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<document id="D172FE7F7DE1E0E99AEDD62F3A7BFCB7" ID-CLB-Dataset="7896" ID-DOI="10.11646/zootaxa.4875.1.1" ID-GBIF-Dataset="a75d6e42-221a-4df9-9a7d-5f33dbff6a56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4578779" ID-ZooBank="278200CE-E16F-45B4-9A89-60C2052415C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610024385246" checkinUser="plazi" docAuthor="Görn, Sebastian" docDate="2020" docId="807787F6B829FFC249A1D00DFA873E41" docLanguage="en" docName="zootaxa.4875.1.1.pdf" docOrigin="Zootaxa 4875 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heptodonta wiesneri, sp. nov." docType="treatment" docVersion="8" lastPageNumber="27" masterDocId="7C4EFF8EB83EFFD84936D36BFFA83A3C" masterDocTitle="Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="24" updateTime="1698923305811" updateUser="ExternalLinkService">
<mods:mods id="A6AF4535B5B5FDF7A438A8067BE8942F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B80137B18FBDF089F34167FD21012F60">
<mods:title id="4DA6C7E1B6C9159CE03F8B339E087473">Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name id="2FD3CD113F9F76F52A53D5935B045AEB" type="personal">
<mods:role id="4EAB418E2D70B239B2F80254C6DC1737">
<mods:roleTerm id="224954094709160CEBFD355F5A620A9E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2ADB1B40A7776E6EA85A8D71CC64473">Görn, Sebastian</mods:namePart>
</mods:name>
<mods:typeOfResource id="26944C66AE56E5D134A38C13302A0DCD">text</mods:typeOfResource>
<mods:relatedItem id="0F78463C290DE351AA8E93BC4A562D1B" type="host">
<mods:titleInfo id="E85C9A81279EE1045F884CDDC2BD0B71">
<mods:title id="441091D8B6E8E4DCD1EFDF1D0C57A94F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B7110DFC2364E8FAB5168C7607015D3C">
<mods:date id="803F3F773CCA23786ADBE7109167AA92">2020</mods:date>
<mods:detail id="D2F56635BC27B9C45E1D682229A51B31" type="pubDate">
<mods:number id="9C59D3E004508349AA78CCFA3827EB0A">2020-11-06</mods:number>
</mods:detail>
<mods:detail id="4CD7854ED5BA678FE1046190D57A80C1" type="volume">
<mods:number id="86304E7AD81B940D44E52A40790222BB">4875</mods:number>
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<mods:detail id="8D85C2E10D743E508DEEB09A03E41470" type="issue">
<mods:number id="0842BA275B49E7451ABA115808814BCC">1</mods:number>
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<mods:start id="F011FF4F7543862584FEBE447F38BED0">1</mods:start>
<mods:end id="68F7D8E56B50A3321090A995A7F71EF0">62</mods:end>
</mods:extent>
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<mods:classification id="C8BD88118EE40047C48C97721446300F">journal article</mods:classification>
<mods:identifier id="E016E31E4D0ED0BEF9E2786CBE311604" type="CLB-Dataset">7896</mods:identifier>
<mods:identifier id="9EC4878A9DEBCC1E59F15FF8376BD6D8" type="DOI">10.11646/zootaxa.4875.1.1</mods:identifier>
<mods:identifier id="4B808243AA5CE5446561D36C4F88EC97" type="GBIF-Dataset">a75d6e42-221a-4df9-9a7d-5f33dbff6a56</mods:identifier>
<mods:identifier id="81FEF5513BC9F2374AB8BC3E17AB47D2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E728EB23CE07A3C5B368AFE64360C106" type="Zenodo-Dep">4578779</mods:identifier>
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</mods:mods>
<treatment id="807787F6B829FFC249A1D00DFA873E41" ID-DOI="http://doi.org/10.5281/zenodo.4579795" ID-GBIF-Taxon="179988680" ID-Zenodo-Dep="4579795" LSID="urn:lsid:plazi:treatment:807787F6B829FFC249A1D00DFA873E41" httpUri="http://treatment.plazi.org/id/807787F6B829FFC249A1D00DFA873E41" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
<subSubSection id="40C4656BB829FFCF49A1D00DFE5E39BD" box="[151,502,870,897]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="086136E0B829FFCF49A1D00DFE5E39BD" blockId="23.[151,502,870,933]" box="[151,502,870,897]" pageId="23" pageNumber="24">
