207 lines
25 KiB
XML
207 lines
25 KiB
XML
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<document id="D169FA044DA32F788C1F842101DFD9B4" ID="10.11646/zootaxa.4126.1.7" ID-CLB-Dataset="38800" ID-DOI="10.11646/zootaxa.4126.1.7" ID-GBIF-Dataset="78917532-4d8e-4d76-a25f-04b29f47e784" ID-ISSN="1175-5326" ID-Zenodo-Dep="260670" ID-ZooBank="8B7F8643-F1BC-467A-B8B3-B7A0DA91C916" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466063277507" checkinUser="plazi" docAuthor="Tang, Yong-Tao, Feng, Chen-Guang, Wanghe, Kun-Yuan, Li, Guo-Gang & Zhao, Kai" docDate="2016" docId="164D8783FF879D0DA9E9068A8E70FEDD" docLanguage="en" docName="zootaxa.4126.1.7.pdf" docOrigin="Zootaxa 4126 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnocypris pengquensis Tang, Feng, Wanghe, Li & Zhao, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="134" masterDocId="EA74FFFBFF8E9D06A97E02098C28FFC1" masterDocTitle="Taxonomic status of a population of Gymoncypris waddelli Regan, 1905 (Cypriniformes: Schizothoracinae) distributed in Pengqu River, Tibet, China" masterLastPageNumber="137" masterPageNumber="123" pageNumber="132" updateTime="1698663010782" updateUser="ExternalLinkService">
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<mods:mods id="00F24A6C738E721692CD1C45510B013D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F65D0030ED99A9BF403479688E708DCE">
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<mods:title id="7FF4754AFA4CF8F96FE3F6AA5A3CE4A4">Taxonomic status of a population of Gymoncypris waddelli Regan, 1905 (Cypriniformes: Schizothoracinae) distributed in Pengqu River, Tibet, China</mods:title>
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</mods:titleInfo>
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<mods:name id="5B4C4D9CDBF85E77E58548126B3CC6B7" type="personal">
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<mods:roleTerm id="4F2E5823D398BBF9FDCC38B06EEC688A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6A35BAABCB36D67BB81E19297D39ADA3">Tang, Yong-Tao</mods:namePart>
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<mods:name id="0C272478F3323121C0FF90F75C5AF71F" type="personal">
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<mods:roleTerm id="D346B88297F864D2B3BA7C597CD0D834">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BCABB3DB5413871D9C7AF58B27353871">Feng, Chen-Guang</mods:namePart>
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</mods:name>
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<mods:name id="B6B80D104DBF9581DF54863B9526B9E1" type="personal">
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<mods:role id="84228A3E6461B66B02E4A7FA95FA72DA">
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<mods:roleTerm id="97711FF761C229BF80AA56D792ADB649">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16A5E7D9A2403DD4E7FE1ACB73271994">Wanghe, Kun-Yuan</mods:namePart>
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</mods:name>
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<mods:name id="1B440794E4F2BAC442A3A3BD8CF10D97" type="personal">
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<mods:roleTerm id="A07628B02AE0220B596284F6F94B8F36">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CE640D6BA87212DF6F698A051D955EBF">Li, Guo-Gang</mods:namePart>
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</mods:name>
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<mods:name id="73C1EFCB123B52E2132E9FD42D4E0B53" type="personal">
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<mods:role id="315C6646F2B13DA36CDE89A4FF16BED7">
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<mods:roleTerm id="CDC6AEC5A3DA090AB410D8588CFF65A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C7035BC01FB56A391BBD2F323B36AA5E">Zhao, Kai</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5D017FF6D5669555B1207A4D01EA226F">text</mods:typeOfResource>
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<mods:titleInfo id="61214F9A7C8E5149A0900AF8980586D4">
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<mods:title id="835FD4156D5D7B0C01C4E2EE44818030">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9CDF6CB3BFD282C9735C34725D67A561">
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<mods:date id="73C3FFBBC2A8C91504AE1B0E574D0CA9">2016</mods:date>
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<mods:detail id="A344423F91E8B8C0CDBC7BACE1DAEFE5" type="volume">
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<mods:number id="F60304520E08782106A0753AF65242D2">4126</mods:number>
