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<document id="05EBCAE0079631E69C3D781C7E37279D" ID-DOI="10.11646/zootaxa.5027.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5448029" ID-ZooBank="CCF10500-01A3-476C-A4BE-57161E164EBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630464259914" checkinUser="plazi" docAuthor="Kompier, Tom, Karube, Haruki, Futahashi, Ryo &amp; Phan, Quoc Toan" docDate="2021" docId="03A687CBFFC7FFF5E0E77577FCC4FC4B" docLanguage="en" docName="zootaxa.5027.1.1.pdf" docOrigin="Zootaxa 5027 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Planaeschna tsuchi Kompier &amp; Karube &amp; Futahashi &amp; Phan 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="12" masterDocId="FF9FFFB3FFC0FFFEE0707207FFC3FFCA" masterDocTitle="The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="8" updateTime="1699280300883" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="03D86AFB1BAB1083CFB0ED8DAE4F1CB2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="290E3806D5C9FEF1F2A20535409C2403">
<mods:title id="E4E25F58328D15957503475E1F241050">The genus Planaeschna McLachlan, 1895 and its subgroupings in Vietnam, with descriptions of three new species (Odonata: Aeshnidae)</mods:title>
</mods:titleInfo>
<mods:name id="EB5F0C426F696285B3CD3507386FBEE5" type="personal">
<mods:role id="F8B8DC874CCE8D180EA6CF228116EA34">
<mods:roleTerm id="DFE287DC028CA8A950BA92BED4D2D115">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB53ADB5860E3F1EB801E5D5C6C23E0B">Kompier, Tom</mods:namePart>
<mods:affiliation id="72F09D4B3253FB934C0BEC2F5BFC33A8">Schoutenstraat 69, 2596 SK Den Haag, the Netherlands.</mods:affiliation>
</mods:name>
<mods:name id="A86CCCED49B840E677B27118577D5A4F" type="personal">
<mods:role id="C19E7EFDF9D602F7235FAFE4B2FECAEA">
<mods:roleTerm id="1686C42882136CDCDFB24A791C57F94F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF09A3BADDB539650AC5AA57D3645068">Karube, Haruki</mods:namePart>
<mods:affiliation id="7AA9B42EDA8726983C595DDB109F896B">Kanagawa Prefectural Museum of Natural History, Odawara, Japan.</mods:affiliation>
</mods:name>
<mods:name id="F2B7D749D5639820AF97C0BB30DFE6A2" type="personal">
<mods:role id="55BC1E576E334FC5D34C63E83EEE418D">
<mods:roleTerm id="2129147B653A1E62A3B6366017EF4B7B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CC80CE0D7BBD9306F34C659D53B676C6">Futahashi, Ryo</mods:namePart>
<mods:affiliation id="6F85257DA252F91B522949A44E96BE59">National Institute of Advanced Industrial Science and Technology (AIST), Tsukuba, Ibaraki, Japan.</mods:affiliation>
</mods:name>
<mods:name id="6B347F38F77169CED5E675E62E4EE832" type="personal">
<mods:role id="537247861919501A9E2535798006EEFA">
<mods:roleTerm id="55DC1E6914948221940E4E51198FF5FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0084ED4CFB4FDB98D77E4863C5D89F9">Phan, Quoc Toan</mods:namePart>
<mods:nameIdentifier id="562ABFE91F0F695CAE96BBD394E7F8AC" type="ORCID">0000-0002-3154-6546</mods:nameIdentifier>
<mods:affiliation id="40F107FDDD21B5EA9F4DE2D2955F380E">The Center for Entomology &amp; Parasitology Research, College of Medicine &amp; Pharmacy, Duy Tan University, 550000, Da Nang, Vietnam. pqtoan 84 @ gmail. com; https: // orcid. org / 0000 - 0002 - 3154 - 6546</mods:affiliation>
<mods:nameIdentifier id="780BAA943E4CA0DBEB9108A4D6FB2D93" type="email">pqtoan84@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="74A7D39A4E1E61CEB06B5B0BA13624FC">text</mods:typeOfResource>
<mods:relatedItem id="25DB138AFA6281A98994D02972C6ED7A" type="host">
<mods:titleInfo id="56D8B94050D2B4A98DBB2C5B6C346336">
<mods:title id="F53A67B29C80941109DDE9F8C8412281">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4BF16ECCE03D3F5E964E65D8229DC13A">
<mods:date id="95874018C39F4030125E58F1298A3230">2021</mods:date>
<mods:detail id="1EC6110B686A87500DB1B47D7B72E597" type="pubDate">
<mods:number id="90FC5E11C676FC7628AE6B60DD93D0BD">2021-08-30</mods:number>
</mods:detail>
<mods:detail id="E1648AAC1C65E5C3FFB290C6AD3DC6B3" type="volume">
<mods:number id="D1724C46BBB25F1846D9794269517287">5027</mods:number>
</mods:detail>
<mods:detail id="E205E68FF6F734A7D68738D275EBD33E" type="issue">
<mods:number id="19C681814173B77D8B19ED95C87850C3">1</mods:number>
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<mods:extent id="052A0A619F258DDC0452C420FE9E605A" unit="page">
