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<document id="B8AA8B4932FF7177BF1A1401AE4F64AE" ID-DOI="10.26107/RBZ-2022-0018" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174164" ID-ZooBank="E3A5AA23-BEAE-445E-981F-93A9E7A79417" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290411274" checkinUser="felipe" docAuthor="Sang, Do Duc &amp; Son, Nguyen Thanh" docDate="2022" docId="03A68780FFCCFFA58D6C07FE7CF4110E" docLanguage="en" docName="RafflesBZool.70.364–375.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pincerna clausa Sang &amp; Son 2022, new species" docType="treatment" docVersion="3" lastPageNumber="373" masterDocId="FF9FFFF8FFC4FFAC8C6000767808166D" masterDocTitle="The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species" masterLastPageNumber="375" masterPageNumber="364" pageNumber="372" updateTime="1699379731378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3557F70154FBD7BF044C2289097EE173">The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species</mods:title>
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<mods:namePart id="AC23BC695E0650F3ECEA0F525A2E8742">Sang, Do Duc</mods:namePart>
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<mods:namePart id="D80BF1262C348114A1F793D33DA56E1B">Son, Nguyen Thanh</mods:namePart>
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<mods:title id="891AF28109838223878BA01453BFA0DE">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="76A45D01AAFFA14D71D1A772FE8BEAA1">2022</mods:date>
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<mods:number id="12186CF8A64A78760B3A59817EE3B516">2022-06-01</mods:number>
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<mods:identifier id="5A308B7C08B5DC43044227E2105C7A17" type="DOI">10.26107/RBZ-2022-0018</mods:identifier>
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<treatment id="03A68780FFCCFFA58D6C07FE7CF4110E" LSID="urn:lsid:plazi:treatment:03A68780FFCCFFA58D6C07FE7CF4110E" httpUri="http://treatment.plazi.org/id/03A68780FFCCFFA58D6C07FE7CF4110E" lastPageId="9" lastPageNumber="373" pageId="8" pageNumber="372">
<subSubSection id="C315651DFFCCFFA48D6C07FE7A5E11CD" box="[268,598,1928,1952]" pageId="8" pageNumber="372" type="nomenclature">
<paragraph id="8BB03696FFCCFFA48D6C07FE7A5E11CD" blockId="8.[216,649,1928,1984]" box="[268,598,1928,1952]" pageId="8" pageNumber="372">
<heading id="D0F881FAFFCCFFA48D6C07FE7A5E11CD" bold="true" box="[268,598,1928,1952]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="372" reason="2">
<emphasis id="B97BEA84FFCCFFA48D6C07FE7A5E11CD" bold="true" box="[268,598,1928,1952]" pageId="8" pageNumber="372">
<taxonomicName id="4C0F4D15FFCCFFA48D6C07FE79C911CD" ID-CoL="24928689-42fd-4ba3-9afa-d02380e9570b" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[268,449,1928,1952]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFCCFFA48D6C07FE79C911CD" bold="true" box="[268,449,1928,1952]" italics="true" pageId="8" pageNumber="372">Pincerna clausa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24857FFFFCCFFA48DB007FE7A5E11CD" box="[464,598,1928,1952]" pageId="8" pageNumber="372" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48CB807DE7A8111AD" box="[216,649,1960,1984]" pageId="8" pageNumber="372" type="description">
<paragraph id="8BB03696FFCCFFA48CB807DE7A8111AD" blockId="8.[216,649,1928,1984]" box="[216,649,1960,1984]" pageId="8" pageNumber="372">
(
<figureCitation id="13342A13FFCCFFA48C8007DE792311AD" box="[224,299,1960,1984]" captionStart="Fig" captionStartId="2.[113,148,1941,1963]" captionTargetBox="[113,1452,311,1926]" captionTargetId="figure-11@2.[113,1452,311,1926]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Pincerna species in Vietnam and Laos. Triangle: Pincerna anceyi; Grey circle: Pincerna costulosus; Black circles: Pincerna mouhoti; Squares: Pincerna vanbuensis; Star: Pincerna clausa, new species. The localities are detailed in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="8" pageNumber="372">Figs. 1</figureCitation>
,
<figureCitation id="13342A13FFCCFFA48D5B07DE797511AD" box="[315,381,1960,1984]" captionStart="Fig" captionStartId="6.[113,148,2063,2085]" captionTargetBox="[126,1440,188,2047]" captionTargetId="figure-11@6.[126,1440,188,2047]" captionTargetPageId="6" captionText="Fig. 4. Shell (apertural view, dorsal view, ventral view, and side view) of Pincerna clausa, new species. A, holotype VNMN-IZ 002.304; B, C, paratypes ZMHU. MOL 044." figureDoi="http://doi.org/10.5281/zenodo.7174172" httpUri="https://zenodo.org/record/7174172/files/figure.png" pageId="8" pageNumber="372">4A–C</figureCitation>
,
<figureCitation id="13342A13FFCCFFA48DEA07DE79C111AD" box="[394,457,1960,1984]" captionStart="Fig" captionStartId="7.[136,171,2063,2085]" captionTargetBox="[148,1462,188,2047]" captionTargetId="figure-8@7.[148,1462,188,2047]" captionTargetPageId="7" captionText="Fig. 5. Details of the shell of Pincerna clausa n. sp. (holotype VNMN-IZ 002.304). A–B, protoconch; C, R2 and sutural tube; D, R3 and boundary between R2 and R3; E, aperture; F, umbilicus." figureDoi="http://doi.org/10.5281/zenodo.7174174" httpUri="https://zenodo.org/record/7174174/files/figure.png" pageId="8" pageNumber="372">5A–F</figureCitation>
,
<figureCitation id="13342A13FFCCFFA48DB607DE79F211AD" box="[470,506,1960,1984]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="8" pageNumber="372">6D</figureCitation>
;
<tableCitation id="C68D032DFFCCFFA48E6707DE7A6C11AD" box="[519,612,1960,1984]" captionStart="Table 1" captionStartId="9.[136,191,189,211]" captionText="Table 1. Shell size variation of Pincerna spp. specimen collections in this study" pageId="8" pageNumber="372">Tables 1</tableCitation>
,
