treatments-xml/data/F3/BB/B5/F3BBB51415CF5E45B1076667E97AEF1B.xml

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<mods:title id="BAB9A83E7C09F35A12ED6738B4B1C921">Four new Cyclopina (Copepoda, Cyclopinidae) from South Korea</mods:title>
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<mods:namePart id="591FC5C958A3D3E743ED66097328492A">Karanovic, Tomislav</mods:namePart>
<mods:affiliation id="D049469829280EC655E7DF58273F9507">Hanyang University, College of Natural Sciences, Department of Life Science, Seoul 04763, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="6685E06528D8CBA5F57387B82CF9900E" type="email">tomislav.karanovic@gmail.com</mods:nameIdentifier>
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<mods:date id="DC3A38B6970BE7F0CC7CDB8097079C8F">2020</mods:date>
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<mods:identifier id="BB1C3D1FADDBCD2E3A590C9BC6A46A13" type="DOI">http://dx.doi.org/10.3897/zookeys.992.54856</mods:identifier>
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<treatment id="F3BBB51415CF5E45B1076667E97AEF1B" ID-GBIF-Taxon="169056257" LSID="urn:lsid:zoobank.org:act:9ACBBC99-22DF-4150-8938-666F7099CC05" httpUri="http://treatment.plazi.org/id/F3BBB51415CF5E45B1076667E97AEF1B" lastPageNumber="59" pageId="0" pageNumber="59" scope_class="Copepoda" scope_family="Cyclopinidae">
<subSubSection id="CDA07B137E269676FFB0FDAFDCBA1F66" pageId="0" pageNumber="59" type="nomenclature">
<paragraph id="51BBEB46B068CFD0F2AB3F23D55EB0E8" pageId="0" pageNumber="59">
<taxonomicName id="C163DFC557AB3C6D4A82C334B43A699D" ID-CoL="7ZG5X" LSID="F3BBB514-15CF-5E45-B107-6667E97AEF1B" authority="Karanovic, 2020" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana" status="sp. nov.">Cyclopina koreana</taxonomicName>
<taxonomicNameLabel id="74626F2F942CB32E93385CEEAA888C82" pageId="0" pageNumber="59">sp. nov.</taxonomicNameLabel>
<figureCitation id="EE2D6BA0BAB683EFA20D5345776459A1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">Figures 7</figureCitation>
<figureCitation id="B729C96D0E5A1ED05807A83D63113A33" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">, 8</figureCitation>
<figureCitation id="14D518AEF4A365F5E448CDFF775D2088" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">, 9</figureCitation>
<figureCitation id="31AF4E8AD426406336E8A78AAF99D97A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">, 10</figureCitation>
<figureCitation id="13343D237D271BDE7EB8E187CB2BCA48" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cyclopina koreana sp. nov., line drawings A-E holotype female F-H allotype male: A third endopodal segment of first swimming leg B third exopodal segment of first swimming leg C third exopodal segment of second swimming leg D third endopodal segment of second swimming leg E fourth swimming leg F antennula G third endopodal segment of fourth swimming leg H third exopodal segment of fourth swimming leg." figureDoi="10.3897/zookeys.992.54856.figure11" httpUri="https://binary.pensoft.net/fig/474559" pageId="0" pageNumber="59">, 11</figureCitation>
<figureCitation id="E0F624911903F6ACB0163BC582BDC7B4" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Light photographs of four new species (not to scale) A Cyclopina busanensis sp. nov., four females (two ovigerous) B Cyclopina koreana sp. nov., three females and four males C Cyclopina curtijeju sp. nov., two females D Cyclopina wido sp. nov., two females (one ovigerous) and one male." figureDoi="10.3897/zookeys.992.54856.figure17" httpUri="https://binary.pensoft.net/fig/474565" pageId="0" pageNumber="59">, 17B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DAD0C8E79CA1A5661FE0A5FF5E73A236" pageId="0" pageNumber="59" type="type locality">
<paragraph id="685530994072B229CDDFFDDE15C40029" pageId="0" pageNumber="59">Type locality.</paragraph>
<paragraph id="FFF58DBD8EB6C3B6523A4F5513D18444" pageId="0" pageNumber="59">
South Korea, East Coast, Gangneung, small beach, intertidal sand, 
<geoCoordinate id="DB5415808E92ACD101DFB897FE9C2FC5" degrees="37" direction="north" minutes="47.824" orientation="latitude" precision="1" value="37.797066">37°47.824'N</geoCoordinate>
, 
<geoCoordinate id="E1263EF5484F271DA6716F8ACF12D71E" degrees="128" direction="east" minutes="55.085" orientation="longitude" precision="1" value="128.91809">128°55.085'E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="7C6A6C78B4DFE241C5194FF1983A04DF" pageId="0" pageNumber="59" type="materials_examined">
<paragraph id="2086EF085B1089CCCF477A551C387C27" pageId="0" pageNumber="59">Specimens examined.</paragraph>
<paragraph id="D83589F0C89A3077D7D7CA600C3B82DB" pageId="0" pageNumber="59">
<emphasis id="08313536AC2FBA3CBDCFA79FBAC8FFE4" bold="true" italics="true" pageId="0" pageNumber="59">Holotype</emphasis>
female dissected on one slide, collected from the type locality, 29 March 2013, leg. T. Karanovic.
</paragraph>
<paragraph id="976CE1787DACEEC549DEAD0B900D5B87" pageId="0" pageNumber="59">
<emphasis id="5BB4C41D498E057CE53B621BA0106067" bold="true" italics="true" pageId="0" pageNumber="59">Paratypes</emphasis>
: two males and one female dissected on one slide each; three males, two females, and four copepodids in alcohol; one male and two females on one SEM stub (together with specimens of other three species described here; row no. 4); all collected from the type locality, 29 March 2013, leg. T. Karanovic.