<heading id="5329818CB829FFCF49A1D00DFE5E39BD" bold="true" box="[151,502,870,897]" fontSize="11" level="2" pageId="23" pageNumber="24" reason="4">
<emphasis id="3AAAEAF2B829FFCF49A1D00DFE5E39BD" bold="true" box="[151,502,870,897]" pageId="23" pageNumber="24">
<taxonomicName id="CFDE4D63B829FFCF49A1D00DFE3939BD" ID-CoL="8M7MJ" authority="Gorn. 2020" box="[151,401,870,897]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="3AAAEAF2B829FFCF49A1D00DFE3939BD" bold="true" box="[151,401,870,897]" italics="true" pageId="23" pageNumber="24">Heptodonta wiesneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21995789B829FFCF48AFD00CFE5E39BD" box="[409,502,871,897]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB829FFCF49A1D0E0FEB33999" box="[151,283,907,933]" pageId="23" pageNumber="24" type="description">
<paragraph id="086136E0B829FFCF49A1D0E0FEB33999" blockId="23.[151,502,870,933]" box="[151,283,907,933]" pageId="23" pageNumber="24">
<figureCitation id="90E52A65B829FFCF49A1D0E0FEB33999" box="[151,283,907,933]" captionStart="FIGURES 62–67" captionStartId="25.[151,264,1931,1955]" captionTargetBox="[152,1435,184,1872]" captionTargetId="figure-15@25.[151,1435,181,1908]" captionTargetPageId="25" captionText="FIGURES 62–67. Heptodonta wiesneri sp. nov., Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="23" pageNumber="24">Figs. 62–67</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB829FFCF49A1D0B8FF5F3EF9" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="086136E0B829FFCF49A1D0B8FD8A39D1" blockId="23.[151,1437,978,2013]" box="[151,546,978,1005]" pageId="23" pageNumber="24">
<materialsCitation id="B8B63CBDB829FFCF49A1D0B8FDB639D1" ID-GBIF-Occurrence="3045076445" box="[151,542,978,1005]" country="Philippines" location="Negros" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
<emphasis id="3AAAEAF2B829FFCF49A1D0B8FE9E39D0" bold="true" box="[151,310,978,1005]" pageId="23" pageNumber="24">
<typeStatus id="D7658842B829FFCF49A1D0B8FF7A39D1" box="[151,210,979,1005]" pageId="23" pageNumber="24">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="70C97670B829FFCF480AD0B8FE1739D1" box="[316,447,979,1005]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
:
<location id="0D01603BB829FFCF48FFD0B8FDB639D1" LSID="urn:lsid:plazi:treatment:807787F6B829FFC249A1D00DFA873E41:0D01603BB829FFCF48FFD0B8FDB639D1" box="[457,542,979,1005]" country="Philippines" name="Negros" pageId="23" pageNumber="24">Negros</location>
</materialsCitation>
.
</paragraph>
<paragraph id="086136E0B829FFCF49F1D09CFF5F3EF9" blockId="23.[151,1437,978,2013]" pageId="23" pageNumber="24">
<emphasis id="3AAAEAF2B829FFCF49F1D09CFEDD3E2C" bold="true" box="[199,373,1014,1041]" pageId="23" pageNumber="24">Type material.</emphasis>
<materialsCitation id="B8B63CBDB829FFCF484BD09CFB7D3E09" ID-GBIF-Occurrence="3045076436" collectorName="Romeo" country="Philippines" determinerName="J. Wiesner" location="Negros oriental" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Negros Oriental" typeStatus="holotype">
<emphasis id="3AAAEAF2B829FFCF484BD09CFE423E2D" bold="true" box="[381,490,1015,1041]" pageId="23" pageNumber="24">
<typeStatus id="D7658842B829FFCF484BD09CFE423E2D" box="[381,490,1015,1041]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="1ED8FD69B829FFCF48CFD09CFDA63E2D" box="[505,526,1015,1041]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
in JWCW, labelled: “
<collectingCountry id="70C97670B829FFCF4A34D09CFC053E2D" box="[770,941,1015,1041]" name="Philippines" pageId="23" pageNumber="24">PHILIPPINES</collectingCountry>
, VII.85, /
<collectingRegion id="CA1AF802B829FFCF4D29D09CFB783E2D" box="[1055,1232,1015,1041]" country="Philippines" name="Negros Oriental" pageId="23" pageNumber="24">Negros oriental</collectingRegion>
, / leg.