<mods:start id="E4E5225A5D0D149A837187B97EAD3661">1</mods:start>
<mods:end id="15D1968EA96EA1EB6111DDE1A48B3274">35</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="5B77BEB00C0CDD43D62C01F3AA488A20">journal article</mods:classification>
<mods:identifier id="DCDCE88BEED86B311DEFF2D8EFD835DA" type="DOI">10.11646/zootaxa.5027.1.1</mods:identifier>
<mods:identifier id="5134A8005F7C5F70483E89F281954AD9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="10937C6FF2B6427D6EFBDFD64184BBFC" type="Zenodo-Dep">5448029</mods:identifier>
<mods:identifier id="7AD7BD843A78FB78628FB876E397F2FC" type="ZooBank">CCF10500-01A3-476C-A4BE-57161E164EBD</mods:identifier>
</mods:mods>
<treatment id="03A687CBFFC7FFF5E0E77577FCC4FC4B" ID-DOI="http://doi.org/10.5281/zenodo.5492686" ID-Zenodo-Dep="5492686" LSID="urn:lsid:plazi:treatment:03A687CBFFC7FFF5E0E77577FCC4FC4B" httpUri="http://treatment.plazi.org/id/03A687CBFFC7FFF5E0E77577FCC4FC4B" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8">
<subSubSection id="C3156556FFC7FFF9E0E77577FDC4F840" box="[151,519,1903,1930]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BB036DDFFC7FFF9E0E77577FDC4F840" blockId="7.[151,519,1903,1967]" box="[151,519,1903,1930]" pageId="7" pageNumber="8">
<heading id="D0F881B1FFC7FFF9E0E77577FDC4F840" bold="true" box="[151,519,1903,1930]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B97BEACFFFC7FFF9E0E77577FDC4F840" bold="true" box="[151,519,1903,1930]" pageId="7" pageNumber="8">
3.
<taxonomicName id="4C0F4D5EFFC7FFF9E0C57568FE62F840" authority="Kompier &amp; Karube &amp; Futahashi &amp; Phan, 2021" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[181,417,1903,1930]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tsuchi" status="sp. nov.">
<emphasis id="B97BEACFFFC7FFF9E0C57568FE62F840" bold="true" box="[181,417,1903,1930]" italics="true" pageId="7" pageNumber="8">Planaeschna tsuchi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24857B4FFC7FFF9E1D97577FDC4F840" box="[425,519,1904,1930]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFC7FFF9E0E77593FF28F865" box="[151,235,1940,1967]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BB036DDFFC7FFF9E0E77593FF28F865" blockId="7.[151,519,1903,1967]" box="[151,235,1940,1967]" pageId="7" pageNumber="8">
(
<figureCitation id="13342A58FFC7FFF9E0D07593FF21F865" box="[160,226,1940,1967]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFC7FFF4E0E775DBFE7BFF33" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BB036DDFFC7FFF9E0E775DBFBAAF83C" blockId="7.[151,1129,2012,2039]" box="[151,1129,2012,2039]" pageId="7" pageNumber="8">
<materialsCitation id="3B673C80FFC7FFF9E0E775DBFBAAF83C" box="[151,1129,2012,2039]" collectingDate="2016-10-22" collectorName="Xuan Son NP" latitude="21.139" location="Phu Tho Prov." longLatPrecision="75" longitude="104.938" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Phu Tho" typeStatus="holotype">
<emphasis id="B97BEACFFFC7FFF9E0E775DBFEC8F83C" bold="true" box="[151,267,2012,2038]" pageId="7" pageNumber="8">
<typeStatus id="54B4887FFFC7FFF9E0E775DBFEC5F83C" box="[151,262,2012,2038]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D09FD54FFC7FFF9E16275DAFEEAF83D" box="[274,297,2013,2039]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
,
<collectorName id="26FA530BFFC7FFF9E14575DAFE0CF83D" box="[309,463,2012,2039]" pageId="7" pageNumber="8">Xuan Son NP</collectorName>
,
<collectingRegion id="49CBF83FFFC7FFF9E1AA75DBFDB8F83D" box="[474,635,2012,2039]" country="Vietnam" name="Phu Tho" pageId="7" pageNumber="8">Phu Tho Prov.</collectingRegion>
(
<geoCoordinate id="EE3B501AFFC7FFF9E2F975DBFD35F83C" box="[649,758,2012,2038]" degrees="21.139" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="55" value="21.139">N 21.139</geoCoordinate>
,
<geoCoordinate id="EE3B501AFFC7FFF9E37175DAFCBBF83C" box="[769,888,2012,2039]" degrees="104.938" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="55" value="104.938">E 104.938</geoCoordinate>
),
<date id="FFB1101DFFC7FFF9E3FB75DBFBCAF83C" box="[907,1033,2012,2038]" pageId="7" pageNumber="8" value="2016-10-22">
<collectingDate id="EFF5E9F5FFC7FFF9E3FB75DBFBCAF83C" box="[907,1033,2012,2038]" pageId="7" pageNumber="8" value="2016-10-22">22-X-2016</collectingDate>
</date>
, TK leg.