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</mods:detail>
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<mods:detail id="456419DE04AAE9D19342B209F335278A" type="issue">
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<mods:number id="6223E1F964319687600B97F8B7C58419">1</mods:number>
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<mods:start id="8E13C2B00593D954914E413A877C6254">123</mods:start>
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<mods:end id="B4F368732E21AC94C57D4353666C2C0D">137</mods:end>
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<mods:classification id="7E7010CB411DA365FFF72F7BDE3D54EF">journal article</mods:classification>
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<mods:identifier id="20B37AEA1CF656855B169503914C5494" type="CLB-Dataset">38800</mods:identifier>
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<mods:identifier id="01336259D4174605C939C9FEB3693BCA" type="DOI">10.11646/zootaxa.4126.1.7</mods:identifier>
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<mods:identifier id="F3741D71F7ACC784FFCA58174DA9A59B" type="GBIF-Dataset">78917532-4d8e-4d76-a25f-04b29f47e784</mods:identifier>
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<mods:identifier id="97573A613411E40D5E94993265F1A8E9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="705AA8E44F62C5A05973922E685737CA" type="Zenodo-Dep">260670</mods:identifier>
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<mods:identifier id="5274E10C6FE2A08BE1BAE03C19AE29E0" type="ZooBank">8B7F8643-F1BC-467A-B8B3-B7A0DA91C916</mods:identifier>
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</mods:mods>
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<treatment id="164D8783FF879D0DA9E9068A8E70FEDD" ID-DOI="http://doi.org/10.5281/zenodo.5612474" ID-GBIF-Taxon="120828805" ID-Zenodo-Dep="5612474" LSID="urn:lsid:plazi:treatment:164D8783FF879D0DA9E9068A8E70FEDD" httpUri="http://treatment.plazi.org/id/164D8783FF879D0DA9E9068A8E70FEDD" lastPageId="11" lastPageNumber="134" pageId="9" pageNumber="132">
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<subSubSection id="D6FE651EFF879D0FA9E9068A8D22FB7E" pageId="9" pageNumber="132" type="nomenclature">
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<paragraph id="9E5B3695FF879D0FA9E9068A8E17FB5F" blockId="9.[151,575,1155,1216]" box="[151,575,1155,1182]" pageId="9" pageNumber="132">
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<heading id="C51381F9FF879D0FA9E9068A8E17FB5F" bold="true" box="[151,575,1155,1182]" fontSize="11" level="1" pageId="9" pageNumber="132" reason="1">
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<emphasis id="AC90EA87FF879D0FA9E9068A8E17FB5F" bold="true" box="[151,575,1155,1182]" pageId="9" pageNumber="132">
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<taxonomicName id="59E44D16FF879D0FA9E9068A8DFBFB5C" box="[151,467,1155,1181]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="pengquensis" status="sp. nov.">
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<emphasis id="AC90EA87FF879D0FA9E9068A8DFBFB5C" bold="true" box="[151,467,1155,1181]" italics="true" pageId="9" pageNumber="132">Gymnocypris pengquensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="B7A357FCFF879D0FA89C068D8E17FB5F" box="[482,575,1156,1182]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9E5B3695FF879D0FA9E906AE8D22FB7E" blockId="9.[151,575,1155,1216]" box="[151,266,1191,1216]" pageId="9" pageNumber="132">
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(
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<figureCitation id="06DF2A10FF879D0FA9E106AE8CCFFB01" box="[159,231,1191,1216]" captionStart="FIGURE 7" captionStartId="9.[151,250,1914,1936]" captionTargetBox="[160,1436,1486,1880]" captionTargetId="figure@9.[160,1445,1486,1880]" captionTargetPageId="9" captionText="FIGURE 7. Gymnocypris pengquensis sp. nov., holotype, NWIPB 1161025, 138 mm SL. Color pattern of specimen preserved in 95 % alcohol (right side; image laterally inverted)." httpUri="https://zenodo.org/record/260677/files/figure.png" pageId="9" pageNumber="132">Figs.7</figureCitation>
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,
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<figureCitation id="06DF2A10FF879D0FA98D06AE8D29FB7E" box="[243,257,1191,1215]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="9" pageNumber="132">8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D6FE651EFF879D0FA9E906E68D3BFA59" pageId="9" pageNumber="132" type="materials_examined">