<collectorName id="A52B5336B829FFCF4C2DD09CFAC43E2D" box="[1307,1388,1015,1041]" pageId="23" pageNumber="24">Romeo</collectorName>
Lu- mawig” [yellow margin, printed] // “melanopyga / Schaum / det.
<determinerName id="FDBDDA24B829FFCF4A44D770FC4F3E09" box="[882,999,1051,1077]" pageId="23" pageNumber="24">J. Wiesner</determinerName>
1985” [handwritten]
</materialsCitation>
.
<emphasis id="3AAAEAF2B829FFCF4DD6D770FAF03E09" bold="true" box="[1248,1368,1051,1077]" pageId="23" pageNumber="24">
<typeStatus id="D7658842B829FFCF4DD6D770FAF03E09" box="[1248,1368,1051,1077]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
(2):
<specimenCount id="1ED8FD69B829FFCF4CB8D770FF043E65" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
in JWCW,
<materialsCitation id="B8B63CBDB829FFCF4807D754FF5B3EF9" ID-GBIF-Occurrence="3045076585" collectorName="Romeo Lumawig" country="Philippines" determinerName="J. Wiesner" location="Negros oriental" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Negros Oriental" typeStatus="holotype">
<specimenCount id="1ED8FD69B829FFCF4807D754FEF43E65" box="[305,348,1087,1113]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
in SMNS: “
<collectingCountry id="70C97670B829FFCF48D1D754FD3A3E65" box="[487,658,1087,1113]" name="Philippines" pageId="23" pageNumber="24">PHILIPPINES</collectingCountry>
, 8.VI.85, /
<collectingRegion id="CA1AF802B829FFCF4A24D754FC6B3E65" box="[786,963,1087,1113]" country="Philippines" name="Negros Oriental" pageId="23" pageNumber="24">Negros oriental</collectingRegion>
, / leg.
<collectorName id="A52B5336B829FFCF4D26D754FB7F3E65" box="[1040,1239,1087,1113]" pageId="23" pageNumber="24">Romeo Lumawig</collectorName>
” [yellow margin, printed] // “
<taxonomicName id="CFDE4D63B829FFCF4821D708FE353E41" authorityName="Analis. Hope" authorityYear="1838" box="[279,413,1123,1149]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ melanopyga / Schaum / det.
<determinerName id="FDBDDA24B829FFCF4BC4D708FCCE3E41" box="[754,870,1123,1149]" pageId="23" pageNumber="24">J. Wiesner</determinerName>
1985” [handwritten]. All type specimens labelled: “
<typeStatus id="D7658842B829FFCF4996D7ECFE9F3E9D" box="[160,311,1159,1185]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
(or
<typeStatus id="D7658842B829FFCF485AD7ECFE5F3E9D" box="[364,503,1159,1185]" pageId="23" pageNumber="24" type="paratype">PARATYPE</typeStatus>
respectively) /
<taxonomicName id="CFDE4D63B829FFCF4B9BD7ECFC983E9D" authorityName="Analis. Hope" authorityYear="1838" box="[685,816,1159,1185]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ wiesneri
<taxonomicNameLabel id="21995789B829FFCF4A82D7ECFBA53E9D" box="[948,1037,1159,1185]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
/ ded. Sebastian Görn 2020” [red, printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB829FFCF49F1D7A5FBEF3F45" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="086136E0B829FFCF49F1D7A5FBEF3F45" blockId="23.[151,1437,978,2013]" pageId="23" pageNumber="24">
<emphasis id="3AAAEAF2B829FFCF49F1D7A5FEEA3ED4" bold="true" box="[199,322,1230,1256]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
Resembling bluish-coloured specimens of
<taxonomicName id="CFDE4D63B829FFCF4A00D7A4FC4C3ED5" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[822,996,1231,1257]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B829FFCF4A00D7A4FC4C3ED5" box="[822,996,1231,1257]" italics="true" pageId="23" pageNumber="24">H. melanopyga</emphasis>
</taxonomicName>
, but aedeagus compact and triangular shaped, with distinctly Y-shaped sclerite. Further, distinguished by the rather smooth, globular pronotum with laterobasal curvature, and the labrum slightly triangular in its median part. In contrast to