</materialsCitation>
</paragraph>
<caption id="DF706655FFC8FFF6E0E7759FFAD4F832" ID-DOI="http://doi.org/10.5281/zenodo.5448039" ID-Zenodo-Dep="5448039" httpUri="https://zenodo.org/record/5448039/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1944,1968]" targetBox="[247,1338,181,1920]" targetPageId="8">
<paragraph id="8BB036DDFFC8FFF6E0E7759FFAD4F832" blockId="8.[151,1436,1944,2040]" pageId="8" pageNumber="9">
<emphasis id="B97BEACFFFC8FFF6E0E7759FFED5F87A" bold="true" box="[151,278,1944,1968]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
<taxonomicName id="4C0F4D5EFFC8FFF6E16F759FFDD2F87A" authorityName=": Asahina" authorityYear="1996" box="[287,529,1944,1968]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tomokunii">
<emphasis id="B97BEACFFFC8FFF6E16F759FFDD2F87A" box="[287,529,1944,1968]" italics="true" pageId="8" pageNumber="9">Planaeschna tomokunii</emphasis>
</taxonomicName>
[A–C] lateral habitus alive, including face, [D–E] scans of dorsal habitus, [F–G] anal appendages of male in lateral (F) and dorsal (G) view. (A, F–G) male, Tam Dao NP, Vinh Phuc Prov., 3-VII-2016, (B) immature male, same location, 4-VI-2016, (C, E) female, same location, 24-IX-2015, (D) male, same location, 16-VI-2016.
</paragraph>
</caption>
<caption id="DF706655FFC9FFF7E0E77590FC49F83D" ID-DOI="http://doi.org/10.5281/zenodo.5448041" ID-Zenodo-Dep="5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1943,1967]" targetBox="[225,1363,181,1920]" targetPageId="9">
<paragraph id="8BB036DDFFC9FFF7E0E77590FC49F83D" blockId="9.[151,1436,1943,2040]" pageId="9" pageNumber="10">
<emphasis id="B97BEACFFFC9FFF7E0E77590FED5F87A" bold="true" box="[151,278,1943,1968]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
<taxonomicName id="4C0F4D5EFFC9FFF7E16F759FFE2BF87A" authority="Kompier &amp; Karube &amp; Futahashi &amp; Phan, 2021" authorityName="Kompier &amp; Karube &amp; Futahashi &amp; Phan" authorityYear="2021" box="[287,488,1944,1968]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="tsuchi" status="sp. nov.">
<emphasis id="B97BEACFFFC9FFF7E16F759FFE2BF87A" box="[287,488,1944,1968]" italics="true" pageId="9" pageNumber="10">Planaeschna tsuchi</emphasis>
</taxonomicName>
<emphasis id="B97BEACFFFC9FFF7E180759FFD80F87A" bold="true" box="[496,579,1944,1968]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A24857B4FFC9FFF7E180759FFD80F87A" box="[496,579,1944,1968]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus
<emphasis id="B97BEACFFFC9FFF7E09875BBFEEAF819" box="[232,297,1980,2003]" italics="true" pageId="9" pageNumber="10">in situ</emphasis>
, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015.