<tableCitation id="C68D032DFFCCFFA48E1307DE7A8911AD" box="[627,641,1960,1984]" captionStart="Table 2" captionStartId="9.[136,191,1135,1157]" captionText="Table 2. Comparison of diagnostic conchological characters among Pincerna species in this study." pageId="8" pageNumber="372">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48C11079E7D1813ED" pageId="8" pageNumber="372" type="materials_examined">
<paragraph id="8BB03696FFCCFFA48C11079E7D1813ED" blockId="8.[112,753,2024,2115]" lastBlockId="8.[813,1452,1256,1408]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48C11079E79111E6D" bold="true" box="[113,281,2024,2048]" pageId="8" pageNumber="372">Type material.</emphasis>
<materialsCitation id="3B673CCBFFCCFFA48D7F079E7CA7130D" collectingDate="2018-12-08" collectorName="Na U Commune &amp; D. S. Do &amp; C. Vilachark" country="Vietnam" county="Tay Trang" elevation="814" latitude="21.226112" location="Di en Bien Province" longLatPrecision="21" longitude="102.9675" municipality="Di en Bien District" pageId="8" pageNumber="372" specimenCode="VNMN-IZ 002.304" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B48834FFCCFFA48D7F079E798A1E6D" box="[287,386,2024,2048]" pageId="8" pageNumber="372" type="holotype">Holotype</typeStatus>
(1 shell, height 5.2 mm, width 5.0 mm;
<figureCitation id="13342A13FFCCFFA48CC7087E790A1E4D" box="[167,258,2056,2080]" captionStart="Fig" captionStartId="6.[113,148,2063,2085]" captionTargetBox="[126,1440,188,2047]" captionTargetId="figure-11@6.[126,1440,188,2047]" captionTargetPageId="6" captionText="Fig. 4. Shell (apertural view, dorsal view, ventral view, and side view) of Pincerna clausa, new species. A, holotype VNMN-IZ 002.304; B, C, paratypes ZMHU. MOL 044." figureDoi="http://doi.org/10.5281/zenodo.7174172" httpUri="https://zenodo.org/record/7174172/files/figure.png" pageId="8" pageNumber="372">Figs. 4A</figureCitation>
,
<figureCitation id="13342A13FFCCFFA48D6B087E79241E4D" box="[267,300,2056,2080]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="8" pageNumber="372">6D</figureCitation>
) (
<specimenCode id="DBA99EEDFFCCFFA48D5F087E7A191E4D" box="[319,529,2056,2080]" collectionCode="VNMN-IZ" pageId="8" pageNumber="372">VNMN-IZ 002.304</specimenCode>
),
<collectingCountry id="F3187606FFCCFFA48E41087E7A761E4D" box="[545,638,2056,2080]" name="Vietnam" pageId="8" pageNumber="372">Vietnam</collectingCountry>
,
<location id="8ED0604DFFCCFFA48EE6087E78DB1E2D" LSID="urn:lsid:plazi:treatment:03A68780FFCCFFA58D6C07FE7CF4110E:8ED0604DFFCCFFA48EE6087E78DB1E2D" country="Vietnam" county="Tay Trang" latitude="21.226112" longLatPrecision="21" longitude="102.9675" municipality="Di en Bien District" name="Di en Bien Province" pageId="8" pageNumber="372">Điện Biên Province</location>
,
<collectingMunicipality id="6BD4ACECFFCCFFA48C83085E79BD1E2D" box="[227,437,2088,2112]" pageId="8" pageNumber="372">Điện Biên District</collectingMunicipality>
,
<collectorName id="26FA5340FFCCFFA48DA2085F7A761E2D" box="[450,638,2088,2115]" pageId="8" pageNumber="372">Na Ư Commune</collectorName>
, near
<collectingCounty id="62D14E1AFFCCFFA48EA6085E7B65136C" box="[710,877,1257,2112]" pageId="8" pageNumber="372">Tây Trang</collectingCounty>
international border gate at the
<collectingCountry id="F3187606FFCCFFA488DC049E7D10136D" box="[1212,1304,1256,1280]" name="Vietnam" pageId="8" pageNumber="372">Vietnam</collectingCountry>
-
<collectingCountry id="F3187606FFCCFFA4894C049F7D68136D" box="[1324,1376,1257,1280]" name="Laos" pageId="8" pageNumber="372">Laos</collectingCountry>
border region, exposed limestone with little soil and vegetation of herbs,
<elevation id="0022D1A5FFCCFFA48F16055E7BF0132D" box="[886,1016,1320,1344]" metricMagnitude="2" metricUnit="m" metricValue="8.14" pageId="8" pageNumber="372" unit="m" value="814.0">
<quantity id="4CF79B73FFCCFFA48F16055E7BB5132D" box="[886,957,1320,1344]" metricMagnitude="2" metricUnit="m" metricValue="8.14" pageId="8" pageNumber="372" unit="m" value="814.0">814 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE3B5051FFCCFFA4886705517C80132D" box="[1031,1160,1319,1344]" degrees="21" direction="north" minutes="13" orientation="latitude" pageId="8" pageNumber="372" precision="15" seconds="34" value="21.226112">21°13′34″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFCCFFA488F605517D1C132D" box="[1174,1300,1319,1344]" degrees="102" direction="east" minutes="58" orientation="longitude" pageId="8" pageNumber="372" precision="15" seconds="3" value="102.9675">102°58′3″E</geoCoordinate>
, col.
<collectorName id="26FA5340FFCCFFA48932055E7DA4132D" box="[1362,1452,1320,1344]" pageId="8" pageNumber="372">D.S. Do</collectorName>
&amp;
<collectorName id="26FA5340FFCCFFA48F29053E7BDC130D" box="[841,980,1352,1376]" pageId="8" pageNumber="372">C. Vilachark</collectorName>
,
<date id="FFB11056FFCCFFA48FBF053E7CA7130D" box="[991,1199,1352,1376]" pageId="8" pageNumber="372" value="2018-12-08">
<collectingDate id="EFF5E9BEFFCCFFA48FBF053E7CA7130D" box="[991,1199,1352,1376]" pageId="8" pageNumber="372" value="2018-12-08">08 December 2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B673CCBFFCCFFA488DA053F7D0413ED" collectingDate="2018-12-08" collectionCode="ZMHU" collectorName="Na U Commune &amp; D. S. Do &amp; C. Vilachark" country="Vietnam" county="Tay Trang" elevation="814" latitude="21.226112" location="Di en Bien Province" longLatPrecision="21" longitude="102.9675" municipality="Di en Bien District" pageId="8" pageNumber="372" specimenCode="MOL 044" specimenCount="3" typeStatus="paratype">
<typeStatus id="54B48834FFCCFFA488DA053F7D2E130C" box="[1210,1318,1353,1377]" pageId="8" pageNumber="372" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D09FD1FFFCCFFA48951053E7DA3130C" box="[1329,1451,1352,1377]" pageId="8" pageNumber="372" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
(
<collectionCode id="ED1EAE53FFCCFFA48F54051E7B8F13ED" box="[820,903,1384,1408]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="372">ZMHU</collectionCode>
.
