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<document id="069B7E0A57005A73F641AB29BD8D4F31" ID-CLB-Dataset="107849" ID-DOI="10.5252/geodiversitas2022v44a25" ID-GBIF-Dataset="d0184974-bea3-4f97-a2f8-98887ec4f87e" ID-ISSN="1638-9395" ID-Zenodo-Dep="6902033" ID-ZooBank="urn:lsid:zoobank.org:pub:EA12DCB7-A5BE-4763-B805-25087EBD726D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658759598753" checkinUser="felipe" docAuthor="Allain, Ronan, Vullo, Romain, Rozada, Lee, Anquetin, Jérémy, Bourgeais, Renaud, Goedert, Jean, Lasseron, Maxime, Martin, Jeremy E., Pérez-García, Adán, Fabrègues, Claire Peyre De, Royo-Torres, Rafael, Augier, Dominique &amp; Bailly, Gilles" docDate="2022" docId="03C887B9FF99FFDA76DEA40AFE1CFC01" docLanguage="en" docName="Geodiversitas.44.25.683-752.pdf.imf" docOrigin="Geodiversitas 44 (25)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pinheirodontidae Hahn &amp; Hahn 1999" docType="treatment" docVersion="7" lastPageNumber="730" masterDocId="FFF1FFC1FFB6FFEB753AA238FFEBFFB1" masterDocTitle="Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary" masterLastPageNumber="752" masterPageNumber="683" pageNumber="728" updateTime="1699361923064" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="9CF9E4B7FB428726DA6932F145136CC7">Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J / K boundary</mods:title>
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<treatment id="03C887B9FF99FFDA76DEA40AFE1CFC01" ID-DOI="http://doi.org/10.5281/zenodo.6929064" ID-GBIF-Taxon="197866767" ID-Zenodo-Dep="6929064" LSID="urn:lsid:plazi:treatment:03C887B9FF99FFDA76DEA40AFE1CFC01" httpUri="http://treatment.plazi.org/id/03C887B9FF99FFDA76DEA40AFE1CFC01" lastPageId="49" lastPageNumber="730" pageId="47" pageNumber="728">
<subSubSection id="C37B6524FF99FFC476DEA40AFB13F9FD" box="[996,1272,1586,1612]" pageId="47" pageNumber="728" type="nomenclature">
<paragraph id="8BDE36AFFF99FFC476DEA40AFB13F9FD" blockId="47.[996,1272,1586,1645]" box="[996,1272,1586,1612]" pageId="47" pageNumber="728">
<taxonomicName id="4C614D2CFF99FFC476DEA40AFB47F9FD" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[996,1196,1586,1612]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="47" pageNumber="728" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet.
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<subSubSection id="C37B6524FF99FFDA713AA46AFD72FD61" lastPageId="49" lastPageNumber="730" pageId="47" pageNumber="728" type="description">
<paragraph id="8BDE36AFFF99FFC4713AA46AFB37F9DC" blockId="47.[996,1272,1586,1645]" box="[1024,1244,1618,1645]" pageId="47" pageNumber="728">
(
<figureCitation id="135A2A2AFF99FFC47130A46AFBB1F9DD" box="[1034,1114,1618,1644]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="47" pageNumber="728">Fig. 33</figureCitation>
A-H, M-P)
</paragraph>
<paragraph id="8BDE36AFFF99FFC47617A4ABFC2BF91C" blockId="47.[811,1455,1683,2028]" box="[813,960,1683,1709]" pageId="47" pageNumber="728">DESCRIPTION</paragraph>
<paragraph id="8BDE36AFFF99FFC47617A48BFAF8F83D" blockId="47.[811,1455,1683,2028]" pageId="47" pageNumber="728">
Most of the multituberculate teeth discovered in Angeac-Charente can be referred to the family
<taxonomicName id="4C614D2CFF99FFC471DAA4EAFA41F95D" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[1248,1450,1746,1772]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="47" pageNumber="728" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
, but it has not been possible to refer them to any existing genera. The material figured here includes a left p4 (ANG M-72), a left P1 (ANG M-03), a right P2 (ANG M-06), a left P3 (ANG M-22), a left?m2 (ANG M-105) and a left M2 (ANG M-32) (
<figureCitation id="135A2A2AFF99FFC47103A549FB61F83A" box="[1081,1162,1905,1931]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="47" pageNumber="728">Fig. 33</figureCitation>
A-H, M-P).