</paragraph>
</subSubSection>
<subSubSection id="D19C2ADF349E55E546CD2951BCD72417" pageId="0" pageNumber="59" type="etymology">
<paragraph id="EB9CB605085EE357A58B32F1E51FE312" pageId="0" pageNumber="59">Etymology.</paragraph>
<paragraph id="EEDFD449FAD042590C439A237EDF87CE" pageId="0" pageNumber="59">The species name refers to South Korea. It is an adjective, agreeing in gender with the feminine genus name.</paragraph>
<caption id="CE7B9EF6581E6A8A11B9D9D337D43A0F" doi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59" start="Figure 7" startId="F7">
<paragraph id="3071244DAA61CAD0E5C518E4ED211DB0" pageId="0" pageNumber="59">
<emphasis id="EA8156D0196BCE2895F229FC67F2D781" bold="true" pageId="0" pageNumber="59">Figure 7.</emphasis>
<taxonomicName id="F58A011E363B4087AFD1BC9423E6D252" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="E480A4EDBE27A5B08DD579A8E842AAFB" italics="true" pageId="0" pageNumber="59">Cyclopina koreana</emphasis>
</taxonomicName>
sp. nov., paratype female 1, SEM photographs, all in dorsal view 
<emphasis id="8874ECBEC4DE850F1DAF97DC30FAFE59" bold="true" pageId="0" pageNumber="59">A</emphasis>
habitus 
<emphasis id="E875928CE00B315A07BC6AB5F693C98A" bold="true" pageId="0" pageNumber="59">B</emphasis>
second pedigerous somite 
<emphasis id="1098BCF7955FBDD57EF125865DAB58CF" bold="true" pageId="0" pageNumber="59">C</emphasis>
third pedigerous somite 
<emphasis id="400F8E100C1EA39B84802C593E84D3F6" bold="true" pageId="0" pageNumber="59">D</emphasis>
first urosomite 
<emphasis id="6B5955748FCF99C8F73BA5578116B94E" bold="true" pageId="0" pageNumber="59">E</emphasis>
genital double-somite and fourth urosomite 
<emphasis id="322369583B8D3DB31049794A509E0A14" bold="true" pageId="0" pageNumber="59">F</emphasis>
anterior medial pore on genital double-somite 
<emphasis id="E840E545D91185969F83F985F46C47BF" bold="true" pageId="0" pageNumber="59">G</emphasis>
fifth and sixth urosomites 
<emphasis id="A2D8ECC8D506CDDC99B4241985877E22" bold="true" pageId="0" pageNumber="59">H</emphasis>
caudal ramus.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6F44CF99185E42CCC35FF2844B155BB1" pageId="0" pageNumber="59" type="description">
<paragraph id="554FD40B0CAD415C42D5BA48219B8648" pageId="0" pageNumber="59">Description.</paragraph>
<paragraph id="8AC17BD10FCD0B48ACA76A0AF388F049" pageId="0" pageNumber="59">
<emphasis id="4A79CE592152D3C9ADD9592DF397EEF6" bold="true" pageId="0" pageNumber="59">Female</emphasis>
(based on holotype and three paratypes). 
<emphasis id="156F934DA27DBB85CF51730DFC04384A" bold="true" italics="true" pageId="0" pageNumber="59">Body length</emphasis>
from 620 to 635 
<normalizedToken id="B496F3DBEEB49A4B0FD64F96730C863B" originalValue="μm">μm</normalizedToken>
. 
<emphasis id="955C81E5F96060B3EE99C3B5BC327F2A" bold="true" italics="true" pageId="0" pageNumber="59">Colour</emphasis>
of preserved specimens yellowish, nauplius eye not visible (Fig. 
<figureCitation id="8CED4E18B7B0C7AE1467BE4064A5C446" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Light photographs of four new species (not to scale) A Cyclopina busanensis sp. nov., four females (two ovigerous) B Cyclopina koreana sp. nov., three females and four males C Cyclopina curtijeju sp. nov., two females D Cyclopina wido sp. nov., two females (one ovigerous) and one male." figureDoi="10.3897/zookeys.992.54856.figure17" httpUri="https://binary.pensoft.net/fig/474565" pageId="0" pageNumber="59">17B</figureCitation>
). Integument on all somites (Figs 
<figureCitation id="40A5817D3B13AC2DA7C4CC25C2A8B867" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7</figureCitation>
, 
<figureCitation id="4AE11DF57005E9BE73FEBDDFB543BD20" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8</figureCitation>
) smooth, with light bacterial cover, cuticular pores on all somites, spinules only on genital somite and caudal rami, and sensilla on all but penultimate somite; hyaline fringes of prosomites smooth, of urosomites serrated. 
<emphasis id="AE4209BADFF237FC32717827D1FE5C07" bold="true" italics="true" pageId="0" pageNumber="59">Habitus</emphasis>
(Fig. 
<figureCitation id="D5B4A79DD09015A12A83B9B5A8EE038D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7A</figureCitation>
) ca. 2.6 
<normalizedToken id="B9BD4A862A5F8FF6958471F009A6009A" originalValue="×">x</normalizedToken>
as long as wide in dorsal view, with pronounced distinction between prosome and urosome; prosome ovoid, ca. 1.5 
<normalizedToken id="50C1BF0E32461561ED9CCD715B96FC35" originalValue="×">x</normalizedToken>
as long as wide in dorsal view, nearly 1.4 
<normalizedToken id="3ABABF1B0DFBDCFF9BBCB40120EBC595" originalValue="×">x</normalizedToken>
as long and 2.6 
<normalizedToken id="13D5A361733E9646B51FE1ACF705F283" originalValue="×">x</normalizedToken>
as wide as urosome, its greatest width at posterior end of first pedigerous somite; urosome nearly cylindrical, ca. 3 
<normalizedToken id="064DB5347DA03425BE364936F0D23EB2" originalValue="×">x</normalizedToken>
as long as wide, its greatest width at posterior end of fifth pedigerous somite (first urosomite). 