<taxonomicName id="CFDE4D63B829FFCF4DB4D67CFA943F0D" authorityName="Cassola" authorityYear="2000" box="[1154,1340,1303,1329]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="mindoroensis">
<emphasis id="3AAAEAF2B829FFCF4DB4D67CFA943F0D" box="[1154,1340,1303,1329]" italics="true" pageId="23" pageNumber="24">H. mindoroensis</emphasis>
</taxonomicName>
, femora without extended dark apical areas. Due to the lack of mediolateral and anteroapical elytral impressions, it is easily distinguishable from
<taxonomicName id="CFDE4D63B829FFCF48B3D634FDA63F45" box="[389,526,1375,1401]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="abasileia" status="sp. nov.">
<emphasis id="3AAAEAF2B829FFCF48B3D634FDA63F45" box="[389,526,1375,1401]" italics="true" pageId="23" pageNumber="24">H. abasileia</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B829FFCF4B23D634FDC73F45" bold="true" box="[533,623,1375,1401]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="21995789B829FFCF4B23D634FDC73F45" box="[533,623,1375,1401]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="CFDE4D63B829FFCF4B4BD634FC833F44" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[637,811,1375,1400]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B829FFCF4B4BD634FC833F44" box="[637,811,1375,1400]" italics="true" pageId="23" pageNumber="24">H. nigrosericea</emphasis>
</taxonomicName>
, and
<taxonomicName id="CFDE4D63B829FFCF4A5FD634FC4F3F45" box="[873,999,1375,1401]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="20" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
H.
<emphasis id="3AAAEAF2B829FFCF4ABDD634FC4F3F45" box="[907,999,1375,1401]" italics="true" pageId="23" pageNumber="24">halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B829FFCF4AD8D634FBEF3F45" bold="true" box="[1006,1095,1375,1401]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="21995789B829FFCF4AD8D634FBEF3F45" box="[1006,1095,1375,1401]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB829FFCF49F1D6E8FEB23FFD" pageId="23" pageNumber="24" type="etymology">
<paragraph id="086136E0B829FFCF49F1D6E8FEB23FFD" blockId="23.[151,1437,978,2013]" pageId="23" pageNumber="24">
<emphasis id="3AAAEAF2B829FFCF49F1D6E8FEE63FA1" bold="true" box="[199,334,1411,1437]" pageId="23" pageNumber="24">Etymology.</emphasis>
This new species is dedicated to Jürgen Wiesner for his comprehensive support with material and knowledge.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB829FFC249F1D6A1FF6F3E2D" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24" type="description">
<paragraph id="086136E0B829FFCF49F1D6A1FEAF3C35" blockId="23.[151,1437,978,2013]" pageId="23" pageNumber="24">
<emphasis id="3AAAEAF2B829FFCF49F1D6A1FEF13FD8" bold="true" box="[199,345,1482,1508]" pageId="23" pageNumber="24">Description.</emphasis>
Body size: Length 11.6–12.0 mm (without labrum), width
<quantity id="CF269B05B829FFCF4D3DD6A0FB3D3FD9" box="[1035,1173,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="23" pageNumber="24" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.7–3.9 mm</quantity>
(
<figureCitation id="90E52A65B829FFCF4D91D6A0FA9C3FDA" box="[1191,1332,1483,1510]" captionStart="FIGURES 62–67" captionStartId="25.[151,264,1931,1955]" captionTargetBox="[152,1435,184,1872]" captionTargetId="figure-15@25.[151,1435,181,1908]" captionTargetPageId="25" captionText="FIGURES 62–67. Heptodonta wiesneri sp. nov., Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="23" pageNumber="24">Figs. 62–63</figureCitation>
). Female unknown.