</paragraph>
<paragraph id="8BDE36AFFF99FFDA7679A5A9FE26FEE0" blockId="47.[811,1455,1683,2028]" lastBlockId="49.[130,776,215,944]" lastPageId="49" lastPageNumber="730" pageId="47" pageNumber="728">
The most mesial part of the blade-like p4 (ANG M-72) is missing (
<figureCitation id="135A2A2AFF99FFC476B4A589FC04F87A" box="[910,1007,1969,1995]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="47" pageNumber="728">Fig. 33A</figureCitation>
). It possesses at least six serrations, and no basal labial cusp, a diagnostic feature of
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<emphasis id="B915EABDFF99FFC471F2A5E9FAADF85A" box="[1224,1350,2001,2027]" italics="true" pageId="47" pageNumber="728">Pinheirodon</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF04B5EFF99FFDA706FA5E9FEE8FF40" author="Hahn &amp; Hahn" firstAuthor="Hahn" lastPageId="49" lastPageNumber="730" pageId="47" pageNumber="728" pagination="77 - 222" refId="ref40980" refString="HAHN G. &amp; HAHN R. 1999. - Pinheirodontidae n. fam. (Multituberculata) (Mammalia) aus der tiefen Unter-Kreide Portugals. Palaeontographica Abteilung A: 77 - 222." type="book chapter" year="1999">Hahn &amp; Hahn 1999</bibRefCitation>
; Kielan-Jaworowska
<emphasis id="B915EABDFF87FFDA74D1A2E0FDCBFF40" box="[491,544,215,241]" italics="true" pageId="49" pageNumber="730">et al.</emphasis>
2004). Nevertheless, the Angeac-Charente specimen has a straight, vertical, distal margin (
<figureCitation id="135A2A2AFF87FFDA75DFA32FFEA3FE83" box="[229,328,279,306]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33B</figureCitation>
), while the latter is strongly convex just above the root in
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<emphasis id="B915EABDFF87FFDA747FA30FFE2DFEE0" box="[325,454,311,337]" italics="true" pageId="49" pageNumber="730">Pinheirodon</emphasis>
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.
</paragraph>
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FIG. 33. — Multituberculate mammal teeth from Angeac-Charente:
<emphasis id="B915EABDFF86FFDB7797A55FFD51F8C9" bold="true" box="[685,698,1895,1912]" pageId="48" pageNumber="729">A</emphasis>
,
<emphasis id="B915EABDFF86FFDB77F9A55FFD3BF8C9" bold="true" box="[707,720,1895,1912]" pageId="48" pageNumber="729">B</emphasis>
, left p4 of
<taxonomicName id="4C614D2CFF86FFDB7612A55FFC5EF8C9" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[808,949,1895,1912]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M-72) in lingual (
<emphasis id="B915EABDFF86FFDB719DA55FFB5FF8C9" bold="true" box="[1191,1204,1895,1912]" pageId="48" pageNumber="729">A</emphasis>
) and labial (
<emphasis id="B915EABDFF86FFDB702DA55FFACFF8C9" bold="true" box="[1303,1316,1895,1912]" pageId="48" pageNumber="729">B</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB705CA55FFA9FF8C9" bold="true" box="[1382,1396,1895,1912]" pageId="48" pageNumber="729">C</emphasis>
,
<emphasis id="B915EABDFF86FFDB7044A55FFA67F8C9" bold="true" box="[1406,1420,1895,1912]" pageId="48" pageNumber="729">D</emphasis>
, left P1 of
<taxonomicName id="4C614D2CFF86FFDB758CA545FEA8F83F" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[182,323,1917,1934]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M-03) in occlusal (
<emphasis id="B915EABDFF86FFDB7772A545FDBDF83F" bold="true" box="[584,598,1917,1934]" pageId="48" pageNumber="729">C</emphasis>
) and distolingual (
<emphasis id="B915EABDFF86FFDB77D4A545FD17F83F" bold="true" box="[750,764,1917,1934]" pageId="48" pageNumber="729">D</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB767AA545FCA7F83F" bold="true" box="[832,844,1917,1934]" pageId="48" pageNumber="729">E</emphasis>
,
<emphasis id="B915EABDFF86FFDB766CA545FC8AF83F" bold="true" box="[854,865,1917,1934]" pageId="48" pageNumber="729">F</emphasis>
, right P2 of
<taxonomicName id="4C614D2CFF86FFDB76F2A545FBBEF83F" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[968,1109,1917,1934]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M-06) in occlusal (
<emphasis id="B915EABDFF86FFDB7060A545FA8DF83F" bold="true" box="[1370,1382,1917,1934]" pageId="48" pageNumber="729">E</emphasis>