<emphasis id="33E9BCE510EBE7C943392688B9D6B5EC" bold="true" italics="true" pageId="0" pageNumber="59">First pedigerous somite</emphasis>
(Fig. 
<figureCitation id="B095F7B5063B8CFC71EB319F184BAB07" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7A</figureCitation>
) not fused to cephalothorax, but its tergites partly covered with posterior extensions of cephalothoracic shield as in 
<taxonomicName id="BFFDA0594DB7BA099CB8CB4180F16444" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="66F04979599BD4312B1AB1671459ED46" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
. 
<emphasis id="D9315F22730819FF99F2D8F1D0D8C1D4" bold="true" italics="true" pageId="0" pageNumber="59">Cephalothorax</emphasis>
(Fig. 
<figureCitation id="D4E0D6F5F1AFF6482A0C6CC9D51EE7AA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7A</figureCitation>
) broader in anterior part than in 
<taxonomicName id="05C101B1126E6022D77CACFC53C4626C" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="9533435D18ADA466B15F22DCD80FB0A1" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, ca. 1.2 
<normalizedToken id="4507C1F3EF8432F4A679903FDBDDA06E" originalValue="×">x</normalizedToken>
as long as wide, and twice as long as free prosomites combined. Second to fourth free prosomites (Figs 
<figureCitation id="ACBDB441473B40AD3A2C43CC8132ECD7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7A-C</figureCitation>
, 
<figureCitation id="1255494E0C63DCC6593F75CB2444C2F7" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8F</figureCitation>
) progressively shorter and narrower towards posterior end, and with fewer cuticular organs; not many prosomal cuticular organs clearly homologous to those in previous species (compare Figs 
<figureCitation id="6F8185397A7EB4D61EC1E9FD2A7523DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cyclopina busanensis sp. nov., paratype female 1, SEM photographs, all in dorsal view A tergite of fourth pedigerous somite B fifth leg C genital double-somite (= fused second and third urosomites) D sixth leg E detail of posterior part of genital double-somite F fourth and fifth urosomites G sixth urosomite (= anal somite) and caudal ramus H detail of sixth urosomite." figureDoi="10.3897/zookeys.992.54856.figure2" httpUri="https://binary.pensoft.net/fig/474550" pageId="0" pageNumber="59">2A</figureCitation>
, 
<figureCitation id="54A3741F043099E74BE2AC4EA689D334" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8F</figureCitation>
), except dorsal medial pores and several posterior sensilla (Fig. 
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).
</paragraph>
<caption id="304AF5E58D4748C77801ADC5F318BA35" doi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59" start="Figure 8" startId="F8">
<paragraph id="A9FAF83AE3C50E463FEB58E71CADCCC4" pageId="0" pageNumber="59">
<emphasis id="0A72114B6153135811FAB4A037EE7D0E" bold="true" pageId="0" pageNumber="59">Figure 8.</emphasis>
<taxonomicName id="970672DB61F7DB415E6D87C704E7264B" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="E5C4BA12AB52B9FEB46DC74517C3EDAC" italics="true" pageId="0" pageNumber="59">Cyclopina koreana</emphasis>
</taxonomicName>
sp. nov., SEM photographs 
<emphasis id="0D9BD95E73D64DD62EFEA5B232AB7B3E" bold="true" pageId="0" pageNumber="59">A-E</emphasis>
paratype male 1, ventral view 
<emphasis id="3E1184C57827C7B831FE0B7C05CC5FDF" bold="true" pageId="0" pageNumber="59">F-H</emphasis>
paratype female 2, lateral view: 
<emphasis id="F7546ADFAC388FFEAF76B148C71D25A4" bold="true" pageId="0" pageNumber="59">A</emphasis>
habitus 
<emphasis id="4E8454166EF472E97A1A658910EED096" bold="true" pageId="0" pageNumber="59">B</emphasis>
distal part of antennula 
<emphasis id="6546B22066B7CBC291E55C30DC6087B7" bold="true" pageId="0" pageNumber="59">C</emphasis>
fourth swimming leg 
<emphasis id="79DE72C4E98BBF284B38AD5A758C056D" bold="true" pageId="0" pageNumber="59">D</emphasis>
fourth and fifth urosomites 
<emphasis id="FDB9982D2B83D2A5FEF277396D4CB9A8" bold="true" pageId="0" pageNumber="59">E</emphasis>
sixth urosomite 
<emphasis id="AF8E510F00B02DCAA1D978326C4115CC" bold="true" pageId="0" pageNumber="59">F</emphasis>
tergite of fourth pedigerous somite 
<emphasis id="6A8D8A276E87428C59E2E959DCEBBF8B" bold="true" pageId="0" pageNumber="59">G</emphasis>
anterior part of urosomite, with fifth and sixth legs 
<emphasis id="5853D9E53EA9F3A3A8257CA37D0C1533" bold="true" pageId="0" pageNumber="59">H</emphasis>
fifth and sixth urosomites.
</paragraph>
</caption>
<paragraph id="778A363F434D9112069AA59A08AA8325" pageId="0" pageNumber="59">
<emphasis id="F5701C0B03DF2C2BE8E7BE535B51A772" bold="true" italics="true" pageId="0" pageNumber="59">First urosomite</emphasis>
(Figs 
<figureCitation id="3869DB82819351E97F547130164C5B7C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7D</figureCitation>
, 
<figureCitation id="6DBFC5591E36AFBD2EC6C82B3FD9D783" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8G</figureCitation>
, 
<figureCitation id="BC3A801C1BC4F940FC8B591F7EDF99CC" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) short, slightly laterally expanded in posterior part, with four dorsal sensilla and single dorsal medial pore.
