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<document id="D2F3049C3ED3D4DBB2B9C14FA13F2953" ID-DOI="10.5281/zenodo.14662376" ID-ISSN="1942-1354" ID-Zenodo-Dep="14662376" ID-ZooBank="1F9254BA-4B88-4FAC-B417-072BF29F76F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736984950950" checkinUser="felipe" docAuthor="Marek, William A. Shear Paul E." docDate="2024" docId="C2099450FFE08B5E3580FAC3FE95FCE1" docLanguage="en" docName="InsectaMundi.2024.1090.1-21.pdf" docOrigin="Insecta Mundi 2024 (90)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Ochrogramma Gardener and Shelley 1989" docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="3E30EC28FFE88B573540FFA0FFC1FFF8" masterDocTitle="Additions to the millipede family Caseyidae Verhoeff, 1909. III. Proposal of the new subfamily Ochrogrammatinae, new records of Vasingtona irritans (Chamberlin, 1910), established species of Ochrogramma Gardener and Shelley, 1989 a new species of Ochrogramma from Washington state, USA" masterLastPageNumber="21" masterPageNumber="1" pageNumber="7" updateTime="1736986191955" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="288474B5A54283D062E432EF51726063">Additions to the millipede family Caseyidae Verhoeff, 1909. III. Proposal of the new subfamily Ochrogrammatinae, new records of Vasingtona irritans (Chamberlin, 1910), established species of Ochrogramma Gardener and Shelley, 1989 a new species of Ochrogramma from Washington state, USA</mods:title>
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<mods:namePart id="A68D4C97D83D44956CA54D04233B0C50">Marek, William A. Shear Paul E.</mods:namePart>
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<mods:date id="07E0667E26738550BFD65FC4BE529F12">2024</mods:date>
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<treatment id="C2099450FFE08B5E3580FAC3FE95FCE1" ID-DOI="http://doi.org/10.5281/zenodo.14662390" ID-Zenodo-Dep="14662390" LSID="urn:lsid:plazi:treatment:C2099450FFE08B5E3580FAC3FE95FCE1" httpUri="http://treatment.plazi.org/id/C2099450FFE08B5E3580FAC3FE95FCE1" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<subSubSection id="02BA76CDFFE08B5F3580FAC3FD0DFA79" box="[192,716,1379,1409]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="4A1F2546FFE08B5F3580FAC3FD0DFA79" blockId="8.[192,716,1379,1409]" box="[192,716,1379,1409]" pageId="8" pageNumber="7">
<heading id="1157922AFFE08B5F3580FAC3FD0DFA79" box="[192,716,1379,1409]" fontSize="12" level="2" pageId="8" pageNumber="7" reason="2">
<taxonomicName id="8DA05EC5FFE08B5F3580FAC3FD0DFA79" ID-CoL="7PMVS" authority="Gardner and Shelley, 1989" authorityName="Gardener and Shelley" authorityYear="1989" box="[192,716,1379,1409]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F3580FAC3FD0DFA79" bold="true" box="[192,716,1379,1409]" pageId="8" pageNumber="7">
<emphasis id="78D4F954FFE08B5F3580FAC3FEB6FA78" bold="true" box="[192,375,1379,1408]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
<bibRefCitation id="2E3158B7FFE08B5F343EFAC4FD0DFA79" author="Gardner MR &amp; Shelley RM" box="[382,716,1380,1409]" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley, 1989</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="02BA76CDFFE08B5F35A0FA31FD70FA51" box="[224,689,1425,1449]" pageId="8" pageNumber="7" type="reference_group">
<paragraph id="4A1F2546FFE08B5F35A0FA31FD70FA51" blockId="8.[224,689,1425,1449]" box="[224,689,1425,1449]" pageId="8" pageNumber="7">
<treatmentCitationGroup id="6AB00268FFE08B5F35A0FA31FD70FA51" box="[224,689,1425,1449]" pageId="8" pageNumber="7">
<treatmentCitation id="CB010357FFE08B5F35A0FA31FD6CFA51" author="Gardner MR &amp; Shelley RM" box="[224,685,1425,1449]" page="194" pageId="8" pageNumber="7" year="1989">
<taxonomicName id="8DA05EC5FFE08B5F35A0FA31FD6CFA51" ID-CoL="7PMVS" authority="Gardner and Shelley 1989: 194" authorityName="Gardener and Shelley" authorityPageNumber="194" authorityYear="1989" box="[224,685,1425,1449]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F35A0FA31FEB1FA51" box="[224,368,1425,1449]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
