id="D48A2A261C4EA31BFD98FE71FD36FE68" box="[598,690,384,411]" captionStart="Fig" captionStartId="10.[189,232,1948,1974]" captionTargetBox="[189,1398,302,1867]" captionTargetId="figure@10.[189,1398,302,1867]" captionTargetPageId="10" captionText="Fig. 5. Habitat of Lacon mertliki sp. nov. in Iran, Mazandaran province. A. Old-growth forest. B. Advanced brown rotten log." httpUri="https://zenodo.org/record/3338054/files/figure.png" pageId="7" pageNumber="8">Fig. 5B</figureCitation>
) of 
<taxonomicName id="8BB14D201C4EA31BFD26FE71FBEDFE69" authority="C. A. Mey." authorityName="C. A. Mey." box="[744,1129,384,410]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="castaneifolia">
<emphasis id="7EC5EAB11C4EA31BFD26FE71FC68FE69" box="[744,1004,384,410]" italics="true" pageId="7" pageNumber="8">Quercus castaneifolia</emphasis>
C.A.Mey.
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4EA31BFB69FE71FEB7FE4D" authority="C. A. Mey." authorityName="C. A. Mey." class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="subcordata">
<emphasis id="7EC5EAB11C4EA31BFB69FE71FAF3FE69" box="[1191,1399,384,410]" italics="true" pageId="7" pageNumber="8">Alnus subcordata</emphasis>
C.A.Mey.
</taxonomicName>
, along with other beetle species like 
<taxonomicName id="8BB14D201C4EA31BFD30FE52FB39FE4E" authority="Reitter, 1889" authorityName="Reitter" authorityYear="1889" box="[766,1213,419,445]" class="Insecta" family="Carabidae" genus="Clinidium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="marginicolle">
<emphasis id="7EC5EAB11C4EA31BFD30FE52FB97FE4E" box="[766,1043,419,445]" italics="true" pageId="7" pageNumber="8">Clinidium marginicolle</emphasis>
Reitter, 1889
</taxonomicName>
, 
<taxonomicName id="8BB14D201C4EA31BFB03FE52FD97FE13" authority="(Ganglbauer, 1891)" authorityName="Ganglbauer" authorityYear="1891" baseAuthorityName="Ganglbauer" baseAuthorityYear="1891" class="Insecta" family="Carabidae" genus="Omoglymmius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="germari">
<emphasis id="7EC5EAB11C4EA31BFB03FE52FE98FE13" italics="true" pageId="7" pageNumber="8">Omoglymmius germari</emphasis>
(Ganglbauer, 1891)
</taxonomicName>
(both 
<taxonomicName id="8BB14D201C4EA31BFDA4FE37FD7DFE12" authorityName="Laporte" authorityYear="1840" box="[618,761,454,481]" class="Insecta" family="Rhysodidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Rhysodidae</taxonomicName>
), 
<taxonomicName id="8BB14D201C4EA31BFCDCFE36FAF6FE12" authority="Fleischer, 1919" authorityName="Fleischer" authorityYear="1919" box="[786,1394,454,481]" class="Insecta" family="Prostomidae" genus="Prostomis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="elburica">
<emphasis id="7EC5EAB11C4EA31BFCDCFE36FC73FE13" box="[786,1015,454,480]" italics="true" pageId="7" pageNumber="8">Prostomis elburica</emphasis>
Fleischer, 1919 (Prostomidae)
</taxonomicName>
, 
<taxonomicName id="8BB14D201C4EA31BFF73FE18FC8DFDF7" authority="Ganglbauer, 1887" authorityName="Ganglbauer" authorityYear="1887" box="[189,777,489,516]" class="Insecta" family="Lucanidae" genus="Aesalus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ulanowskyi">
<emphasis id="7EC5EAB11C4EA31BFF73FE18FE25FDF0" box="[189,417,489,515]" italics="true" pageId="7" pageNumber="8">Aesalus ulanowskyi</emphasis>
Ganglbauer, 1887 (Lucanidae)
</taxonomicName>
, 
<taxonomicName id="8BB14D201C4EA31BFCDCFE18FAF6FDF7" authority="(Schonherr, 1817)" authorityName="Schonherr" authorityYear="1817" baseAuthorityName="Schonherr" baseAuthorityYear="1817" box="[786,1394,489,516]" class="Insecta" family="Elateridae" genus="Ampedus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="7EC5EAB11C4EA31BFCDCFE18FB8DFDF0" box="[786,1033,489,515]" italics="true" pageId="7" pageNumber="8">Ampedus elegantulus</emphasis>
(Schönherr, 1817) (Elateridae)
</taxonomicName>
, and 
<taxonomicName id="8BB14D201C4EA31BFF21FDFCFCC4FDDB" authority="(Linnaeus, 1758)" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[239,832,524,552]" class="Insecta" family="Tenebrionidae" genus="Uloma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="culinaris">
<emphasis id="7EC5EAB11C4EA31BFF21FDFCFE2AFDD4" box="[239,430,525,551]" italics="true" pageId="7" pageNumber="8">Uloma culinaris</emphasis>
(Linnaeus, 1758) (Tenebrionidae)
</taxonomicName>
(J. Müller, pers. com.).
