treatments-xml/data/26/CB/A4/26CBA40789005DD7BEC91DD96238EE9E.xml

<document id="BD45B48D9EC7C9A3BF11B9BD63AE5B4F" ID-DOI="10.3897/zookeys.1209.125897" ID-publisher-id="125897" URI-arpha="6ED9C06E-4DBB-5A79-A98B-FE427E12ED58" URI-zoobank="http://zoobank.org/CEE78803-61EB-40CC-8D63-46142E6383A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1723117912200" checkinUser="pensoft" docAuthor="Harden, Curt W. &amp; Caterino, Michael S." docDate="2024" docId="26CBA40789005DD7BEC91DD96238EE9E" docLanguage="en" docName="ZooKeys 1209: 69-197" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/125897/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Anillinus seneca Harden &amp; Caterino, 2024, sp. nov." docType="treatment" docUuid="3ECB8AD7-7F13-46A2-8500-F72D6A9C2B32" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="197" masterDocId="6ED9C06E4DBB5A79A98BFE427E12ED58" masterDocTitle="Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" masterLastPageNumber="197" masterPageNumber="69" pageNumber="69" updateTime="1723146111653" updateUser="admin">
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<mods:title id="AF77FABEC3FE6963D5B11BD96A389456">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
</mods:titleInfo>
<mods:name id="43F57CF56BC1844E2390C62A9F2F2613" type="personal">
<mods:role id="2F6FE43CE6480C50A58FBF9302F1D5E0">
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<mods:namePart id="9A8387120AC00616D1E688D61BF179C0">Harden, Curt W.</mods:namePart>
<mods:nameIdentifier id="4BD7C75EAC0777C9FF8B85193024C7E9" type="ORCID">0000-0002-5226-8725</mods:nameIdentifier>
<mods:affiliation id="8BEC2A11CD9226C13CD5890B19A2CE08">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
</mods:name>
<mods:name id="19E043B162E2AA2E1F44B7EB4B6CED1E" type="personal">
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</mods:role>
<mods:namePart id="894E2DA9DD82EB9679230FCB58FBD9EB">Caterino, Michael S.</mods:namePart>
<mods:nameIdentifier id="67FD544E0D7451AC56DAC5A6E5D62E48" type="ORCID">0000-0002-2597-5707</mods:nameIdentifier>
<mods:affiliation id="BD974202D5C30F562B900FDA50E14F09">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
</mods:name>
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<mods:title id="DCA3F9171E9351318716F835F48D17AA">ZooKeys</mods:title>
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<mods:part id="2EC74A5A9EF678623AE4EA7B422A3F40">
<mods:date id="B5F807637C89AF2A0FDDE1F1A8CC6E59">2024</mods:date>
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<mods:number id="72FB9E782EB2AEB8EF4FD89ABE70AB63">2024-08-08</mods:number>
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<mods:number id="59D680BF1059824EE1FA0C586AA0F6C2">1209</mods:number>
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<mods:classification id="BD682D243E882594B4647A1DE0576AD9">journal article</mods:classification>
<mods:identifier id="AA93A263B376321EEB8EAB843A487F09" type="DOI">10.3897/zookeys.1209.125897</mods:identifier>
<mods:identifier id="4FB72FD8CE79BF214163DE93A1CB9961" type="ZooBank">CEE78803-61EB-40CC-8D63-46142E6383A1</mods:identifier>
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<treatment id="26CBA40789005DD7BEC91DD96238EE9E" ID-DOI="http://doi.org/10.5281/zenodo.13270403" ID-Zenodo-Dep="13270403" ID-arpha="26CBA407-8900-5DD7-BEC9-1DD96238EE9E" ID-zoobank="https://zoobank.org/3ECB8AD7-7F13-46A2-8500-F72D6A9C2B32" LSID="urn:lsid:zoobank.org:act:3ECB8AD7-7F13-46A2-8500-F72D6A9C2B32" LSID-ZBK="urn:lsid:zoobank.org:act:3ECB8AD7-7F13-46A2-8500-F72D6A9C2B32" httpUri="http://treatment.plazi.org/id/26CBA40789005DD7BEC91DD96238EE9E">
<subSubSection id="C285A01BCD24606534A8852D0ADCC192" type="nomenclature">
<paragraph id="9904ACC2E2670D6A08D36D669131B0D3">
<taxonomicName id="A13943ADF89C67ED5D5325AB9DA7C1CB" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seneca" status="sp. nov.">
<emphasis id="604F030D228D5D6A6A33D56D2AC8DEA6" italics="true">Anillinus seneca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4E815C2E853D984386A2A1BB70AB76ED" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="5398F51E9F5CD1B0D10369C347B00298" type="description">
<paragraph id="B77DC0FC065A254ED824213073873C88">
<figureCitation id="2073460761EF5F5E4A2E4895A766B461" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. SEM micrographs of male left metafemora of Anillinus species. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure12" httpUri="https://binary.pensoft.net/fig/1110351">Figs 12 D</figureCitation>
, 
<figureCitation id="2FCE7BFCA5CE5D1E1AA2AD68D025B514" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 K</figureCitation>
, 
<figureCitation id="F633EE71603A3F61AC2F04BAD805C4B4" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1209.125897.figure24" httpUri="https://binary.pensoft.net/fig/1110363">24 J</figureCitation>
, 
<figureCitation id="C023CE36338B2CA744DB96219F2BFC8E" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure34" httpUri="https://binary.pensoft.net/fig/1110373">34 D</figureCitation>
, 
<figureCitation id="B31C1A94628425E57986932AFC663D5B" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure35" httpUri="https://binary.pensoft.net/fig/1110374">35 D – F</figureCitation>
, 
<figureCitation id="D13974DFEA967839187FFA6A698FC528" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." figureDoi="10.3897/zookeys.1209.125897.figure36" httpUri="https://binary.pensoft.net/fig/1110375">36</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3SFM" type="type material">
<paragraph id="FBD2A6E21576CDEF9D3E9E288AC30B90">
<heading id="E2F6FBF5442458800DC3B098B39DF252" reason="title">Type material.</heading>
</paragraph>
<paragraph id="5B8D9C82ED682A95212FACE89017AB60">
<materialsCitation id="140D0D9E7FB8E811AB100214DDD60E3B" collectedFrom="Under embedded rocks in oak-hickory woods near lake" collectingDate="2020-01-11" collectingDateMax="2020-05-13" collectingDateMin="2020-01-11" collectionCode="USNM" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6389" location="Lake Hartwell, Martins Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Euparal" specimenCode="CUAC 000168244" specimenCount="1" stateProvince="SOUTH CAROLINA" typeStatus="holotype">
<emphasis id="7F365AD94DEBED4C3152C1D0F5B8942B" bold="true">
<emphasis id="57AE874CAF5ECA35A1DFE27362046E0B" bold="true" italics="true">
<typeStatus id="66606EB2EE3F61679FC41B8B5324761E" type="holotype">Holotype</typeStatus>
male
</emphasis>
(
<named-content id="4ABF6516B971FAE0DADA938EBA1A75C6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">
<collectionCode id="DACA126C715BF65AF17B020EB7D58A16" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</named-content>
)
</emphasis>
: point mounted, with abdominal ventrites glued to point and genitalia in Euparal on microslide pinned beneath specimen. Original labels: “ 
<collectingCountry id="637D31B40DBC13019324FA9E8DE1FEB8" name="United States of America">USA</collectingCountry>
: 
<collectingRegion id="F0A8B23CF8FD9BEE6CF3B7D405306EF3" country="United States of America" name="South Carolina">SOUTH CAROLINA</collectingRegion>
, 
<collectingCounty id="255F094237FC99549A02600C0C706645">Oconee Co.</collectingCounty>
<location id="F8AFF8DCE9602605A096729FB18EDBB3" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:F8AFF8DCE9602605A096729FB18EDBB3" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Euparal" name="Lake Hartwell, Martins Creek Landing" stateProvince="SOUTH CAROLINA">Lake Hartwell, Martins Creek Landing</location>
, 
<locationDeviation id="16B1AFDA3B25667BEA62BF7B7E27AA75">SW of Clemson</locationDeviation>
. 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOTFM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="F8C4FB479EA711B10FC56C43BA0B035C" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="5DB0DFA00F4A0B1C606A721E87F94A15" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
. 
<date id="7D709ED007FCE25527E6CDCDB0864FD2" value="2020-01-11">
<collectingDate id="5AB8E79A416C4C913DF30ACA9DBF3897" value="2020-01-11">11 - January- 2020</collectingDate>
</date>
. 
<collectorName id="9BFA61BAC0A54B280AE1A22F1469B80C">C. W. Harden</collectorName>
. 
<collectedFrom id="070ED8B1AB1AEE68836ADFA8DD46D01D">Under embedded rocks in oak-hickory woods near lake</collectedFrom>
. ” “ [QR Code] CLEMSON-ENT 
<specimenCode id="B874C00D0EA68B0787B9C1D0650B7C74" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168244</specimenCode>
” “ Harden DNA Voucher CWH- 121 Anill. sp n ‘ martincrk’ Ext. 
<date id="14ECCA4A35E3BBBB0DBFF0867A4A3D3E" value="2020-05-13">
<collectingDate id="1DDD3890B1527A9E4C46E5DDC8039D84" value="2020-05-13">13 / May / 2020</collectingDate>
</date>
[green-bordered cardstock] ” “ HOLOTYPE 
<taxonomicName id="C63B27FBA95FC0C744CBE40632FEDD1A" authority="Harden &amp; Caterino" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seneca">
<originalDetermination id="212C747CD11C7D3F5FAB2875FF207866">
<emphasis id="B4828C75F8D29A12332961BB5D254F0E" italics="true">Anillinus seneca</emphasis>
Harden &amp; Caterino
</originalDetermination>
</taxonomicName>
[orange cardstock] ”
</materialsCitation>
</paragraph>
<paragraph id="9EB199F9831AABE0786EFE9BB6278B88">
GenBank: 
<ext-link id="69EE81501759B8CD633291E3EFD45A24" ext-link-type="gen" xlink_href="OR853323" xlink_type="simple">OR 853323</ext-link>
, 
<ext-link id="175A65C9CEB505386DAB983921DDE022" ext-link-type="gen" xlink_href="OR839466" xlink_type="simple">OR 839466</ext-link>
.