</paragraph>
<paragraph id="086136E0B829FFCF49F1D578FEC93D85" blockId="23.[151,1437,978,2013]" pageId="23" pageNumber="24">
Dorsal surface of head iridescent bluish-green with yellow-orange lustre on frons and occiput, temples blue. Frons irregularly rugose. Vertex median with parallel longitudinal striations converging in transition to the orbital plates. Orbital plates with shallow longitudinal striae, two setae on each side. Occiput with transverse grooves, anteromedian irregularly rugose. Genae glabrous and shallowly grooved, violet-blue posterior and yellowish-green anterior. Clypeus glabrous, iridescent bluish-green, with two dark-blue frontal impressions. Labrum pale-testaceous, with four setae, wider (
<quantity id="CF269B05B829FFCF4897D5ADFDE03CDD" box="[417,584,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="1.4649999999999999" metricValueMax="1.5" metricValueMin="1.43" pageId="23" pageNumber="24" unit="mm" value="1.4649999999999999" valueMax="1.5" valueMin="1.43">1.43–1.50 mm</quantity>
) than long (
<quantity id="CF269B05B829FFCF4BE5D5ADFCD23CDD" box="[723,890,1734,1761]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="6.5" metricValueMin="6.1" pageId="23" pageNumber="24" unit="mm" value="0.63" valueMax="0.65" valueMin="0.61">0.61–0.65 mm</quantity>
) (
<figureCitation id="90E52A65B829FFCF4AA5D5ACFC403CDE" box="[915,1000,1735,1762]" captionStart="FIGURES 62–67" captionStartId="25.[151,264,1931,1955]" captionTargetBox="[152,1435,184,1872]" captionTargetId="figure-15@25.[151,1435,181,1908]" captionTargetPageId="25" captionText="FIGURES 62–67. Heptodonta wiesneri sp. nov., Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="23" pageNumber="24">Fig. 64</figureCitation>
), slightly antero-triangular, five apical teeth, and one lateral tooth on each side, acute median tooth with a dark tip, third apical teeth reduced. Mandibles pale-testaceous on the lateral and rufous-testaceous on the inner side, teeth dark reddish-brown to black. Labial and maxillary palpi pale-testaceous with dark-brown apices of terminal palpomeres. Antennae slender, extending back over the middle of the elytra, dark brown, scape and antennomere 3 dark-testaceous on underside, pedicel testaceous at base, scape with a single apical seta, antennomeres 3–4 with few scattered setae, antennomeres 5–11 finely and evenly pubescent.
</paragraph>
<paragraph id="086136E0B829FFC249F1D4A8FEE03B59" blockId="23.[151,1437,978,2013]" lastBlockId="26.[151,1437,150,1149]" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
Thorax entirely glabrous. Pronotum with irregularly rugose orange spot on disc, bordered yellow, followed by iridescent bluish-green, and finally a blue lateral margin, slightly longer (
<quantity id="CF269B05B824FFC24A8BD3FDFBCA3A8D" box="[957,1122,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.55" metricValueMin="2.45" pageId="26" pageNumber="27" unit="mm" value="2.5" valueMax="2.55" valueMin="2.45">2.45–2.55 mm</quantity>
) than wide (
<quantity id="CF269B05B824FFC24DDFD3FCFA383A8D" box="[1257,1424,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.36" metricValueMax="2.41" metricValueMin="2.31" pageId="26" pageNumber="27" unit="mm" value="2.36" valueMax="2.41" valueMin="2.31">2.31–2.41 mm</quantity>
), rather shallow transverse grooves, running concentrically on disc, strong median line, pronounced anterior and posterior sulci, anterior lobe wider than posterior, lateral margins of the median lobe slightly converging to the base, but sloping stronger posteriorly, giving the median lobe a specific globular shape. Episterna violet-blue, metepisternum with blue-green lustre. Prosternum, posterior mesosternum, metasternum, and epimera iridescent yellowish-green to bluish-green.
</paragraph>
<caption id="5CA16668B826FFC049A1D4EAFAF33DDD" ID-DOI="http://doi.org/10.5281/zenodo.4578806" ID-Zenodo-Dep="4578806" httpUri="https://zenodo.org/record/4578806/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,264,1921,1945]" targetBox="[152,1435,184,1870]" targetPageId="24">
<paragraph id="086136E0B826FFC049A1D4EAFAF33DDD" blockId="24.[151,1437,1921,2018]" pageId="24" pageNumber="25">
<emphasis id="3AAAEAF2B826FFC049A1D4EAFEFE3DA5" bold="true" box="[151,342,1921,1945]" pageId="24" pageNumber="25">FIGURES 55–61.</emphasis>
<taxonomicName id="CFDE4D63B826FFC04868D4EAFC8F3DA5" authority="Cassola, 2000: 55" authorityName="Cassola" authorityPageNumber="55" authorityYear="2000" box="[350,807,1921,1946]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mindoroensis">
<emphasis id="3AAAEAF2B826FFC04868D4EAFDCD3DA5" box="[350,613,1921,1945]" italics="true" pageId="24" pageNumber="25">Heptodonta mindoroensis</emphasis>
<bibRefCitation id="6C4F4B11B826FFC04B5BD4EAFC8F3DA5" author="Cassola, F." box="[621,807,1921,1946]" pageId="24" pageNumber="25" pagination="491 - 509" refId="ref37970" refString="Cassola, F. (2000) Studies on tiger beetles 102. The Cicindelidae collected by Roland A. Muller in the Philippine Islands, with description of three new species (Coleoptera: Cicindelidae). Zoologische Mededelingen, 73, 491 - 509." type="journal article" year="2000">Cassola, 2000: 55</bibRefCitation>
</taxonomicName>
, 57, 59–61—♂, Philippines, Mindoro, Mt. Halcon (JWCW): 55—habitus, 57—head, 59—lateral body, 60—aedeagus in left lateral aspect, 61—aedeagus in right lateral aspect; 56, 58—♀, holotype, Philippines, Mindoro, Mt. Halcon (RMNH): 56—habitus, 58—head. Scales: 55–59 = 1 mm, 60–61 = 0.5 mm.