</paragraph>
<paragraph id="4C0E36A31C4EA31BFF73FDBFFC9CFD9A" blockId="7.[189,792,590,617]" box="[189,792,590,617]" pageId="7" pageNumber="8">
<heading id="174681CF1C4EA31BFF73FDBFFC9CFD9A" bold="true" box="[189,792,590,617]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7EC5EAB11C4EA31BFF73FDBFFC9CFD9A" bold="true" box="[189,792,590,617]" pageId="7" pageNumber="8">
Taxonomic remarks on 
<taxonomicName id="8BB14D201C4EA31BFE2BFDBFFDE8FD9A" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[485,620,590,617]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFE2BFDBFFDE8FD9A" bold="true" box="[485,620,590,617]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4EA31BFD64FDBFFC9CFD9A" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[682,792,590,617]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFD64FDBFFC9CFD9A" bold="true" box="[682,792,590,617]" italics="true" pageId="7" pageNumber="8">L. nadaii</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04AB65281C4EA31BFF73FD8CFE09FB39" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="4C0E36A31C4EA31BFF73FD8CFE09FB39" blockId="7.[189,1399,637,1226]" pageId="7" pageNumber="8">
<taxonomicName id="8BB14D201C4EA31BFF73FD8CFEF3FD64" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[189,375,637,663]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFF73FD8CFEF3FD64" box="[189,375,637,663]" italics="true" pageId="7" pageNumber="8">Lacon unicolor</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="34A676331C4EA31BFDBAFD8CFD21FD64" box="[628,677,637,663]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
(
<bibRefCitation id="28204B521C4EA31BFD7BFD8CFCE0FD64" author="Candeze E. C. A." box="[693,868,637,663]" firstAuthor="Candeze" journalOrPublisher="Memoires de la Societe royale des Sciences de Liege" pageId="7" pageNumber="8" pagination="1 - 218" part="2" refId="ref5330" refString="Candeze E. C. A. 1874. Revision de la Monographie des Elaterides. Fasc. 1. Memoires de la Societe royale des Sciences de Liege 2: 1 - 218. https: // doi. org / 10.5962 / bhl. title. 47120" title="Revision de la Monographie des Elaterides. Fasc. 1" type="journal article" year="1874">Candèze 1874</bibRefCitation>
) and later reported from 
<collectingCountry id="34A676331C4EA31BFB5BFD8CFB6FFD64" box="[1173,1259,637,663]" name="Turkey" pageId="7" pageNumber="8">Turkey</collectingCountry>
(
<bibRefCitation id="28204B521C4EA31BFB33FD8CFEC8FD49" author="Mertlik J. &amp; Platia G." firstAuthor="Mertlik" journalOrPublisher="Elateridarium" pageId="7" pageNumber="8" pagination="1 - 40" part="2" refId="ref6035" refString="Mertlik J. &amp; Platia G. 2008. Catalogue of the family Cebrionidae, Elateridae, Lissomidae, Melasidae and Throscidae (Coleoptera) from Turkey. Elateridarium 2: 1 - 40." title="Catalogue of the family Cebrionidae, Elateridae, Lissomidae, Melasidae and Throscidae (Coleoptera) from Turkey" type="journal article" year="2008">Mertlik &amp; Platia 2008</bibRefCitation>
). 