<bibRefCitation id="2E3158B7FFE08B5F3436FA31FD6CFA51" author="Gardner MR &amp; Shelley RM" box="[374,685,1425,1449]" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley 1989: 194</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="02BA76CDFFE08B5E3580FA19FE95FCE1" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7" type="type_taxon">
<paragraph id="4A1F2546FFE08B5F3580FA19FB73FA2A" blockId="8.[192,1202,1464,1491]" box="[192,1202,1464,1491]" pageId="8" pageNumber="7">
<emphasis id="78D4F954FFE08B5F3580FA19FE98FA2A" bold="true" box="[192,345,1464,1491]" pageId="8" pageNumber="7">
<typeStatus id="951B9BE4FFE08B5F3580FA19FF38FA2B" box="[192,249,1465,1491]" pageId="8" pageNumber="7">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8DA05EC5FFE08B5F341FFA18FC61FA2B" authority="Gardner and Shelley, 1989" authorityName="Gardner and Shelley" authorityYear="1989" box="[351,928,1464,1491]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="formulosa">
<emphasis id="78D4F954FFE08B5F341FFA18FDB3FA2A" box="[351,626,1464,1490]" italics="true" pageId="8" pageNumber="7">Ochrogramma formulosa</emphasis>
<bibRefCitation id="2E3158B7FFE08B5F3739FA18FC61FA2B" author="Gardner MR &amp; Shelley RM" box="[633,928,1464,1491]" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley, 1989</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="4A1F2546FFE08B5E3580FA42FD50FEBB" blockId="8.[192,1441,1506,1902]" lastBlockId="9.[191,1442,230,793]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="78D4F954FFE08B5F3580FA42FECAFA04" bold="true" box="[192,267,1506,1532]" pageId="8" pageNumber="7">Notes.</emphasis>
<taxonomicName id="8DA05EC5FFE08B5F3453FA42FE75FA04" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[275,436,1506,1532]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F3453FA42FE75FA04" box="[275,436,1506,1532]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="8DA05EC5FFE08B5F37A1FA43FC9CFA04" ID-CoL="7Q67B" authorityName="Chamberlin" authorityYear="1941" box="[737,861,1507,1532]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F37A1FA43FC9CFA04" box="[737,861,1507,1532]" italics="true" pageId="8" pageNumber="7">Vasingtona</emphasis>
</taxonomicName>
. Males of the species of
<taxonomicName id="8DA05EC5FFE08B5F313BFA42FADDFA04" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[1147,1308,1506,1532]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F313BFA42FADDFA04" box="[1147,1308,1506,1532]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
</taxonomicName>
are usually smaller (
<quantity id="8D5888A3FFE08B5F3465F9A3FE64F9E6" box="[293,421,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="7" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
) than those of
<taxonomicName id="8DA05EC5FFE08B5F371AF9A4FCF3F9E5" ID-CoL="7FHZ3" baseAuthorityName="Chamberlin" baseAuthorityYear="1910" box="[602,818,1540,1565]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="irritans">
<emphasis id="78D4F954FFE08B5F371AF9A4FCF3F9E5" box="[602,818,1540,1565]" italics="true" pageId="8" pageNumber="7">Vasingtona irritans</emphasis>
</taxonomicName>
(
<quantity id="8D5888A3FFE08B5F3604F9A3FC06F9E6" box="[836,967,1539,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="8" pageNumber="7" unit="mm" value="19.0" valueMax="22.0" valueMin="16.0">16–22 mm</quantity>
). In our new interpretation of the gonopods in this subfamily,
<taxonomicName id="8DA05EC5FFE08B5F348BF985FDADF9C7" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[459,620,1573,1599]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F348BF985FDADF9C7" box="[459,620,1573,1599]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
</taxonomicName>
is distinct from
<taxonomicName id="8DA05EC5FFE08B5F3669F986FC64F9C7" ID-CoL="7Q67B" authorityName="Chamberlin" authorityYear="1941" box="[809,933,1574,1599]" class="Diplopoda" family="Caseyidae" genus="Vasingtona" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F3669F986FC64F9C7" box="[809,933,1574,1599]" italics="true" pageId="8" pageNumber="7">Vasingtona</emphasis>