<specimenCode id="DBA99EEDFFCCFFA48FF3051E7C0B13ED" box="[915,1027,1384,1408]" collectionCode="MOL" country="Peru" lsid="urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" pageId="8" pageNumber="372" type="Herbarium">MOL 044</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48F4D05DE7CEE138D" pageId="8" pageNumber="372" type="etymology">
<paragraph id="8BB03696FFCCFFA48F4D05DE7CEE138D" blockId="8.[813,1452,1448,1504]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48F4D05DE7BA713AD" bold="true" box="[813,943,1448,1472]" pageId="8" pageNumber="372">Etymology.</emphasis>
This new species is named for its conspicuous closed umbilicus (Latin: clausa = close).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA48F4D067E7CCE10ED" pageId="8" pageNumber="372" type="diagnosis">
<paragraph id="8BB03696FFCCFFA48F4D067E7CCE10ED" blockId="8.[813,1451,1544,1664]" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48F4D067E7BAA104D" bold="true" box="[813,930,1544,1568]" pageId="8" pageNumber="372">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D15FFCCFFA48FCB067F7C55104D" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[939,1117,1544,1568]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="372" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFCCFFA48FCB067F7C55104D" box="[939,1117,1544,1568]" italics="true" pageId="8" pageNumber="372">Pincerna clausa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24857FFFFCCFFA4880E067F7CFC104D" box="[1134,1268,1544,1568]" pageId="8" pageNumber="372" rank="species">new species</taxonomicNameLabel>
, is characterised by a small sized, globular shell with clearly spiral striation on R1 and a closed umbilicus, which is completely obscured by the reflected columellar extension.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCCFFA58F4D06DE798811C9" lastPageId="9" lastPageNumber="373" pageId="8" pageNumber="372" type="description">
<paragraph id="8BB03696FFCCFFA58F4D06DE7B001109" blockId="8.[812,1452,1704,2112]" lastBlockId="9.[136,776,1580,1892]" lastPageId="9" lastPageNumber="373" pageId="8" pageNumber="372">
<emphasis id="B97BEA84FFCCFFA48F4D06DE7BBE10AD" bold="true" box="[813,950,1704,1728]" pageId="8" pageNumber="372">Description.</emphasis>
Shell globose, off-white, rather glossy, medium sized (height 5.1–5.3 mm, width 4.9–5.0 mm). Spire high, slightly obtuse (spire height 1.3–1.4, spire width 1.8–2.0 mm). Whorls 4.25–4.5, the periphery rounded. Suture rather deep. Protoconch smooth, glossy, without spiral striation, 1.25–1.75 whorls. R1 of 2.0 whorls, with rather regular, indistinct and low ribs, and clearly spiral striation; on the first part of R1 the ribs are rather weak and irregular, on the last part, however, they are close together and sharp, with well visible spiral striation (
<figureCitation id="13342A13FFCCFFA488FC07BE7CE3118D" box="[1180,1259,1992,2016]" captionStart="Fig" captionStartId="6.[113,148,2063,2085]" captionTargetBox="[126,1440,188,2047]" captionTargetId="figure-11@6.[126,1440,188,2047]" captionTargetPageId="6" captionText="Fig. 4. Shell (apertural view, dorsal view, ventral view, and side view) of Pincerna clausa, new species. A, holotype VNMN-IZ 002.304; B, C, paratypes ZMHU. MOL 044." figureDoi="http://doi.org/10.5281/zenodo.7174172" httpUri="https://zenodo.org/record/7174172/files/figure.png" pageId="8" pageNumber="372">Figs. 4</figureCitation>
,
<figureCitation id="13342A13FFCCFFA4889907BE7D16118D" box="[1273,1310,1992,2016]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="8" pageNumber="372">6D</figureCitation>
). R1 and R2 boundary conspicuous due to denser and slender R2 ribs. R2 and breathing tube relatively long (breathing tube length ca 1.2–1.4 mm), consisting of 34–41 stripes, which are slender and slightly elevated from the surface, overall surface slightly wrinkled. R3 slightly longer than R2; boundary between R2 and R3 distinct due to constriction and change in sculpture. R3 smooth, glossy, with some widely spaced, irregular and weak ribs on the terminal part. Aperture circular, moderately expanded (aperture diameter 3.0–3.1 mm). Peristome double, slightly thickened; outer peristome sharp, expanded but not reflected; inner peristome protruding, sharp; outer and inner peristomes are clearly separated. Umbilicus narrow and completely obscured by the reflected columellar extension.
</paragraph>
<caption id="DF70661EFFCDFFA58CE800CB7B9116BE" box="[136,921,189,211]" pageId="9" pageNumber="373" startId="9.[136,191,189,211]">
<paragraph id="8BB03696FFCDFFA58CE800CB7B9116BE" blockId="9.[136,921,189,211]" box="[136,921,189,211]" pageId="9" pageNumber="373">
Table 1. Shell size variation of
<taxonomicName id="4C0F4D15FFCDFFA58DDF00C87A4F16BE" box="[447,583,190,211]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B97BEA84FFCDFFA58DDF00C87A1F16BE" box="[447,535,190,211]" italics="true" pageId="9" pageNumber="373">Pincerna</emphasis>
spp.
</taxonomicName>
specimen collections in this study
</paragraph>
</caption>
<paragraph id="8BB03696FFCDFFA58CF701577D96124D" pageId="9" pageNumber="373">
<table id="F90FC436FFCD00538CF701737DA2124F" box="[151,1450,261,1058]" gridcols="6" gridrows="21" pageId="9" pageNumber="373">
<tr id="353F34D4FFCD00538CF701737DA21739" box="[151,1450,261,340]" gridrow="0" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF701737ADA1739" box="[151,722,261,340]" gridcol="0" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58CF70157796B175A" bold="true" box="[151,355,289,311]" pageId="9" pageNumber="373">Species and locality</emphasis>
</th>
<th id="76EE5DA8FFCD00538E9401737B541739" box="[756,860,261,340]" gridcol="1" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58F6601657B54172B" bold="true" pageId="9" pageNumber="373">No. of specimens</emphasis>
</th>
<th id="76EE5DA8FFCD00538F1B01737BF51739" box="[891,1021,261,340]" gridcol="2" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58F1D01657BF5172B" bold="true" pageId="9" pageNumber="373">Shell height (SH, in mm)</emphasis>
</th>
<th id="76EE5DA8FFCD0053887E01737CAD1739" box="[1054,1189,261,340]" gridcol="3" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA5884501657CAD172B" bold="true" pageId="9" pageNumber="373">Shell width (SW, in mm)</emphasis>
</th>