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<paragraph id="9E5B3695FF879D0FA9E906E68CFDFA91" blockId="9.[151,1436,1263,1432]" pageId="9" pageNumber="132">
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<emphasis id="AC90EA87FF879D0FA9E906E68D23FAC9" bold="true" box="[151,267,1263,1288]" pageId="9" pageNumber="132">
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<typeStatus id="415F8837FF879D0FA9E906E68D2EFAC9" box="[151,262,1263,1288]" pageId="9" pageNumber="132" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="F8F5AE50FF879D0FA86906F98D59FAC6" box="[279,369,1264,1287]" pageId="9" pageNumber="132">NWIPB</collectionCode>
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<quantity id="591C9B70FF879D0FA80306E68E7AFAC6" box="[381,594,1263,1288]" metricMagnitude="6" metricUnit="m" metricValue="1.161025167" pageId="9" pageNumber="132" unit="mm" value="1.161025167E9">1161025, 167 mm</quantity>
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SL, Pengqu River, Zhaguo Town, Nielamu County, Tibet Autonomous Region,
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<collectingCountry id="E6F37605FF879D0FA987071A8D14FAED" box="[249,316,1299,1324]" name="China" pageId="9" pageNumber="132">China</collectingCountry>
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(
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<geoCoordinate id="FBD05052FF879D0FA831071A8DC3FAED" box="[335,491,1299,1324]" direction="north" orientation="latitude" pageId="9" pageNumber="132" precision="1" value="28.600084">28°36'0.30"N</geoCoordinate>
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,
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<geoCoordinate id="FBD05052FF879D0FA884071A8E88FAED" box="[506,672,1299,1324]" direction="east" orientation="longitude" pageId="9" pageNumber="132" precision="1" value="86.91984">86°55'11.43"E</geoCoordinate>
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),
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<date id="EA5A1055FF879D0FABC9071A8F7DFAEA" box="[695,853,1299,1323]" pageId="9" pageNumber="132" value="2011-06-30">30 June 2011</date>
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, collected by Kai Zhao, Guogang Li and Yongtao Tang.
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</paragraph>
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<paragraph id="9E5B3695FF879D0FA9B907528D3BFA59" blockId="9.[151,1436,1263,1432]" pageId="9" pageNumber="132">
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<emphasis id="AC90EA87FF879D0FA9B907528D6EFAB5" bold="true" box="[199,326,1371,1396]" pageId="9" pageNumber="132">
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<typeStatus id="415F8837FF879D0FA9B907528D69FAB5" box="[199,321,1371,1396]" pageId="9" pageNumber="132" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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<collectionCode id="F8F5AE50FF879D0FA82E07558D82FAB2" box="[336,426,1372,1395]" pageId="9" pageNumber="132">NWIPB</collectionCode>
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1161012–24, 1161026–34, collected with the
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<typeStatus id="415F8837FF879D0FAABD07528801FAB5" box="[963,1065,1371,1396]" pageId="9" pageNumber="132" type="holotype">holotype</typeStatus>
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.
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<quantity id="591C9B70FF879D0FAD48075288E6FAB2" box="[1078,1230,1371,1396]" metricMagnitude="-1" metricUnit="m" metricValue="1.435" metricValueMax="1.86" metricValueMin="1.01" pageId="9" pageNumber="132" unit="mm" value="143.5" valueMax="186.0" valueMin="101.0">101–186 mm</quantity>
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SL. 22 examined specimens.
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</paragraph>
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</subSubSection>
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<caption id="CA9B661DFF879D0FA9E905738E83F86E" httpUri="https://zenodo.org/record/260677/files/figure.png" pageId="9" pageNumber="132" targetBox="[160,1436,1486,1880]" targetPageId="9">
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<paragraph id="9E5B3695FF879D0FA9E905738E83F86E" blockId="9.[151,1435,1914,1967]" pageId="9" pageNumber="132">
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<emphasis id="AC90EA87FF879D0FA9E905738D25F851" bold="true" box="[151,269,1914,1936]" pageId="9" pageNumber="132">FIGURE 7</emphasis>
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.