</taxonomicName>
in having many fewer arborescent processes on the gonopod (
<figureCitation id="D29B39C3FFE08B5F34C8F9E7FE18F999" box="[392,473,1607,1633]" captionStart="Figures 17–20" captionStartId="11.[192,271,1609,1633]" captionTargetBox="[192,1440,230,1580]" captionTargetId="figure-11@11.[192,1440,230,1580]" captionTargetPageId="11" captionText="Figures 17–20. Male Ochrogramma bentona (Chamberlin, 1952). 17) Tip of coxal lobe of leg 3, ventral view. 18) Legpair 7, posterior view. 19) Gonopods, anterior view. 20) Gonopods, posterior view. Abbreviations: aac, anterior angiocoxites; c, gonopod coxa; cc, colpocoxite; cx7, coxae of leg 7; pac, posterior angiocoxite, t, telopodite of gonopod." figureDoi="http://doi.org/10.5281/zenodo.14662386" httpUri="https://zenodo.org/record/14662386/files/figure.png" pageId="8" pageNumber="7">Fig. 19</figureCitation>
,
<figureCitation id="D29B39C3FFE08B5F34A6F9E7FDEAF999" box="[486,555,1607,1633]" captionStart="Figures 21–24" captionStartId="13.[192,271,1764,1788]" captionTargetBox="[192,1440,229,1734]" captionTargetId="figure-60@13.[192,1440,229,1734]" captionTargetPageId="13" captionText="Figures 21–24. Male Ochrogramma bentona (Chamberlin, 1952). 21) Left gonopod, mesal view. 22) Left gonopod, posterior view. 23) Tip of left gonopod anterior angiocoxite, posterior view. 24) Arborescent processes of gonopod, mesoanterior view. Abbreviations: aac, anterior angiocoxite; c, coxa, cc, colpocoxite; f, flagellocoxite; pac, posterior angiocoxite; t, gonopod telopodite." figureDoi="http://doi.org/10.5281/zenodo.14662388" httpUri="https://zenodo.org/record/14662388/files/figure.png" pageId="8" pageNumber="7">23, 24</figureCitation>
)—two species of
<taxonomicName id="8DA05EC5FFE08B5F37B8F9E7FC58F999" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[760,921,1607,1633]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE08B5F37B8F9E7FC58F999" box="[760,921,1607,1633]" italics="true" pageId="8" pageNumber="7">Ochrogramma</emphasis>
</taxonomicName>
may have none—in the reduced and perhaps sometimes absent telopodite (
<emphasis id="78D4F954FFE08B5F3752F9C9FDDAF97B" bold="true" box="[530,539,1641,1667]" pageId="8" pageNumber="7">t</emphasis>
,
<figureCitation id="D29B39C3FFE08B5F3769F9C8FDB8F97B" box="[553,633,1640,1667]" captionStart="Figures 17–20" captionStartId="11.[192,271,1609,1633]" captionTargetBox="[192,1440,230,1580]" captionTargetId="figure-11@11.[192,1440,230,1580]" captionTargetPageId="11" captionText="Figures 17–20. Male Ochrogramma bentona (Chamberlin, 1952). 17) Tip of coxal lobe of leg 3, ventral view. 18) Legpair 7, posterior view. 19) Gonopods, anterior view. 20) Gonopods, posterior view. Abbreviations: aac, anterior angiocoxites; c, gonopod coxa; cc, colpocoxite; cx7, coxae of leg 7; pac, posterior angiocoxite, t, telopodite of gonopod." figureDoi="http://doi.org/10.5281/zenodo.14662386" httpUri="https://zenodo.org/record/14662386/files/figure.png" pageId="8" pageNumber="7">Fig. 20</figureCitation>
), having a rodlike posterior angiocoxite (
<emphasis id="78D4F954FFE08B5F310AF9C9FBB2F97B" bold="true" box="[1098,1139,1641,1667]" pageId="8" pageNumber="7">pac</emphasis>
,
<figureCitation id="D29B39C3FFE08B5F31C0F9C8FB11F97B" box="[1152,1232,1640,1667]" captionStart="Figures 17–20" captionStartId="11.[192,271,1609,1633]" captionTargetBox="[192,1440,230,1580]" captionTargetId="figure-11@11.[192,1440,230,1580]" captionTargetPageId="11" captionText="Figures 17–20. Male Ochrogramma bentona (Chamberlin, 1952). 17) Tip of coxal lobe of leg 3, ventral view. 18) Legpair 7, posterior view. 19) Gonopods, anterior view. 20) Gonopods, posterior view. Abbreviations: aac, anterior angiocoxites; c, gonopod coxa; cc, colpocoxite; cx7, coxae of leg 7; pac, posterior angiocoxite, t, telopodite of gonopod." figureDoi="http://doi.org/10.5281/zenodo.14662386" httpUri="https://zenodo.org/record/14662386/files/figure.png" pageId="8" pageNumber="7">Fig. 20</figureCitation>