<th id="76EE5DA8FFCD005388A301737D2C1739" box="[1219,1316,261,340]" gridcol="4" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA588AD01657D03172B" bold="true" pageId="9" pageNumber="373">SH/SW ratio</emphasis>
</th>
<th id="76EE5DA8FFCD0053892B01737DA21739" box="[1355,1450,261,340]" gridcol="5" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA5892B01737DAD1739" bold="true" box="[1355,1450,261,340]" pageId="9" pageNumber="373">Aperture diameter (in mm)</emphasis>
</th>
</tr>
<tr id="353F34D4FFCD00538CF701027DA217E4" box="[151,1450,372,393]" gridrow="1" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF701027DA217E4" box="[151,1450,372,393]" colspan="6" colspanRight="5" gridcol="0" gridrow="1" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF70102793117E4" baseAuthorityName="Mabille" baseAuthorityYear="1887" box="[151,313,372,393]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFCDFFA58CF70102793117E4" bold="true" box="[151,313,372,393]" italics="true" pageId="9" pageNumber="373">Pincerna anceyi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F34D4FFCD00538CF701E27DA217C7" box="[151,1450,404,426]" gridrow="2" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF701E27ADA17C7" box="[151,722,404,426]" gridcol="0" gridrow="2" pageId="9" pageNumber="373">
1. Tonkin, Vietnam (
<bibRefCitation id="EF9E4B67FFCDFFA58D0501E279F817C4" author="Mabille J" box="[357,496,404,426]" pageId="9" pageNumber="373" pagination="73 - 164" refId="ref6829" refString="Mabille J (1887) On some molluscs from Northern Vietnam. Bulletin of the Malacological Society of France, 4: 73 - 164." type="journal article" year="1887">Mabille, 1887</bibRefCitation>
)
</th>
<td id="76EE5DA8FFCD00538E9401E27B5417C7" box="[756,860,404,426]" gridcol="1" gridrow="2" pageId="9" pageNumber="373">–</td>
<td id="76EE5DA8FFCD00538F1B01E27BF517C7" box="[891,1021,404,426]" gridcol="2" gridrow="2" pageId="9" pageNumber="373">5.0</td>
<td id="76EE5DA8FFCD0053887E01E27CAD17C7" box="[1054,1189,404,426]" gridcol="3" gridrow="2" pageId="9" pageNumber="373">3.5–5.0</td>
<td id="76EE5DA8FFCD005388A301E27D2C17C7" box="[1219,1316,404,426]" gridcol="4" gridrow="2" pageId="9" pageNumber="373">1.00</td>
<td id="76EE5DA8FFCD0053892B01E27DA217C7" box="[1355,1450,404,426]" gridcol="5" gridrow="2" pageId="9" pageNumber="373">–</td>
</tr>
<tr id="353F34D4FFCD00538CF701B47DA217B5" box="[151,1450,450,472]" gridrow="3" pageId="9" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76EE5DA8FFCD00538CF701B47ADA17B5" box="[151,722,450,472]" gridcol="0" gridrow="3" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF701B5796817B5" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[151,352,450,472]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFCDFFA58CF701B5796817B5" bold="true" box="[151,352,450,472]" italics="true" pageId="9" pageNumber="373">Pincerna costulosus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F34D4FFCD00538CF701947DA21797" box="[151,1450,482,506]" gridrow="4" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF701947ADA1797" box="[151,722,482,506]" gridcol="0" gridrow="4" pageId="9" pageNumber="373">2. Khổng Lào, Phong Thổ, Lai Châu, Vietnam</th>
<td id="76EE5DA8FFCD00538E9401947B541797" box="[756,860,482,506]" gridcol="1" gridrow="4" pageId="9" pageNumber="373">4</td>
<td id="76EE5DA8FFCD00538F1B01947BF51797" box="[891,1021,482,506]" gridcol="2" gridrow="4" pageId="9" pageNumber="373">4.8–5.3</td>
<td id="76EE5DA8FFCD0053887E01947CAD1797" box="[1054,1189,482,506]" gridcol="3" gridrow="4" pageId="9" pageNumber="373">4.6–5.2</td>
<td id="76EE5DA8FFCD005388A301947D2C1797" box="[1219,1316,482,506]" gridcol="4" gridrow="4" pageId="9" pageNumber="373">1.02–1.04</td>
<td id="76EE5DA8FFCD0053892B01947DA21797" box="[1355,1450,482,506]" gridcol="5" gridrow="4" pageId="9" pageNumber="373">2.4–2.8</td>
</tr>
<tr id="353F34D4FFCD00538CF702677DA2144A" box="[151,1450,529,551]" gridrow="5" pageId="9" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76EE5DA8FFCD00538CF702677ADA144A" box="[151,722,529,551]" gridcol="0" gridrow="5" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF702677944144A" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[151,332,529,551]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFCDFFA58CF702677944144A" bold="true" box="[151,332,529,551]" italics="true" pageId="9" pageNumber="373">Pincerna mouhoti</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F34D4FFCD00538CF702477DA2142A" box="[151,1450,561,583]" gridrow="6" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702477ADA142A" box="[151,722,561,583]" gridcol="0" gridrow="6" pageId="9" pageNumber="373">
3. Ngoy, Luang Phrabang, Laos (
<bibRefCitation id="EF9E4B67FFCDFFA58DBE02477AB5142B" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[478,701,561,583]" pageId="9" pageNumber="373" pagination="1 - 166" refId="ref6528" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
)
</th>
<td id="76EE5DA8FFCD00538E9402477B54142A" box="[756,860,561,583]" gridcol="1" gridrow="6" pageId="9" pageNumber="373">–</td>
<td id="76EE5DA8FFCD00538F1B02477BF5142A" box="[891,1021,561,583]" gridcol="2" gridrow="6" pageId="9" pageNumber="373">5.25</td>
<td id="76EE5DA8FFCD0053887E02477CAD142A" box="[1054,1189,561,583]" gridcol="3" gridrow="6" pageId="9" pageNumber="373">6.3–7.7</td>
<td id="76EE5DA8FFCD005388A302477D2C142A" box="[1219,1316,561,583]" gridcol="4" gridrow="6" pageId="9" pageNumber="373">0.68</td>
<td id="76EE5DA8FFCD0053892B02477DA2142A" box="[1355,1450,561,583]" gridcol="5" gridrow="6" pageId="9" pageNumber="373">–</td>
</tr>
<tr id="353F34D4FFCD00538CF702267DA2140B" box="[151,1450,592,614]" gridrow="7" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702267ADA140B" box="[151,722,592,614]" gridcol="0" gridrow="7" pageId="9" pageNumber="373">4. Hang Kia – Pà Cò Nature Reserve, Hòa Bình, Vietnam</th>
<td id="76EE5DA8FFCD00538E9402267B54140B" box="[756,860,592,614]" gridcol="1" gridrow="7" pageId="9" pageNumber="373">7</td>
<td id="76EE5DA8FFCD00538F1B02267BF5140B" box="[891,1021,592,614]" gridcol="2" gridrow="7" pageId="9" pageNumber="373">6.2–6.5</td>
<td id="76EE5DA8FFCD0053887E02267CAD140B" box="[1054,1189,592,614]" gridcol="3" gridrow="7" pageId="9" pageNumber="373">6.5–6.9</td>
<td id="76EE5DA8FFCD005388A302267D2C140B" box="[1219,1316,592,614]" gridcol="4" gridrow="7" pageId="9" pageNumber="373">0.92–0.97</td>