</paragraph>
</caption>
<paragraph id="8BB036DDFFCAFFF4E0B77290FE7BFF33" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<materialsCitation id="3B673C80FFCAFFF4E0B77290FC7DFF7B" box="[199,958,151,177]" collectingDate="2014-10-26" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B97BEACFFFCAFFF4E0B77290FE85FF7B" bold="true" box="[199,326,151,177]" pageId="10" pageNumber="11">
<typeStatus id="54B4887FFFCAFFF4E0B77290FE82FF7B" box="[199,321,151,177]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D09FD54FFCAFFF4E13B7290FEB5FF7B" box="[331,374,151,177]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
, same location and collector as
<typeStatus id="54B4887FFFCAFFF4E2A17290FCF5FF7B" box="[721,822,151,177]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
,
<date id="FFB1101DFFCAFFF4E34F7290FC7DFF7B" box="[831,958,151,177]" pageId="10" pageNumber="11" value="2014-10-26">
<collectingDate id="EFF5E9F5FFCAFFF4E34F7290FC7DFF7B" box="[831,958,151,177]" pageId="10" pageNumber="11" value="2014-10-26">26-X-2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFCAFFF4E3B77290FF00FF1F" collectingDate="2015-10-13" collectorName="H. Karube" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D09FD54FFCAFFF4E3B77290FC31FF7B" box="[967,1010,151,177]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
, same location,
<date id="FFB1101DFFCAFFF4E4D27291FADCFF7B" box="[1186,1311,150,177]" pageId="10" pageNumber="11" value="2015-10-13">
<collectingDate id="EFF5E9F5FFCAFFF4E4D27291FADCFF7B" box="[1186,1311,150,177]" pageId="10" pageNumber="11" value="2015-10-13">13-X-2015</collectingDate>
</date>
,
<collectorName id="26FA530BFFCAFFF4E5597290FA5FFF7B" box="[1321,1436,151,177]" pageId="10" pageNumber="11">H. Karube</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B673C80FFCAFFF4E0BE72BCFC9FFF1F" box="[206,860,186,213]" collectingDate="2015-10-15" collectorName="H. Karube" pageId="10" pageNumber="11" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<specimenCount id="9D09FD54FFCAFFF4E0BE72BCFECDFF1F" box="[206,270,187,213]" pageId="10" pageNumber="11" type="female">2 ♀♀</specimenCount>
, same location and collector as
<typeStatus id="54B4887FFFCAFFF4E21E72BCFD10FF1F" box="[622,723,187,213]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
,
<date id="FFB1101DFFCAFFF4E2AD72BDFC9FFF1F" box="[733,860,186,213]" pageId="10" pageNumber="11" value="2015-10-15">
<collectingDate id="EFF5E9F5FFCAFFF4E2AD72BDFC9FFF1F" box="[733,860,186,213]" pageId="10" pageNumber="11" value="2015-10-15">15-X-2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673C80FFCAFFF4E31672BCFE77FF33" collectingDate="2015-10-24" collectorName="H. Karube" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D09FD54FFCAFFF4E31672BCFC4CFF1F" box="[870,911,187,213]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
(specimen used for description), same location and collector,
<date id="FFB1101DFFCAFFF4E14572D9FE77FF33" box="[309,436,222,249]" pageId="10" pageNumber="11" value="2015-10-24">
<collectingDate id="EFF5E9F5FFCAFFF4E14572D9FE77FF33" box="[309,436,222,249]" pageId="10" pageNumber="11" value="2015-10-24">24-X-2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFCAFFF4E0B77304FBD6FE8B" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BB036DDFFCAFFF4E0B77304FBD6FE8B" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77304FE8DFED7" bold="true" box="[199,334,259,285]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species takes its name from the pattern on the dorsum of S2, which is reminiscent of the Japanese character for ground “±”, pronounced “tsuchi”. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFCAFFF5E0B7734DFBC1FEAF" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
<paragraph id="8BB036DDFFCAFFF4E0B7734DFD4CFEAF" blockId="10.[151,1437,150,2013]" box="[199,655,330,357]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B7734DFE1DFEAF" bold="true" box="[199,478,330,357]" pageId="10" pageNumber="11">
Description of
<typeStatus id="54B4887FFFCAFFF4E107734CFE1DFEAF" box="[375,478,331,357]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="13342A58FFCAFFF4E19D734CFD40FEAF" box="[493,643,330,357]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="10" pageNumber="11">Fig. 5C, E–F</figureCitation>
).