) and lingual (
<emphasis id="B915EABDFF86FFDB758BA5ACFF57F814" bold="true" box="[177,188,1940,1957]" pageId="48" pageNumber="729">F</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB75C5A5ACFEE6F814" bold="true" box="[255,269,1940,1957]" pageId="48" pageNumber="729">G</emphasis>
,
<emphasis id="B915EABDFF86FFDB7422A5ACFECDF814" bold="true" box="[280,294,1940,1957]" pageId="48" pageNumber="729">H</emphasis>
, left P3 of
<taxonomicName id="4C614D2CFF86FFDB7445A5ACFDE7F814" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[383,524,1940,1957]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M-22) in occlusal (
<emphasis id="B915EABDFF86FFDB7634A5ACFCF7F814" bold="true" box="[782,796,1940,1957]" pageId="48" pageNumber="729">G</emphasis>
) and lingual (
<emphasis id="B915EABDFF86FFDB76B0A5ACFC73F814" bold="true" box="[906,920,1940,1957]" pageId="48" pageNumber="729">H</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB76E1A5ACFC0AF814" bold="true" box="[987,993,1940,1957]" pageId="48" pageNumber="729">I</emphasis>
,
<emphasis id="B915EABDFF86FFDB76D1A5ACFC1EF814" bold="true" box="[1003,1013,1940,1957]" pageId="48" pageNumber="729">J</emphasis>
, left?P4 of
<taxonomicName id="4C614D2CFF86FFDB7163A5ACFB33F814" box="[1113,1240,1940,1957]" class="Mammalia" family="Plagiaulacidae" genus="Sunnyodon" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B915EABDFF86FFDB7163A5ACFB53F814" box="[1113,1208,1940,1957]" italics="true" pageId="48" pageNumber="729">Sunnyodon</emphasis>
sp.
</taxonomicName>
(ANG M-04) in occlusal (
<emphasis id="B915EABDFF86FFDB709FA5ACFA40F814" bold="true" box="[1445,1451,1940,1957]" pageId="48" pageNumber="729">I</emphasis>
) and labial (
<emphasis id="B915EABDFF86FFDB75E5A593FF02F80D" bold="true" box="[223,233,1963,1980]" pageId="48" pageNumber="729">J</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB7417A593FED1F80D" bold="true" box="[301,314,1963,1980]" pageId="48" pageNumber="729">K</emphasis>
,
<emphasis id="B915EABDFF86FFDB747FA593FEBBF80D" bold="true" box="[325,336,1963,1980]" pageId="48" pageNumber="729">L</emphasis>
, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (
<emphasis id="B915EABDFF86FFDB76BFA593FC79F80D" bold="true" box="[901,914,1963,1980]" pageId="48" pageNumber="729">K</emphasis>
) and occluso-labial (
<emphasis id="B915EABDFF86FFDB717AA593FBA0F80D" bold="true" box="[1088,1099,1963,1980]" pageId="48" pageNumber="729">L</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB71B5A593FB4BF80D" bold="true" box="[1167,1184,1963,1980]" pageId="48" pageNumber="729">M</emphasis>
,
<emphasis id="B915EABDFF86FFDB7191A593FB52F80D" bold="true" box="[1195,1209,1963,1980]" pageId="48" pageNumber="729">N</emphasis>
, left?m2 of
<taxonomicName id="4C614D2CFF86FFDB7018A593FA44F80D" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[1314,1455,1963,1980]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M- 105) in occlusal (
<emphasis id="B915EABDFF86FFDB74ADA5FAFE43F862" bold="true" box="[407,424,1986,2003]" pageId="48" pageNumber="729">M</emphasis>
) and labial (
<emphasis id="B915EABDFF86FFDB7734A5FAFDF7F862" bold="true" box="[526,540,1986,2003]" pageId="48" pageNumber="729">N</emphasis>
) views;
<emphasis id="B915EABDFF86FFDB775BA5FAFD9BF862" bold="true" box="[609,624,1986,2003]" pageId="48" pageNumber="729">O</emphasis>
,
<emphasis id="B915EABDFF86FFDB7741A5FAFD6CF862" bold="true" box="[635,647,1986,2003]" pageId="48" pageNumber="729">P</emphasis>
, right M2 of
<taxonomicName id="4C614D2CFF86FFDB77CEA5FAFC6AF862" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[756,897,1986,2003]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="729" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
indet. (ANG M-32) in occlusal (
<emphasis id="B915EABDFF86FFDB71B0A5FAFB72F862" bold="true" box="[1162,1177,1986,2003]" pageId="48" pageNumber="729">O</emphasis>
) and lingual (
<emphasis id="B915EABDFF86FFDB7033A5FAFAFEF862" bold="true" box="[1289,1301,1986,2003]" pageId="48" pageNumber="729">P</emphasis>
) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm.
