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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1184.106698" ID-Pensoft-Pub="1313-2970-1184-301" ID-Pensoft-UUID="283A8869D1C3500A8A3C075E14584C90" ID-ZooBank="4EB4BEE3865E4CF29F7B6477D2DEF4AF" ModsDocID="1313-2970-1184-301" checkinTime="1700745464390" checkinUser="pensoft" docAuthor="Romero-Ortiz, Catalina, Sarmiento, Carlos E. &amp; Harvey, Mark S." docDate="2023" docId="51B6BA6598825A12BB5F24F108170BDD" docLanguage="en" docName="ZooKeys 1184: 301-326" docOrigin="ZooKeys 1184" docPubDate="2023-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1184.106698" docTitle="Oligowithius achagua Romero-Ortiz &amp; Sarmiento &amp; Harvey 2023, sp. nov." docType="treatment" docUuid="AF485E46-4657-4F22-8462-6CEFDB467C5C" docUuidSource="ZooBank" docVersion="1" id="283A8869D1C3500A8A3C075E14584C90" lastPageNumber="301" masterDocId="283A8869D1C3500A8A3C075E14584C90" masterDocTitle="A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia" masterLastPageNumber="326" masterPageNumber="301" pageNumber="301" updateTime="1700745464390" updateUser="pensoft">
<mods:mods id="CDEA9EBE3C845D95F304964FD29542E8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6E6CA3F38645691E4D3AF61A01A36E1">
<mods:title id="D3FCD7BE5B558FE3CC5C3B7259639E4E">A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia</mods:title>
</mods:titleInfo>
<mods:name id="2B07A783A89FB6F15F75B30D2A481D4F" type="personal">
<mods:role id="C82D7DC653E4CDACE56B653C4FF975C9">
<mods:roleTerm id="B8B24413115DD9B30A5C28A9AB4C21F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6EB7ABF246AC42ACEDB82B2DF07B94E">Romero-Ortiz, Catalina</mods:namePart>
<mods:nameIdentifier id="D9F7343CE4A1C27073F50B79E8DE7AB1" type="ORCID">https://orcid.org/0000-0001-8939-7814</mods:nameIdentifier>
<mods:affiliation id="70D0009DA897A6436CFFE6528F017F95">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
<mods:nameIdentifier id="3DD083B0E4E2021571B2FE128F764FFE" type="email">icromeroo@unal.edu.co</mods:nameIdentifier>
</mods:name>
<mods:name id="CCC78C57D53A2485C48555193996C7CB" type="personal">
<mods:role id="6B33E82534A78C55A24B2124FD618103">
<mods:roleTerm id="3F45DA6361159C47997E9B43879A409F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97B511F1C5D904B838CB647934C135CD">Sarmiento, Carlos E.</mods:namePart>
<mods:nameIdentifier id="039935BB8FE68EEA1DB81FDD006A35BA" type="ORCID">https://orcid.org/0000-0003-4012-8108</mods:nameIdentifier>
<mods:affiliation id="AEC2A3AF7DC7B44FED7B6B4D998A8053">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
</mods:name>
<mods:name id="D44CCA6214A5B2462B6CB00DEFB80497" type="personal">
<mods:role id="D66999E5A2DD991631945CD5606E8D5F">
<mods:roleTerm id="AF97E4693FFA2B30C944FB2BA6D9B9BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FEBD2BB369143FE702CF3A19D3BE265D">Harvey, Mark S.</mods:namePart>
<mods:nameIdentifier id="E4ED4485EFE1C331F4AE7890DE7DB76A" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
<mods:affiliation id="48C36D7FB20B5A27FBC6920AD3418511">Collections and Research, Western Australian Museum, Welshpool, Western Australia 6106, Australia &amp; School of Biological Sciences, The University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="D899F318F5812A53EF89B0393DDF8689">text</mods:typeOfResource>
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<mods:title id="B1A04B4A89C5D06013C487BBE4EB27C3">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="2C94B3A88D064EB84289305952C6D53C">
<mods:date id="34002ADF391B5F6763613985FDBED0B9">2023</mods:date>
<mods:detail id="D75587B670AAFB14E239040CF07A26DD" type="pubDate">
<mods:number id="A21EE5196685A4B856739A1D1C727DF7">2023-11-22</mods:number>
</mods:detail>
<mods:detail id="A7165A1A24A5BD6A630DFD0B6AD248CF" type="volume">
<mods:number id="21556599E979DA3D4CA044C3B6F3FA98">1184</mods:number>
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<mods:start id="78D6A7B91DA07CA77A164EC845A1E7F3">301</mods:start>
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<mods:url id="F058FFC41262D31A0E9D2CEA29ACF3CE">http://dx.doi.org/10.3897/zookeys.1184.106698</mods:url>
</mods:location>
<mods:classification id="861743A63D13FF29FC267688836830E5">journal article</mods:classification>
<mods:identifier id="D1BB3BDB6D98E1C22B165A72C2427ED6" type="DOI">http://dx.doi.org/10.3897/zookeys.1184.106698</mods:identifier>
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<subSection id="54F7D1291920B2F94542E4BCF05D7D77" pageId="0" pageNumber="301" type="taxonomic account">