</paragraph>
</caption>
<caption id="5CA16668B827FFC149A1D4E0FD9E3DD7" ID-DOI="http://doi.org/10.5281/zenodo.4578808" ID-Zenodo-Dep="4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,264,1931,1955]" targetBox="[152,1435,184,1872]" targetPageId="25">
<paragraph id="086136E0B827FFC149A1D4E0FD9E3DD7" blockId="25.[151,1437,1931,2028]" pageId="25" pageNumber="26">
<emphasis id="3AAAEAF2B827FFC149A1D4E0FEFC3D98" bold="true" box="[151,340,1931,1956]" pageId="25" pageNumber="26">FIGURES 62–67.</emphasis>
<taxonomicName id="CFDE4D63B827FFC1486FD4E7FD843D9F" box="[345,556,1932,1956]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="3AAAEAF2B827FFC1486FD4E7FD843D9F" box="[345,556,1932,1956]" italics="true" pageId="25" pageNumber="26">Heptodonta wiesneri</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B827FFC14B04D4E7FD2A3D98" bold="true" box="[562,642,1932,1956]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="21995789B827FFC14B04D4E7FD2A3D98" box="[562,642,1932,1956]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="086136E0B824FFC249F1D204FDE43801" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">
Elytra elongate, length
<quantity id="CF269B05B824FFC248E7D205FDF33BB5" box="[465,603,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.5" metricValueMin="7.1" pageId="26" pageNumber="27" unit="mm" value="7.3" valueMax="7.5" valueMin="7.1">7.1–7.5 mm</quantity>
, slightly dilated laterally, maximal width posterior, iridescent colouration bronze along the sutures from base to posterior declivity, yellow transition to bluish-green sublateral area, passing to blue margin and violet juxtaepipleural area, discal and juxtahumeral impression moderate, as well as impression on posterior declivity, apical impression deep, basodiscal convexity and posterior gibbosity moderate; deep blue punctures irregularly distributed, large in front, decreasing in size to apex, particularly anteriorly and along the sutures, slightly fused to transverse lines.
</paragraph>
<paragraph id="086136E0B824FFC249F1D12CFB9438CD" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">Coxae dark-testaceous with iridescent green lustre, pro- and mesocoxae anteriorly, metacoxae laterally densely covered with long white setae. Trochanters and femora testaceous. Femora distally sparsely black-coloured dorsoapically and on apical margin, black colouration slightly extended at metafemoral apex. Tibiae for the most part metallic-black, but testaceous in proximal half of protibiae, proximal third of mesotibiae and proximal apex of metatibiae. Tarsi metallic-black with a slight violet lustre. Claws rufous-testaceous.</paragraph>
<paragraph id="086136E0B824FFC249F1D190FA343905" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">
Abdominal sternites glabrous (apart from few long setae at posterior margins, and some minute setae on posterior margin of sternite 5), dark-brown with violet-blue reflections, anal sternite almost completely brown (
<figureCitation id="90E52A65B824FFC24C0DD074FA383905" box="[1339,1424,798,825]" captionStart="FIGURES 62–67" captionStartId="25.[151,264,1931,1955]" captionTargetBox="[152,1435,184,1872]" captionTargetId="figure-15@25.[151,1435,181,1908]" captionTargetPageId="25" captionText="FIGURES 62–67. Heptodonta wiesneri sp. nov., Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="26" pageNumber="27">Fig. 65</figureCitation>
).