</paragraph>
</caption>
<paragraph id="8BDE36AFFF87FFDA75A1A36FFE19FDA0" blockId="49.[130,776,215,944]" pageId="49" pageNumber="730">
The?m2 (ANG M-105) has a well-developed central basin. Labial cusps are missing and there are only two lingual cusps as in
<taxonomicName id="4C614D2CFF87FFDA75FFA3AEFEBBFE01" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[197,336,406,432]" class="Mammalia" family="Pinheirodontidae" genus="Bernardodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="730" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF87FFDA75FFA3AEFEBBFE01" box="[197,336,406,432]" italics="true" pageId="49" pageNumber="730">Bernardodon</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A2AFF87FFDA745BA3AEFE20FE00" box="[353,459,406,433]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33N</figureCitation>
), whereas there are three in
<taxonomicName id="4C614D2CFF87FFDA75BCA38EFDEAFE60" authority="(Hahn &amp; Hahn 1999)" baseAuthorityName="Hahn &amp; Hahn" baseAuthorityYear="1999" box="[134,513,438,465]" class="Mammalia" family="Pinheirodontidae" genus="Pinheirodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="730" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF87FFDA75BCA38EFEECFE61" box="[134,263,438,464]" italics="true" pageId="49" pageNumber="730">Pinheirodon</emphasis>
(
<bibRefCitation id="EFF04B5EFF87FFDA742FA38EFE1EFE60" author="Hahn &amp; Hahn" box="[277,501,438,465]" firstAuthor="Hahn" pageId="49" pageNumber="730" pagination="77 - 222" refId="ref40980" refString="HAHN G. &amp; HAHN R. 1999. - Pinheirodontidae n. fam. (Multituberculata) (Mammalia) aus der tiefen Unter-Kreide Portugals. Palaeontographica Abteilung A: 77 - 222." type="book chapter" year="1999">Hahn &amp; Hahn 1999</bibRefCitation>
)
</taxonomicName>
. Nevertheless, the tooth shape is considerably longer than wide, and precludes assigning it to
<taxonomicName id="4C614D2CFF87FFDA75D9A3CEFE80FDA1" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[227,363,502,528]" class="Mammalia" family="Pinheirodontidae" genus="Bernardodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="730" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF87FFDA75D9A3CEFE80FDA1" box="[227,363,502,528]" italics="true" pageId="49" pageNumber="730">Bernardodon</emphasis>
</taxonomicName>
(
<figureCitation id="135A2A2AFF87FFDA7441A3CEFE0EFDA0" box="[379,485,502,529]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33M</figureCitation>
).