<treatment id="51B6BA6598825A12BB5F24F108170BDD" LSID="urn:lsid:zoobank.org:act:AF485E46-4657-4F22-8462-6CEFDB467C5C" httpUri="http://treatment.plazi.org/id/51B6BA6598825A12BB5F24F108170BDD" lastPageNumber="301" pageId="0" pageNumber="301">
<subSubSection id="6BD191C841910BE7990FFB2015A133D4" pageId="0" pageNumber="301" type="nomenclature">
<paragraph id="EC5A0D552F91309304803BD02137548E" pageId="0" pageNumber="301">
<taxonomicName id="520F445B2BAC6BA746DEAF5836E2141E" LSID="https://zoobank.org/AF485E46-4657-4F22-8462-6CEFDB467C5C" authority="Romero-Ortiz &amp; Sarmiento &amp; Harvey, 2023" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua" status="sp. nov.">Oligowithius achagua</taxonomicName>
<taxonomicNameLabel id="F4826A51003089294FF4BC8F28CE2AB6" pageId="0" pageNumber="301">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8205FE13198940C1B57B3192F26AE988" pageId="0" pageNumber="301" type="description">
<paragraph id="85CDB85377FB9E07BADD7C2AF9C41287" pageId="0" pageNumber="301">
<figureCitation id="039A57DDEE179B758AE777577EAA3ED9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">Figs 1D</figureCitation>
<figureCitation id="40CECC730C638F732011BFC9A16E038C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the new species and the new record of Withiidae from Colombia." figureDoi="10.3897/zookeys.1184.106698.figure2" httpUri="https://binary.pensoft.net/fig/940871" pageId="0" pageNumber="301">, 2</figureCitation>
<figureCitation id="915B011FF37B0DF725E04883D4EAEEC5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">, 8</figureCitation>
<figureCitation id="7F821CDA58032BA77CF58AFE0F8BA798" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">, 9E, F</figureCitation>
<figureCitation id="04F3B7155501914C90337C7E56799BA5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Oligowithius achagua sp. nov. (male paratype, ICN-APs- 388) A right galea B sternites VII-VII C patch of glandular setae on sternite VIII D detail of the glandular setae on sternite VIII. Scale bars: 100 µm (A, B); 10 µm (C); 5 µm (D)." figureDoi="10.3897/zookeys.1184.106698.figure10" httpUri="https://binary.pensoft.net/fig/940879" pageId="0" pageNumber="301">, 10</figureCitation>
<figureCitation id="705E14ADA83B3AE6C95FBBD4F9CBB75F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Light micrograph of male genitalia of Oligowithius achagua sp. nov. (holotype, ICN-Aps- 388). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1184.106698.figure11" httpUri="https://binary.pensoft.net/fig/940880" pageId="0" pageNumber="301">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E42218B26676826E7458744837C547C8" pageId="0" pageNumber="301" type="materials_examined">
<paragraph id="48A68C0327F612CBB3AF8E599CA8A0F0" pageId="0" pageNumber="301">Material examined.</paragraph>
<paragraph id="7F4703FB89F4A6C8FF998015D4EF9F87" pageId="0" pageNumber="301">
<materialsCitation id="FF7BF6BE451D50E1719AF94F3A9B8304" collectingDate="2012-04-22" collectorName="D. Martinez" country="Colombia" elevation="160" location="Carimagua" municipality="Puerto Gaitan" specimenCount="♂" stateProvince="Meta" typeStatus="Holotype">
<emphasis id="9D07BDA4060367100688B032D003BF34" bold="true" pageId="0" pageNumber="301">
<emphasis id="F649382C8C8F517CACE0E84401C14AA2" italics="true" pageId="0" pageNumber="301">
<typeStatus id="48147C1057391A1CF711B196BAC721AE">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="A33F4C533F1884553D21B67068F85EBB" name="Colombia">Colombia</collectingCountry>
•
<specimenCount id="CAB90A9EBB665B9AF119741631FAF6A5">♂</specimenCount>
;
<collectingRegion id="27E323959FA60C9055992CD9E4CCC252" country="Colombia" name="Meta">Meta</collectingRegion>
,
<collectingMunicipality id="3B5F4FB790577999637DE604C7B48D87">
Puerto
<normalizedToken id="51AADB4DFCF04096F01F2BCE06BABEE8" originalValue="Gaitán">Gaitan</normalizedToken>
</collectingMunicipality>
,
<location id="34684555872AEDDE51A0898B4806526C" LSID="urn:lsid:plazi:treatment:51B6BA6598825A12BB5F24F108170BDD:34684555872AEDDE51A0898B4806526C" country="Colombia" municipality="Puerto Gaitan" name="Carimagua" stateProvince="Meta">Carimagua</location>
;
<quantity id="BB1B7C68C869CE72938C6FEA76D700D6" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">
<elevation id="3605C69C208AA93584034A56656D24D8" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">160 m</elevation>
</quantity>
;
<collectingDate id="9253B9203CF6DB78D488B0B80A5EE39C" value="2012-04-22">22 Apr. 2012</collectingDate>
;
<collectorName id="639BC65570D38AB94D0D61146D30C9D1">D. Martinez</collectorName>
leg.; estuary on the eastern plains; ICN-APs-388.