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77368FB45FDD3" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77368FEC8FE43" box="[199,267,367,393]" italics="true" pageId="10" pageNumber="11">Head.</emphasis>
Labium and base of mandibles orange. Upper half of labrum yellow, lower half orange, anteclypeus orange-brown, darker medially, postclypeus yellow with two orange dimples, and covered in sparse dark hairs. Antefrons yellow along lower margin and at sides, with large brown mark on central part, covered in dark hairs. Postfrons blackish brown, covered in denser dark hairs. Antennae blackish brown. Vertex and occiput black. Compound eyes green with some bluish and grey tones, yellow at lower posterior margin.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77024FEA3FCF3" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77024FEDBFDF7" box="[199,280,547,573]" italics="true" pageId="10" pageNumber="11">Thorax</emphasis>
. Prothorax dark brown, somewhat paler at the sides. Pterothorax brown with greenish yellow markings as follows: two green dorsal stripes on the mesepisternum (but lacking markings on the antealar sinus), mesepimeron dark brown along humeral suture, but largely covered by greenish yellow broad lateral stripe. Mesinfraepisternum pale brown. Metepisternum with small greenish yellow triangle near base of wing. Metepimeron greenish yellow, except for brown adjacent to metapleural suture. Metinfraepisternum with posterior fourth yellow. Venter brownish. Trochanters brown, but hind pair with some yellow posteriorly. Legs including spikes and claws blackish brown, but base of fore and middle femora orange-brown on the outer surface and femora of fore legs yellow for proximal half on the inside.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77144FABDFC4B" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77144FEC8FC96" box="[199,267,835,860]" italics="true" pageId="10" pageNumber="11">Wings</emphasis>
hyaline, veins black. Nodal index: 23–26:29–22/24–20:21–24; Pt blackish brown covering 3–4 underlying cells in FW and HW. Triangle 6-celled in FW, 4–5 celled in HW. Anal triangle 3-celled. Anal loop 12–14 celled.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B7718CFC80FB6B" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B7718CFEF2FC6F" box="[199,305,907,933]" italics="true" pageId="10" pageNumber="11">Abdomen</emphasis>
(
<figureCitation id="13342A58FFCAFFF4E14F718CFE50FC6F" box="[319,403,906,933]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
) blackish brown with markings as follows: S1 laterally with lower two-thirds yellow, invaded from the anterior margin by a large brown mark. S2 laterally lower half yellow, with yellow auricle and blackish brownish blotch on posterior half. Dorsally with clear green wedge-shaped AD mark, the point facing posteriorly, a green stripe over posterior part of mid-dorsal line, leaving centre and posterior margin free, two MD spots in the form of wedges with apices pointing outward, and two triangular PD markings along the posterior margin, expanding laterally a little more than the MD wedges. S3 with yellow AL mark, two triangular green MD and two oval yellow PD spots, and a small greenish yellow PL spot. S4–8 with green MD spots progressively reduced until tiny on S8, and laterally with a yellow AL spot. S9–10 black.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B776ACFCD6FB23" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B776ACFE64FB0F" box="[199,423,1195,1221]" italics="true" pageId="10" pageNumber="11">Secondary genitalia</emphasis>
apical segment of vesica spermalis of usual
<typeStatus id="54B4887FFFCAFFF4E3E076ACFC03FB0F" box="[912,960,1195,1221]" pageId="10" pageNumber="11">type</typeStatus>
with two somewhat triangular spheres, but not much separated by apical cleft, and with ears at base.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B776F4FCD1FAB3" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B776F4FE4FFAC7" box="[199,396,1267,1293]" italics="true" pageId="10" pageNumber="11">Anal appendages</emphasis>
(
<figureCitation id="13342A58FFCAFFF4E1EF76F4FDD7FAC7" box="[415,532,1266,1293]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="10" pageNumber="11">Fig. 5E–F</figureCitation>
) black. Cerci in dorsal view with apical two-thirds clearly paddle-shaped and with prominent dorsal carina, in lateral view slightly upcurving, apically pointed and with a slight basal swelling. Epiproct extending just beyond midpoint of cerci, in lateral view slender at base, quickly thickening towards middle and tapering again, apex dorsally with two prominences.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77784FC00FA57" blockId="10.[151,1437,150,2013]" box="[199,963,1410,1437]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77784FEB5FA57" bold="true" box="[199,374,1411,1437]" pageId="10" pageNumber="11">Measurements</emphasis>