<td id="76EE5DA8FFCD0053892B02267DA2140B" box="[1355,1450,592,614]" gridcol="5" gridrow="7" pageId="9" pageNumber="373">3.5–3.8</td>
</tr>
<tr id="353F34D4FFCD00538CF702067DA214E5" box="[151,1450,624,648]" gridrow="8" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702067ADA14E5" box="[151,722,624,648]" gridcol="0" gridrow="8" pageId="9" pageNumber="373">5. Pa Cốp, Vân Hồ, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E9402067B5414E5" box="[756,860,624,648]" gridcol="1" gridrow="8" pageId="9" pageNumber="373">10</td>
<td id="76EE5DA8FFCD00538F1B02067BF514E5" box="[891,1021,624,648]" gridcol="2" gridrow="8" pageId="9" pageNumber="373">6.2–6.5</td>
<td id="76EE5DA8FFCD0053887E02067CAD14E5" box="[1054,1189,624,648]" gridcol="3" gridrow="8" pageId="9" pageNumber="373">6.2–7.0</td>
<td id="76EE5DA8FFCD005388A302067D2C14E5" box="[1219,1316,624,648]" gridcol="4" gridrow="8" pageId="9" pageNumber="373">0.94–1.00</td>
<td id="76EE5DA8FFCD0053892B02067DA214E5" box="[1355,1450,624,648]" gridcol="5" gridrow="8" pageId="9" pageNumber="373">3.4–3.8</td>
</tr>
<tr id="353F34D4FFCD00538CF702E67DA214CA" box="[151,1450,656,679]" gridrow="9" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702E67ADA14CA" box="[151,722,656,679]" gridcol="0" gridrow="9" pageId="9" pageNumber="373">6. Thẳm Bó Cave, MƯờng La, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E9402E67B5414CA" box="[756,860,656,679]" gridcol="1" gridrow="9" pageId="9" pageNumber="373">17</td>
<td id="76EE5DA8FFCD00538F1B02E67BF514CA" box="[891,1021,656,679]" gridcol="2" gridrow="9" pageId="9" pageNumber="373">6.8–7.1</td>
<td id="76EE5DA8FFCD0053887E02E67CAD14CA" box="[1054,1189,656,679]" gridcol="3" gridrow="9" pageId="9" pageNumber="373">6.7–6.8</td>
<td id="76EE5DA8FFCD005388A302E67D2C14CA" box="[1219,1316,656,679]" gridcol="4" gridrow="9" pageId="9" pageNumber="373">1.00–1.04</td>
<td id="76EE5DA8FFCD0053892B02E67DA214CA" box="[1355,1450,656,679]" gridcol="5" gridrow="9" pageId="9" pageNumber="373">3.8–4.0</td>
</tr>
<tr id="353F34D4FFCD00538CF702C67DA214AA" box="[151,1450,688,711]" gridrow="10" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702C67ADA14AA" box="[151,722,688,711]" gridcol="0" gridrow="10" pageId="9" pageNumber="373">7. Lằn, MƯờng Do, Phù Yên, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E9402C67B5414AA" box="[756,860,688,711]" gridcol="1" gridrow="10" pageId="9" pageNumber="373">2</td>
<td id="76EE5DA8FFCD00538F1B02C67BF514AA" box="[891,1021,688,711]" gridcol="2" gridrow="10" pageId="9" pageNumber="373">6.9–7.0</td>
<td id="76EE5DA8FFCD0053887E02C67CAD14AA" box="[1054,1189,688,711]" gridcol="3" gridrow="10" pageId="9" pageNumber="373">6.7–6.8</td>
<td id="76EE5DA8FFCD005388A302C67D2C14AA" box="[1219,1316,688,711]" gridcol="4" gridrow="10" pageId="9" pageNumber="373">1.02–1.03</td>
<td id="76EE5DA8FFCD0053892B02C67DA214AA" box="[1355,1450,688,711]" gridcol="5" gridrow="10" pageId="9" pageNumber="373">3.7–3.8</td>
</tr>
<tr id="353F34D4FFCD00538CF702B97DA2148A" box="[151,1450,719,743]" gridrow="11" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF702B97ADA148A" box="[151,722,719,743]" gridcol="0" gridrow="11" pageId="9" pageNumber="373">8. Copia Nature Reserve, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E9402B97B54148A" box="[756,860,719,743]" gridcol="1" gridrow="11" pageId="9" pageNumber="373">3</td>
<td id="76EE5DA8FFCD00538F1B02B97BF5148A" box="[891,1021,719,743]" gridcol="2" gridrow="11" pageId="9" pageNumber="373">6.0–6.2</td>
<td id="76EE5DA8FFCD0053887E02B97CAD148A" box="[1054,1189,719,743]" gridcol="3" gridrow="11" pageId="9" pageNumber="373">6.2–6.3</td>
<td id="76EE5DA8FFCD005388A302B97D2C148A" box="[1219,1316,719,743]" gridcol="4" gridrow="11" pageId="9" pageNumber="373">0.95–1.00</td>
<td id="76EE5DA8FFCD0053892B02B97DA2148A" box="[1355,1450,719,743]" gridcol="5" gridrow="11" pageId="9" pageNumber="373">3.2–3.4</td>
</tr>
<tr id="353F34D4FFCD00538CF702887DA21579" box="[151,1450,766,788]" gridrow="12" pageId="9" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76EE5DA8FFCD00538CF702887ADA1579" box="[151,722,766,788]" gridcol="0" gridrow="12" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF7028879601579" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[151,360,766,788]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFCDFFA58CF7028879601579" bold="true" box="[151,360,766,788]" italics="true" pageId="9" pageNumber="373">Pincerna vanbuensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353F34D4FFCD00538CF703687DA21558" box="[151,1450,798,821]" gridrow="13" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF703687ADA1558" box="[151,722,798,821]" gridcol="0" gridrow="13" pageId="9" pageNumber="373">9. Copia Nature Reserve, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E9403687B541558" box="[756,860,798,821]" gridcol="1" gridrow="13" pageId="9" pageNumber="373">20</td>
<td id="76EE5DA8FFCD00538F1B03687BF51558" box="[891,1021,798,821]" gridcol="2" gridrow="13" pageId="9" pageNumber="373">5.4–5.6</td>
<td id="76EE5DA8FFCD0053887E03687CAD1558" box="[1054,1189,798,821]" gridcol="3" gridrow="13" pageId="9" pageNumber="373">5.7–5.8</td>
<td id="76EE5DA8FFCD005388A303687D2C1558" box="[1219,1316,798,821]" gridcol="4" gridrow="13" pageId="9" pageNumber="373">0.93–0.97</td>
<td id="76EE5DA8FFCD0053892B03687DA21558" box="[1355,1450,798,821]" gridcol="5" gridrow="13" pageId="9" pageNumber="373">3.1–3.3</td>
</tr>
<tr id="353F34D4FFCD00538CF7034B7DA21538" box="[151,1450,829,853]" gridrow="14" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7034B7ADA1538" box="[151,722,829,853]" gridcol="0" gridrow="14" pageId="9" pageNumber="373">10. Chi Đảy Cave, Yên Châu, SƠn La, Vietnam</th>
<td id="76EE5DA8FFCD00538E94034B7B541538" box="[756,860,829,853]" gridcol="1" gridrow="14" pageId="9" pageNumber="373">5</td>
<td id="76EE5DA8FFCD00538F1B034B7BF51538" box="[891,1021,829,853]" gridcol="2" gridrow="14" pageId="9" pageNumber="373">4.5–4.8</td>
<td id="76EE5DA8FFCD0053887E034B7CAD1538" box="[1054,1189,829,853]" gridcol="3" gridrow="14" pageId="9" pageNumber="373">5.0–5.2</td>
<td id="76EE5DA8FFCD005388A3034B7D2C1538" box="[1219,1316,829,853]" gridcol="4" gridrow="14" pageId="9" pageNumber="373">0.90–0.93</td>