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<taxonomicName id="59E44D16FF879D0FA86705728E0BF851" box="[281,547,1915,1936]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="132" phylum="Chordata" rank="species" species="pengquensis" status="sp. nov.">
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<emphasis id="AC90EA87FF879D0FA86705728E0BF851" box="[281,547,1915,1936]" italics="true" pageId="9" pageNumber="132">Gymnocypris pengquensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B7A357FCFF879D0FAB5405728E53F851" box="[554,635,1915,1936]" pageId="9" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, NWIPB 1161025, 138 mm SL. Color pattern of specimen preserved in 95% alcohol (right side; image laterally inverted).
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</paragraph>
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</caption>
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<caption id="CA9B661DFF849D0CA9E901BE88FBFBCA" httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133" targetBox="[340,1247,155,930]" targetPageId="10">
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<paragraph id="9E5B3695FF849D0CA9E901BE88FBFBCA" blockId="10.[151,1436,951,1035]" pageId="10" pageNumber="133">
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<emphasis id="AC90EA87FF849D0CA9E901BE8D25FC0D" bold="true" box="[151,269,951,973]" pageId="10" pageNumber="133">FIGURE 8</emphasis>
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.
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<taxonomicName id="59E44D16FF849D0CA86401B18E0CFC0F" box="[282,548,952,974]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="pengquensis">
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<emphasis id="AC90EA87FF849D0CA86401B18E0CFC0F" box="[282,548,952,974]" italics="true" pageId="10" pageNumber="133">Gymnocypris pengquensis</emphasis>
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</taxonomicName>
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, paratype, NWIPB 1161026, 172mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine.
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</paragraph>
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</caption>
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<subSubSection id="D6FE651EFF849D0CA9B9063B8E3BFB1A" pageId="10" pageNumber="133" type="diagnosis">
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<paragraph id="9E5B3695FF849D0CA9B9063B8E3BFB1A" blockId="10.[151,1437,1074,2035]" pageId="10" pageNumber="133">
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<emphasis id="AC90EA87FF849D0CA9B9063B8D69FB8A" bold="true" box="[199,321,1074,1099]" pageId="10" pageNumber="133">Diagnosis.</emphasis>
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<taxonomicName id="59E44D16FF849D0CA831063A8DD7FB8B" box="[335,511,1075,1098]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="pengquensis">
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<emphasis id="AC90EA87FF849D0CA831063A8DD7FB8B" box="[335,511,1075,1098]" italics="true" pageId="10" pageNumber="133">G. pengquensis</emphasis>
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</taxonomicName>
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is cleary distinguished from congeners by the combination of the following characters: mouth subinferior, nearly straight oral fissure, arched; inner side of lower jaw without a horny layer; dorsal-fin origin nearly at midway of the body length. Commencement of ventral fin under the 4–5th branched ray of dorsal fin; last unbranched dorsal-fin ray weak; dorsal fin spines 8–15, minutely serrated; gill rakers on first arch: ONG = 9–14, ING = 13–21.
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</paragraph>
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</subSubSection>
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<subSubSection id="D6FE651EFF849D0CA9B906EF8FE0F832" pageId="10" pageNumber="133" type="description">
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<paragraph id="9E5B3695FF849D0CA9B906EF8F92FB3E" blockId="10.[151,1437,1074,2035]" box="[199,954,1254,1279]" pageId="10" pageNumber="133">
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<emphasis id="AC90EA87FF849D0CA9B906EF8D71FB3E" bold="true" box="[199,345,1254,1279]" pageId="10" pageNumber="133">Description.</emphasis>
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D iv, 8; P i, 15–19; V i, 8–9; A iii, 5; vertebrae: 4+47.