</paragraph>
<caption id="C91E164A432F0A84FBE8AB19E3C8335A" doi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59" start="Figure 9" startId="F9">
<paragraph id="E740BC607192E53854ACC9781A959FC5" pageId="0" pageNumber="59">
<emphasis id="533E945FF967977358109E5F20B7C5E9" bold="true" pageId="0" pageNumber="59">Figure 9.</emphasis>
<taxonomicName id="5D8099ACAA3D2A8C9601734F781BC536" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="CBD557131876F86F9F0AB4C18539DE27" italics="true" pageId="0" pageNumber="59">Cyclopina koreana</emphasis>
</taxonomicName>
sp. nov., line drawings 
<emphasis id="587F6601D99D3DF23D3963823719CB09" bold="true" pageId="0" pageNumber="59">A</emphasis>
holotype female, urosome, ventral view 
<emphasis id="F7689709BBC14A5874205163C287760A" bold="true" pageId="0" pageNumber="59">B</emphasis>
holotype female, antennula 
<emphasis id="6928960546A96968582601ADEB101555" bold="true" pageId="0" pageNumber="59">C</emphasis>
allotype male, urosome, ventral view.
</paragraph>
</caption>
<paragraph id="D35F13A4E5FDDC0D8A9DD276B3F6A4F0" pageId="0" pageNumber="59">
<emphasis id="643646C9BA38BC6AB512D5F89D8B4972" bold="true" italics="true" pageId="0" pageNumber="59">Genital double-somite</emphasis>
(Figs 
<figureCitation id="AB15825E79B7BB9C1FA6FACA24ABEBFF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7E, F</figureCitation>
, 
<figureCitation id="F81C336793F5EB6C6B4029FCD2EFE858" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8G</figureCitation>
, 
<figureCitation id="C12CDEA7055F477ECD40702AC0CD2711" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) ca. 0.9 
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as long as wide in dorsal view, laterally expanded anterior part only ca. 1.1 
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as wide as posterior margin; sensilla and pores as in 
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<emphasis id="6001601DDEA8E254BED65B98BC8945B4" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
. Copulatory pore, copulatory duct, seminal receptacles, oviducts, and genital apertures as in 
<taxonomicName id="D1574A2C9C63D11B255E6222435A258E" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="C5BC9EBCB8F0C2BCEB6DDB7E08099403" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, except first part of copulatory duct slightly wider. Fourth urosomite (Figs 
<figureCitation id="FD9665CD8EE0A4BDD773CF00958B5A9D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7E</figureCitation>
, 
<figureCitation id="58EAAA19290A9DC559D28E4617766243" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) ca. 0.6 
<normalizedToken id="08F058875AF9CBC15754BCCDFC1FFD61" originalValue="×">x</normalizedToken>
as long as genital double-somite, with sensilla and pores as in 
<taxonomicName id="F7EB7495F6512ABA681C41C8424E3540" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="EB3560C8F4EF4A6F082C87C5A2EF4248" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
. Fifth urosomite (Figs 
<figureCitation id="EA4AEFA3C644C88EC90DB8DE80731E16" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7G</figureCitation>
, 
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, 
<figureCitation id="CA7E27C3B4477B8B26EE1C3153E9285A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) 0.7 
<normalizedToken id="A9C05EF1D3A31B8982AF6A8CF2AD5F00" originalValue="×">x</normalizedToken>
as long as fourth urosomite, with medial dorsal pore and one pair of widely spaced ventral pores as in 
<taxonomicName id="DE1368D0DB6ADEAA29969E24B8F1AADA" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="43F084AE6D5B9171EFF9858AEB6E6A71" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
. Sixth urosomite (Figs 
<figureCitation id="FD4B1496A98EDC193494AAFC9B6E5311" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7G</figureCitation>
, 
<figureCitation id="AF763CDA6B36BF76C41B447C61DABD4F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8H</figureCitation>
, 
<figureCitation id="041BD3224B8694A8A5C2B04D7DF26D3A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) 1.2 
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as long as fifth urosomite, with one pair of dorsal sensilla, two pairs of dorsal pores, and single pair of ventral pores; no spinules on fringes of anal sinus; anal operculum smooth, short, broad, slightly convex, situated in first third, represents 62% of 
<normalizedToken id="EB9A43883E7130EF9D4EF1E27AC40744" originalValue="somite’s">somite's</normalizedToken>
width.
</paragraph>
<paragraph id="251CC05DBA44EE6246CDFAA98BA9697F" pageId="0" pageNumber="59">
<emphasis id="F14E91936BF67002149031E73622BF68" bold="true" italics="true" pageId="0" pageNumber="59">Caudal rami</emphasis>
(Figs 
<figureCitation id="77FCEB6F8401FBD3BA3066332171A863" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7H</figureCitation>
, 
<figureCitation id="1C23404B6DB0052523BA0A5FDFF7BAF0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8H</figureCitation>
, 
<figureCitation id="D796B79C7974B60D7824450510923301" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9A</figureCitation>
) cylindrical, ca. 3.5 
<normalizedToken id="B50F8644F5B5638AA115FAAEA78A72F4" originalValue="×">x</normalizedToken>
as long as wide and 1.5 
<normalizedToken id="E068BB3B66ADC5BF6590D6F88C1F67BD" originalValue="×">x</normalizedToken>
as long as anal somite, very widely spaced on anal somite, diverging posteriorly; armed as in 
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<emphasis id="5924858FABB85F95FC139D1DC025B5B8" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
; ornamented with single sensilla near proximal lateral seta, row of small spinules at base of proximal lateral seta, and posterior ventral row of spinules. Proximal lateral seta inserted at ca. two fifths of 
<normalizedToken id="DF1F9A06D6637779BC843F538DE03956" originalValue="ramus’">ramus'</normalizedToken>
length; medial terminal seta nearly 0.9 
<normalizedToken id="19E29AA6280EF96DA9F01FFF6323EEDE" originalValue="×">x</normalizedToken>
as long as caudal ramus, 1.6 
<normalizedToken id="35646A9907BF179DDC93772044615911" originalValue="×">x</normalizedToken>
as long as lateral terminal seta, 0.8 
<normalizedToken id="35B3B9B86D065A17F10253EAF2549AC3" originalValue="×">x</normalizedToken>
as long as dorsal seta, and 2.3 
<normalizedToken id="68462813A6156F3F789FCABE07AC3280" originalValue="×">x</normalizedToken>
as long as proximal lateral seta.