<td id="76EE5DA8FFCD0053892B034B7DA21538" box="[1355,1450,829,853]" gridcol="5" gridrow="14" pageId="9" pageNumber="373">2.5–2.7</td>
</tr>
<tr id="353F34D4FFCD00538CF7032B7DA21518" box="[151,1450,861,885]" gridrow="15" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7032B7ADA1518" box="[151,722,861,885]" gridcol="0" gridrow="15" pageId="9" pageNumber="373">11. Tân SƠn, Mai Châu, Hòa Bình, Vietnam</th>
<td id="76EE5DA8FFCD00538E94032B7B541518" box="[756,860,861,885]" gridcol="1" gridrow="15" pageId="9" pageNumber="373">4</td>
<td id="76EE5DA8FFCD00538F1B032B7BF51518" box="[891,1021,861,885]" gridcol="2" gridrow="15" pageId="9" pageNumber="373">4.9–5.0</td>
<td id="76EE5DA8FFCD0053887E032B7CAD1518" box="[1054,1189,861,885]" gridcol="3" gridrow="15" pageId="9" pageNumber="373">5.0–5.1</td>
<td id="76EE5DA8FFCD005388A3032B7D2C1518" box="[1219,1316,861,885]" gridcol="4" gridrow="15" pageId="9" pageNumber="373">0.90–0.94</td>
<td id="76EE5DA8FFCD0053892B032B7DA21518" box="[1355,1450,861,885]" gridcol="5" gridrow="15" pageId="9" pageNumber="373">2.5–2.6</td>
</tr>
<tr id="353F34D4FFCD00538CF7030B7DA215F9" box="[151,1450,893,916]" gridrow="16" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7030B7ADA15F9" box="[151,722,893,916]" gridcol="0" gridrow="16" pageId="9" pageNumber="373">12. Cúc PhƯƠng N.P., Ninh Bình, Vietnam</th>
<td id="76EE5DA8FFCD00538E94030B7B5415F9" box="[756,860,893,916]" gridcol="1" gridrow="16" pageId="9" pageNumber="373">9</td>
<td id="76EE5DA8FFCD00538F1B030B7BF515F9" box="[891,1021,893,916]" gridcol="2" gridrow="16" pageId="9" pageNumber="373">4.8–5.1</td>
<td id="76EE5DA8FFCD0053887E030B7CAD15F9" box="[1054,1189,893,916]" gridcol="3" gridrow="16" pageId="9" pageNumber="373">4.9–5.0</td>
<td id="76EE5DA8FFCD005388A3030B7D2C15F9" box="[1219,1316,893,916]" gridcol="4" gridrow="16" pageId="9" pageNumber="373">0.96–1.02</td>
<td id="76EE5DA8FFCD0053892B030B7DA215F9" box="[1355,1450,893,916]" gridcol="5" gridrow="16" pageId="9" pageNumber="373">2.4–2.6</td>
</tr>
<tr id="353F34D4FFCD00538CF703EB7DA215DE" box="[151,1450,925,947]" gridrow="17" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF703EB7ADA15DE" box="[151,722,925,947]" gridcol="0" gridrow="17" pageId="9" pageNumber="373">13. Nam Phan, Khoune, Xieng Khaong, Laos</th>
<td id="76EE5DA8FFCD00538E9403EB7B5415DE" box="[756,860,925,947]" gridcol="1" gridrow="17" pageId="9" pageNumber="373">26</td>
<td id="76EE5DA8FFCD00538F1B03EB7BF515DE" box="[891,1021,925,947]" gridcol="2" gridrow="17" pageId="9" pageNumber="373">5.0–5.4</td>
<td id="76EE5DA8FFCD0053887E03EB7CAD15DE" box="[1054,1189,925,947]" gridcol="3" gridrow="17" pageId="9" pageNumber="373">5.3–5.7</td>
<td id="76EE5DA8FFCD005388A303EB7D2C15DE" box="[1219,1316,925,947]" gridcol="4" gridrow="17" pageId="9" pageNumber="373">0.93–0.95</td>
<td id="76EE5DA8FFCD0053892B03EB7DA215DE" box="[1355,1450,925,947]" gridcol="5" gridrow="17" pageId="9" pageNumber="373">2.8–3.0</td>
</tr>
<tr id="353F34D4FFCD00538CF703CA7DA215BE" box="[151,1450,956,979]" gridrow="18" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF703CA7ADA15BE" box="[151,722,956,979]" gridcol="0" gridrow="18" pageId="9" pageNumber="373">14. Xiengkho, Houaphanh, Laos</th>
<td id="76EE5DA8FFCD00538E9403CA7B5415BE" box="[756,860,956,979]" gridcol="1" gridrow="18" pageId="9" pageNumber="373">18</td>
<td id="76EE5DA8FFCD00538F1B03CA7BF515BE" box="[891,1021,956,979]" gridcol="2" gridrow="18" pageId="9" pageNumber="373">5.0–5.1</td>
<td id="76EE5DA8FFCD0053887E03CA7CAD15BE" box="[1054,1189,956,979]" gridcol="3" gridrow="18" pageId="9" pageNumber="373">4.9–5.2</td>
<td id="76EE5DA8FFCD005388A303CA7D2C15BE" box="[1219,1316,956,979]" gridcol="4" gridrow="18" pageId="9" pageNumber="373">0.95–1.04</td>
<td id="76EE5DA8FFCD0053892B03CA7DA215BE" box="[1355,1450,956,979]" gridcol="5" gridrow="18" pageId="9" pageNumber="373">2.8–3.1</td>
</tr>
<tr id="353F34D4FFCD00538CF7039D7DA2126C" box="[151,1450,1003,1025]" gridrow="19" pageId="9" pageNumber="373" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76EE5DA8FFCD00538CF7039D7ADA126C" box="[151,722,1003,1025]" gridcol="0" gridrow="19" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58CF7039D79B7126C" bold="true" box="[151,447,1003,1025]" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF7039D7931126C" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[151,313,1003,1025]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFCDFFA58CF7039D7931126C" bold="true" box="[151,313,1003,1025]" italics="true" pageId="9" pageNumber="373">Pincerna clausa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24857FFFFCDFFA58D27039D79B7126C" box="[327,447,1003,1025]" pageId="9" pageNumber="373" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="353F34D4FFCD00538CF7047D7DA2124F" box="[151,1450,1035,1058]" gridrow="20" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7047D7ADA124F" box="[151,722,1035,1058]" gridcol="0" gridrow="20" pageId="9" pageNumber="373">15. Na Ư, Điện Biên, Điện Biên, Vietnam</th>
<td id="76EE5DA8FFCD00538E94047D7B54124F" box="[756,860,1035,1058]" gridcol="1" gridrow="20" pageId="9" pageNumber="373">4</td>
<td id="76EE5DA8FFCD00538F1B047D7BF5124F" box="[891,1021,1035,1058]" gridcol="2" gridrow="20" pageId="9" pageNumber="373">5.1–5.3</td>
<td id="76EE5DA8FFCD0053887E047D7CAD124F" box="[1054,1189,1035,1058]" gridcol="3" gridrow="20" pageId="9" pageNumber="373">4.9–5.0</td>
<td id="76EE5DA8FFCD005388A3047D7D2C124F" box="[1219,1316,1035,1058]" gridcol="4" gridrow="20" pageId="9" pageNumber="373">1.04–1.06</td>
<td id="76EE5DA8FFCD0053892B047D7DA2124F" box="[1355,1450,1035,1058]" gridcol="5" gridrow="20" pageId="9" pageNumber="373">3.0–3.1</td>
</tr>
</table>
</paragraph>
<caption id="DF70661EFFCDFFA58CE804197C5912E8" box="[136,1105,1135,1157]" pageId="9" pageNumber="373" startId="9.[136,191,1135,1157]">
<paragraph id="8BB03696FFCDFFA58CE804197C5912E8" blockId="9.[136,1105,1135,1157]" box="[136,1105,1135,1157]" pageId="9" pageNumber="373">
Table 2. Comparison of diagnostic conchological characters among
<taxonomicName id="4C0F4D15FFCDFFA58F4404067BC112E8" box="[804,969,1135,1157]" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58F4404067B7412E8" box="[804,892,1136,1157]" italics="true" pageId="9" pageNumber="373">Pincerna</emphasis>
species
</taxonomicName>
in this study.