(in mm). HW 46.0; abdomen incl. appendages 54.0.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B777A1FE5CF9E7" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B777A1FDCDFA0B" bold="true" box="[199,526,1446,1473]" pageId="10" pageNumber="11">
Variation in male
<typeStatus id="54B4887FFFCAFFF4E1D377A0FDCDFA0B" box="[419,526,1447,1473]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
</emphasis>
(
<figureCitation id="13342A58FFCAFFF4E26F77A0FDB8FA0B" box="[543,635,1446,1473]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
). Virtually identical to
<materialsCitation id="3B673C80FFCAFFF4E3F377A0FE5FF9E7" collectionCode="PD, MD" pageId="10" pageNumber="11" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B4887FFFCAFFF4E3F377A0FC2AFA0B" box="[899,1001,1447,1473]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
. Dorsal stripes on pterothorax slightly wider. Yellow
<collectionCode id="ED1EAE18FFCAFFF4E14B77CCFE9CFA2F" box="[315,351,1483,1509]" country="Netherlands" name="Dutch Plant Protection Service, Culture Collection of Plant Pathogenic Bacteria" pageId="10" pageNumber="11">PD</collectionCode>
spots on S2 smaller, but
<collectionCode id="ED1EAE18FFCAFFF4E20C77CCFD6AFA2F" box="[636,681,1483,1509]" country="Germany" name="Museum Donaueschingen" pageId="10" pageNumber="11">MD</collectionCode>
spots on S8 slightly more prominent. Ax in FW 26–30 and in HW 19–23, Px in FW 22–23 and in HW 23–26. Anal loop 10–13 celled. Triangles 6–7 celled in FW, 5–6 celled in HW. Pt covering 3–3.5 cells
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77430FB82F99B" blockId="10.[151,1437,150,2013]" box="[199,1089,1590,1617]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77430FEB5F99B" bold="true" box="[199,374,1591,1617]" pageId="10" pageNumber="11">Measurements</emphasis>
(in mm). HW 47.3–48.1; abdomen incl. appendages 55.4–57.8.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B7745DFB80F9BF" blockId="10.[151,1437,150,2013]" box="[199,1091,1626,1653]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B7745DFDFBF9BF" bold="true" box="[199,568,1626,1653]" pageId="10" pageNumber="11">
Description of female
<typeStatus id="54B4887FFFCAFFF4E1BD745CFDFBF9BF" box="[461,568,1627,1653]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
</emphasis>
(
<figureCitation id="13342A58FFCAFFF4E238745CFD62F9BF" box="[584,673,1626,1653]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="10" pageNumber="11">Fig. 5D</figureCitation>
). As
<typeStatus id="54B4887FFFCAFFF4E2AC745CFCFDF9BF" box="[732,830,1627,1653]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
male, unless specified.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77478FE04F977" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77478FEC8F953" box="[199,267,1663,1689]" italics="true" pageId="10" pageNumber="11">Head.</emphasis>
Yellow areas of labrum, postclypeus and antefrons with greenish wash. Brown central mark on antefrons extending to lower margin.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B774C0FEC0F8E3" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B774C0FEC8F92A" box="[199,267,1735,1760]" italics="true" pageId="10" pageNumber="11">Wings</emphasis>
amber at base, extending to the level of the third Ax. Wings with somewhat smoky-brown wash. Nodal index 21–30:28–20/23–19:18–23, triangle in FW 6-celled and in HW 4–5 celled, anal loop 11–12 celled, Pt covering 3–4 cells.
</paragraph>
<paragraph id="8BB036DDFFCAFFF4E0B77534FD06F817" blockId="10.[151,1437,150,2013]" pageId="10" pageNumber="11">
<emphasis id="B97BEACFFFCAFFF4E0B77534FEFAF887" box="[199,313,1843,1869]" italics="true" pageId="10" pageNumber="11">Abdomen.</emphasis>
S1 with only a triangular yellow PL spot. S2 laterally with lower half yellowish green area invaded by a broad dark brown stripe extending from the anteroventral corner obliquely upward. Dorsal pattern of S2 with somewhat longer MD wedges and shorter PD triangles. S3 with hardly discernible PD spots. S4 with greenishyellow PL spot narrowly connected to yellow kidney-shaped AL mark, S5–6 with kidney-shaped AL mark and small greenish-yellow PL spot. S7–8 lacking MD spots.
</paragraph>
<paragraph id="8BB036DDFFCBFFF5E0B77290FACBFF7B" blockId="11.[151,1437,151,897]" box="[199,1288,151,177]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B77290FE4AFF7B" box="[199,393,151,177]" italics="true" pageId="11" pageNumber="12">Anal appendages</emphasis>
. Cerci black and as long as S8 (shorter than S9–10), ovipositor blackish brown.