</paragraph>
<paragraph id="7158140D9436B90240C8790FFCC0F88F" pageId="0" pageNumber="59">
<emphasis id="C3A8430816B00EB7C269E49FD5535C5F" bold="true" italics="true" pageId="0" pageNumber="59">Antennula</emphasis>
(Fig. 
<figureCitation id="5C2BEFB5D70A8930E518569190907353" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9B</figureCitation>
) segmentation and most armature as in 
<taxonomicName id="C9413F4DFDA16E306EFCA83E90312D63" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="D658F6A37EF3E3080FA3743ECC7E33D2" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but proximal half stouter and distal half slenderer; armature formula 3.6.8.4.5.6.4.2.2.7+ae; apical aesthetasc significantly shorter than in 
<taxonomicName id="9A1B7C4D91003A8271F86DDA2EED33B5" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="AD532756B4ECBA297A7BCBE8DDF11A84" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
and fifth segment with two short setae; sixth segment longest, ca. 3 
<normalizedToken id="11243D70EA1B5441D60B85567F36B9A9" originalValue="×">x</normalizedToken>
as long as wide, and nearly 0.9 
<normalizedToken id="33F21A22C7E33989E0E39168D19C1A6F" originalValue="×">x</normalizedToken>
as long as subsequent four segments combined; tenth segment nearly twice as long as wide.
</paragraph>
<paragraph id="854E3B22D648CA845C396CB87769F30B" pageId="0" pageNumber="59">
<emphasis id="F59781793C20FA7014FC31BCE4F91F00" bold="true" italics="true" pageId="0" pageNumber="59">Antenna</emphasis>
(Fig. 
<figureCitation id="1BD8A076DA412718C6EDB009A1C4DB0D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10A</figureCitation>
) as in 
<taxonomicName id="4615C5A0FB446E9307C585C179745794" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="925B824115D57232D5268DACA88D452B" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but another small exopodal seta present and second endopodal segment slightly longer.
</paragraph>
<caption id="F1665B74CE65B9F505C33C86E40DD69E" doi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59" start="Figure 10" startId="F10">
<paragraph id="69C3157B5D8F389B584207FFF5040CFE" pageId="0" pageNumber="59">
<emphasis id="8412C1C1854B6EF7CA7A8F8285340E3E" bold="true" pageId="0" pageNumber="59">Figure 10.</emphasis>
<taxonomicName id="BF41506ADC1BDAD3C990EC15F815F2EE" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="751DAE4D18895095D676FA21D9810FA9" italics="true" pageId="0" pageNumber="59">Cyclopina koreana</emphasis>
</taxonomicName>
sp. nov., line drawings, holotype female 
<emphasis id="EBC3ABE5A9D3C53EA63CC36D9C83A6A5" bold="true" pageId="0" pageNumber="59">A</emphasis>
antenna 
<emphasis id="A32D38C77B538CF52D6CD363814908B2" bold="true" pageId="0" pageNumber="59">B</emphasis>
mandibula 
<emphasis id="229DB8A3B286156303EFEE85E95F7577" bold="true" pageId="0" pageNumber="59">C</emphasis>
maxillula 
<emphasis id="8E3485BFA5FBC22A2C70F68A2FFB3F29" bold="true" pageId="0" pageNumber="59">D</emphasis>
maxilla 
<emphasis id="C201240953061D58A020EF3D22E2FD29" bold="true" pageId="0" pageNumber="59">E</emphasis>
maxilliped.
</paragraph>
</caption>
<paragraph id="C1B252B4BFBEF07968DD6F5AAA098B44" pageId="0" pageNumber="59">
<taxonomicName id="CEA3E47D75B71660EECA7C6B6DA2C363" genus="Mandibula" lsidName="Mandibula" pageId="0" pageNumber="59" rank="genus">
<emphasis id="FF5877B55CCC6B1E358FED34482BFBFC" bold="true" italics="true" pageId="0" pageNumber="59">Mandibula</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="EC70ABF8721B47A5F76836E46DFF49D8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10B</figureCitation>
) as in 
<taxonomicName id="7E45383592F83ADA080033AD1DAC9941" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="6F5EF5A903A2C41BB1C7F326593D6BB4" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, except second endopodal segment with six setae, apical setae on fourth exopodal segment of markedly different lengths (outer one twice as long as inner one), and additional row of minute spinules at base of unicuspid teeth.
</paragraph>
<paragraph id="EF0784B28872DBD449339EF4F5F4D962" pageId="0" pageNumber="59">
<emphasis id="09454B61DF994E6A0FBC0241CA34E22E" bold="true" italics="true" pageId="0" pageNumber="59">Maxillula</emphasis>
(Fig. 
<figureCitation id="2D97B1356A79A1210F9CE35494650663" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10C</figureCitation>
) segmentation and armature formula as in 
<taxonomicName id="8ABA5C78216CCEE92C3FF1943017FDCB" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="F14AE9E16952070CE6FAE044ABA9AA60" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but only one minute seta on praecoxal arthrite smooth, one seta on endopod markedly shorter than other endopodal setae, and both endopod and exopod slightly slenderer.