</materialsCitation>
</paragraph>
<paragraph id="CF56369B03E753475793C399495A1D61" pageId="0" pageNumber="301">
<materialsCitation id="C5049D65EAEF8EDFD62F18F57E8D6D47" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="D. Martinez" country="Colombia" elevation="160" location="Meta" municipality="Puerto Gaitan" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Meta" typeStatus="Paratypes">
<emphasis id="977115F8C98BA6DBA3AA61AC84698B0F" bold="true" pageId="0" pageNumber="301">
<emphasis id="AEB7126397AC95592AE7E4351D474268" italics="true" pageId="0" pageNumber="301">
<typeStatus id="BD933D1E7321759CCD4AAC3F6A85803A">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="7A8F6E7D8D3481177C111112899EEC62" name="Colombia">Colombia</collectingCountry>
•
<specimenCount id="725F18379DFC8BDE5DBE4646C50E7EB0" type="male">1 ♂</specimenCount>
and
<specimenCount id="7FDFCD1E1A5796E041C52350CB9E8817" type="female">1 ♀</specimenCount>
;
<collectingRegion id="A8D071C5BDBD2917074846B227802547" country="Colombia" name="Meta">Meta</collectingRegion>
, same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D91A2AD9BD0CF317FC246A3BF3F96E3A" pageId="0" pageNumber="301" type="diagnosis">
<paragraph id="AB22D09C69BC8DA5FC114AC2D940E94E" pageId="0" pageNumber="301">Diagnosis.</paragraph>
<paragraph id="6F527478EF55492179644AE773A715C3" pageId="0" pageNumber="301">
<taxonomicName id="3E5689266181DEC48AC7BBF303D6DCA3" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="3D55516137BCD9E1DDC06C8DCE99CB81" italics="true" pageId="0" pageNumber="301">Oligowithius achagua</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName id="09DD469E053F8B244717CEC2AF9FA145" lsidName="O. abnormis" pageId="0" pageNumber="301" rank="species" species="abnormis">
<emphasis id="58249D3EC485DBC7B5C02DA0A11DC317" italics="true" pageId="0" pageNumber="301">O. abnormis</emphasis>
</taxonomicName>
by its smaller size (1.7 mm vs 2.18 mm in
<taxonomicName id="5C4BC4E305C7FA7EB7DBCC31040B340A" lsidName="O. achagua" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="A624D2773C53C276BD7D12E68F6BA7C2" italics="true" pageId="0" pageNumber="301">O. achagua</emphasis>
</taxonomicName>
), more slender patella in
<taxonomicName id="969BEBA7643E5447580D7395D2D6E185" lsidName="O. achagua" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="5F73C225DAA38FEA60FDFB434FE08300" italics="true" pageId="0" pageNumber="301">O. achagua</emphasis>
</taxonomicName>
(3.05
<normalizedToken id="647F3F02D278E85B2090DB4E4243D5E7" originalValue="×">x</normalizedToken>
longer than broad) than in
<taxonomicName id="D6CCC5A737DC0BACFBA14CA29A4C901D" lsidName="O. abnormis" pageId="0" pageNumber="301" rank="species" species="abnormis">
<emphasis id="9FB151B6B64D68004436DA6263F21B23" italics="true" pageId="0" pageNumber="301">O. abnormis</emphasis>
</taxonomicName>
(2.74
<normalizedToken id="755920B05B216EB331328479CA5072FE" originalValue="×">x</normalizedToken>
), and a stouter chela in
<taxonomicName id="6D7E22A6415F131DBD432F109DE46237" lsidName="O. abnormis" pageId="0" pageNumber="301" rank="species" species="abnormis">
<emphasis id="9162FFA460C044BF828D8F0A38806236" italics="true" pageId="0" pageNumber="301">O. abnormis</emphasis>
</taxonomicName>
(3.70
<normalizedToken id="44BDDD0BDF405684E299D11EB0670325" originalValue="×">x</normalizedToken>
longer than broad) than in
<taxonomicName id="CC99BE641F7F9CDAF51B9F29AFCBB563" lsidName="O. achagua" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="F24D4D330629979ED75CEE7FD728A1EB" italics="true" pageId="0" pageNumber="301">O. achagua</emphasis>
</taxonomicName>
(4.05
<normalizedToken id="82B555A3B568102C4D1036A6281EC875" originalValue="×">x</normalizedToken>
longer than broad).