</paragraph>
<paragraph id="42309C226F9F35AB4F5E0C78FB91CE9C">
<materialsCitation id="BB42C3B70356505D90D6D33FEE35DFB1" collectedFrom="Under embedded rocks in oak-hickory woods near lake" collectingDate="2020-01-11" collectingDateMax="2020-05-13" collectingDateMin="2020-01-11" collectionCode="CUAC" collectorName="C. W. Harden. Under" country="United States of America" county="Oconee Co." latitude="34.6389" location="South Carolina" longLatPrecision="6" longitude="-82.8656" municipality="Euparal" specimenCode="CUAC 000168247, CUAC 000168248, CUAC 000168250, CUAC 000168242" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="South Carolina" typeStatus="paratype">
<emphasis id="EF6115F370B64FE6217158CC17BA00C1" bold="true" italics="true">
<typeStatus id="DBF39C8AF795E9FE8718577ABBE36280" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="5239557EABAD96C6E2C8E4F97CDC573A" italics="true">n</emphasis>
= 27; 
<named-content id="F0284E68224AD46A71C434F7944574A9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">CMNH</named-content>
, 
<named-content id="8DEDBDC5B004C325915B185FC02753BA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">CUAC</named-content>
, 
<named-content id="42DBBAC02D16A1F7DFD1B6A40EAE4917" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/louisiana-state-arthropod-museum" xlink_title="Louisiana State Arthropod Museum">LSAM</named-content>
, 
<named-content id="9EAF5D442B249CA4911B04A3C198648C" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">USNM</named-content>
, 
<named-content id="60026FAEFD56424F33415C30894B537B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/virginia-museum-natural-history" xlink_title="Virginia Museum of Natural History">VMNH</named-content>
). 
<emphasis id="B3AE7B653FBEAA901DD45C637C1DE9DE" bold="true">
<collectingCountry id="7F0F01FDF08420C6DA6E79472D8272AC" name="United States of America">USA</collectingCountry>
</emphasis>
• 
<emphasis id="18B291F5A77867A7F4EF13413F2178BE" bold="true">
<collectingRegion id="1D84AFA5AE843DFE2B5953387631138E" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="D08A8D7B1EAF8F8F425BE2382F8BC713" count="2" type="male">2 ♂</specimenCount>
, 
<specimenCount id="A3690606E0F599293836F9A284768E47" count="2" type="female">2 ♀</specimenCount>
; same data as holotype; 
<specimenCode id="DCCAF63118B9A338954C66974BEF0011" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168247</specimenCode>
, 
<specimenCode id="3B1954D3C229DFA45A67A6D589605B70" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168248</specimenCode>
, 
<specimenCode id="32E903CC75727532D0C36C157B01A541" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168250</specimenCode>
, 
<specimenCode id="E09B1A66CD1E596F33E4880F8BC54B95" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168242</specimenCode>
; CWH- 118, CWH- 120, CWH- 122, CWH- 124; 
<named-content id="6073094611801C705EA708255D7E2EC2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="B50AC87465E5EA28BF201975185E3F21" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="48E04FCD33908FF5A168F915887F959C" collectedFrom="Under embedded rocks in oak-hickory woods near lake" collectingDate="2020-01-04" collectingDateMax="2020-05-13" collectingDateMin="2020-01-11" collectingMethod="soilwash flotation Berlese" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6389" location="Original" longLatPrecision="6" longitude="-82.8656" municipality="Euparal" specimenCode="CUAC 000168239, CUAC 000169241" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="8CE038B85EFC879DA1CFDECEB8EED549" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="CC4664F0FAE8F6B53B15269A38C834B7" count="2" type="female">2 ♀</specimenCount>
; same locality as holotype; 
<date id="8111E24DB95D3CECEED4CEA7F2BB551A" value="2020-01-04">
<collectingDate id="542FD43780D634C7373343DB795C5B7C" value="2020-01-04">4 Jan. 2020</collectingDate>
</date>
; 
<collectorName id="9909361C943659C08FA3724D2E7EB8B8">C. W. Harden</collectorName>
&amp; 
<collectorName id="E835A5EAD9F91496B460C1D1B7CDC889">L. M. Thompson</collectorName>
leg.; 
<collectingMethod id="18837C4ADD7425E8781BF45C4931F0A2">soilwash flotation Berlese</collectingMethod>
; CWH- 115 to CWH- 117, 
<specimenCode id="2B32F315F8445062E86546959A09B472" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168239</specimenCode>
to 
<specimenCode id="FAA47C9E5AC8969F434A929BD6C4D546" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000169241</specimenCode>
; 
<named-content id="4228BA365915DC71F6853C2EA5BFEFDE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="23F375E2EA4303FE9574EA1048F48A52" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="6C67ED7AC9FD544411782076A56D39CC" collectingDate="2022-05-15" collectingMethod="soilwashing" collectionCode="CMNH, USNM" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6388" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8655" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="71B8E9B3528A0C724DF6A311BF4B5253" count="2" type="male">2 ♂</specimenCount>
, 
<specimenCount id="93C4315A5A621ADC5E85D776DCA964CD" count="5" type="female">5 ♀</specimenCount>
; 
<collectingCounty id="D85D54413F95582E82DA749F94129E23">
<emphasis id="5CF4F33D5F64F93CC99328BFD74944C5" bold="true">Oconee Co</emphasis>
.
</collectingCounty>
; 
<location id="FD83BC96B6AD7DA705EB9832160FC603" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:FD83BC96B6AD7DA705EB9832160FC603" country="United States of America" county="Oconee Co." latitude="34.6388" longLatPrecision="6" longitude="-82.8655" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYVFM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865500,34.638800]}">
<geoCoordinate id="A4737C289DE297FC96FE60094D242B60" degrees="34.6388" direction="north" orientation="latitude" precision="5" value="34.6388">34.6388</geoCoordinate>
, 
<geoCoordinate id="0978ACE3BDBFD7B84D27B86478E54F6E" degrees="82.8655" direction="west" orientation="longitude" precision="5" value="-82.8655">- 82.8655</geoCoordinate>
</named-content>
; 
<date id="5483B15A24C1CBAB6D7552A0CF0C7EFC" value="2022-05-15">
<collectingDate id="4A4B64CD87A8E93B01345701BDE0EA1B" value="2022-05-15">15 May 2022</collectingDate>
</date>
; 
<collectorName id="EC109F87A5E8FE133D230CFBAFF22A4D">C. W. Harden</collectorName>
and 
<collectorName id="50EE6E904973B69E7B6078EE923A9CB7">L. M. Thompson</collectorName>
leg.; 
<collectingMethod id="9F3CB9A551A31D7EA51DF09ED92D01B5">soilwashing</collectingMethod>
; 
<named-content id="4D696347EB04528786031F1DAA6B57CC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="B927A9BC31CA3B3D14BF0B1F94219EFD" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
, 
<named-content id="0B8814F09B5FCEB8791B7714C246F610" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">
<collectionCode id="2131CFEB8874B8469FFED4BF57E5167A" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="B2B1D7FFAE6A426E54EDCF2E38571EB2" collectedFrom="under rock" collectingDate="2021-10-22" collectionCode="LSAM, VMNH" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.639" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8638" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="85FB08365D86EE3F3D1BBF6A91509B82" count="3" type="male">3 ♂</specimenCount>
, 
<specimenCount id="F4F364C10B48E66745BD3E3B0B89E903" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="7D2530BAA0BEFF4AC95B8A653BF0F012" bold="true">
<collectingCounty id="93551EECA2AF76F022427F4CDF81C3F3">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="F4B67D4C9E72CB3E78A96492DE7D520A" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:F4B67D4C9E72CB3E78A96492DE7D520A" country="United States of America" county="Oconee Co." latitude="34.639" longLatPrecision="6" longitude="-82.8638" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKWFM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.863800,34.639000]}">
<geoCoordinate id="8165CBB49ACEFFB0B7F341E663C6783B" degrees="34.6390" direction="north" orientation="latitude" precision="5" value="34.639">34.6390</geoCoordinate>
, 
<geoCoordinate id="76625E31F86B333C2D7739B569052DE7" degrees="82.8638" direction="west" orientation="longitude" precision="5" value="-82.8638">- 82.8638</geoCoordinate>
</named-content>
; 
<date id="1915DE2DDCB0C532CFD7297B608F72C7" value="2021-10-22">
<collectingDate id="6C672DC9D4A79CC0111DFFAE549B3365" value="2021-10-22">22 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="C2BD2087239D696A5CBD987EB4BC3FAE">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="0581B239F823DB841B6A19EB162B2AB7">under rock</collectedFrom>
; 
<named-content id="2E02ECA2B8F7B444CDF490CDC4E03795" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/louisiana-state-arthropod-museum" xlink_title="Louisiana State Arthropod Museum">
<collectionCode id="034364E0555B58C4EE71A424E5119DF2" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" type="">LSAM</collectionCode>
</named-content>
, 
<named-content id="6AF8B9A3F157E50539156085CD8AB558" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/virginia-museum-natural-history" xlink_title="Virginia Museum of Natural History">
<collectionCode id="9F12258B080D75918D341788EEC04FA8" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" type="">VMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="F733F68B037D710C26ED203A03184CAD" collectedFrom="wood, pine stump near fishing pier" collectingDate="2020-01-30" collectingMethod="Berlese, sifted soil" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.71165" location="South Cove Park" longLatPrecision="1" longitude="-82.96594" specimenCode="CUAC 000168255, CUAC 000168262" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="9FE4ACD5384353C4FD348786B12E862B" count="4" type="male">4 ♂</specimenCount>
, 
<specimenCount id="1E7DD3023FBA4866DF8296E0F8625018" count="4" type="female">4 ♀</specimenCount>
; 
<emphasis id="D9803A5372F3E3DC7F1CD25042D2DCD7" bold="true">
<collectingCounty id="62BC5A50E53DDF7C32FEC8B7B5D95257">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="547F1AF76F9F7272EB73CA01AB88602D" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:547F1AF76F9F7272EB73CA01AB88602D" country="United States of America" county="Oconee Co." latitude="34.71165" longLatPrecision="1" longitude="-82.96594" name="South Cove Park" stateProvince="South Carolina">South Cove Park</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3WFM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.965940,34.711650]}">
<geoCoordinate id="DB28CA81869EFAA4C815844C4D8DE1B5" degrees="34.71165" direction="north" orientation="latitude" precision="1" value="34.71165">34.71165</geoCoordinate>
, 
<geoCoordinate id="9147A10377F4BD1E009570C3B09F92EF" degrees="82.96594" direction="west" orientation="longitude" precision="1" value="-82.96594">- 82.96594</geoCoordinate>
</named-content>
; 
<date id="C58A79363CBC8028991EF2D1B7D9674F" value="2020-01-30">
<collectingDate id="F28C6AF83D5CF59F91E34DB31B9AB614" value="2020-01-30">30 Jan. 2020</collectingDate>
</date>
; 