</paragraph>
<paragraph id="8BDE36AFFF87FFDA75A1A02EFD72FD61" blockId="49.[130,776,215,944]" pageId="49" pageNumber="730">
The M2 (ANG M-32) shows a prominent anterolingual shelf (
<figureCitation id="135A2A2AFF87FFDA75F2A00EFEC5FDE1" box="[200,302,566,592]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33O</figureCitation>
), like that observed in the molars of
<emphasis id="B915EABDFF87FFDA77F7A00FFF25FDC1" italics="true" pageId="49" pageNumber="730">Bernarodon</emphasis>
and
<taxonomicName id="4C614D2CFF87FFDA7436A06EFCEAFDC1" authority="(Kielan-Jaworowska et al. 2004)" baseAuthorityName="Kielan-Jaworowska" baseAuthorityYear="2004" box="[268,769,598,624]" class="Mammalia" family="Pinheirodontidae" genus="Pinheirodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="730" phylum="Chordata" rank="genus">
<emphasis id="B915EABDFF87FFDA7436A06EFE64FDC1" box="[268,399,598,624]" italics="true" pageId="49" pageNumber="730">Pinheirodon</emphasis>
(Kielan-Jaworowska
<emphasis id="B915EABDFF87FFDA7747A06FFD5FFDC1" box="[637,692,598,624]" italics="true" pageId="49" pageNumber="730">et al.</emphasis>
2004)
</taxonomicName>
. As in the latter two genera there are three lingual cusps, but ANG M-32 differs from other pinheirodontids in lacking the labial cusps (
<figureCitation id="135A2A2AFF87FFDA7400A08DFE72FD61" box="[314,409,693,720]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33P</figureCitation>
;
<bibRefCitation id="EFF04B5EFF87FFDA749FA08DFD63FD61" author="Hahn &amp; Hahn" box="[421,648,693,720]" firstAuthor="Hahn" pageId="49" pageNumber="730" pagination="77 - 222" refId="ref40980" refString="HAHN G. &amp; HAHN R. 1999. - Pinheirodontidae n. fam. (Multituberculata) (Mammalia) aus der tiefen Unter-Kreide Portugals. Palaeontographica Abteilung A: 77 - 222." type="book chapter" year="1999">Hahn &amp; Hahn 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37B6524FF87FFDA75A1A0EDFE1CFC01" pageId="49" pageNumber="730" type="discussion">
<paragraph id="8BDE36AFFF87FFDA75A1A0EDFE1CFC01" blockId="49.[130,776,215,944]" pageId="49" pageNumber="730">
The anterior upper premolars P1 (ANG M-03), P2 (ANG M-06) and P3 (ANG M-22) have 3-4 ribbed cusps arranged in two rows like in other “plagiaulacidans” (
<figureCitation id="135A2A2AFF87FFDA7741A12DFD20FC9E" box="[635,715,789,815]" captionStart="FIG" captionStartId="48.[132,143,1895,1912]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-21@48.[132,1455,195,1858]" captionTargetPageId="48" captionText="FIG. 33. — Multituberculate mammal teeth from Angeac-Charente: A, B, left p4 of Pinheirodontidae indet. (ANG M-72) in lingual (A) and labial (B) views; C, D, left P1 of Pinheirodontidae indet. (ANG M-03) in occlusal (C) and distolingual (D) views; E, F, right P2 of Pinheirodontidae indet. (ANG M-06) in occlusal (E) and lingual (F) views; G, H, left P3 of Pinheirodontidae indet. (ANG M-22) in occlusal (G) and lingual (H) views; I, J, left?P4 of Sunnyodon sp. (ANG M-04) in occlusal (I) and labial (J) views; K, L, left?P5 of Multituberculata indet. (ANG M-106) in mesio-occlusal (K) and occluso-labial (L) views; M, N, left?m2 of Pinheirodontidae indet. (ANG M- 105) in occlusal (M) and labial (N) views; O, P, right M2 of Pinheirodontidae indet. (ANG M-32) in occlusal (O) and lingual (P) views. Scale bar: A, B, 1 mm; C, D, 750 µm; E-P, 500 µm." figureDoi="http://doi.org/10.5281/zenodo.6902115" httpUri="https://zenodo.org/record/6902115/files/figure.png" pageId="49" pageNumber="730">Fig. 33</figureCitation>
C-H; Kielan-Jaworowska
<emphasis id="B915EABDFF87FFDA7462A10EFE61FCFE" box="[344,394,821,847]" italics="true" pageId="49" pageNumber="730">et al.</emphasis>
2004). They are not very diagnostic and only tentatively assigned here to
<taxonomicName id="4C614D2CFF87FFDA773EA16DFD2AFCDE" authorityName="Hahn &amp; Hahn" authorityYear="1999" box="[516,705,853,879]" class="Mammalia" family="Pinheirodontidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="49" pageNumber="730" phylum="Chordata" rank="family">Pinheirodontidae</taxonomicName>
, based on their similarities with the pinheirodontid teeth described by
<bibRefCitation id="EFF04B5EFF87FFDA759CA1ADFE78FC1E" author="Hahn &amp; Hahn" box="[166,403,917,943]" firstAuthor="Hahn" pageId="49" pageNumber="730" pagination="77 - 222" refId="ref40980" refString="HAHN G. &amp; HAHN R. 1999. - Pinheirodontidae n. fam. (Multituberculata) (Mammalia) aus der tiefen Unter-Kreide Portugals. Palaeontographica Abteilung A: 77 - 222." type="book chapter" year="1999">Hahn &amp; Hahn (1999</bibRefCitation>
: fig. 58).
</paragraph>
</subSubSection>
</treatment>
</document>
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