</paragraph>
</subSubSection>
<subSubSection id="70E28D2B8EB057D02F7184D6A892E484" pageId="0" pageNumber="301" type="description">
<paragraph id="6EEA6C9CA45DE876B8917F6F2B79FE7F" pageId="0" pageNumber="301">Description.</paragraph>
<paragraph id="FE1C3CAD0AC5EA653B0C2717FD3AC066" pageId="0" pageNumber="301">
<emphasis id="D0539B20A6153450FDBE171DAB2251C7" bold="true" pageId="0" pageNumber="301">Adults.</emphasis>
Color: yellowish brown; carapace and pedipalp reddish brown; legs yellowish, paler than the abdomen, darker in the edges; carapace metazone without paired pale spots (Fig.
<figureCitation id="6299978C40A2FD9023938A54C7C81ABC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">1D</figureCitation>
).
</paragraph>
<paragraph id="38CAC9F9CBF59D68630A972BA14302E9" pageId="0" pageNumber="301">
<emphasis id="644A80C7DD2ED3BEC3F209919090F1FA" bold="true" italics="true" pageId="0" pageNumber="301">Dimensions</emphasis>
(mm): male: holotype (followed by male paratype): body length 2.18 (2.10). Pedipalps: trochanter 0.33/0.16 (0.27/0.15), femur 0.58/0.15 (0.54/0.15), patella 0.54/0.18 (0.50/0.16), chela (with pedicel) 0.94/0.23 (0.88/0.22), chela (without pedicel) 0.88 (0.84), hand (without pedicel) length 0.48 (0.43), movable finger length 0.45 (0.43). Chelicera 0.19, movable finger length 0.16. Carapace 0.74/0.58 (0.68/0.56) (width at medial area); eyespots diameter 0.08. Leg I: femur 0.14/0.14, patella 0.26/0.14, tibia 0.26/0.08, tarsus 0.26/0.06. Leg IV: femur + patella 0.47/0.16, tibia 0.34/0.11, tarsus 0.33/0.06, TS 0.76.
</paragraph>
<paragraph id="5357E25063A6EA69DC028DC466867EB1" pageId="0" pageNumber="301">
<emphasis id="D1BCF215D3943290A7B06E10B2EBEEA2" bold="true" pageId="0" pageNumber="301">Female</emphasis>
: paratype: body length 2.36. Pedipalps: trochanter 0.28/0.16, femur 0.54/0.15, patella 0.52/0.18, chela (with pedicel) 0.94/0.25, chela (without pedicel) 0.88, hand (without pedicel) length 0.48, movable finger length 0.44. Carapace 0.73/0.58.
</paragraph>
<paragraph id="537DAC55BDC5A658BC626410E04E48F9" pageId="0" pageNumber="301">
<emphasis id="AA9E1EF9F9EF768F96250E3D1DEA1EAE" bold="true" italics="true" pageId="0" pageNumber="301">Carapace</emphasis>
(Fig.
<figureCitation id="7713BA71F245E2C3AC2DFCCFA136C77E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8A</figureCitation>
): 1.28 (1.20) (♂), 1.07 (♀)
<normalizedToken id="AAA800BE1BAD70CA74E7A1BB957D43F4" originalValue="×">x</normalizedToken>
longer than broad; posterior lateral margins slightly widened; with two non-corneate eyes; with 55 (♂) setae, including 2 (♂) near anterior margin, 11 (♂) near posterior margin, 21 in the medial zone and 23 in the anterior region; with two distinct furrows; posterior furrow slightly closer to posterior carapace margin than to median furrow (Fig.