<collectorName id="4E7E933BFDBB4FE6F21B1DDE4F7E6EA3">C. W. Harden</collectorName>
&amp; 
<collectorName id="3D3D05986B4DEB765D6012087F111609">L. M. Thompson</collectorName>
leg.; 
<collectingMethod id="8845C644D8AA12A933907D1B083F904A">Berlese, sifted soil</collectingMethod>
/ 
<collectedFrom id="060AA805583802F3F5A3BF00755BD8E2">wood, pine stump near fishing pier</collectedFrom>
; CWH- 338 to CWH- 345, 
<specimenCode id="F4B44DA0760B568AEFBE52C44E30033D" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168255</specimenCode>
to 
<specimenCode id="8D8370354A0E7ADDBCB34D12B2E6615D" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168262</specimenCode>
; 
<named-content id="200F0078B7656306EED94AF866440CB6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="D91AF9C37224A4719731DE9A9C4054E8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="37958FCE04C39E4D77C43D1098EA0991">
GenBank accession numbers for 
<typeStatus id="963CD6C67AEC68829EBA213A7E138313" type="paratype">paratypes</typeStatus>
: 
<ext-link id="5B924D2CF16095BCAC9274B9F50E2175" ext-link-type="gen" xlink_href="OR839463" xlink_type="simple">OR 839463</ext-link>
, 
<ext-link id="568CF48878B9EA651C37EFC1F71FEDDB" ext-link-type="gen" xlink_href="OR839465" xlink_type="simple">OR 839465</ext-link>
, 
<ext-link id="CF1AF3D480801DDEBBCD8120D87FB276" ext-link-type="gen" xlink_href="OR839467" xlink_type="simple">OR 839467</ext-link>
, 
<ext-link id="33EA77FAEED9FA30489B18577DB39B4F" ext-link-type="gen" xlink_href="OR853324" xlink_type="simple">OR 853324</ext-link>
, 
<ext-link id="E290D26B7DDFB2945DC30E9465CD5A02" ext-link-type="gen" xlink_href="OR839469" xlink_type="simple">OR 839469</ext-link>
, 
<ext-link id="333A5DF389347E42027E1BF5FE2CEEBB" ext-link-type="gen" xlink_href="OR839460" xlink_type="simple">OR 839460</ext-link>
, 
<ext-link id="1FD09395C95597E5DB2F6B70EB06C44F" ext-link-type="gen" xlink_href="OR839461" xlink_type="simple">OR 839461</ext-link>
, 
<ext-link id="732101B2EECF4DC13DCDAE9D7DE18C68" ext-link-type="gen" xlink_href="OR839462" xlink_type="simple">OR 839462</ext-link>
, 
<ext-link id="FFD8C2746C28F43D31688A25390E10AE" ext-link-type="gen" xlink_href="OR853327" xlink_type="simple">OR 853327</ext-link>
, 
<ext-link id="D14DD38EABFDBE83A8844315C4636C62" ext-link-type="gen" xlink_href="OR839630" xlink_type="simple">OR 839630</ext-link>
, 
<ext-link id="B8521A10A32DE795461C32ADCE9DAF2E" ext-link-type="gen" xlink_href="OR838053" xlink_type="simple">OR 838053</ext-link>
, 
<ext-link id="34CF0D27A540D3159A93720C3E1432F5" ext-link-type="gen" xlink_href="OR853328" xlink_type="simple">OR 853328</ext-link>
, 
<ext-link id="EA6F73C69D8817972DC9128E4955B82E" ext-link-type="gen" xlink_href="OR839631" xlink_type="simple">OR 839631</ext-link>
, 
<ext-link id="6F594F662B7BEFA92E8B0DD76BBCE548" ext-link-type="gen" xlink_href="OR839632" xlink_type="simple">OR 839632</ext-link>
, 
<ext-link id="DFF6DC05BB5E449FC97A490AF79CF900" ext-link-type="gen" xlink_href="OR839633" xlink_type="simple">OR 839633</ext-link>
, 
<ext-link id="CE74B558C5E41928F8332957460C6FA2" ext-link-type="gen" xlink_href="OR853326" xlink_type="simple">OR 853326</ext-link>
, 
<ext-link id="0D69A3CD0BB597CE76BAE978276D1A46" ext-link-type="gen" xlink_href="OR839621" xlink_type="simple">OR 839621</ext-link>
, 
<ext-link id="882A4A7D0175FDC92194D7FC360FD1D7" ext-link-type="gen" xlink_href="OR838050" xlink_type="simple">OR 838050</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EB1FM" type="material">
<paragraph id="AE99385C4261FB658E1B2BC14E3113DC">
<heading id="C782E8CFDCD2FE14E0A4E935C438BDB0" reason="title">Other material</heading>
</paragraph>
<paragraph id="F4B8593ADC7AC62481A6E91534373DCD">
<emphasis id="E58C716F2F356DC7F7775D3CF9CA622A" bold="true">
(
<emphasis id="714783343427FBAEFB697D6CF883EF9F" bold="true" italics="true">n</emphasis>
= 40 
<named-content id="F84AF3C54CB597DB3A425CF7F0266F77" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="E41908B0A9BC1A964CEE41F2D79A236F" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
, 
<abbrev id="ABBRID0EP1FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="F192B44946BE6E99A1DF231588758B26">CWHc</collectionCode>
</abbrev>
). 
<materialsCitation id="456CC05E1CA6A18A9AA347E0F9FFCF49" collectedFrom="Under embedded rock in clay soil" collectingDate="2020-05-21" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Anderson Co." latitude="34.60262" location="Big Oaks area near Lake Hartwell" longLatPrecision="1" longitude="-82.83858" municipality="Clemson Experimental Forest" specimenCode="CUAC 000168245, CUAC 000168251" specimenCount="2" specimenCount-male="2" stateProvince="South Carolina">
<collectingCountry id="CAE78B1B8F7F9FD9FEEAB64729107040" name="United States of America">USA</collectingCountry>
• 
<emphasis id="FA9B34C122A45C4CD4F5B70F35C28112" bold="true">
<collectingRegion id="CA7AF33AC59B1FC439BE697E3209A7E7" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="A3E3B7E48AC13882306AC57A686D10E8" count="2" type="male">2 ♂</specimenCount>
; 
<emphasis id="CE8995F22A2C0BF77361DB0291FC64DB" bold="true">
<collectingCounty id="8FCC5270F782E9531658D30FB6264731">Anderson Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="9053C13845F157828217086AA9B9E70E">Clemson Experimental Forest</collectingMunicipality>
, 
<location id="6AC0CF4B9053EE8B09353A903343B3AC" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:6AC0CF4B9053EE8B09353A903343B3AC" country="United States of America" county="Anderson Co." latitude="34.60262" longLatPrecision="1" longitude="-82.83858" municipality="Clemson Experimental Forest" name="Big Oaks area near Lake Hartwell" stateProvince="South Carolina">Big Oaks area near Lake Hartwell</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E21FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.838580,34.602620]}">
<geoCoordinate id="D414DDD2E28F8D246B628D40D3EA283C" degrees="34.60262" direction="north" orientation="latitude" precision="1" value="34.60262">34.60262</geoCoordinate>
, 
<geoCoordinate id="5B9AED9F26A3B42B988E8C821E86085F" degrees="82.83858" direction="west" orientation="longitude" precision="1" value="-82.83858">- 82.83858</geoCoordinate>
</named-content>
; 
<date id="597760A60AE61F6264E9A4552C2AEF9E" value="2020-05-21">
<collectingDate id="0F200C6D2BA229538FB3150A6971616D" value="2020-05-21">21 May 2020</collectingDate>
</date>
; 
<collectorName id="F65C85AE7EEE6049E0895FB69AB4C2E6">C. W. Harden</collectorName>
; 
<collectedFrom id="F8A55E384AAC533C77D768D680DD414E">Under embedded rock in clay soil</collectedFrom>
; CWH- 161 and CWH- 162, 
<specimenCode id="D771E6DA819CB392F1F121A590A19DA5" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168245</specimenCode>
and 
<specimenCode id="490F1FEB0BE7160DC8C9AA834D14BD85" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168251</specimenCode>
; 
<named-content id="8C628A819C8E422C3250A8D25318698E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="9097361FF77CA2CA1C91A0355637D141" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
</emphasis>
; • 
<materialsCitation id="B136D05D6747477DA74668908C10670C" collectedFrom="deep soil by large stump near lake, after flood" collectingDate="2020-05-21" collectingMethod="Berlese" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Anderson Co." latitude="34.60225" location="Big Oaks area" longLatPrecision="1" longitude="-82.83893" municipality="Clemson Experimental Forest" specimenCode="CUAC 000168252, CUAC 000168253" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="64E03F7B54A18DF4711BC3B64352C901" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="0AE29DAE51F053185CF8B70B56F9F06C" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="B5EE0C72EBE50B34F9E97C627BC1C4C3" bold="true">
<collectingCounty id="26F08F8252D8BCDA9E8FEDE6A231999C">Anderson Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="567CD77296016BDCE320B497123C15BA">Clemson Experimental Forest</collectingMunicipality>
, 
<location id="C7809D3E0F8A6D664BFCD9E360856007" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:C7809D3E0F8A6D664BFCD9E360856007" country="United States of America" county="Anderson Co." latitude="34.60225" longLatPrecision="1" longitude="-82.83893" municipality="Clemson Experimental Forest" name="Big Oaks area" stateProvince="South Carolina">Big Oaks area</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJ2FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.838930,34.602250]}">
<geoCoordinate id="CE061AF53FF6095B9AEDF6046C5F9828" degrees="34.60225" direction="north" orientation="latitude" precision="1" value="34.60225">34.60225</geoCoordinate>
, 
<geoCoordinate id="233D976CDA5D67D783E1B3A0CB069114" degrees="82.83893" direction="west" orientation="longitude" precision="1" value="-82.83893">- 82.83893</geoCoordinate>
</named-content>
; 
<date id="DD1660206C392B967B5A2A022E60A5A3" value="2020-05-21">
<collectingDate id="A1825A043BEE50BD7B5A43A5B1AD9211" value="2020-05-21">21 May 2020</collectingDate>
</date>
; 
<collectorName id="6B484B44E4A6A9415BE62767DE0327D1">C. W. Harden</collectorName>
&amp; 
<collectorName id="DC94D6BD134FC5725E7417C0FADBEB17">L. M. Thompson</collectorName>
; 
<collectingMethod id="67981D8A7761697D7E91E7E51CD537D2">Berlese</collectingMethod>
, 
<collectedFrom id="AE1D62BF5368EE4679F48F61E764E5DC">deep soil by large stump near lake, after flood</collectedFrom>
; CWH- 163 and CWH- 164, 
<specimenCode id="14B7D8AD904CEEB2A62810AB269A2F8C" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168252</specimenCode>
and 
<specimenCode id="CC1B3C816EC92014080D8F615240DC4A" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168253</specimenCode>
; 
<named-content id="4FE27DD1A12973A74676A71CFFA85C0B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="8C94FA5538D6208D63046156027AC258" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="554AC5B5584591705EB03B129330996C" collectedFrom="sifted pine stump soil, near lake" collectingDate="2021-03-19" collectingMethod="Berlese" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Anderson Co." latitude="34.47497" location="River Forks Recreation Area" longLatPrecision="1" longitude="-82.80914" specimenCode="CUAC 000168254" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="DA91970CD0B1C2EADEFD1DAFCEABAFBA" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="AE455FFC0D4F081065BD13E5DEAC9840" bold="true">
<collectingCounty id="6F0EACD9C37280D9C05F935EAC3296A6">Anderson Co.</collectingCounty>
</emphasis>
; 
<location id="CF1C34018BDAD1D70059A9664C7544CC" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:CF1C34018BDAD1D70059A9664C7544CC" country="United States of America" county="Anderson Co." latitude="34.47497" longLatPrecision="1" longitude="-82.80914" name="River Forks Recreation Area" stateProvince="South Carolina">River Forks Recreation Area</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX2FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.809140,34.474970]}">