), and a prominent posterior lobe of the colpocoxite. The hypothesis that the colpocoxite originates from the coxal gland of the eighth legpair is supported by the relatively poorly sclerotized colpocoxites emerging from distinct sockets in this genus (
<emphasis id="78D4F954FFE08B5F3404F96DFE9DF91F" bold="true" box="[324,348,1741,1767]" pageId="8" pageNumber="7">cc</emphasis>
,
<figureCitation id="D29B39C3FFE08B5F342AF96DFE7AF91F" box="[362,443,1741,1767]" captionStart="Figures 17–20" captionStartId="11.[192,271,1609,1633]" captionTargetBox="[192,1440,230,1580]" captionTargetId="figure-11@11.[192,1440,230,1580]" captionTargetPageId="11" captionText="Figures 17–20. Male Ochrogramma bentona (Chamberlin, 1952). 17) Tip of coxal lobe of leg 3, ventral view. 18) Legpair 7, posterior view. 19) Gonopods, anterior view. 20) Gonopods, posterior view. Abbreviations: aac, anterior angiocoxites; c, gonopod coxa; cc, colpocoxite; cx7, coxae of leg 7; pac, posterior angiocoxite, t, telopodite of gonopod." figureDoi="http://doi.org/10.5281/zenodo.14662386" httpUri="https://zenodo.org/record/14662386/files/figure.png" pageId="8" pageNumber="7">Fig. 20</figureCitation>
). Gonopod structure is similar among the species, but two of them,
<taxonomicName id="8DA05EC5FFE08B5F3185F96EFE78F8F1" authority="Gardner and Shelley, 1989" authorityName="Gardner and Shelley" authorityYear="1989" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="formulosa">
<emphasis id="78D4F954FFE08B5F3185F96EFA97F91F" box="[1221,1366,1741,1768]" italics="true" pageId="8" pageNumber="7">O. formulosa</emphasis>
<bibRefCitation id="2E3158B7FFE08B5F301DF96DFE78F8F1" author="Gardner MR &amp; Shelley RM" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley, 1989</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="8DA05EC5FFE08B5F34BAF94FFC12F8F1" ID-CoL="6SF56" authority="Gardner and Shelley, 1989" authorityName="Gardner and Shelley" authorityYear="1989" box="[506,979,1775,1801]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="heterogona">
<emphasis id="78D4F954FFE08B5F34BAF94FFD59F8F1" box="[506,664,1775,1801]" italics="true" pageId="8" pageNumber="7">O. heterogona</emphasis>
<bibRefCitation id="2E3158B7FFE08B5F37E1F94FFC12F8F1" author="Gardner MR &amp; Shelley RM" box="[673,979,1775,1801]" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley, 1989</bibRefCitation>
</taxonomicName>
, seem to lack the arborescent processes (“spicules” of Gardner and Shelley) on the anterior surfaces of the anterior angiocoxites. We have not seen specimens of these species.
<taxonomicName id="8DA05EC5FFE08B5F3741F892FCC5F8B4" ID-CoL="74F57" baseAuthorityName="Chamberlin" baseAuthorityYear="1952" box="[513,772,1842,1868]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="bentona">
<emphasis id="78D4F954FFE08B5F3741F892FCC5F8B4" box="[513,772,1842,1868]" italics="true" pageId="8" pageNumber="7">Ochrogramma bentona</emphasis>
</taxonomicName>
has the most complex gonopods in the genus while those of
<emphasis id="78D4F954FFE08B5F359EF8F4FE14F896" box="[222,469,1876,1902]" italics="true" pageId="8" pageNumber="7">
<taxonomicName id="8DA05EC5FFE08B5F359EF8F4FEF8F896" ID-CoL="48DYD" authorityName="Gardner and Shelley" authorityYear="1989" box="[222,313,1876,1902]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="haigi">O. haigi</taxonomicName>
,
<taxonomicName id="8DA05EC5FFE08B5F3404F8F4FE14F896" authorityName="Gardner and Shelley" authorityYear="1989" box="[324,469,1876,1902]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="formulosa">O. formulosa</taxonomicName>
</emphasis>
and
<taxonomicName id="8DA05EC5FFE08B5F374FF8F4FD6BF896" ID-CoL="6SF56" authorityName="Gardner and Shelley" authorityYear="1989" box="[527,682,1876,1902]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="heterogona">
<emphasis id="78D4F954FFE08B5F374FF8F4FD6BF896" box="[527,682,1876,1902]" italics="true" pageId="8" pageNumber="7">O. heterogona</emphasis>
</taxonomicName>
are progressively simpler.