<figureCitation id="CBC1CC8D6D657AB34F1391832484637D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8A</figureCitation>
).
</paragraph>
<paragraph id="7F3581D814C8718680E1449003A5231B" pageId="0" pageNumber="301">
<emphasis id="BCD67E0FFC701AE23C2E6B692BF023C1" bold="true" italics="true" pageId="0" pageNumber="301">Chelicera</emphasis>
(Fig.
<figureCitation id="67626C4EF2A1FC36E25ECD22B924332E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8B</figureCitation>
): with five setae on hand,
<emphasis id="F0138A98D7BD359F81DCD4A6B30ECB17" italics="true" pageId="0" pageNumber="301">sb</emphasis>
missing,
<emphasis id="45E93556D30750C3BF5CF7049C701846" italics="true" pageId="0" pageNumber="301">b</emphasis>
slightly denticulate, all others acuminate; movable finger with one subdistal seta; galea of male broken in holotype, galea with multiple rami in paratype (Fig.
<figureCitation id="08012845EB7AE9A8A781503B5A32D6F3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Oligowithius achagua sp. nov. (male paratype, ICN-APs- 388) A right galea B sternites VII-VII C patch of glandular setae on sternite VIII D detail of the glandular setae on sternite VIII. Scale bars: 100 µm (A, B); 10 µm (C); 5 µm (D)." figureDoi="10.3897/zookeys.1184.106698.figure10" httpUri="https://binary.pensoft.net/fig/940879" pageId="0" pageNumber="301">10A</figureCitation>
); rallum of four blades, the most distal blade with several serrations on leading edge, other blades smooth (Fig.
<figureCitation id="D1CE3E5830737EFCDB3B06482DCD65AB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8E</figureCitation>
); two lyrifissures on dorsal side.
</paragraph>
<paragraph id="57A1F1510CBD91C2A02FD322CA5AC624" pageId="0" pageNumber="301">
<emphasis id="FC67EE0929B87DD663BB2B135FC41232" bold="true" italics="true" pageId="0" pageNumber="301">Pedipalp</emphasis>
(Fig.
<figureCitation id="57786C86BDF3C29F0945A94CA6A349F3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8C, H</figureCitation>
): trochanter, femur, patella and dorsal chelal hand granulate, ventral chelal hand and fingers smooth; dorsal setae clavate and denticulate; trochanter 2.05
<normalizedToken id="75DA36C57C1921AA8C724458D33BC4E1" originalValue="×">x</normalizedToken>
(1.79) (♂), 1.75
<normalizedToken id="DA40DF3A2FBFBB5951808DD849703FA9" originalValue="×">x</normalizedToken>
(♀), femur 3.79
<normalizedToken id="DD7A2479101A67362A44EDE82D86E9F0" originalValue="×">x</normalizedToken>
(3.58) (♂), 3.53
<normalizedToken id="DE1AC2BBAAB0B18363FC8460CC789C5A" originalValue="×">x</normalizedToken>
(♀), patella 3.05
<normalizedToken id="57A8DC08B729AD393BCE497910BDA5FA" originalValue="×">x</normalizedToken>
(3.10) (♂), 2.95
<normalizedToken id="8FEEF82D844FAA3C28A28AE88D679090" originalValue="×">x</normalizedToken>
(♀), chela (with pedicel) 4.05
<normalizedToken id="0E82FCD46CA385A932FED3B8384B500B" originalValue="×">x</normalizedToken>
(4.07) (♂), 3.79
<normalizedToken id="39A0774907770B4706207604A5417AE6" originalValue="×">x</normalizedToken>
(♀), chela (without pedicel) 3.79
<normalizedToken id="F5C286A5B7D3F676CBD9F6EF04DC745F" originalValue="×">x</normalizedToken>
(3.89) (♂), 3.55
<normalizedToken id="AF95124B6E77E012D2F38E616112E49F" originalValue="×">x</normalizedToken>
(♀), hand 2.07
<normalizedToken id="CA2ABE08F0C7CE217613C57892C0764D" originalValue="×">x</normalizedToken>
(2.0) (♂), 1.94
<normalizedToken id="05833A37BA872950B76CFFBC330B942D" originalValue="×">x</normalizedToken>
(♀)longer than broad, movable finger 0.93
<normalizedToken id="13E623BC9C9D938A8BBE64735BFFCCB1" originalValue="×">x</normalizedToken>
(1.00) (♂), 0.92
<normalizedToken id="3E92A4B68F437A3ADD582EAD63E594B4" originalValue="×">x</normalizedToken>
(♀)longer than hand. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria (Fig.