<taxonomicName id="8BB14D201C4EA31BFE91FD50FD84FD49" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[351,512,672,698]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFE91FD50FD84FD49" box="[351,512,672,698]" italics="true" pageId="7" pageNumber="8">Lacon nadaii</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="34A676331C4EA31BFD35FD51FCA8FD49" box="[763,812,672,698]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
and compared to 
<taxonomicName id="8BB14D201C4EA31BFBCDFD50FB0EFD49" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[1027,1162,672,698]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFBCDFD50FB0EFD49" box="[1027,1162,672,698]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
due to their similar habitus and size (
<bibRefCitation id="28204B521C4EA31BFE59FD32FD32FD2E" author="Platia G. &amp; Nemeth T." box="[407,694,707,733]" firstAuthor="Platia" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="7" pageNumber="8" pagination="65 - 106" part="103" refId="ref6180" refString="Platia G. &amp; Nemeth T. 2011. On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 103: 65 - 106." title="On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae)" type="journal article" year="2011">Platia &amp; Németh 2011</bibRefCitation>
). 
<taxonomicName id="8BB14D201C4EA31BFD03FD35FCF5FD2E" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[717,881,707,733]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFD03FD35FCF5FD2E" box="[717,881,707,733]" italics="true" pageId="7" pageNumber="8">Lacon nadaii</emphasis>
</taxonomicName>
was distinguished from the latter species by “the black piceous colour, shagreened dorsal surface giving a dull appearance and shorter third antennomere“ (
<bibRefCitation id="28204B521C4EA31BFEB9FCF8FD3FFCD7" author="Platia G. &amp; Nemeth T." box="[375,699,777,804]" firstAuthor="Platia" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="7" pageNumber="8" pagination="65 - 106" part="103" refId="ref6180" refString="Platia G. &amp; Nemeth T. 2011. On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 103: 65 - 106." title="On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae)" type="journal article" year="2011">Platia &amp; Németh 2011: 67</bibRefCitation>
). 
<bibRefCitation id="28204B521C4EA31BFD1FFCF8FB88FCD7" author="Mertlik J. &amp; Nemeth T." box="[721,1036,777,804]" firstAuthor="Mertlik" journalOrPublisher="Elateridarium" pageId="7" pageNumber="8" pagination="61 - 66" part="8" refId="ref6004" refString="Mertlik J. &amp; Nemeth T. 2014. Distributional notes on Lacon nadaii and L. unicolor (Coleoptera: Elateridae). Elateridarium 8: 61 - 66." title="Distributional notes on Lacon nadaii and L. unicolor (Coleoptera: Elateridae)" type="journal article" year="2014">Mertlik &amp; Németh (2014)</bibRefCitation>
reviewed distribution of both species and found out that 
<taxonomicName id="8BB14D201C4EA31BFDC0FCDCFD13FCB4" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[526,663,813,839]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFDC0FCDCFD13FCB4" box="[526,663,813,839]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
occurs only in 
<collectingCountry id="34A676331C4EA31BFC92FCDCFC09FCB4" box="[860,909,813,839]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
and 
<taxonomicName id="8BB14D201C4EA31BFC02FCDCFBB9FCB4" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[972,1085,813,839]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFC02FCDCFBB9FCB4" box="[972,1085,813,839]" italics="true" pageId="7" pageNumber="8">L. nadaii</emphasis>
</taxonomicName>
in 
<collectingCountry id="34A676331C4EA31BFBA9FCDCFB3BFCB4" box="[1127,1215,813,839]" name="Turkey" pageId="7" pageNumber="8">Turkey</collectingCountry>
, 
<collectingCountry id="34A676331C4EA31BFB00FCDCFB7BFCB4" box="[1230,1279,813,839]" name="Iraq" pageId="7" pageNumber="8">Iraq</collectingCountry>
and 
<collectingCountry id="34A676331C4EA31BFAF0FCDCFAF7FCB4" box="[1342,1395,813,839]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
. However, they also reported two intermediate specimens (both in the coloration and the length of antennomere III) from 
<collectingCountry id="34A676331C4EA31BFE1BFC82FDAFFC7E" box="[469,555,883,909]" name="Turkey" pageId="7" pageNumber="8">Turkey</collectingCountry>
and 
<collectingCountry id="34A676331C4EA31BFDA9FC85FD18FC7D" box="[615,668,884,910]" name="Iraq" pageId="7" pageNumber="8">Iraq</collectingCountry>
. 