<geoCoordinate id="7E569ED3D48B895072BBE7EC1E2EDA81" degrees="34.47497" direction="north" orientation="latitude" precision="1" value="34.47497">34.47497</geoCoordinate>
, 
<geoCoordinate id="583C8AA3BFB0AAE71E05735212393A6F" degrees="82.80914" direction="west" orientation="longitude" precision="1" value="-82.80914">- 82.80914</geoCoordinate>
</named-content>
; 
<date id="793772CE0AE305BE27F049F863B2892D" value="2021-03-19">
<collectingDate id="B62F459B4C4F837ACCFDCC8127AD1E4E" value="2021-03-19">19 Mar. 2021</collectingDate>
</date>
; 
<collectorName id="217AA8F0D4D9C97E8ABFF375B3769CA0">C. W. Harden</collectorName>
&amp; 
<collectorName id="86447A6B0FFF9275B39573C7F01C4849">L. M. Thompson</collectorName>
; 
<collectingMethod id="268781405010F970A25A988DF38827EC">Berlese</collectingMethod>
, 
<collectedFrom id="5EEF425472FB18DA2207798FF68D3340">sifted pine stump soil, near lake</collectedFrom>
; CWH- 327, 
<specimenCode id="2415BC10C18F06006E09875C40A086BB" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168254</specimenCode>
; 
<named-content id="D41DA92A646C6B0205F0E8C6D0832F3A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="F448E33636D51984477053E37B129B21" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="9EC55F5A99521C46E8C9011A5FDBA7FF" collectingDate="2020-05-21" collectingMethod="deep soil Berlese" collectionCode="CWHc" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Anderson Co." latitude="34.60225" location="Big Oaks Area" longLatPrecision="1" longitude="-82.83893" municipality="Clemson Experimental Forest" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="South Carolina">
<specimenCount id="D8B75F8051436F336A7424982CD91347" count="2" type="male">2 ♂</specimenCount>
, 
<specimenCount id="041E62E1CF5679F6A8F8E073065FEEC1" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="341D99F36FF08393FAE119E4BBE4A978" bold="true">
<collectingCounty id="25C3571EFCD6082428C3ECA811683426">Anderson Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="E245EE69A6A11BA5FA34008C86209EE7">Clemson Experimental Forest</collectingMunicipality>
, 
<location id="589443AD29B2EC3896418B367A1B4914" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:589443AD29B2EC3896418B367A1B4914" country="United States of America" county="Anderson Co." latitude="34.60225" longLatPrecision="1" longitude="-82.83893" municipality="Clemson Experimental Forest" name="Big Oaks Area" stateProvince="South Carolina">Big Oaks Area</location>
near 
<location id="7E94B629C3ED4FEF8C687ECEAF4564D9" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:7E94B629C3ED4FEF8C687ECEAF4564D9" country="United States of America" county="Anderson Co." latitude="34.60225" longLatPrecision="1" longitude="-82.83893" municipality="Clemson Experimental Forest" name="Lake Hartwell" stateProvince="South Carolina">Lake Hartwell</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EF3FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.838930,34.602250]}">
<geoCoordinate id="C2AA9FC188890F26A5CAA718BCB5B27C" degrees="34.60225" direction="north" orientation="latitude" precision="1" value="34.60225">34.60225</geoCoordinate>
, 
<geoCoordinate id="E6C02450B5CC919F1324251EC8C2D9B6" degrees="82.83893" direction="west" orientation="longitude" precision="1" value="-82.83893">- 82.83893</geoCoordinate>
</named-content>
; 
<date id="3BB269F13A89FF537748E4D99DD7F372" value="2020-05-21">
<collectingDate id="2452B85671965BA71943A71691789F2F" value="2020-05-21">21 May 2020</collectingDate>
</date>
; 
<collectorName id="86AF3A8CF90F3000C71E24B74B142146">C. W. Harden</collectorName>
and 
<collectorName id="C89875AFD7F2B3DB3B9A7EFE1D05CF4D">L. M. Thompson</collectorName>
leg.; 
<collectingMethod id="684E3C1153475F7D24E7FFAAA367B6AE">deep soil Berlese</collectingMethod>
; 
<abbrev id="ABBRID0EK3FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="9BB36A2CB296A81CD63BF18A7E778625">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="98688F234E296163FA9A75191B95016A" collectedFrom="deep soil and litter Berlese" collectingDate="2020-03-25" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Anderson Co." latitude="34.61559" location="Big Oaks" longLatPrecision="1" longitude="-82.83061" municipality="Clemson Experimental Forest" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="0944CAD92CA8E78AD30013421D3F36C7" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="82E5FBBADC7C58EC3FF8438AB06E19FE" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="FF4FDB65511C58E1CC97F5651E2151B8" bold="true">
<collectingCounty id="D6EB582227E3823FAE0B0E82856B6BFE">Anderson Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="C245574420ECBABAFAD3373C700CD0E0">Clemson Experimental Forest</collectingMunicipality>
, 
<location id="8FF963814C51A48213140B35261E8565" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:8FF963814C51A48213140B35261E8565" country="United States of America" county="Anderson Co." latitude="34.61559" longLatPrecision="1" longitude="-82.83061" municipality="Clemson Experimental Forest" name="Big Oaks" stateProvince="South Carolina">Big Oaks</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET3FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.830610,34.615590]}">
<geoCoordinate id="F7DB1571D2BADE8130C8F90335E3F50D" degrees="34.61559" direction="north" orientation="latitude" precision="1" value="34.61559">34.61559</geoCoordinate>
, 
<geoCoordinate id="19BA5B27D1D1449BC4324188C7AE12E3" degrees="82.83061" direction="west" orientation="longitude" precision="1" value="-82.83061">- 82.83061</geoCoordinate>
</named-content>
; 
<date id="5A6B1CD0523A7E5AEE2E5E25615DCBC0" value="2020-03-25">
<collectingDate id="091A57B4FE4AD15BCA13CBBE5A717369" value="2020-03-25">25 Mar 2020</collectingDate>
</date>
; 
<collectorName id="4337FAED3A2E9B6517D54C4E82FA6147">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="3C55A69E6C1C7A6CA26B9891AD30456A">deep soil and litter Berlese</collectedFrom>
; 
<abbrev id="ABBRID0EY3FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="DD0AC542CD31044F633E7568FB06DECF">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="2BA6206F0782584A17A967C056B044F2" collectedFrom="soilwash flotation Berlese" collectingDate="2020-01-04" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCode="CUAC 000168237, CUAC 000168238" specimenCount="2" specimenCount-male="2" stateProvince="South Carolina">
<specimenCount id="E7E6EFC26BBC4D8B573B760B4933EEB0" count="2" type="male">2 ♂</specimenCount>
; 
<emphasis id="739FD553BE4477A9E5ABED2B128AD1CA" bold="true">
<collectingCounty id="D936528D303D241B6D49048ABAD35AFB">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="42E13ADCD5ED252B80989183E031C66D">Lake Hartwell</collectingMunicipality>
, 
<location id="965CD036FE77A90404139912DB463721" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:965CD036FE77A90404139912DB463721" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
, 
<locationDeviation id="5E3BC636BA1904E14209444FA1890A86" location="Clemson">SW of Clemson</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EB4FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="7A4F3C81A40A37D2B078C7CE0089863E" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="146C3447D4103D2B9B9CC5841179804F" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="9826D77CE06B540B6612525F226BB0BF" value="2020-01-04">
<collectingDate id="EAD0D6D8B302DC4BBD295924644E1407" value="2020-01-04">4 Jan. 2020</collectingDate>
</date>
; 
<collectorName id="24D1DA3D2EE73BA14C9BCC34E1B8C48D">C. W. Harden</collectorName>
&amp; 
<collectorName id="998618C3A40352CA40176DE3A2D8AFE0">L. M. Thompson</collectorName>
leg.; 
<collectedFrom id="77F948FED3EA884B539287C3F144E0FD">soilwash flotation Berlese</collectedFrom>
; 
<specimenCode id="0D4398701997F871D50B219B7E37A928" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168237</specimenCode>
and 
<specimenCode id="ED4C6997D732E95F4DE53D07A0715B47" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168238</specimenCode>
; CWH- 070 and CWH- 114; 
<named-content id="ABD54C3B6C8C41CF99BC7426F0CBF027" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="9AADB230D68A51311603597ADAB8EC7C" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="AE826197F42B90E200233FABE0F6696D" collectedFrom="soilwash flotation Berlese" collectingDate="2020-01-04" collectionCode="CWHc" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="833F46ADEDB57D075CC970D88F2476C9" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="7D3EBDFE99D342AB8B736C5DD1F60F49" count="2" type="female">2 ♀</specimenCount>
; same data as previous; 
<abbrev id="ABBRID0EK4FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="F6A8A12538FEE1A4D7609F7645EFC834">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="6E6A1ADA8143C4727796CD7C2D7FCCF5" collectedFrom="On underside of deeply embedded rocks" collectingDate="2020-01-04" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCode="CUAC 000168243, CUAC 000168233" specimenCount="2" specimenCount-male="2" stateProvince="South Carolina">
<specimenCount id="5BFC998ED25148CE67D8900C2A6C7DC6" count="2" type="male">2 ♂</specimenCount>
; 
<emphasis id="6A3E7F6F548A692803A75E90D0BBA45E" bold="true">
<collectingCounty id="F6EE0355ECF2ABF5780DE869FC64A9F3">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="B1847961383973180AE11A0514C7CFCB">Lake Hartwell</collectingMunicipality>
, 
<location id="4C5E3223BE181FFC25FE11D390AFEE0B" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:4C5E3223BE181FFC25FE11D390AFEE0B" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
, 
<locationDeviation id="FAAB3B70A44BC7950343EA869B3EDBE0" location="Clemson">SW of Clemson</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET4FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="3213245BD5CADAB7670F486B3F5065F4" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="BDE5607C55B88E1F53F20F5D763AC297" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="407B36DBB631D7A9B6E323DF4F126003" value="2020-01-04">
<collectingDate id="4AA05A41A14A95B97F7EB04C08E188BE" value="2020-01-04">4 Jan. 2020</collectingDate>
</date>
; 
<collectorName id="12E96794FC007079C4A13343391D1389">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="36740A4790C3DB7F874B7C5E4CF62B8A">On underside of deeply embedded rocks</collectedFrom>
; 
<specimenCode id="3F14106B3E43C1E7E217FD0B81DCA94B" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168243</specimenCode>
and 
<specimenCode id="C9EBB8638B35D48572DE31982E5EE64F" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168233</specimenCode>
; CWH- 062 and CWH- 063; 
<named-content id="8D162AC1F586DF6A79CCD6D85F309D1F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="86557DAD797D35AE04297105F9821CFC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="0A9E86153FA048BCB6911F4CF5DDA38E" collectedFrom="On underside of deeply embedded rocks" collectingDate="2019-12-31" collectionCode="CUAC" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCode="CUAC 000168234, CUAC 000168236" specimenCount="3" specimenCount-male="3" stateProvince="South Carolina">