</paragraph>
<paragraph id="8BB036DDFFCBFFF5E0B772BDFCBDFE8B" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B772BDFE73FF1E" bold="true" box="[199,432,186,212]" pageId="11" pageNumber="12">Variation in female</emphasis>
<materialsCitation id="3B673C80FFCBFFF5E1C972BCFCB9FE8B" collectionCode="MD" pageId="11" pageNumber="12" specimenCount="1" typeStatus="paratype">
<emphasis id="B97BEACFFFCBFFF5E1C972BCFDECFF1F" bold="true" box="[441,559,187,213]" pageId="11" pageNumber="12">
<typeStatus id="54B4887FFFCBFFF5E1C972BCFDECFF1F" box="[441,559,187,213]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
</emphasis>
(
<figureCitation id="13342A58FFCBFFF5E23072BCFD59FF1F" box="[576,666,186,213]" captionStart="FIGURE 5" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[225,1363,181,1920]" captionTargetId="figure-17@9.[225,1363,181,1920]" captionTargetPageId="9" captionText="FIGURE 5. Planaeschna tsuchi sp. nov. [A–B] lateral habitus alive, including face, [C] scan of dorsal habitus, [D] dorsal habitus in situ, [E–F] anal appendages of male in dorsal (E) and lateral (F) view. (A) male paratype, 26-X-2014, (B) female paratype, 15-X-2015, (C, E–F) holotype, (D) female paratype, 24-X-2015." figureDoi="http://doi.org/10.5281/zenodo.5448041" httpUri="https://zenodo.org/record/5448041/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
). Little variation, dorsal pattern on S2 with
<collectionCode id="ED1EAE18FFCBFFF5E4E272BCFB7CFF1F" box="[1170,1215,187,213]" country="Germany" name="Museum Donaueschingen" pageId="11" pageNumber="12">MD</collectionCode>
wedges reduced in both individuals. Amber in wing extending as far as arculus in
<specimenCount id="9D09FD54FFCBFFF5E32872D8FC36FF33" box="[856,1013,223,249]" pageId="11" pageNumber="12" type="generic">one specimen</specimenCount>
, brownish wash absent from wings in the other. Ax in FW 28–30 and in HW 19–23, Px in FW 19–23 and in HW 21–23. Anal loop 10–12 celled. Triangles 5–7 celled in FW and 4–6 celled in HW. Pt covering 2.5–3.5 cells
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036DDFFCBFFF5E0B7734CFBC1FEAF" blockId="11.[151,1437,151,897]" box="[199,1026,330,357]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B7734CFEB5FEAF" bold="true" box="[199,374,331,357]" pageId="11" pageNumber="12">Measurements</emphasis>
(in mm). HW 47.0; abdomen incl. appendages 53.0–56.5.
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFCBFFF5E0B77369FD58FD63" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BB036DDFFCBFFF5E0B77369FCB6FE3F" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B77369FE10FE42" bold="true" box="[199,467,366,392]" pageId="11" pageNumber="12">Differential Diagnosis.</emphasis>
Both the combination of the pattern on S2 and lack of a longitudinal line over S3 and the robust paddle-shaped cerci are unique among the other known Vietnamese
<taxonomicName id="4C0F4D5EFFCBFFF5E47A7394FB39FE67" box="[1034,1274,403,429]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BEACFFFCBFFF5E47A7394FB58FE67" box="[1034,1179,403,429]" italics="true" pageId="11" pageNumber="12">Planaeschna</emphasis>
species.
</taxonomicName>
Together with the long epiproct of the male, extending beyond the middle of the cerci, and the cerci of the female being shorter than S9+S10, this is enough the conclusively identify the species.