</paragraph>
</caption>
<paragraph id="8BB03696FFCDFFA58CF704B07D7313B0" pageId="9" pageNumber="373">
<table id="F90FC436FFCD00538CF704C17DB813B0" box="[151,1456,1207,1501]" gridcols="6" gridrows="6" pageId="9" pageNumber="373">
<tr id="353F34D4FFCD00538CF704C17DB81287" box="[151,1456,1207,1258]" gridrow="0" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF704C179161287" box="[151,286,1207,1258]" gridcol="0" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58CF704B078EA12B1" bold="true" box="[151,226,1222,1244]" pageId="9" pageNumber="373">Species</emphasis>
</th>
<th id="76EE5DA8FFCD00538D1604C17A121287" box="[374,538,1207,1258]" gridcol="1" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58D1604B07A1212B6" bold="true" box="[374,538,1222,1244]" pageId="9" pageNumber="373">Sculpture of R1</emphasis>
</th>
<th id="76EE5DA8FFCD00538E0004C17AFC1287" box="[608,756,1207,1258]" gridcol="2" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58E0004B07AFC12B1" bold="true" box="[608,756,1222,1244]" pageId="9" pageNumber="373">No. of R2 ribs</emphasis>
</th>
<th id="76EE5DA8FFCD00538F5B04C17BEB1287" box="[827,995,1207,1258]" gridcol="3" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58F3A04C17BEB1284" bold="true" pageId="9" pageNumber="373">R2 length compared to R3</emphasis>
</th>
<th id="76EE5DA8FFCD0053885B04C17CAB1287" box="[1083,1187,1207,1258]" gridcol="4" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA5885B04B07CAB12B1" bold="true" box="[1083,1187,1222,1244]" pageId="9" pageNumber="373">Umbilicus</emphasis>
</th>
<th id="76EE5DA8FFCD0053889204C17DB81287" box="[1266,1456,1207,1258]" gridcol="5" gridrow="0" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA5889204B07DB812B6" bold="true" box="[1266,1456,1222,1244]" pageId="9" pageNumber="373">Number of whorls</emphasis>
</th>
</tr>
<tr id="353F34D4FFCD00538CF7057A7DB8134F" box="[151,1456,1292,1314]" gridrow="1" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7057A7916134F" box="[151,286,1292,1314]" gridcol="0" gridrow="1" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF7057A78FA134C" baseAuthorityName="Mabille" baseAuthorityYear="1887" box="[151,242,1292,1313]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFCDFFA58CF7057A78FA134C" box="[151,242,1292,1313]" italics="true" pageId="9" pageNumber="373">P. anceyi</emphasis>
</taxonomicName>
</th>
<td id="76EE5DA8FFCD00538D16057A7A12134F" box="[374,538,1292,1314]" gridcol="1" gridrow="1" pageId="9" pageNumber="373">weakly ribbed</td>
<td id="76EE5DA8FFCD00538E00057A7AFC134F" box="[608,756,1292,1314]" gridcol="2" gridrow="1" pageId="9" pageNumber="373">26</td>
<td id="76EE5DA8FFCD00538F5B057A7BEB134F" box="[827,995,1292,1314]" gridcol="3" gridrow="1" pageId="9" pageNumber="373">≤ R3</td>
<td id="76EE5DA8FFCD0053885B057A7CAB134F" box="[1083,1187,1292,1314]" gridcol="4" gridrow="1" pageId="9" pageNumber="373">narrow</td>
<td id="76EE5DA8FFCD00538892057A7DB8134F" box="[1266,1456,1292,1314]" gridcol="5" gridrow="1" pageId="9" pageNumber="373">4.5</td>
</tr>
<tr id="353F34D4FFCD00538CF7054C7DB8133C" box="[151,1456,1338,1361]" gridrow="2" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7054C7916133C" box="[151,286,1338,1361]" gridcol="0" gridrow="2" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF7054D791F133D" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[151,279,1339,1360]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFCDFFA58CF7054D791F133D" box="[151,279,1339,1360]" italics="true" pageId="9" pageNumber="373">P. costulosus</emphasis>
</taxonomicName>
</th>
<td id="76EE5DA8FFCD00538D16054C7A12133C" box="[374,538,1338,1361]" gridcol="1" gridrow="2" pageId="9" pageNumber="373">strongly ribbed</td>
<td id="76EE5DA8FFCD00538E00054C7AFC133C" box="[608,756,1338,1361]" gridcol="2" gridrow="2" pageId="9" pageNumber="373">14–20</td>
<td id="76EE5DA8FFCD00538F5B054C7BEB133C" box="[827,995,1338,1361]" gridcol="3" gridrow="2" pageId="9" pageNumber="373">≤ R3</td>
<td id="76EE5DA8FFCD0053885B054C7CAB133C" box="[1083,1187,1338,1361]" gridcol="4" gridrow="2" pageId="9" pageNumber="373">narrow</td>
<td id="76EE5DA8FFCD00538892054C7DB8133C" box="[1266,1456,1338,1361]" gridcol="5" gridrow="2" pageId="9" pageNumber="373">4.5–5.0</td>
</tr>
<tr id="353F34D4FFCD00538CF7051F7DB81312" box="[151,1456,1385,1407]" gridrow="3" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF7051F79161312" box="[151,286,1385,1407]" gridcol="0" gridrow="3" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF7051C790A1313" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[151,258,1385,1407]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFCDFFA58CF7051C790A1313" box="[151,258,1385,1407]" italics="true" pageId="9" pageNumber="373">P. mouhoti</emphasis>
</taxonomicName>
</th>
<td id="76EE5DA8FFCD00538D16051F7A121312" box="[374,538,1385,1407]" gridcol="1" gridrow="3" pageId="9" pageNumber="373">strongly ribbed</td>
<td id="76EE5DA8FFCD00538E00051F7AFC1312" box="[608,756,1385,1407]" gridcol="2" gridrow="3" pageId="9" pageNumber="373">35–76</td>
<td id="76EE5DA8FFCD00538F5B051F7BEB1312" box="[827,995,1385,1407]" gridcol="3" gridrow="3" pageId="9" pageNumber="373">≥ R3</td>
<td id="76EE5DA8FFCD0053885B051F7CAB1312" box="[1083,1187,1385,1407]" gridcol="4" gridrow="3" pageId="9" pageNumber="373">wide</td>
<td id="76EE5DA8FFCD00538892051F7DB81312" box="[1266,1456,1385,1407]" gridcol="5" gridrow="3" pageId="9" pageNumber="373">4.5</td>
</tr>
<tr id="353F34D4FFCD00538CF705EE7DB813C3" box="[151,1456,1432,1454]" gridrow="4" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF705EE791613C3" box="[151,286,1432,1454]" gridcol="0" gridrow="4" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF705EF791613C0" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[151,286,1432,1454]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFCDFFA58CF705EF791613C0" box="[151,286,1432,1454]" italics="true" pageId="9" pageNumber="373">P. vanbuensis</emphasis>
</taxonomicName>
</th>