</paragraph>
<paragraph id="135DFED33324781401E7FF92C3504BE5" pageId="0" pageNumber="59">
<emphasis id="785E3A3F55860A874B213BF2BA4165AA" bold="true" italics="true" pageId="0" pageNumber="59">Maxilla</emphasis>
(Fig. 
<figureCitation id="FE8D2E2C54F20B0425C4F547F4A53951" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10D</figureCitation>
) as in 
<taxonomicName id="FB878ED89F7F4ADFD762FE28E30C3006" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="73BB1F82A7A314EBDB1149BF019752AA" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but with only three setae on proximal syncoxal endite, proximal basal endite less mobile and with one seta minute, endopodal claw smooth, and endopod four-segment.
</paragraph>
<paragraph id="27D43313F769F61B5CAD9B763F0ECD68" pageId="0" pageNumber="59">
<emphasis id="AB1677C701B99515E0800635EEA37781" bold="true" italics="true" pageId="0" pageNumber="59">Maxilliped</emphasis>
(Fig. 
<figureCitation id="AFD233FE0F17CA785F1D3735A8BA03AE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10E</figureCitation>
) segmentation and armature formula as in 
<taxonomicName id="2668EAFD5E9D7EEC32843EF17E6627D7" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="72C3E19D24F25A8EC9E4E99AB9475C4D" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but with longer syncoxa, shorter first endopodal segment, and two long setae on ultimate endopodal segment.
</paragraph>
<paragraph id="3CFB76F51099906CBF9F46B19CFC8D7B" pageId="0" pageNumber="59">
<emphasis id="114B68E642DE98E2DF1838B57D48AFA2" bold="true" italics="true" pageId="0" pageNumber="59">Swimming legs</emphasis>
(Fig. 
<figureCitation id="082B805AA2F983EA0F3011D698BF109E" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cyclopina koreana sp. nov., line drawings, holotype female A antenna B mandibula C maxillula D maxilla E maxilliped." figureDoi="10.3897/zookeys.992.54856.figure10" httpUri="https://binary.pensoft.net/fig/474558" pageId="0" pageNumber="59">10A-E</figureCitation>
) shape, segmentation, and ornamentation as in 
<taxonomicName id="F0EC7B6CF8BEED4DB7984F0BE5341708" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="4CFFF279ADA252F4065EE5484928FE7F" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
; armature formula as in 
<taxonomicName id="69CE8209644C146F3822EB51DD6EFB81" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="C10EC3D6E0331D54B5069763CB33C2CD" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, except third exopodal segment of fourth leg with only four setae; all spines lanceolate; three setae on endopod of fourth leg also lanceolate, other setae slender; third endopodal segments seta formula 6.6.6.5; third exopodal segment seta formula 4.5.5.4 and spine formula 4.4.4.3; third endopodal segment of fourth leg 1.7 
<normalizedToken id="75C40048A3EB425AEACF3374901FF376" originalValue="×">x</normalizedToken>
as long as wide and third exopodal segment of fourth leg ca. 1.6 
<normalizedToken id="00E4EBBDDF8847D7304DCEA3EBED3D12" originalValue="×">x</normalizedToken>
as long as wide.
</paragraph>
<paragraph id="49D9B4D7845EF28624B74AE3140E8F37" pageId="0" pageNumber="59">
<emphasis id="5EB74ACB28AA5D9463AB6467B1F6F088" bold="true" italics="true" pageId="0" pageNumber="59">Fifth leg</emphasis>
(Fig. 
<figureCitation id="DB03D713E312792629AF07DDD9C736C4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8A, G</figureCitation>
) shape, segmentation, armature formula, and ornamentation as in 
<taxonomicName id="70A5F85F964A8D2B110EFB5923E8C559" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="A634802A1E1C1A27359B02F466582A6A" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but first segment slightly shorter (ca. 0.6 
<normalizedToken id="88192E5130C6B924F7AC65D82BEF095A" originalValue="×">x</normalizedToken>
as long as wide) and lateral spine on second segment also proportionately shorter (approximately as long as second segment and 1.3 
<normalizedToken id="02F86CF629158B69A11FD56FAD41658D" originalValue="×">x</normalizedToken>
as long as medial spine).
</paragraph>
<paragraph id="DFC7952DB256525664BACE737D7CA266" pageId="0" pageNumber="59">
<emphasis id="5FC0B51DE236AFE12920750792E02FAB" bold="true" italics="true" pageId="0" pageNumber="59">Sixth leg</emphasis>
(Fig. 
<figureCitation id="313E171E3C7C331F900D9B0326A81DCF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8G</figureCitation>
) as in 
<taxonomicName id="CD4BB311264C82BA55C4D0D27F595B69" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="C8C288B948E0F68532C93B59EDD8C6FA" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3A8BB2EA97809B5F5C00C7293DDC693B" pageId="0" pageNumber="59">
<emphasis id="DE95F906AA86F029A1F69C42E615DF6B" italics="true" pageId="0" pageNumber="59">Male</emphasis>
(based on allotype and two other paratypes). 
<emphasis id="03EFB0784C1C1672175431EB162C8B4D" bold="true" italics="true" pageId="0" pageNumber="59">Body</emphasis>
length from 440 to 500 
<normalizedToken id="81576A0FE6F9EDE7B2F6A843ACA1F87D" originalValue="μm">μm</normalizedToken>
. Habitus (Fig. 
<figureCitation id="957779E755EF9F9660E9CC52382CE003" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8A</figureCitation>
) similar to female, but slenderer. 