<figureCitation id="982DE5CBA8E7DF0BF1FAB523001AA509" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8H</figureCitation>
):
<emphasis id="8E0B75D801D8958ECE88014C4CEFDD4C" italics="true" pageId="0" pageNumber="301">eb</emphasis>
and
<emphasis id="953CC19DF2E6D762A482CCDAD1085F5F" italics="true" pageId="0" pageNumber="301">esb</emphasis>
situated basally, as well as
<emphasis id="0EB80C077F02E1FF3BBBDD5B0E1FB2D1" italics="true" pageId="0" pageNumber="301">ib</emphasis>
and
<emphasis id="BF0810FB097A5FDB0FACAF5CFD55DF79" italics="true" pageId="0" pageNumber="301">ist</emphasis>
;
<emphasis id="084515C24644D76A2685A10A1094E13C" italics="true" pageId="0" pageNumber="301">est</emphasis>
situated midway between
<emphasis id="B1B32EA3911A8F8D0465185D6AF53C31" italics="true" pageId="0" pageNumber="301">it</emphasis>
and
<emphasis id="19B0369C7E02F00C7F83B75597E05644" italics="true" pageId="0" pageNumber="301">isb</emphasis>
in the middle of the finger,
<emphasis id="E0B1DB7B91CBEBDD5043AA30A893F6BE" italics="true" pageId="0" pageNumber="301">et</emphasis>
close to fingertip;
<emphasis id="569D6DD7758F3241D67AFCA608442F72" italics="true" pageId="0" pageNumber="301">b</emphasis>
and
<emphasis id="35C6D98AA45DBB0EB36AAC948716FE5A" italics="true" pageId="0" pageNumber="301">sb</emphasis>
situated near one another,
<emphasis id="812948CD8686C1958629E1A2147F29E8" italics="true" pageId="0" pageNumber="301">st</emphasis>
located midway between
<emphasis id="ADED79072A3069114E8C4ACA61F03565" italics="true" pageId="0" pageNumber="301">sb</emphasis>
and
<emphasis id="9A1AB5E74D361489B8BC23CF2ED4CE14" italics="true" pageId="0" pageNumber="301">t</emphasis>
,
<emphasis id="351BC4793875F383DA12CE084F29BCF6" italics="true" pageId="0" pageNumber="301">t</emphasis>
parallel to
<emphasis id="287DC5270103640CB5E7390F96603C64" italics="true" pageId="0" pageNumber="301">it</emphasis>
. Venom apparatus not visible, venom ducts not visible in ♂; nodus ramosus not visible. Retrolateral margin of fixed finger with two double sense-spots (Fig.
<figureCitation id="41EF38F11FFB91DB37699139B0338BE4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8H</figureCitation>
) situated midway between
<emphasis id="D99DE348F5D429A42BA7CAE899ABBFE3" italics="true" pageId="0" pageNumber="301">esb</emphasis>
and
<emphasis id="F23B374FBA780ED42D70AB15DC801640" italics="true" pageId="0" pageNumber="301">est</emphasis>
, movable finger with one double sense-spot close to
<emphasis id="5A6E5C748F83C15362A1DEA9E7838BD1" italics="true" pageId="0" pageNumber="301">sb</emphasis>
. Chelal teeth rounded; fixed finger with 40 (♂) teeth; movable finger with 43 (♂) teeth; accessory teeth absent.
</paragraph>
<paragraph id="A97B9BA2E6638806610F8B8ADEDB0A47" pageId="0" pageNumber="301">
<emphasis id="8B473388304F84278F29875D0718492A" bold="true" italics="true" pageId="0" pageNumber="301">Coxal region</emphasis>
: coxal chaetotaxy: ♂, 6: 7: 9: 18; maxilla with 19 setae including two apical setae and one very small internal, sub-oral seta; median maxillary lyrifissure medial-anterior in position, posterior lyrifissure present.
</paragraph>
<paragraph id="6F149AF4EB56F085AFEA3E9550595925" pageId="0" pageNumber="301">
<emphasis id="A301AD11387CB4D0AB4C9DDE0D3176DA" bold="true" italics="true" pageId="0" pageNumber="301">Legs</emphasis>
(Fig.
<figureCitation id="891225990CD4792F7D0AC90AB2716371" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8F, G</figureCitation>
): junction between femora and patellae I and II parallel, junction in legs III and IV oblique; tarsal tactile seta of leg IV situated distally, 0.76 (♂) of tarsus length (Fig.