<bibRefCitation id="28204B521C4EA31BFD66FC82FCF3FC7E" author="Prosvirov A. S." box="[680,887,883,909]" firstAuthor="Prosvirov" journalOrPublisher="Entomological Review" pageId="7" pageNumber="8" pagination="1120 - 1136" part="97" refId="ref6313" refString="Prosvirov A. S. 2017. New data on the fauna and systematics of Palaearctic species of the click-beetle subfamily Agrypninae Candeze, 1857 (Coleoptera, Elateridae). Entomological Review 97: 1120 - 1136. https: // doi. org / 10.1134 / S 0013873817080127" title="New data on the fauna and systematics of Palaearctic species of the click-beetle subfamily Agrypninae Candeze, 1857 (Coleoptera, Elateridae)" type="journal article" year="2017">Prosvirov (2017)</bibRefCitation>
recorded 
<taxonomicName id="8BB14D201C4EA31BFC3FFC85FBFDFC7E" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[1009,1145,883,909]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFC3FFC85FBFDFC7E" box="[1009,1145,883,909]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
from 
<collectingCountry id="34A676331C4EA31BFB0AFC85FB71FC7D" box="[1220,1269,884,910]" name="Iraq" pageId="7" pageNumber="8">Iraq</collectingCountry>
and found out that the external morphological characters of this species vary greatly but the shape of aedeagus is more or less uniform among the different populations. We examined the name-bearing 
<typeStatus id="930A88011C4EA31BFADAFC4BFAD0FC27" box="[1300,1364,954,980]" pageId="7" pageNumber="8">types</typeStatus>
of both 
<taxonomicName id="8BB14D201C4EA31BFF30FC2CFE0CFC04" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[254,392,989,1015]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFF30FC2CFE0CFC04" box="[254,392,989,1015]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4EA31BFE04FC2CFDBFFC04" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[458,571,989,1015]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFE04FC2CFDBFFC04" box="[458,571,989,1015]" italics="true" pageId="7" pageNumber="8">L. nadaii</emphasis>
</taxonomicName>
as well as a long series of specimens from 
<collectingCountry id="34A676331C4EA31BFBA5FC2CFB1BFC04" box="[1131,1183,989,1015]" name="Iran" pageId="7" pageNumber="8">Iran</collectingCountry>
, 
<collectingCountry id="34A676331C4EA31BFB60FC2CFB5BFC04" box="[1198,1247,989,1015]" name="Iraq" pageId="7" pageNumber="8">Iraq</collectingCountry>
and 
<collectingCountry id="34A676331C4EA31BFAEFFC2CFAF3FC04" box="[1313,1399,989,1015]" name="Turkey" pageId="7" pageNumber="8">Turkey</collectingCountry>
from the MNHN, HNHM, NHMW, PCRK and PCJM collections (
<figureCitation id="D48A2A261C4EA31BFC25FBF1FBB0FBE9" box="[1003,1076,1024,1050]" captionStart="Fig" captionStartId="8.[189,232,1883,1909]" captionTargetBox="[189,1397,278,1778]" captionTargetId="figure@8.[189,1398,273,1838]" captionTargetPageId="8" captionText="Fig. 4. Habitus images of Lacon spp., dorsal view. A. L. lepidopterus (Panzer, 1801), ♀. B. L. punctatus (Herbst, 1779), ♀. C. L. modestus (Boisduval, 1835), lectotype, ♀. D. L. mekrani (Candèze, 1889), ♀. E. L. unicolor (Candèze, 1874), lectotype, ♀. F. L. unicolor, ♂. G. L. nadaii Platia &amp; Németh, 2011, holotype, ♂. H. L. funebris (Solsky, 1881), ♂. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/3338052/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
E–G), and we can confirm the findings of 