<specimenCount id="85324B4B9D9E45691644572325DF4737" count="3" type="male">3 ♂</specimenCount>
; 
<emphasis id="CCFCBF7667B3D9B4612120411708EA8A" bold="true">
<collectingCounty id="023B9D1A5EFF889B8E0711759DBE465B">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="69ACE5913DACC4DC0469C5A515CB2DCB">Lake Hartwell</collectingMunicipality>
, 
<location id="C964F102B14DC3303F1CD0B8914DF6FF" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:C964F102B14DC3303F1CD0B8914DF6FF" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
, 
<locationDeviation id="462EF34512BB5EE04BCA1C733913BF1E" location="Clemson">SW of Clemson</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EB5FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="AAAF813A4AE2D93C22EBC5ECE9150838" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="6EC67A4DAC5BB967808A2BB0A3F8B600" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="EA82A2CA98860CB2C9F38865B5771373" value="2019-12-31">
<collectingDate id="301B1D8E46DA1E75B6D79BFF4F3EE5E7" value="2019-12-31">31 Dec. 2019</collectingDate>
</date>
; 
<collectorName id="7856070583E3FA41A4E4157A4E553C88">C. W. Harden</collectorName>
&amp; 
<collectorName id="C0E587AF58D7480BC3FAD7D7A4F13EB3">L. M. Thompson</collectorName>
leg.; 
<collectedFrom id="3CA3D1EC35F785F20872DD1F887E344C">On underside of deeply embedded rocks</collectedFrom>
; 
<specimenCode id="EBD6F830598807257195D85EA72C09A5" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168234</specimenCode>
to 
<specimenCode id="2B18926655EB423905D60C1CAC174C4C" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168236</specimenCode>
; CWH- 064 to CWH- 066; 
<named-content id="48B7ACB43CBD2C0D9770CA7FDE36796A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="1A82AE81B8EB78D0107FEDA70CE61A71" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="DD68F729A4DE35A4BEF5EFE7EC00EF43" collectedFrom="Under embedded rocks" collectingDate="2020-01-11" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCode="CUAC 000168246, CUAC 000168249" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="19D4F1E7EB64865892AF25428D2E7C30" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="4AB5D18428241CD8D19DB0A44F526BA9" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="D07AB5A3BFA89C348A3F273A64121AAF" bold="true">
<collectingCounty id="D0F6BFDC945D47F79CA3373F70D18D33">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="7E5D79DC81AF88E658D28C9B3FEC72A4">Lake Hartwell</collectingMunicipality>
, 
<location id="2A0ADF6E2186A63AC7535520E09AB311" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:2A0ADF6E2186A63AC7535520E09AB311" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
, 
<locationDeviation id="61D846212CD857B38C293A557791F225" location="Clemson">SW of Clemson</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EP5FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="5D5344C605401E4F0F9C077B1CAFB270" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="4280AAADED0BC44ABD4D07CE7B4BA9D6" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="B0516DEA5641F6A8CC09F2D3DE63B8B0" value="2020-01-11">
<collectingDate id="0BADFFBC08B0F012E5B7EC74989911C6" value="2020-01-11">11 Jan. 2020</collectingDate>
</date>
; 
<collectorName id="9EC6C3D23807A7A85684A811FDBFDD90">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="A55CB481799DED3918503D8E54F77075">Under embedded rocks</collectedFrom>
; 
<specimenCode id="9955FD3E9427D6EE7CDA86D434DB1387" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168246</specimenCode>
and 
<specimenCode id="A9721616C5A7E01F29DED5411CA9A15E" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168249</specimenCode>
; CWH- 119 and CWH- 123; 
<named-content id="E4AF1384C1C4D2F39CF1F03D4444861F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="A879122289F8925D455BF1B9FDD87629" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="927E6253DA1CB4367DCB24157012EB53" collectingDate="2022-05-15" collectingMethod="soilwashing" collectionCode="CWHc" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6388" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8655" specimenCount="1" specimenCount-larva="1" stateProvince="South Carolina">
<specimenCount id="14A309DEC38B133D0A6BFD8C225C5034" count="1" type="larva">1 larva</specimenCount>
; 
<emphasis id="985B56DCBE344ED612D467B00B74B0FD" bold="true">
<collectingCounty id="CF9D6170490DB867B7B6FDC7CBD1F6F3">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="26D9B14374587FFA78BDAF8217EBEFDC" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:26D9B14374587FFA78BDAF8217EBEFDC" country="United States of America" county="Oconee Co." latitude="34.6388" longLatPrecision="6" longitude="-82.8655" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E45FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865500,34.638800]}">
<geoCoordinate id="350C06A4F32E47EB0CCE588084942663" degrees="34.6388" direction="north" orientation="latitude" precision="5" value="34.6388">34.6388</geoCoordinate>
, 
<geoCoordinate id="8662BA8ABDEB85A6C80690C19F0CC5DB" degrees="82.8655" direction="west" orientation="longitude" precision="5" value="-82.8655">- 82.8655</geoCoordinate>
</named-content>
; 
<date id="069C765734D8404AB5529C03921B8561" value="2022-05-15">
<collectingDate id="23DDAE1889C1F70427B13BE8C8D77A9E" value="2022-05-15">15 May 2022</collectingDate>
</date>
; 
<collectorName id="A40A7B01D42907972FC55E17812B27C1">C. W. Harden</collectorName>
and 
<collectorName id="425169B29FF2A45BC1AAC1847ED43816">L. M. Thompson</collectorName>
leg.; 
<collectingMethod id="35FF263210B36C492169845BAFEA19B1">soilwashing</collectingMethod>
; 
<abbrev id="ABBRID0EC6FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="9C908851179CA64A97A87A24589FCA76">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="38BBC2193AB64D7ECE50A1A0A79B4102" collectedFrom="under rock" collectingDate="2021-10-22" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6388" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8644" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="South Carolina">
<specimenCount id="743D33C93181D3A30B3E9E7EF5707441" count="2" type="male">2 ♂</specimenCount>
, 
<specimenCount id="0E59A42575550D316F85BC7B3E510A9E" count="6" type="female">6 ♀</specimenCount>
; 
<emphasis id="9EFD4D6CF7F738A4BAC3773EB12DB336" bold="true">
<collectingCounty id="138234C43DC9723149EC3F92C293DB71">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="CD861C314A8ADCE6FECFC162C8489F8D" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:CD861C314A8ADCE6FECFC162C8489F8D" country="United States of America" county="Oconee Co." latitude="34.6388" longLatPrecision="6" longitude="-82.8644" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EL6FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.864400,34.638800]}">
<geoCoordinate id="657BDB81D36700CA4F23F01D40159A1D" degrees="34.6388" direction="north" orientation="latitude" precision="5" value="34.6388">34.6388</geoCoordinate>
, 
<geoCoordinate id="FFCC248B887D5DD1707DC9979EEA15EF" degrees="82.8644" direction="west" orientation="longitude" precision="5" value="-82.8644">- 82.8644</geoCoordinate>
</named-content>
; 
<date id="BA15A858A76B3CBABECAD051FC3F322C" value="2021-10-22">
<collectingDate id="0CE3822DFF13741A519711AC6AA4E955" value="2021-10-22">22 Oct 2021</collectingDate>
</date>
; 
<collectorName id="50E12FF921B2F416BDCBAE54FF49756D">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="7F237E74D18B736D91C113E73806FB08">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EQ6FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="F82E8A7B339AB0F4445E993EB1D7C0BB">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="09CE25750D0B3D7B0831DFBD4154C380" collectedFrom="under rock" collectingDate="2021-10-22" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.639" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8638" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="7A30AA04A35321FFBC89FDE40D7F3044" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="0760CBD2B02515CC1D92F30A797C0D4C" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="CF69154DAFDA985AFFA8C72F388AA139" bold="true">
<collectingCounty id="4B8CA328F432B8E10905B02C1F64B19D">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="357B6AA45BF062C8386C44E2556084F1" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:357B6AA45BF062C8386C44E2556084F1" country="United States of America" county="Oconee Co." latitude="34.639" longLatPrecision="6" longitude="-82.8638" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZ6FM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.863800,34.639000]}">
<geoCoordinate id="6F3BDDDA8CA3CE9D987C3DF14229BA6C" degrees="34.6390" direction="north" orientation="latitude" precision="5" value="34.639">34.6390</geoCoordinate>
, 
<geoCoordinate id="C69FFBE06CDFA24A36A7947724DD8660" degrees="82.8638" direction="west" orientation="longitude" precision="5" value="-82.8638">- 82.8638</geoCoordinate>
</named-content>
; 
<date id="3A71E8F338FF0F0C5FD5751EE4411153" value="2021-10-22">
<collectingDate id="74A2C876EBD4455B23BEAD3B626AB002" value="2021-10-22">22 Oct 2021</collectingDate>
</date>
; 
<collectorName id="BDB8263DA95992F8E99E1746DBD8B11F">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="D6DC855FCC071961A24E234A04532EB6">under rock</collectedFrom>
; 
<abbrev id="ABBRID0E56FM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="D160F2381C56B76E9BE373D139DC7EC3">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="417FFB930222CC099629859876E836DE" collectedFrom="under rock" collectingDate="2021-10-22" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6391" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8636" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina">
<specimenCount id="AC696F918743DAFDB9BD9054AFA78C9F" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="59FB4797CEE768651C7F741078464A91" bold="true">
<collectingCounty id="F91F38C00878BBCA61B90220E17CE64C">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="46286E0E16B3DF0D99D2CB49555D3A04" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:46286E0E16B3DF0D99D2CB49555D3A04" country="United States of America" county="Oconee Co." latitude="34.6391" longLatPrecision="6" longitude="-82.8636" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHAGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.863600,34.639100]}">
<geoCoordinate id="C55642E9DBD686FB2C7E868596F75523" degrees="34.6391" direction="north" orientation="latitude" precision="5" value="34.6391">34.6391</geoCoordinate>
, 
<geoCoordinate id="5A424C1FB7CB3EF592DD10ADDE89F2F1" degrees="82.8636" direction="west" orientation="longitude" precision="5" value="-82.8636">- 82.8636</geoCoordinate>
</named-content>
; 
<date id="1962F0258DC426CBA7E4865F2F28EB62" value="2021-10-22">
<collectingDate id="A06D2D8CFD1FE0EF9A26A6E8AACF631C" value="2021-10-22">22 Oct 2021</collectingDate>
</date>
; 