<figureCitation id="0AB4BA9983C5B3C19C068BE92F92789B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8G</figureCitation>
); arolium slightly shorter than claws. Ratios: leg I: femur 1
<normalizedToken id="ACBD6C3BCA6745F2E631E3E1DF445DBC" originalValue="×">x</normalizedToken>
, patella 1.94
<normalizedToken id="99EF60851B22B8939E4E52401D805775" originalValue="×">x</normalizedToken>
, tibia 3.30
<normalizedToken id="8B35A454997F6233A1FE8105047FBF46" originalValue="×">x</normalizedToken>
, tarsus 4.72
<normalizedToken id="5CB6257C55FA145321A4EE6AA7D45437" originalValue="×">x</normalizedToken>
deeper than broad; Leg IV: femur + patella 2.85
<normalizedToken id="D397EFD85271E22A231E901F38536087" originalValue="×">x</normalizedToken>
, tibia 3.00
<normalizedToken id="3A50ED221E7E8C0F9B020C4AF115CD9F" originalValue="×">x</normalizedToken>
, tarsus 5.13
<normalizedToken id="6FD38F5CBB592B54AA348A807740E250" originalValue="×">x</normalizedToken>
deeper than broad.
</paragraph>
<paragraph id="17C9243A1C2975F728C6DF8A2E5D81D2" pageId="0" pageNumber="301">
<emphasis id="711C9F596E29507C7F695867FC53C368" bold="true" italics="true" pageId="0" pageNumber="301">Abdomen</emphasis>
(Fig.
<figureCitation id="75AC1A425DC6A2F5B3EA318AE2F21F22" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Oligowithius achagua sp. nov. (male holotype, ICN-Aps- 388) A carapace B chelicera C left pedipalp D abdomen, sternites E rallum F left leg I G left leg IV H right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 8 µm (A, C, D, F-H); 2 µm (B, E). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure." figureDoi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301">8D</figureCitation>
): first four tergites entire, the others with faint medial suture. Tergal chaetotaxy: ♂, 12: 14: 13: 18: 18: 19: 19: 18: 19: 15: 6: 2, 2; all setae foliate; mostly uniseriate but some tergites with a few setae placed anteriorly. All sternites entire, except for the last three with a faint medial suture. Sternal chaetotaxy: ♂, 10: (1) 6 (1): (1) 8 (1): 14: 12: 8 + 13/13
<emphasis id="CE0CC79D460A17C1913D0D2C7A7715F5" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 8 + 18/16
<emphasis id="703705B5919C5E19FB54023B3C1F46D5" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 8: 8: 6 (including two clavate setae): 2; sternites VII-VIII of ♂ with two small patch circles of glandular setae each (Figs
<figureCitation id="72DBC1C4CE53CE29262E4F604D5FD18F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">9F</figureCitation>
,
<figureCitation id="8D4DD9F1B4A983A2F040B8028122ACC0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Oligowithius achagua sp. nov. (male paratype, ICN-APs- 388) A right galea B sternites VII-VII C patch of glandular setae on sternite VIII D detail of the glandular setae on sternite VIII. Scale bars: 100 µm (A, B); 10 µm (C); 5 µm (D)." figureDoi="10.3897/zookeys.1184.106698.figure10" httpUri="https://binary.pensoft.net/fig/940879" pageId="0" pageNumber="301">10B-D</figureCitation>
), paratypes with one circular patch of glandular setae each (Fig.
<figureCitation id="D41B96B55E719C04FCCE8FCAD098464A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">9E</figureCitation>
); setae uniseriate and acuminate; ♂ without paired invaginations on anterior margins of sternites.