<emphasis id="40ED7C8DF735222F8AE563B3F4779733" bold="true" italics="true" pageId="0" pageNumber="59">Urosome</emphasis>
(Figs 
<figureCitation id="410122862052C20BCB3BEB0CAF0B33AE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8D, E</figureCitation>
, 
<figureCitation id="0174C9B2BADA31C5E0346CA2F6C35164" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9C</figureCitation>
) also slenderer than in female, and second and third urosomites fully articulated as in 
<taxonomicName id="A96ED9AF4FDEB2A9D423E25ECB03F8F4" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="0E940909304BA96F0200C00095ECE1A6" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
; ornamentation as in female.
</paragraph>
<paragraph id="14AE73BA3997394EE00981285A4AC4C3" pageId="0" pageNumber="59">
<emphasis id="A3978872CABC85F825AAC78096DC8BEF" bold="true" italics="true" pageId="0" pageNumber="59">Caudal rami</emphasis>
(Fig. 
<figureCitation id="5D9B5AA3DAA62017315054290731E449" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9C</figureCitation>
) slightly less widely spaced than in female, but armature and ornamentation without significant differences (perhaps dorsal seta somewhat shorter).
</paragraph>
<paragraph id="FA339CC724087908492B75FE35307661" pageId="0" pageNumber="59">
<emphasis id="DD8DC53EA40518F973029D389CF1A4AA" bold="true" italics="true" pageId="0" pageNumber="59">Antennula</emphasis>
(Figs 
<figureCitation id="57F6BD1820B9839A6D9BD7779BE2B96B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8B</figureCitation>
, 
<figureCitation id="C7E4BD4563F169CE820350CF966CC9E7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cyclopina koreana sp. nov., line drawings A-E holotype female F-H allotype male: A third endopodal segment of first swimming leg B third exopodal segment of first swimming leg C third exopodal segment of second swimming leg D third endopodal segment of second swimming leg E fourth swimming leg F antennula G third endopodal segment of fourth swimming leg H third exopodal segment of fourth swimming leg." figureDoi="10.3897/zookeys.992.54856.figure11" httpUri="https://binary.pensoft.net/fig/474559" pageId="0" pageNumber="59">11F</figureCitation>
) shape, geniculation, segmentation, ornamentation, and almost all armature as in 
<taxonomicName id="466C3BD9AF3BEB593124F101ED731D40" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="BCCD108B221E7831342300C51E87AC5A" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but fifth segment with traces of additional segmentation, both aesthetascs longer, and tenth segment with additional short seta (armature formula therefore: 2.5.4.2.6.1.1.2.2.2+ae.2.1.2.1.11+ae).
</paragraph>
<caption id="442E077396DF252AA0E041D332C33A7E" doi="10.3897/zookeys.992.54856.figure11" httpUri="https://binary.pensoft.net/fig/474559" pageId="0" pageNumber="59" start="Figure 11" startId="F11">
<paragraph id="ADF2609720482A4415B8387A97B4D3F6" pageId="0" pageNumber="59">
<emphasis id="519E1913CE1671CD1FD3B15BABA7DA68" bold="true" pageId="0" pageNumber="59">Figure 11.</emphasis>
<taxonomicName id="7AE7C0AF5D22694A40783DAD34004931" authorityName="Karanovic" authorityYear="2020" class="Copepoda" family="Cyclopinidae" genus="Cyclopina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclopina koreana" order="Cyclopoida" pageId="0" pageNumber="59" phylum="Arthropoda" rank="species" species="koreana">
<emphasis id="FE5B74D46AF09C952D1C19F8194B8D72" italics="true" pageId="0" pageNumber="59">Cyclopina koreana</emphasis>
</taxonomicName>
sp. nov., line drawings 
<emphasis id="77082F1BF1EC5FA176937E4D40DC19A1" bold="true" pageId="0" pageNumber="59">A-E</emphasis>
holotype female 
<emphasis id="A21B5D7732301129B170039F3C057070" bold="true" pageId="0" pageNumber="59">F-H</emphasis>
allotype male: 
<emphasis id="98E88B5443A759C44733D89F22E5923C" bold="true" pageId="0" pageNumber="59">A</emphasis>
third endopodal segment of first swimming leg 
<emphasis id="09A9C41C5D8469AEFEC89DC21388F79C" bold="true" pageId="0" pageNumber="59">B</emphasis>
third exopodal segment of first swimming leg 
<emphasis id="AE93DE5139B1B4FDFF5448EA9A659E7E" bold="true" pageId="0" pageNumber="59">C</emphasis>
third exopodal segment of second swimming leg 
<emphasis id="2EAE7E363455C2F2235B7B69FC15A3C9" bold="true" pageId="0" pageNumber="59">D</emphasis>
third endopodal segment of second swimming leg 
<emphasis id="7FC007D07881625408B237BEF1BDFA42" bold="true" pageId="0" pageNumber="59">E</emphasis>
fourth swimming leg 
<emphasis id="AACCE9C125B6F08623D42F277E1BEBE4" bold="true" pageId="0" pageNumber="59">F</emphasis>
antennula 
<emphasis id="D0CD91D1505DBA219D63FA789B53C027" bold="true" pageId="0" pageNumber="59">G</emphasis>
third endopodal segment of fourth swimming leg 
<emphasis id="91A6EB203D5EBD3B08F1D4E0A3D7F6F9" bold="true" pageId="0" pageNumber="59">H</emphasis>
third exopodal segment of fourth swimming leg.
</paragraph>
</caption>
<paragraph id="83737D513F6D3EA8145ADA85831D054F" pageId="0" pageNumber="59">
<emphasis id="3CB523482AB5B4D327466024ECEE659E" bold="true" italics="true" pageId="0" pageNumber="59">Antenna</emphasis>
, mandibula, maxillula, maxilla, maxilliped, and all four swimming legs (Fig. 