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</paragraph>
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<paragraph id="9E5B3695FF849D0CA9B907028872F8DA" blockId="10.[151,1437,1074,2035]" pageId="10" pageNumber="133">
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Counts and proportional measurements are given in
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<tableCitation id="D366032EFF849D0CAA6407038F59FAE2" box="[794,881,1290,1315]" captionStart="TABLE 2" captionStartId="3.[151,239,620,643]" captionTargetBox="[159,1366,670,1932]" captionTargetPageId="3" captionText="TABLE 2. Traditional measurement data for Gymnocypris waddelli." httpUri="http://table.plazi.org/id/CA9B661DFF8D9D05A9E900658F5AFD42" pageId="10" pageNumber="133" tableUuid="CA9B661DFF8D9D05A9E900658F5AFD42">Table 2</tableCitation>
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. Body elongate, somewhat compressed. Greatest depth at origin of dorsal fin, dorsal profile arched, ventral profile less curved. Anal opening near origin of anal fin. Caudal forked. Head somewhat large, conical. Snout obtuse; No barbels; anterior border of the mouth below level of lower margin of eye. Eyes moderate, round. Nostrils nearer to eye than to tip of snout. Mouth subinferior, nearly straight oral fissure, inner side of lower jaw without horny layer. Ventral view of margin of lower jaw horseshoeshaped; lip of lower jaw underdeveloped, thin; lower lip divided into two lateral lobes, postlabial groove discontinued (
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<figureCitation id="06DF2A10FF849D0CA84507EB8DAAFA3A" box="[315,386,1506,1531]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133">Fig. 8</figureCitation>
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a). Body entirely naked, except 2–3 rows of scales above the pectoral axil and a sheath consisting of 20–24 enlarged scales at the base of anal fin behind ventral fin. Dorsal origin nearly at midpoint of body length. Commencement of ventral fin under 4 or 5th branched ray of dorsal; last unbranched dorsal-fin ray weak; dorsal-fin spines with 8–15 small, almost indiscernible serrations. Lateral line complete, running along middle of flank and caudal peduncle. Gill rakers small, short, thick; those on outside somewhat sparse set; those on inside slightly close set (
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<figureCitation id="06DF2A10FF849D0CA8CC049F8DDEF96E" box="[434,502,1686,1711]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133">Fig. 8</figureCitation>
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b,c). Pharyngeal teeth in 2 row 3,4/4,3, rod-like, hooked at tip; first tooth in main row small, second tooth in main row well developed (
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<figureCitation id="06DF2A10FF849D0CAB8D04B38F1EF912" box="[755,822,1722,1747]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133">Fig. 8</figureCitation>
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d). Air bladder composed of 2 chambers, posterior one is 2.2 times longer than anterior one (
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<figureCitation id="06DF2A10FF849D0CAB3904D78EA4F936" box="[583,652,1758,1783]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133">Fig. 8</figureCitation>
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e). Digestive canal short: intestine exhibits three rostro-caudad loops before descending to anus (
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<figureCitation id="06DF2A10FF849D0CA8B4050B8E24F8DA" box="[458,524,1794,1819]" captionStart="FIGURE 8" captionStartId="10.[151,250,951,973]" captionTargetBox="[340,1247,155,930]" captionTargetId="figure@10.[340,1248,155,930]" captionTargetPageId="10" captionText="FIGURE 8. Gymnocypris pengquensis, paratype, NWIPB 1161026, 172 mm SL. (a) Ventral view of head; (b) Lateral view of outside of first gill arch from right side; (c) Lateral view of inside of first gill arch from right side; (d) Dorsomedial view of left side pharyngeal tooth plate; (e) Ventral view of air bladder; (f) Ventral view of the coiling type of intestine." httpUri="https://zenodo.org/record/260678/files/figure.png" pageId="10" pageNumber="133">Fig. 8</figureCitation>
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f). Total length of intestinal tract about 1.3 times SL.
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</paragraph>
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<paragraph id="9E5B3695FF849D0CA9B9052F8F2DF8A3" blockId="10.[151,1437,1074,2035]" pageId="10" pageNumber="133">
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<emphasis id="AC90EA87FF849D0CA9B9052F8D5AF8FE" bold="true" box="[199,370,1830,1855]" pageId="10" pageNumber="133">Color pattern.</emphasis>
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Preserved specimens in alcohol back-gray, abdomen yellowish, whole body with large or small black spots; dorsal and caudal fin also with black spots.