<collectorName id="33C392C0E669CB3DF3AC3C23DBF9C5B1">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="4AB25EF9109B4D6805381CCAAB3D62B2">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EMAGM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="272A17F9E18D17EF7E3CE7CE4B320BFE">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="7F0F363D8770E3CA7B12781044D803BB" collectedFrom="under rock" collectingDate="2021-10-22" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6393" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8633" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina">
<specimenCount id="E68237688AC1DA53C218F3C9CB54F6BE" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="8E09442FC31970CB97651C8BBEAC5045" bold="true">
<collectingCounty id="33640623EFBB724C5A667ED0EDDA6FF4">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="408069C8F1CA9816F2FA58DC1A743582" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:408069C8F1CA9816F2FA58DC1A743582" country="United States of America" county="Oconee Co." latitude="34.6393" longLatPrecision="6" longitude="-82.8633" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVAGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.863300,34.639300]}">
<geoCoordinate id="2D377938ABC8959802609D26B4951AC3" degrees="34.6393" direction="north" orientation="latitude" precision="5" value="34.6393">34.6393</geoCoordinate>
, 
<geoCoordinate id="EAF90F37FF4C147FD1C2810F9F0C9420" degrees="82.8633" direction="west" orientation="longitude" precision="5" value="-82.8633">- 82.8633</geoCoordinate>
</named-content>
; 
<date id="46137F13B5C38703E3953DD8C0FA110B" value="2021-10-22">
<collectingDate id="5D4696EF5FFCD12F2AE571EB03528603" value="2021-10-22">22 Oct 2021</collectingDate>
</date>
; 
<collectorName id="A821746526A8696E774BF0BEF3A5E668">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="66FFDA34150502F03103CCCE1E69C55D">under rock</collectedFrom>
; 
<abbrev id="ABBRID0E1AGM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="37BF96EE3A895B681B174D9D2E85408F">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="598E3FF7718B2DB55768AB6563DD3DF4" collectingDate="2020-01-04" collectionCode="CWHc" collectorName="C. W. Harden &amp; L. M. Thompson" country="United States of America" county="Oconee Co." latitude="34.6389" location="Martin Creek Landing" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" specimenCount="2" specimenCount-female="2" stateProvince="South Carolina">
<specimenCount id="C7B833A0D4159183F79583DEAFDFA377" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="B38DEDDB19AC6544ED268B9347D3ACC8" bold="true">
<collectingCounty id="EC8EEA86AA4A2E951FA222B55B29C589">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="2BAA30700BE494CBA9A8B236405660C9">Lake Hartwell</collectingMunicipality>
, 
<location id="54D8978B62DCC6842C87991498CC1C26" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:54D8978B62DCC6842C87991498CC1C26" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Lake Hartwell" name="Martin Creek Landing" stateProvince="South Carolina">Martin Creek Landing</location>
, 
<locationDeviation id="987D8C017D2590332F1A65A98A0A822F">southwest of Clemson</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDBGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="EFFFC32945A6D679CC721387E59F0FBB" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="52DBD040288CC3979BCA719243EF6E58" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="8837DD37CE81AF00B8330F586859F890" value="2020-01-04">
<collectingDate id="5211D136DAD24C5DF0DF138B32A9186D" value="2020-01-04">4 Jan 2020</collectingDate>
</date>
; 
<collectorName id="B16E94BD78247C86A3713B963EFDC881">C. W. Harden</collectorName>
and 
<collectorName id="43F6A94225E5FD860F35626D0C1A6CB5">L. M. Thompson</collectorName>
leg.; 
<abbrev id="ABBRID0EIBGM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="F607769AF3ECB9D57FA9FD702A4D6C7D">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="1020396999C4CA5FEC51F4E9A70FD7B3" collectedFrom="under rock" collectingDate="2023-02-13" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="34.6389" location="southwest of Clemson" longLatPrecision="6" longitude="-82.8656" municipality="Martin Creek Landing" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina">
<specimenCount id="768391C9D358225320F67ACE633D61FA" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="8FFCEECE626FF4DF98D020FF297185D2" bold="true">
<collectingCounty id="8EAD87DE162751F6B0BE482F28CF4BE7">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="B297D562756F80EB1BCE6B564FCB92F4">Martin Creek Landing</collectingMunicipality>
, 
<location id="99F79FE20870A03F574949B2DD1AB737" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:99F79FE20870A03F574949B2DD1AB737" country="United States of America" county="Oconee Co." latitude="34.6389" longLatPrecision="6" longitude="-82.8656" municipality="Martin Creek Landing" name="southwest of Clemson" stateProvince="South Carolina">southwest of Clemson</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERBGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-82.865600,34.638900]}">
<geoCoordinate id="5FB1E44133CA5EB136C4A97ABED1CE77" degrees="34.6389" direction="north" orientation="latitude" precision="5" value="34.6389">34.6389</geoCoordinate>
, 
<geoCoordinate id="FDF2829A29A26DFCF508C432632C3CA9" degrees="82.8656" direction="west" orientation="longitude" precision="5" value="-82.8656">- 82.8656</geoCoordinate>
</named-content>
; 
<date id="EAF69773CE6C811B0433206CFB1FB3BF" value="2023-02-13">
<collectingDate id="AA2C2A5900327072D449E5291B2DA4DC" value="2023-02-13">13 Feb. 2023</collectingDate>
</date>
; 
<collectorName id="A80599D391068D05C6F6F45494AC35F1">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="E3ABCF2C2089E884AA1DC99763CC5ABC">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EWBGM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="944444C3BDC664D34B002EB25D05F1AA">CWHc</collectionCode>
</abbrev>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1BGM" type="diagnosis">
<paragraph id="8109E1923E51982093F9917A1CABEBED">
<heading id="D7B38F6C8DCDF14FDA2DD05C23A7BD10" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="82CF582F56481E4DB18EB3AA735F7124">
Males with metafemora not heavily modified, with a median tuberculate area on posterior margin (Fig. 
<figureCitation id="BC730D029FED0362F90EA8FF79FB0CF4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. SEM micrographs of male left metafemora of Anillinus species. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure12" httpUri="https://binary.pensoft.net/fig/1110351">12 D</figureCitation>
), sometimes with a small blunt tooth. Male genitalia distinctive (Fig. 
<figureCitation id="4795AABB7A14229E51226A1AC42F28D6" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure35" httpUri="https://binary.pensoft.net/fig/1110374">35 D – F</figureCitation>
): median lobe of aedeagus with straight and narrow ventral margin, internal sac without groups of spines, flagellum relatively large and evenly curved.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EICGM" type="description">
<paragraph id="16AF205503C46C3ABCA4B467341BA875">
<heading id="47A072A3A3364981D3980A243C6C8844" reason="title">Description.</heading>
</paragraph>
<paragraph id="52A5626BFEEDA24A897D70467DC56B32">
<emphasis id="508128E9E159C508D99C57973EAC703D" bold="true" italics="true">Habitus</emphasis>
Widely variable in body size (
<abbrev id="ABBRID0ETCGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 
<quantity id="73A3E170D0BE4B0B0F34F3B735930A32" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.78" metricValueMin="1.46" unit="mm" value="1.62" valueMax="1.78" valueMin="1.46">1.46–1.78 mm</quantity>
, average = 1.63 ± 0.08) and shape, with smaller specimens being more convex and ovoid (Fig. 
<figureCitation id="80200C8A2B975C52A96C7D38A3C939F8" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure34" httpUri="https://binary.pensoft.net/fig/1110373">34 D</figureCitation>
) and larger specimens being flatter and more parallel sided (similar to Fig. 
<figureCitation id="37E2224F1AD983D10CA296F06A7632B1" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Dorsal habitus of Anillinus species in the elongatus group and sinuaticollis group A Anillinus montrex B Anillinus arenicollis C Anillinus choestoea sp. nov. D Anillinus seneca sp. nov. E Anillinus mica sp. nov. F Anillinus micamicus sp. nov. Abdomens removed in B – F for DNA extraction. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure34" httpUri="https://binary.pensoft.net/fig/1110373">34 E</figureCitation>
). Males and females similarly variable in size (
<abbrev id="ABBRID0E6CGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 
<quantity id="7518163C5D0ED9A853BA0548999F18B7" metricMagnitude="-3" metricUnit="m" metricValue="1.645" metricValueMax="1.78" metricValueMin="1.51" unit="mm" value="1.645" valueMax="1.78" valueMin="1.51">1.51–1.78 mm</quantity>
and 
<quantity id="B57E98EF0B13A977DEA42C07E6700278" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.78" metricValueMin="1.46" unit="mm" value="1.62" valueMax="1.78" valueMin="1.46">1.46–1.78 mm</quantity>
, respectively). Not markedly narrow (average 
<abbrev id="ABBRID0EDDGM" xlink_title="Maximum elytral width">EW</abbrev>
/ 
<abbrev id="ABBRID0EHDGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.36). 
<emphasis id="A948E227A4E8D890D9657E7EEDE5A8C6" bold="true" italics="true">Integument</emphasis>
Dorsal surface of head fully microsculptured; microsculpture coverage of pronotum varying: usually lacking from disc and present across entire anterior margin and along sides, sometimes absent anteriorly except for anterior angles; sometimes distinct across entire anterior 1 / 4, including part of disc. 
<emphasis id="F34D220B17E9862731B689B3F7003AFD" bold="true" italics="true">Head</emphasis>
<abbrev id="ABBRID0ERDGM" xlink_title="Maximum head width">HW</abbrev>
/ 
<abbrev id="ABBRID0EVDGM" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.74–79, antennomeres IV – X moniliform; frontoclypeal horn small, but distinct in lateral view; three pairs of supraorbital setae present, outer posterior pair smaller than other two. 
<emphasis id="133E42C8D98DB65F26A04CE8B2C2B436" bold="true" italics="true">Pronotum</emphasis>
Size and shape variable: 
<abbrev id="ABBRID0E3DGM" xlink_title="Pronotal length, measured along midline">PL</abbrev>
/ 
<abbrev id="ABBRID0EAEGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.22–0.24, 
<abbrev id="ABBRID0EEEGM" xlink_title="Maximum pronotal width">PW</abbrev>
/ 
<abbrev id="ABBRID0EIEGM" xlink_title="Maximum elytral width">EW</abbrev>
= 0.78–0.87, 
<abbrev id="ABBRID0EMEGM" xlink_title="Pronotal basal width, measured at posterior angles and not including external denticles">PbW</abbrev>
/ 
<abbrev id="ABBRID0EQEGM" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.73–0.79. Sides either convergent or distinctly sinuate before posterior angles, which are either obtuse or nearly right. 