<figureCitation id="FC23409D5E7DE975183C7C859EEE8063" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Cyclopina koreana sp. nov., SEM photographs A-E paratype male 1, ventral view F-H paratype female 2, lateral view: A habitus B distal part of antennula C fourth swimming leg D fourth and fifth urosomites E sixth urosomite F tergite of fourth pedigerous somite G anterior part of urosomite, with fifth and sixth legs H fifth and sixth urosomites." figureDoi="10.3897/zookeys.992.54856.figure8" httpUri="https://binary.pensoft.net/fig/474556" pageId="0" pageNumber="59">8A, C</figureCitation>
) as in female. Third endopodal segment of fourth leg (Fig. 
<figureCitation id="000C8A0AD9DA3AA4A27CF162D5DD47DA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cyclopina koreana sp. nov., line drawings A-E holotype female F-H allotype male: A third endopodal segment of first swimming leg B third exopodal segment of first swimming leg C third exopodal segment of second swimming leg D third endopodal segment of second swimming leg E fourth swimming leg F antennula G third endopodal segment of fourth swimming leg H third exopodal segment of fourth swimming leg." figureDoi="10.3897/zookeys.992.54856.figure11" httpUri="https://binary.pensoft.net/fig/474559" pageId="0" pageNumber="59">11G</figureCitation>
) ca. 1.6 
<normalizedToken id="7FFFC00D754A1F5D7D4655ECC188F709" originalValue="×">x</normalizedToken>
as long as wide, with proximal medial seta lanceolate along both sides; third exopodal segment (Fig. 
<figureCitation id="F5AF7F82302752073D5F64D773AE386A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Cyclopina koreana sp. nov., line drawings A-E holotype female F-H allotype male: A third endopodal segment of first swimming leg B third exopodal segment of first swimming leg C third exopodal segment of second swimming leg D third endopodal segment of second swimming leg E fourth swimming leg F antennula G third endopodal segment of fourth swimming leg H third exopodal segment of fourth swimming leg." figureDoi="10.3897/zookeys.992.54856.figure11" httpUri="https://binary.pensoft.net/fig/474559" pageId="0" pageNumber="59">11H</figureCitation>
) with only four setae as in female, ca. 1.6 
<normalizedToken id="7D120EFA68B95377489ACB63A5C76CC7" originalValue="×">x</normalizedToken>
as long as wide.
</paragraph>
<paragraph id="53D0230495D3987650FBE715B9D1F7B3" pageId="0" pageNumber="59">
<emphasis id="BA8CAF9A3BCC9E4BE6E7566EE697A703" bold="true" italics="true" pageId="0" pageNumber="59">Fifth leg</emphasis>
(Fig. 
<figureCitation id="5D4B52C569BF9DDB76898546A1D1F963" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9C</figureCitation>
) segmentation, ornamentation, and armature formula as in 
<taxonomicName id="10C5AAC950D396F7178FB62BCC4EF56E" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="C94487703C8EFBC6274521770222DA57" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, i.e., with two medial setae on second segment; proximal segment as in female; lateral spine ca. 0.7 
<normalizedToken id="26DB26AFE2AE100223815A6E9ECBA46F" originalValue="×">x</normalizedToken>
as long as second segment and 1.1 
<normalizedToken id="63FC66C03EADCB154AFF8F839FEBB0B9" originalValue="×">x</normalizedToken>
as long as medial spine.
</paragraph>
<paragraph id="53A836B0E797544D3F8277C3E11C83C1" pageId="0" pageNumber="59">
<emphasis id="176000F69E069FA0981A93B6BD6DF1CA" bold="true" italics="true" pageId="0" pageNumber="59">Sixth leg</emphasis>
(Fig. 
<figureCitation id="FA212F169760BBAF147DC07520EFC77F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cyclopina koreana sp. nov., line drawings A holotype female, urosome, ventral view B holotype female, antennula C allotype male, urosome, ventral view." figureDoi="10.3897/zookeys.992.54856.figure9" httpUri="https://binary.pensoft.net/fig/474557" pageId="0" pageNumber="59">9C</figureCitation>
) as in 
<taxonomicName id="19CFE4473F4876B471047D5C7FD07AC7" lsidName="C. busanensis" pageId="0" pageNumber="59" rank="species" species="busanensis">
<emphasis id="DE82535FD14AA24F85DB0B9CD95FCDF1" italics="true" pageId="0" pageNumber="59">C. busanensis</emphasis>
</taxonomicName>
, but broader and without minute medial spine; lateral seta 1.5 
<normalizedToken id="A01DE47155D2DC81DA46F90B0762B3A1" originalValue="×">x</normalizedToken>
as long as medial seta.
</paragraph>
</subSubSection>
<subSubSection id="90868D75BE3F0D2150B62F70E7EBF175" pageId="0" pageNumber="59" type="variability">
<paragraph id="F5965E8C6B70633DA5B4C6A36779299A" pageId="0" pageNumber="59">Variability.</paragraph>
<paragraph id="95DABB20AEABC3989EE423B267C51201" pageId="0" pageNumber="59">
Except for small differences in body size no other forms of variability were observed, but some specimens were damaged (e.g., with some setae broken off; see Fig. 
<figureCitation id="645A9E59293B7CC23DFD88CA52142362" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cyclopina koreana sp. nov., paratype female 1, SEM photographs, all in dorsal view A habitus B second pedigerous somite C third pedigerous somite D first urosomite E genital double-somite and fourth urosomite F anterior medial pore on genital double-somite G fifth and sixth urosomites H caudal ramus." figureDoi="10.3897/zookeys.992.54856.figure7" httpUri="https://binary.pensoft.net/fig/474555" pageId="0" pageNumber="59">7H</figureCitation>
) so comparisons were somewhat limited.
</paragraph>
</subSubSection>
</treatment>
</document>