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</paragraph>
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<paragraph id="9E5B3695FF849D0CA9B905678FE0F832" blockId="10.[151,1437,1074,2035]" pageId="10" pageNumber="133">
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<emphasis id="AC90EA87FF849D0CA9B905678D03F846" bold="true" box="[199,299,1902,1927]" pageId="10" pageNumber="133">Ecology.</emphasis>
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<taxonomicName id="59E44D16FF849D0CA84B05668E77F847" box="[309,607,1903,1926]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="133" phylum="Chordata" rank="species" species="pengquensis">
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<emphasis id="AC90EA87FF849D0CA84B05668E77F847" box="[309,607,1903,1926]" italics="true" pageId="10" pageNumber="133">Gymnocypris pengquensis</emphasis>
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</taxonomicName>
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inhabits cold highland streams with a substrate of sand or sand-mud at altitudes
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<quantity id="591C9B70FF849D0CA87E059A8D71F86B" box="[256,345,1939,1963]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="133" unit="m" value="4000.0">4000 m</quantity>
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above sea level (
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<figureCitation id="06DF2A10FF849D0CAB57059B8E59F86A" box="[553,625,1938,1963]" captionStart="FIGURE 9" captionStartId="11.[151,250,1071,1093]" captionTargetBox="[340,1246,351,1030]" captionTargetId="figure@11.[340,1247,351,1031]" captionTargetPageId="11" captionText="FIGURE 9. Type locality of Gymnocypris pengquensis, Pengqu River, Zhaguo Town, Nielamu County, Tibet Autonomous Region, China. Photographed by Yongtao Tang." httpUri="https://zenodo.org/record/260679/files/figure.png" pageId="10" pageNumber="133">Fig. 9</figureCitation>
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). It is an omnivores fish, feeding on juvenile fishes, plant fragments, periphytic algae (e.g. diatoms growing on solid substrate, such as stones), benthic invertebrates and organic debris. Mud and sand were found in the gut, suggesting a benthic foraging style.
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</paragraph>
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</subSubSection>
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<subSubSection id="D6FE651EFF859D0DA9B9029E8F83FF15" pageId="11" pageNumber="134" type="distribution">
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<paragraph id="9E5B3695FF859D0DA9B9029E8F83FF15" blockId="11.[151,1436,151,285]" pageId="11" pageNumber="134">
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<emphasis id="AC90EA87FF859D0DA9B9029E8D48FF71" bold="true" box="[199,352,151,176]" pageId="11" pageNumber="134">Distribution.</emphasis>
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<taxonomicName id="59E44D16FF859D0DA81402918EBDFF6E" box="[362,661,152,175]" class="Actinopterygii" family="Cyprinidae" genus="Gymnocypris" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="134" phylum="Chordata" rank="species" species="pengquensis">
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<emphasis id="AC90EA87FF859D0DA81402918EBDFF6E" box="[362,661,152,175]" italics="true" pageId="11" pageNumber="134">Gymnocypris pengquensis</emphasis>
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</taxonomicName>
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is known only from the Pengqu River and its tributaries, Nielamu County and Dingri County, Tibet Autonomous Region,
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<collectingCountry id="E6F37605FF859D0DAA7802B58F61FF14" box="[774,841,188,213]" name="China" pageId="11" pageNumber="134">China</collectingCountry>
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(
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<figureCitation id="06DF2A10FF859D0DAA2602B58FB2FF15" box="[856,922,188,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1191,1213]" captionTargetBox="[241,1346,519,1169]" captionTargetId="figure@1.[241,1346,519,1169]" captionTargetPageId="1" captionText="FIGURE 1. Sampling locations for Gymnocypris waddelli (red square: Pengqu River population; blue square: Lake Yamzhog Yumco population; gray area indicates approximate inferred distribution of G. waddelli)." httpUri="https://zenodo.org/record/260671/files/figure.png" pageId="11" pageNumber="134">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D6FE651EFF859D0DA9B902D68E70FEDD" pageId="11" pageNumber="134" type="etymology">
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<paragraph id="9E5B3695FF859D0DA9B902D68E70FEDD" blockId="11.[151,1436,151,285]" pageId="11" pageNumber="134">
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<emphasis id="AC90EA87FF859D0DA9B902D68D6EFF39" bold="true" box="[199,326,223,248]" pageId="11" pageNumber="134">Etymology</emphasis>
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. The species name ‘pengquensis’ is a combination of
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<emphasis id="AC90EA87FF859D0DAADD02E88FD1FF39" box="[931,1017,225,248]" italics="true" pageId="11" pageNumber="134">Pengqu</emphasis>
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(where the
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<typeStatus id="415F8837FF859D0DAD0302E98885FF39" box="[1149,1197,224,248]" pageId="11" pageNumber="134">type</typeStatus>
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series was collected) and the geographic Latin suffix (-
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<emphasis id="AC90EA87FF859D0DAB71030C8E6FFEDD" box="[527,583,261,284]" italics="true" pageId="11" pageNumber="134">ensis</emphasis>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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