<emphasis id="FD43340AC4A6E4CC1D4E41EB1F11CF4F" bold="true" italics="true">Elytra</emphasis>
Variable in shape, either parallel sided and flat or slightly ovoid and convex. 
<abbrev id="ABBRID0EXEGM" xlink_title="Elytral length, measured from posterior edge of scutellum to apex of elytra">EL</abbrev>
/ 
<abbrev id="ABBRID0E2EGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.52–0.56. 
<emphasis id="57BEDC9A77DA6B31BCB85A840EE4C5E3" bold="true" italics="true">Legs</emphasis>
Profemora of males unmodified. First protarsomere of males dilated and spinose on inner margin, bearing adhesive setae ventrally; second protarsomere of males unmodified. Mesotrochanters of males unmodified. Metafemora of males variable, either slightly enlarged medially with patch of coarse microsculpture (Fig. 
<figureCitation id="DCA788F3B55337AC9B5A5285C258C0A6" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. SEM micrographs of male left metafemora of Anillinus species. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure12" httpUri="https://binary.pensoft.net/fig/1110351">12 D</figureCitation>
) or swollen and bearing a blunt tooth on posterior margin. Females without leg modifications. 
<emphasis id="27C26E065B1AC339D7E266937CB2217E" bold="true" italics="true">Abdominal ventrites</emphasis>
Unmodified in either sex. 
<emphasis id="0F8B15DBDDC1476C2032DB2F999D8AA0" bold="true" italics="true">Male genitalia</emphasis>
Aedeagus small (
<abbrev id="ABBRID0EMFGM" xlink_title="Ring sclerite length">RL</abbrev>
/ 
<abbrev id="ABBRID0EQFGM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.23). Median lobe (Fig. 
<figureCitation id="8E3FE1B79EFDDFBC33605D16FF1FEEA2" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure35" httpUri="https://binary.pensoft.net/fig/1110374">35 F</figureCitation>
) slightly rotated dorsally from plane of basal lobes, also curved towards left side in apical 1 / 2; slightly narrowed apically; ventral margin evenly curved, not expanded, without setae; dorsal margin sclerotized for ~ 2 / 3 its length; apex small, rounded. Internal sac covered in small scales, without spines; flagellum relatively long, not rotated dorsally, sinuate, abruptly narrowed apically, lightly sclerotized, and “ open ” laterally. Right paramere (Fig. 
<figureCitation id="7093192E4FD4AE82DD1240442B6CEC6F" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure35" httpUri="https://binary.pensoft.net/fig/1110374">35 E</figureCitation>
) small, broadly rounded, with four apical setae. Left paramere (Fig. 
<figureCitation id="C9FB009D734B21B9A894C5D9ECEA92A3" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Male genitalia of Anillinus species in the sinuaticollis group A – C Anillinus choestoea sp. nov. D – F Anillinus seneca sp. nov. G – I Anillinus mica sp. nov. J – L Anillinus micamicus sp. nov. Median lobe of aedeagus in right dorsolateral (C, I, L) or right lateral (F) aspect. Left parameres (A, D, G, J) and right parameres (B, E, H, K) in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure35" httpUri="https://binary.pensoft.net/fig/1110374">35 D</figureCitation>
) subtriangular, ventral margin with four apical setae of variable length. 
<emphasis id="B24694126055E54C420A887134008473" bold="true" italics="true">Female genitalia</emphasis>
Spermatheca long, abruptly expanded distally, stem with loose proximal coil, nearly straight before enlarged apex which is strongly curved (Fig. 
<figureCitation id="4768D4AADDAF75CBAC965392D0942A8D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 K</figureCitation>
). Spermathecal duct short, not coiled.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHGGM" type="distribution">
<paragraph id="4CF7C1B5ECF8000FBB7B9BD320DD8FB9">
<heading id="0FD77B05AC0018E27AADBDE5A2A9935D" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="497CC2021CC7354C4E815F00CB8C79D3">
Endemic to 
<collectingRegion id="09DEE3F00F6BE3A09E6D5B2F3C3B7CBE" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, known from Oconee and Anderson Counties on both sides of the former Seneca River (currently Lake Hartwell) from Seneca south to River Forks recreation area (Fig. 
<figureCitation id="5B87E607E73F5C33F27227DA6D7F30A6" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." figureDoi="10.3897/zookeys.1209.125897.figure36" httpUri="https://binary.pensoft.net/fig/1110375">36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERGGM" type="Sympatry">
<paragraph id="A95D5E6D18AAB3EB8F3D0E01BC749C0E">
<heading id="910BACEB9D21DE82DB5DF4B5C459A2C6" reason="title">Sympatry.</heading>
</paragraph>
<paragraph id="95A84A2FBCA3521E1612C814DA6D2553">
At the type locality, 
<taxonomicName id="946E96FDA8A3AFCF4F31747173E59B06" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="45DCB462F2FF6505C9A1B84DD8A6D105" italics="true">S. dunavani</emphasis>
</taxonomicName>
and a large 
<taxonomicName id="FD94A4AAD4E83931FE48805C839CF4B3">
<emphasis id="BE8B5C4949E7FD13EE3C1A8059A9E92F" italics="true">Anillinus</emphasis>
species
</taxonomicName>
belonging to the 
<taxonomicName id="42808AF41F7BA86CC7FCA8E63E05F3AD" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Troglanillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="valentinei">
<emphasis id="0142C792FCC401AEBC9784DE166FECA0" italics="true">valentinei</emphasis>
</taxonomicName>
group (
<taxonomicName id="6853D1C6A3ED5B96197D7516670E8AB6" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="5131A2CC4AED2FF2AC7743D671A94085" italics="true">Anillinus</emphasis>
sp.
</taxonomicName>
“ 
<collectingRegion id="4BBFC83B3BBBEA71FDC09C36A07D5337" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, Waldrop Stone ”) also occur. The other 
<collectingRegion id="EAB101062A4591E86CB85ADE20F01F8D" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
members of the 
<taxonomicName id="135ED7D5EAE84F4F8EC6F6813CEE8810" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="sinuaticollis">
<emphasis id="30EB1CB69779AA1FCFBC695546FB6DF8" italics="true">sinuaticollis</emphasis>
</taxonomicName>
group are apparently allopatric with respect to 
<taxonomicName id="4357E940A0734A71F3D47C97A73FD644" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seneca">
<emphasis id="56A760A289A61D49274EBBAB39A6EE0D" italics="true">A. seneca</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPIGM" type="natural history">
<paragraph id="CAD13EA6240EAB3F39DD797654CBA72A">
<heading id="D352D21C8B97067776D2B064FA839F94" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="70B49233755B4E56964B8D1999B89F34">
Members of this species are endogean, inhabiting mineral soil layers in sandy clay rich soils. They have been collected beneath embedded rocks and through Berlese extraction of soil and washed soil. In samples from the type locality, 
<tp_taxon-name id="91309D937A58AF3670919BAF4C888B28">
<tp_taxon-name-part id="3823E369E4A52CDE8065C0C797853D35" taxon-name-part-type="order">Laboulbeniales</tp_taxon-name-part>
</tp_taxon-name>
fungi were observed on the dorsal surface of pronotum and elytra of several females and on the abdominal apex in a male. Specimens were collected in January, February, March, May, October, and December.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1IGM" type="Species status justification">
<paragraph id="1F05C9AD7591CDDAF8246EBBD221A38C">
<heading id="D97C4B722EFB5F2AA41FBB945E6C0EE7" reason="title">Species status justification.</heading>
</paragraph>
<paragraph id="2CEB9378B7299862339DC9B9BEBD3F01">
The male genitalia are unique within the genus. DNA sequence data indicate that 
<taxonomicName id="9D3004E7EF3046A6E047BCFCA7F80311" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="seneca">
<emphasis id="888A29E48E48659C6F69648580A89A84" italics="true">A. seneca</emphasis>
</taxonomicName>
is sister to the pair of 
<taxonomicName id="D7B78237BCF7A2584FA10F35B91DDA8A" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mica">
<emphasis id="CBFD96C4BBB7D33034F9C70479E12BDF" italics="true">A. mica</emphasis>
</taxonomicName>
and 
<taxonomicName id="D356DE152AFDB3BD7B28DB35C9945150" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="A880B14AE43BCCCDB79EDAE5E8E3CC39" italics="true">A. micamicus</emphasis>
</taxonomicName>
, both of which differ markedly in male genitalic characters.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBKGM" type="Derivation of species name">
<paragraph id="7C4B1FCA388B7A07B418DB93F64A65E3">
<heading id="F70B2FE207724060439AC6EF8B2F6600" reason="title">Derivation of species name.</heading>
</paragraph>
<paragraph id="BD2B2D9D88352AE883F66690066BA8D4">A noun in apposition, named for the former Seneca River. The known localities for this species are all near the former course of this river, which was lost due to the construction of Lake Hartwell in the 1950 s.</paragraph>
</subSubSection>
<subSubSection id="SECID0EGKGM" type="notes">
<paragraph id="BA6342053FD54F2E9E0E83A7AB2141B2">
<heading id="0F0C31D985C98DCC5479D119A2381BE4" reason="title">Notes.</heading>
</paragraph>
<paragraph id="54BBA92638097735CD8B1E8ECE6B9B93">DNA sequences of individuals from the opposite side of the historic course of the Seneca River in Anderson Co. are divergent from Oconee Co. individuals. There are no differences in genitalic morphology, and the divergence is interpreted as recent intraspecific variation. Still, those Anderson Co. individuals are not made part of the type series, in case future study concludes they are specifically distinct.</paragraph>
<paragraph id="D6A402B61D88868E5370C647385C0981">
<title id="A81E419634048C1C4A5EFF43CAF48AA3">
<taxonomicName id="4AD655D6298EE9353ACF4BF9E147E316" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A484F76A56807C1B046DAB403AEBD504" italics="true">Anillinus</emphasis>
sp.
</taxonomicName>
“ 
<collectingRegion id="A3A7CA99B83C690709FF414112476E0C" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, Coon Branch ”
</title>
</paragraph>
<paragraph id="4FBE09F2EBBDCED177AC9D9EA95109A3">
Figs 
<figureCitation id="17A2D7870173062C850420D13E4A9BA4" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 G</figureCitation>
, 
<figureCitation id="2A345C97A35DC76E216C935FC2FD11E2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." figureDoi="10.3897/zookeys.1209.125897.figure25" httpUri="https://binary.pensoft.net/fig/1110364">25 B</figureCitation>
, 
<figureCitation id="86AD6D588032701B657E8798644F9708" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Distribution map of sinuaticollis - group species of Anillinus in South Carolina. Data are from Harden (2024). Blue lines show modern and pre-inundation courses of the Tugaloo River and Seneca River and their major tributaries. The confluence of the two rivers becomes the Savannah River." figureDoi="10.3897/zookeys.1209.125897.figure36" httpUri="https://binary.pensoft.net/fig/1110375">36</figureCitation>
</paragraph>
<paragraph id="A793D32069340CA7E43060D151010EFA">
<emphasis id="0B003C784003B6E2EE0F9184B4E207BA" bold="true">
<collectingCounty id="C608CAD4E3AFEF52F69B9B83EBDC809B">Material</collectingCounty>
examined (
<emphasis id="9DB076EA4DE5FB790B0738D84BF86EF4" bold="true" italics="true">n</emphasis>
= 3). 