</paragraph>
<paragraph id="8BB036DDFFCBFFF5E0B773F8FD58FD63" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="12">
Closest is
<taxonomicName id="4C0F4D5EFFCBFFF5E14C73F8FC84FDD3" authority="Zhang, Yeh &amp; Tong, 2010" authorityName="Zhang, Yeh &amp; Tong" authorityYear="2010" box="[316,839,511,537]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nankunshanensis">
<emphasis id="B97BEACFFFCBFFF5E14C73F8FDDBFDD3" box="[316,536,511,537]" italics="true" pageId="11" pageNumber="12">P. nankunshanensis</emphasis>
<bibRefCitation id="EF9E4B2CFFCBFFF5E25073F8FC84FDD3" author="Zhang, H. &amp; Yeh, W. &amp; Tong, X." box="[544,839,511,537]" pageId="11" pageNumber="12" pagination="51 - 60" refId="ref17893" refString="Zhang, H., Yeh, W. &amp; Tong, X. (2010) Descriptions of two new species of the genus Planaeschna from China (Odonata: Anisoptera: Aeshnidae). Zootaxa, 2674 (1), 51 - 60. https: // doi. org / 10.11646 / zootaxa. 2674.1.4" type="journal article" year="2010">Zhang, Yeh &amp; Tong, 2010</bibRefCitation>
</taxonomicName>
of
<collectingRegion id="49CBF83FFFCBFFF5E31F73F8FB9EFDD3" box="[879,1117,511,537]" country="China" name="Guangdong" pageId="11" pageNumber="12">Guangdong Province</collectingRegion>
in
<collectingCountry id="F318764DFFCBFFF5E4F373F8FB0AFDD3" box="[1155,1225,511,537]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
, which has similar maculation and of which the male likewise has robust paddle-shaped cerci. The male of
<taxonomicName id="4C0F4D5EFFCBFFF5E4027024FA8EFDF7" authorityName="Zhang, Yeh &amp; Tong" authorityYear="2010" box="[1138,1357,547,573]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nankunshanensis">
<emphasis id="B97BEACFFFCBFFF5E4027024FA8EFDF7" box="[1138,1357,547,573]" italics="true" pageId="11" pageNumber="12">P. nankunshanensis</emphasis>
</taxonomicName>
can be distinguished by the curved outer edge of the cerci, which have a less prominent dorsal carina, and the much shorter epiproct, and the female by details of maculation of S2 and particularly the very long cerci, as long as S9 and S10 combined (
<bibRefCitation id="EF9E4B2CFFCBFFF5E1647088FE63FD63" author="Zhang, H." box="[276,416,655,681]" pageId="11" pageNumber="12" refId="ref17812" refString="Zhang, H. (2019) Dragonflies and damselflies of China. Vol. 1. Chongqing University Press, Chongqing, 736 pp." type="book" year="2019">Zhang 2019</bibRefCitation>
: p. 197, pp. 211–212).
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFCBFFF5E0B770B4FB0EFC97" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="8BB036DDFFCBFFF5E0B770B4FB0EFC97" blockId="11.[151,1437,151,897]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B770B4FEEFFD07" bold="true" box="[199,300,691,717]" pageId="11" pageNumber="12">Ecology.</emphasis>
All records are from the same well-vegetated shallow side stream under heavy forest cover springing from karst rocks at Xuan Son NP of several hundred meters at appr.
<quantity id="4CF79B38FFCBFFF5E3F870D0FC0DFD3B" box="[904,974,727,753]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="m" value="380.0">380 m</quantity>
asl. It shares this habitat with at least five other
<taxonomicName id="4C0F4D5EFFCBFFF5E0A970FCFE09FCDF" box="[217,458,763,789]" class="Insecta" family="Aeshnidae" genus="Planaeschna" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97BEACFFFCBFFF5E0A970FCFEA9FCDF" box="[217,362,763,789]" italics="true" pageId="11" pageNumber="12">Planaeschna</emphasis>
species.
</taxonomicName>
Males were caught in the afternoon hovering over small open areas of water amongst the rocks, whereas females were observed either over the stream or hanging up in nearby bushes. Although the area was visited from spring to late autumn in consecutive years, it was only encountered in October.
</paragraph>
</subSubSection>
<subSubSection id="C3156556FFCBFFF5E0B77161FCC4FC4B" box="[199,775,870,897]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BB036DDFFCBFFF5E0B77161FCC4FC4B" blockId="11.[151,1437,151,897]" box="[199,775,870,897]" pageId="11" pageNumber="12">
<emphasis id="B97BEACFFFCBFFF5E0B77161FEA2FC4A" bold="true" box="[199,353,870,896]" pageId="11" pageNumber="12">Distribution.</emphasis>
Endemic to
<collectingCountry id="F318764DFFCBFFF5E19E7160FD8EFC4B" box="[494,589,871,897]" name="Vietnam" pageId="11" pageNumber="12">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF83FFFCBFFF5E22C7160FCC3FC4B" box="[604,768,871,897]" country="Vietnam" name="Phu Tho" pageId="11" pageNumber="12">Phu Tho Prov.</collectingRegion>
)
</paragraph>
</subSubSection>
</treatment>
</document>
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