<bibRefCitation id="28204B521C4EA31BFEB8FBD2FDCEFBCE" author="Prosvirov A. S." box="[374,586,1059,1085]" firstAuthor="Prosvirov" journalOrPublisher="Entomological Review" pageId="7" pageNumber="8" pagination="1120 - 1136" part="97" refId="ref6313" refString="Prosvirov A. S. 2017. New data on the fauna and systematics of Palaearctic species of the click-beetle subfamily Agrypninae Candeze, 1857 (Coleoptera, Elateridae). Entomological Review 97: 1120 - 1136. https: // doi. org / 10.1134 / S 0013873817080127" title="New data on the fauna and systematics of Palaearctic species of the click-beetle subfamily Agrypninae Candeze, 1857 (Coleoptera, Elateridae)" type="journal article" year="2017">Prosvirov (2017)</bibRefCitation>
. What is more, we found out that the differences in coloration and antennal morphology between 
<taxonomicName id="8BB14D201C4EA31BFDF8FBB6FD3AFB93" baseAuthorityName="Candeze" baseAuthorityYear="1874" box="[566,702,1094,1120]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFDF8FBB6FD3AFB93" box="[566,702,1094,1120]" italics="true" pageId="7" pageNumber="8">L. unicolor</emphasis>
</taxonomicName>
and 
<taxonomicName id="8BB14D201C4EA31BFD35FBB6FCEEFB93" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[763,874,1094,1120]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFD35FBB6FCEEFB93" box="[763,874,1094,1120]" italics="true" pageId="7" pageNumber="8">L. nadaii</emphasis>
</taxonomicName>
cannot be used for a reliable identification of these species and that there is a gradual intraspecific variability in these characters. Therefore, we propose that 
<taxonomicName id="8BB14D201C4EA31BFE94FB7CFC9DFB54" authority="Platia &amp; Nemeth, 2011" authorityName="Platia &amp; Nemeth" authorityYear="2011" box="[346,793,1165,1191]" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nadaii">
<emphasis id="7EC5EAB11C4EA31BFE94FB7CFE7FFB54" box="[346,507,1165,1191]" italics="true" pageId="7" pageNumber="8">Lacon nadaii</emphasis>
<bibRefCitation id="28204B521C4EA31BFDCDFB7CFC9DFB54" author="Platia G. &amp; Nemeth T." box="[515,793,1165,1191]" firstAuthor="Platia" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="7" pageNumber="8" pagination="65 - 106" part="103" refId="ref6180" refString="Platia G. &amp; Nemeth T. 2011. On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 103: 65 - 106." title="On some Palaearctic click beetles deposited in the Hungarian Natural History Museum (Coleoptera: Elateridae)" type="journal article" year="2011">Platia &amp; Németh, 2011</bibRefCitation>
</taxonomicName>
is a junior subjective synonym of 
<taxonomicName id="8BB14D201C4EA31BFB73FB7CFE0DFB39" authority="(Candeze, 1874)" authorityName="Candeze" authorityYear="1874" baseAuthorityName="Candeze" baseAuthorityYear="1874" class="Insecta" family="Elateridae" genus="Lacon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="7EC5EAB11C4EA31BFB73FB7CFAF3FB54" box="[1213,1399,1165,1191]" italics="true" pageId="7" pageNumber="8">Lacon unicolor</emphasis>
(
<bibRefCitation id="28204B521C4EA31BFF0BFB41FE04FB39" author="Candeze E. C. A." box="[197,384,1200,1226]" firstAuthor="Candeze" journalOrPublisher="Memoires de la Societe royale des Sciences de Liege" pageId="7" pageNumber="8" pagination="1 - 218" part="2" refId="ref5330" refString="Candeze E. C. A. 1874. Revision de la Monographie des Elaterides. Fasc. 1. Memoires de la Societe royale des Sciences de Liege 2: 1 - 218. https: // doi. org / 10.5962 / bhl. title. 47120" title="Revision de la Monographie des Elaterides. Fasc. 1" type="journal article" year="1874">Candèze, 1874</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>