<collectingCountry id="1803A425654B8D64BC6CBD36CF73A0CF" name="United States of America">USA</collectingCountry>
</emphasis>
• 
<emphasis id="39CBFFF2668E3BC0D0C183F383456E55" bold="true">
<collectingRegion id="736BF358A1E293FA1856B812933F8BE3" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<emphasis id="FDF20A48FF15978E432B561CD6807602" bold="true">
<collectingCounty id="6E41A8DC276EAE84D8E41E7AE54F5845">Oconee Co.</collectingCounty>
</emphasis>
; 
<location id="B3B6C8B496FE4B51E05994416E8D308A" LSID="urn:lsid:plazi:treatment:26CBA40789005DD7BEC91DD96238EE9E:B3B6C8B496FE4B51E05994416E8D308A" country="United States of America" county="Oconee Co." name="Coon Branch Natural Area" stateProvince="South Carolina">Coon Branch Natural Area</location>
; • 
<specimenCount id="7F1365E341DBF9143FCF08AF36FF05AB" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="FB27BD24C05B2E8EAF1C101CC41F0880" count="1" type="female">1 ♀</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESLGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.000000,35.020000]}">
<geoCoordinate id="1379A76270608A2F80BC7BEA7D9ADC59" degrees="35.020" direction="north" orientation="latitude" precision="55" value="35.02">35.020</geoCoordinate>
, 
<geoCoordinate id="E308FF2EBE8B39CA18CFB16B6613DF06" degrees="83.000" direction="west" orientation="longitude" precision="55" value="-83.0">- 83.000</geoCoordinate>
</named-content>
; 
<date id="8B181821AC10E4DA1E4E74369E4B8D08" value="2018-06-21">
<collectingDate id="E19EDF37E68480511C4666FFF1D8CD31" value="2018-06-21">21 Jun. 2018</collectingDate>
</date>
; 
<collectorName id="D68531236D5AA1C944D46871E60E121C">B. Owens</collectorName>
and 
<collectorName id="03CCFA521B68F16AD43DCD7F6DC20AA6">C. Carlton</collectorName>
leg.; soil flotation; 
<specimenCode id="8A06BDEA199885642F749FC8997A59A2" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168231</specimenCode>
and 
<specimenCode id="1D8C62EAE1BDACABD08D72AC58023CC2" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000168232</specimenCode>
, CWH- 102 and CWH- 178; 
<named-content id="49254A0334FF105C787B8A55BEF0BCD8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="8CD9AAA3F6942729B5DC5F6FCD097D7F" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
• 
<specimenCount id="8D95BE6D11939DE779398B120C1E12CE" count="1" type="male">1 ♂</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5LGM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.005000,35.025600]}">
<geoCoordinate id="ACE6678F576663930961C1E3F88004D5" degrees="35.0256" direction="north" orientation="latitude" precision="5" value="35.0256">35.0256</geoCoordinate>
, 
<geoCoordinate id="B8706B4DA6841700B3EBFD52D0140447" degrees="83.0050" direction="west" orientation="longitude" precision="5" value="-83.005">- 83.0050</geoCoordinate>
</named-content>
; 
<date id="58A40760823F1ACA95363BCB50B62F28" value="2021-10-02">
<collectingDate id="CECB79DBBE123EFB5CFF3BAD2DAB364F" value="2021-10-02">2 Oct. 2021</collectingDate>
</date>
; 
<collectorName id="18552D50C7643A320474022C8130F3EE">C. W. Harden</collectorName>
leg.; on soil under large rock.; 
<abbrev id="ABBRID0EDMGM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="E35C07B9268AA79A778CDF70C4F8DEE7">CWHc</collectionCode>
</abbrev>
.
</paragraph>
<paragraph id="FF129DD6E4D23A4F776B2FE879DD3898">
GenBank: 
<ext-link id="C1D3B95FFDCC6BE0EF94A63DECFDE108" ext-link-type="gen" xlink_href="OR853183" xlink_type="simple">OR 853183</ext-link>
, 
<ext-link id="F617AA25B032C0A858E1974DC9F88F94" ext-link-type="gen" xlink_href="OR839228" xlink_type="simple">OR 839228</ext-link>
, 
<ext-link id="41457F1DFA9CCD5F3D5186E3B88B7614" ext-link-type="gen" xlink_href="OR839449" xlink_type="simple">OR 839449</ext-link>
, 
<ext-link id="054DF6DE18E798683329FB4C1B1F2E75" ext-link-type="gen" xlink_href="OR837811" xlink_type="simple">OR 837811</ext-link>
, 
<ext-link id="E440B2CAF12338A511C0931276EE30CB" ext-link-type="gen" xlink_href="OR837968" xlink_type="simple">OR 837968</ext-link>
, 
<ext-link id="4548E29C4D07E0C5E4182B8E3ED32C24" ext-link-type="gen" xlink_href="OR838156" xlink_type="simple">OR 838156</ext-link>
, 
<ext-link id="97FAFECE9E5F9E72E50315C6A1999620" ext-link-type="gen" xlink_href="OR839507" xlink_type="simple">OR 839507</ext-link>
, 
<ext-link id="4253911EBF75A236C5177117C498D004" ext-link-type="gen" xlink_href="OR837843" xlink_type="simple">OR 837843</ext-link>
, 
<ext-link id="F93E5B5F599F3809BFD61E5092F12D02" ext-link-type="gen" xlink_href="OR837990" xlink_type="simple">OR 837990</ext-link>
, 
<ext-link id="D2135BB443749A7B9EBEF314E0F94316" ext-link-type="gen" xlink_href="OR838176" xlink_type="simple">OR 838176</ext-link>
.
</paragraph>
<paragraph id="DCFB09ADD312AC12576C01B3E4E70FB8">
<emphasis id="DD4065A08C0DA11618F425918DF40EBB" bold="true">Diagnosis.</emphasis>
The male genitalia are similar to those of 
<taxonomicName id="9A6EAF667E7CB23939D012960AB77C98" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mica">
<emphasis id="538FE6B0E427134BF74493E73A5546F3" italics="true">A. mica</emphasis>
</taxonomicName>
and 
<taxonomicName id="E7ECDDF1B0D5B9AE93E91BDEAB0EC3AC" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="87301154A8C0E3604FF8AB37E060A275" italics="true">A. micamicus</emphasis>
</taxonomicName>
, but differ in several characters: the shape of the median lobe is more elongate, with the ventral margin less curved and abruptly straightened at apex, which is obtusely angulate; the internal sac has a flagellum that is similar to 
<taxonomicName id="66D0205CAF5ED914AEE33067F5B62BA3" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="5F1CC03A54B58A7C4223931F8D8F2F60" italics="true">A. micamicus</emphasis>
</taxonomicName>
, and has a single group of approximately 12 elongate, well-sclerotized spines on the left side; there is a sclerotized saddle like structure near the ostium, as in 
<taxonomicName id="064D81F652C09B61A2CC1A87B0A7D721" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mica">
<emphasis id="D4A63B3B5385CD8EE2E796A6B7826361" italics="true">A. mica</emphasis>
</taxonomicName>
, but it is situated dorsal to the level of the flagellum; a pair of parallel rows of small sclerotized teeth run along the left side of the ostium, appearing as a pair of dark curved lines below the saddle like structure. The right paramere is encased in a membranous sheath as in 
<taxonomicName id="9CF2B3CC776B21FED55BF5EDE452088D" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mica">
<emphasis id="590E1872B3F1FE490987A9D554916000" italics="true">A. mica</emphasis>
</taxonomicName>
and 
<taxonomicName id="9003EDD86B2628DC95103EE13B050C57" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="7301D8CA42E17762F0E9F2472B6E05FA" italics="true">A. micamicus</emphasis>
</taxonomicName>
. The left paramere is shaped similar to that of 
<taxonomicName id="1D7717505C405E0F7DFB28F790085B41" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="4DECE99225C876269E6D15724E5D2C33" italics="true">A. micamicus</emphasis>
</taxonomicName>
, but the inner two setae on the apical margin are much shorter than the outer two, which are elongate but not as much as in 
<taxonomicName id="6277CF0D50C047A3EAC237B7E8294BB2" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="mica">
<emphasis id="B1664B7E2FC51F917E4FC2043DACD65D" italics="true">A. mica</emphasis>
</taxonomicName>
and 
<taxonomicName id="D0F4E5B70360E868D60DCF1A814B155C" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="DE917167E6D4073F636E47AC2606DB7F" italics="true">A. micamicus</emphasis>
</taxonomicName>
. The spermatheca (Fig. 
<figureCitation id="AFB4EBB7AFA871FF772A44A05B99FF5D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 G</figureCitation>
) has a stem that is coiled evenly in a spiral; it is evenly enlarged distally. The duct has a single sharp bend just behind the junction with the spermatheca, and is otherwise straight and relatively wide, without coils. The bursa has a few lightly sclerotized regions.
</paragraph>
<paragraph id="1BA121EFCBBB9B08055FD64D3BDEBB2A">
<emphasis id="137D9FC57DEF0E804832CA2D9A7AACC8" bold="true">Notes.</emphasis>
The data from morphology and DNA sequences indicate that these individuals might represent a species distinct from 
<taxonomicName id="B6975776B480160C64C8B90BAD077AE9" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="C596EBDE60A41F307E758C413ACD4614" italics="true">A. micamicus</emphasis>
</taxonomicName>
. However, more sampling in the intervening area between Coon Branch and the localities of 
<taxonomicName id="3DB5E172FEC22AE6EC80E78F2C1855A5" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="micamicus">
<emphasis id="EA8B2A1E6FB70D1825533B66DB8A57F9" italics="true">A. micamicus</emphasis>
</taxonomicName>
will be necessary to test this hypothesis.
</paragraph>
<paragraph id="EABD713146C1D55A2DBB97FFB60D1EFA">
<title id="621729B0E108C2A9136689FF3D49712D">
<taxonomicName id="1BBC2021558302B94FA9EC5C88D8CFB5" authorityName="Sokolov &amp; Carlton" authorityYear="2004" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="langdoni">
‘ 
<emphasis id="BD91182B7113DF5CF47776C48B9812D5" italics="true">langdoni</emphasis>
</taxonomicName>
group’
</title>
</paragraph>
</subSubSection>
</treatment>
</document>