</paragraph>
<caption id="09662086855CD1579C819538F55DA2B0" doi="10.3897/zookeys.1184.106698.figure10" httpUri="https://binary.pensoft.net/fig/940879" pageId="0" pageNumber="301" start="Figure 10" startId="F10">
<paragraph id="791CFCF7E87B38492032579A821BBB76" pageId="0" pageNumber="301">
<emphasis id="CDC01C08DBCD4DB03B69A05BE481F402" bold="true" pageId="0" pageNumber="301">Figure 10.</emphasis>
<taxonomicName id="E5E71BB6F61E56FE0B4D9B7482E40DF3" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="247E3874A752CE35683718E689EC58BC" italics="true" pageId="0" pageNumber="301">Oligowithius achagua</emphasis>
</taxonomicName>
sp. nov. (male paratype, ICN-APs-388)
<emphasis id="03192EBA833A246C8214539D3DC56DC8" bold="true" pageId="0" pageNumber="301">A</emphasis>
right galea
<emphasis id="853DCA20FA598819698D34DC56E40AF4" bold="true" pageId="0" pageNumber="301">B</emphasis>
sternites VII-VII
<emphasis id="F27AA8B0570B2ACBCAAFEF2DE9596871" bold="true" pageId="0" pageNumber="301">C</emphasis>
patch of glandular setae on sternite VIII
<emphasis id="BE4BC73171CE512423534980AECC4DB8" bold="true" pageId="0" pageNumber="301">D</emphasis>
detail of the glandular setae on sternite VIII. Scale bars: 100
<normalizedToken id="BC2520DAFFCC930E9A9D4D44BA299E7F" originalValue="µm">µm</normalizedToken>
(
<emphasis id="934E4ED35D38ED7241272ABEF6AC4528" bold="true" pageId="0" pageNumber="301">A, B</emphasis>
); 10
<normalizedToken id="AF4C1A0E90D321A20BFCFDE7C19A22B0" originalValue="µm">µm</normalizedToken>
(
<emphasis id="12B18FB425AC14B7D3F2DE697FAC4F14" bold="true" pageId="0" pageNumber="301">C</emphasis>
); 5
<normalizedToken id="49BAEAA5A7F3AE17201735EC005A3514" originalValue="µm">µm</normalizedToken>
(
<emphasis id="9D8B320C6811336505DCF5AEE1EF6AEB" bold="true" pageId="0" pageNumber="301">D</emphasis>
).
</paragraph>
</caption>
<paragraph id="37E1D1EF7CF717564176FE8D6C6D9A96" pageId="0" pageNumber="301">
<emphasis id="271CFEED71D94BCAC39323E458759F78" bold="true" italics="true" pageId="0" pageNumber="301">Genitalia</emphasis>
: simple structure with most of the components extremely reduced to chitinized lateral apodemes that do not merge with the dorsal apodemes. The level of sclerotization of the ejaculatory channel is weak; however, it is projected from the lateral apodemes, which allows us to classify
<taxonomicName id="300BA7DF1B5CA5131C3766B6E94B0FC1" authorityName="Beier" authorityYear="1936" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="genus">
<emphasis id="BB808169305E03F90F76E8C746166DEE" italics="true" pageId="0" pageNumber="301">Oligowithius</emphasis>
</taxonomicName>
as a
<taxonomicName id="C82D3041271D27CF57C77CEDFA75F92B" lsidName="" pageId="0" pageNumber="301" rank="tribe" tribe="Cacodemoniini">Cacodemoniini</taxonomicName>
(Fig.
<figureCitation id="3C69D8A73514DCFF4BA42F4F87A70293" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Light micrograph of male genitalia of Oligowithius achagua sp. nov. (holotype, ICN-Aps- 388). Scale bar: 100 µm." figureDoi="10.3897/zookeys.1184.106698.figure11" httpUri="https://binary.pensoft.net/fig/940880" pageId="0" pageNumber="301">11</figureCitation>
).
</paragraph>
<caption id="216FF851F556EE413E5518C27B2C6D31" doi="10.3897/zookeys.1184.106698.figure11" httpUri="https://binary.pensoft.net/fig/940880" pageId="0" pageNumber="301" start="Figure 11" startId="F11">
<paragraph id="7462502D3138F25C8A944E541FD278D9" pageId="0" pageNumber="301">
<emphasis id="85DE8582C9DE863944FCCD719D3D7F8F" bold="true" pageId="0" pageNumber="301">Figure 11.</emphasis>
Light micrograph of male genitalia of
<taxonomicName id="A124816ECAECAB98A7BCEAC2501D829A" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="D9C006A6697321D95A8D384853652866" italics="true" pageId="0" pageNumber="301">Oligowithius achagua</emphasis>
</taxonomicName>
sp. nov. (holotype, ICN-Aps-388). Scale bar: 100
<normalizedToken id="AF8918A14C63D37D5BD85A3AB13CC10C" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="24DCAB33B7B61648EB89220843DD4900" pageId="0" pageNumber="301" type="etymology">
<paragraph id="655640D936146D3EF6ECC8B2A1DD32FE" pageId="0" pageNumber="301">Etymology.</paragraph>
<paragraph id="79E7F8D43BAAA5CFC2C740914A36315A" pageId="0" pageNumber="301">This species is named after the indigenous people, original inhabitants of the Meta, Vichada and the Venezuelan Llanos, the Achaguas. The name should be treated as a noun in apposition.</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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