</paragraph>
<paragraph id="086136E0B824FFC249F1D028FF4639F6" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">
Aedeagus robust, length
<quantity id="CF269B05B824FFC248DED028FD383961" box="[488,656,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="2.29" metricValueMax="2.32" metricValueMin="2.26" pageId="26" pageNumber="27" unit="mm" value="2.29" valueMax="2.32" valueMin="2.26">2.26–2.32 mm</quantity>
, ventral side straight, also anterodorsally and medioposterodorsally straight and ascending, forming a generally triangular shape; straight, converging apex with short round tip, short and stout base steeply sloping. Inner sac in left lateral aspect with specifically y-shaped median sclerite (
<figureCitation id="90E52A65B824FFC24C52D0E0FF4A39F6" captionStart="FIGURES 62–67" captionStartId="25.[151,264,1931,1955]" captionTargetBox="[152,1435,184,1872]" captionTargetId="figure-15@25.[151,1435,181,1908]" captionTargetPageId="25" captionText="FIGURES 62–67. Heptodonta wiesneri sp. nov., Philippines, Negros oriental (JWCW): 62, 64–67—♂, holotype: 62—habitus, 64—head, 65—lateral body, 66—aedeagus in left lateral aspect, 67—aedeagus in right lateral aspect; 63—habitus, ♂, paratype. Scales: 62–65 = 1 mm, 66–67 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578808" httpUri="https://zenodo.org/record/4578808/files/figure.png" pageId="26" pageNumber="27">Figs. 66–67</figureCitation>
).
</paragraph>
<paragraph id="086136E0B824FFC249F1D0B9FF6F3E2D" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">
<emphasis id="3AAAEAF2B824FFC249F1D0B9FEE539D0" bold="true" box="[199,333,978,1004]" pageId="26" pageNumber="27">Variability.</emphasis>
Pronotal and aedeagal shape showing some variation without losing the species-specific characters.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB824FFC249F1D771FD4E3E09" box="[199,742,1050,1077]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="086136E0B824FFC249F1D771FD4E3E09" blockId="26.[151,1437,150,1149]" box="[199,742,1050,1077]" pageId="26" pageNumber="27">
<emphasis id="3AAAEAF2B824FFC249F1D771FEC93E08" bold="true" box="[199,353,1050,1076]" pageId="26" pageNumber="27">Distribution.</emphasis>
Central
<emphasis id="3AAAEAF2B824FFC248F7D771FDDC3E08" bold="true" box="[449,628,1050,1076]" pageId="26" pageNumber="27">
<collectingCountry id="70C97670B824FFC248F7D771FDDC3E08" box="[449,628,1050,1076]" name="Philippines" pageId="26" pageNumber="27">PHILIPPINES</collectingCountry>
</emphasis>
(Negros).
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB824FFC249F1D754FA873E41" pageId="26" pageNumber="27" type="discussion">
<paragraph id="086136E0B824FFC249F1D754FA873E41" blockId="26.[151,1437,150,1149]" pageId="26" pageNumber="27">
<emphasis id="3AAAEAF2B824FFC249F1D754FE933E65" bold="true" box="[199,315,1087,1113]" pageId="26" pageNumber="27">Remarks.</emphasis>
Together with the three
<typeStatus id="D7658842B824FFC24B67D754FD293E65" box="[593,641,1087,1113]" pageId="26" pageNumber="27">type</typeStatus>
specimens, Romeo Lumawig collected a specimen of
<taxonomicName id="CFDE4D63B824FFC24DDFD754FA3D3E65" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[1257,1429,1087,1113]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B824FFC24DDFD754FA3D3E65" box="[1257,1429,1087,1113]" italics="true" pageId="26" pageNumber="27">H. melanopyga</emphasis>
</taxonomicName>
, showing the same specific colour pattern of
<taxonomicName id="CFDE4D63B824FFC24BB0D708FCA03E40" box="[646,776,1123,1148]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="3AAAEAF2B824FFC24BB0D708FCA03E40" box="[646,776,1123,1148]" italics="true" pageId="26" pageNumber="27">H. wiesneri</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B824FFC24A39D708FCC03E41" bold="true" box="[783,872,1123,1149]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="21995789B824FFC24A39D708FCC03E41" box="[783,872,1123,1149]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, which is unique within
<taxonomicName id="CFDE4D63B824FFC24D4BD708FA803E41" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[1149,1320,1123,1149]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B824FFC24D4BD708FA803E41" box="[1149,1320,1123,1149]" italics="true" pageId="26" pageNumber="27">H. melanopyga</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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