<td id="76EE5DA8FFCD00538D1605EE7A1213C3" box="[374,538,1432,1454]" gridcol="1" gridrow="4" pageId="9" pageNumber="373">strongly ribbed</td>
<td id="76EE5DA8FFCD00538E0005EE7AFC13C3" box="[608,756,1432,1454]" gridcol="2" gridrow="4" pageId="9" pageNumber="373">65–71</td>
<td id="76EE5DA8FFCD00538F5B05EE7BEB13C3" box="[827,995,1432,1454]" gridcol="3" gridrow="4" pageId="9" pageNumber="373">~ R3</td>
<td id="76EE5DA8FFCD0053885B05EE7CAB13C3" box="[1083,1187,1432,1454]" gridcol="4" gridrow="4" pageId="9" pageNumber="373">narrow</td>
<td id="76EE5DA8FFCD0053889205EE7DB813C3" box="[1266,1456,1432,1454]" gridcol="5" gridrow="4" pageId="9" pageNumber="373">4.25–4.5</td>
</tr>
<tr id="353F34D4FFCD00538CF705B17DB813B0" box="[151,1456,1479,1501]" gridrow="5" pageId="9" pageNumber="373">
<th id="76EE5DA8FFCD00538CF705B1791613B0" box="[151,286,1479,1501]" gridcol="0" gridrow="5" pageId="9" pageNumber="373">
<taxonomicName id="4C0F4D15FFCDFFA58CF705BE78FA13B1" authorityName="Sang &amp; Son" authorityYear="2022" box="[151,242,1479,1501]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="clausa">
<emphasis id="B97BEA84FFCDFFA58CF705BE78FA13B1" box="[151,242,1479,1501]" italics="true" pageId="9" pageNumber="373">P. clausa</emphasis>
</taxonomicName>
</th>
<td id="76EE5DA8FFCD00538D1605B17A1213B0" box="[374,538,1479,1501]" gridcol="1" gridrow="5" pageId="9" pageNumber="373">weakly ribbed</td>
<td id="76EE5DA8FFCD00538E0005B17AFC13B0" box="[608,756,1479,1501]" gridcol="2" gridrow="5" pageId="9" pageNumber="373">34–41</td>
<td id="76EE5DA8FFCD00538F5B05B17BEB13B0" box="[827,995,1479,1501]" gridcol="3" gridrow="5" pageId="9" pageNumber="373">≤ R3</td>
<td id="76EE5DA8FFCD0053885B05B17CAB13B0" box="[1083,1187,1479,1501]" gridcol="4" gridrow="5" pageId="9" pageNumber="373">closed</td>
<td id="76EE5DA8FFCD0053889205B17DB813B0" box="[1266,1456,1479,1501]" gridcol="5" gridrow="5" pageId="9" pageNumber="373">4.25–4.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB03696FFCDFFA58CE807FA798811C9" blockId="9.[136,384,1932,1956]" box="[136,384,1932,1956]" pageId="9" pageNumber="373">Operculum. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCDFFA58CE807BA7A951E49" pageId="9" pageNumber="373" type="distribution">
<paragraph id="8BB03696FFCDFFA58CE807BA7A951E49" blockId="9.[136,775,1996,2084]" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58CE807BA79121189" bold="true" box="[136,282,1996,2020]" pageId="9" pageNumber="373">Distribution.</emphasis>
The new species is thus far known only from the limestone hill in Na Ư Commune, Điện Biên District, Điện Biên Province, northwest
<collectingCountry id="F3187606FFCDFFA58D80087A7A341E49" box="[480,572,2060,2084]" name="Vietnam" pageId="9" pageNumber="373">Vietnam</collectingCountry>
(
<figureCitation id="13342A13FFCDFFA58E2C087A7A851E49" box="[588,653,2060,2084]" captionStart="Fig" captionStartId="2.[113,148,1941,1963]" captionTargetBox="[113,1452,311,1926]" captionTargetId="figure-11@2.[113,1452,311,1926]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Pincerna species in Vietnam and Laos. Triangle: Pincerna anceyi; Grey circle: Pincerna costulosus; Black circles: Pincerna mouhoti; Squares: Pincerna vanbuensis; Star: Pincerna clausa, new species. The localities are detailed in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="9" pageNumber="373">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFCDFFA58F23065D7CF4110E" pageId="9" pageNumber="373" type="discussion">
<paragraph id="8BB03696FFCDFFA58F23065D7CF4110E" blockId="9.[835,1475,1579,1891]" pageId="9" pageNumber="373">
<emphasis id="B97BEA84FFCDFFA58F23065D7BBC102E" bold="true" box="[835,948,1579,1603]" pageId="9" pageNumber="373">Remarks.</emphasis>
This new species has a similar shell shape as
<taxonomicName id="4C0F4D15FFCDFFA58F24063A7C16100E" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[836,1054,1611,1635]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFCDFFA58F24063A7C16100E" box="[836,1054,1611,1635]" italics="true" pageId="9" pageNumber="373">Pincerna vanbuensis</emphasis>
</taxonomicName>
but differs in having a closed umbilicus, stronger spiral striation on R1, and weaker ribs on R2.
<taxonomicName id="4C0F4D15FFCDFFA58F2406FA7BF510CE" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[836,1021,1675,1699]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFCDFFA58F2406FA7BF510CE" box="[836,1021,1675,1699]" italics="true" pageId="9" pageNumber="373">Pincerna clausa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24857FFFFCDFFA5886F06FA7C9310CE" box="[1039,1179,1675,1699]" pageId="9" pageNumber="373" rank="species">new species</taxonomicNameLabel>
, also resembles
<taxonomicName id="4C0F4D15FFCDFFA5893D06FA7BBA10AE" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="costulusus">
<emphasis id="B97BEA84FFCDFFA5893D06FA7BBA10AE" italics="true" pageId="9" pageNumber="373">Pincerna costulusus</emphasis>
</taxonomicName>
in shell shape but differs in its more pronounced spiral striation as well as R1 with denser and weaker ribs. It differs from
<taxonomicName id="4C0F4D15FFCDFFA58F8B069A7C95116E" baseAuthorityName="Mabille" baseAuthorityYear="1887" box="[1003,1181,1772,1795]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFCDFFA58F8B069A7C95116E" box="[1003,1181,1772,1795]" italics="true" pageId="9" pageNumber="373">Pincerna anceyi</emphasis>
</taxonomicName>
by having much stronger, more widely-spaced ribs, well visible spiral striation and closed umbilicus. It differs from
<taxonomicName id="4C0F4D15FFCDFFA588D9075A7D3B112E" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[1209,1331,1835,1859]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="9" pageNumber="373" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFCDFFA588D9075A7D3B112E" box="[1209,1331,1835,1859]" italics="true" pageId="9" pageNumber="373">P. mouhoti</emphasis>
</taxonomicName>
by having a narrow umbilicus and smaller shell size.
</paragraph>
</subSubSection>
</treatment>
</document>
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