<bibRefCitation id="2E3158B7FFE08B5F3697F8F4FAD1F896" author="Gardner MR &amp; Shelley RM" box="[983,1296,1876,1902]" pageId="8" pageNumber="7" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
did not give length measurements for females, but we found that females are typically longer than males (
<quantity id="8D5888A3FFE18B5E31A4FF46FAA1FEF8" box="[1252,1376,230,256]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="9" pageNumber="8" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">17–19 mm</quantity>
in
<taxonomicName id="8DA05EC5FFE18B5E30C7FF46FED8FED9" ID-CoL="74F57" baseAuthorityName="Chamberlin" baseAuthorityYear="1952" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="bentona">
<emphasis id="78D4F954FFE18B5E30C7FF46FED8FED9" italics="true" pageId="9" pageNumber="8">O. bentona</emphasis>
</taxonomicName>
). The dorsal yellow stripe typical of the subfamily may be suppressed entirely in
<taxonomicName id="8DA05EC5FFE18B5E31EAFEA7FA61FED9" box="[1194,1440,263,289]" pageId="9" pageNumber="8">
<emphasis id="78D4F954FFE18B5E31EAFEA7FA8AFED9" box="[1194,1355,263,289]" italics="true" pageId="9" pageNumber="8">Ochrogramma</emphasis>
species
</taxonomicName>
males, or present only on posterior rings.
</paragraph>
<paragraph id="4A1F2546FFE18B5E3440FEEAFDA5FE13" blockId="9.[191,1442,230,793]" pageId="9" pageNumber="8">
The gonopod telopodite is seen in only two of the species,
<taxonomicName id="8DA05EC5FFE18B5E36C3FEEBFC3AFE9C" ID-CoL="74F57" baseAuthorityName="Chamberlin" baseAuthorityYear="1952" box="[899,1019,330,357]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="bentona">
<emphasis id="78D4F954FFE18B5E36C3FEEBFC3AFE9C" box="[899,1019,330,357]" italics="true" pageId="9" pageNumber="8">O. bentona</emphasis>
</taxonomicName>
and
<emphasis id="78D4F954FFE18B5E3170FEEBFB07FE9C" box="[1072,1222,330,357]" italics="true" pageId="9" pageNumber="8">
<taxonomicName id="8DA05EC5FFE18B5E3170FEEBFB03FE9C" authorityName="Gardner and Shelley" authorityYear="1989" box="[1072,1218,330,357]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="formulosa">O. formulosa</taxonomicName>
.
</emphasis>
It is highly reduced and rodlike (
<emphasis id="78D4F954FFE18B5E340FFECCFE99FE7E" bold="true" box="[335,344,364,390]" pageId="9" pageNumber="8">t</emphasis>
,
<figureCitation id="D29B39C3FFE18B5E3424FECCFE73FE7E" box="[356,434,364,390]" captionStart="Figures 17–20" captionStartId="11.[192,271,1609,1633]" captionTargetBox="[192,1440,230,1580]" captionTargetId="figure-11@11.[192,1440,230,1580]" captionTargetPageId="11" captionText="Figures 17–20. Male Ochrogramma bentona (Chamberlin, 1952). 17) Tip of coxal lobe of leg 3, ventral view. 18) Legpair 7, posterior view. 19) Gonopods, anterior view. 20) Gonopods, posterior view. Abbreviations: aac, anterior angiocoxites; c, gonopod coxa; cc, colpocoxite; cx7, coxae of leg 7; pac, posterior angiocoxite, t, telopodite of gonopod." figureDoi="http://doi.org/10.5281/zenodo.14662386" httpUri="https://zenodo.org/record/14662386/files/figure.png" pageId="9" pageNumber="8">Fig. 20</figureCitation>
), articulating about one-third of the way distal on the anterior angiocoxite and terminating in several filaments which are likely modified setae. The posterior angiocoxite, evidently missing in
<taxonomicName id="8DA05EC5FFE18B5E3041FE2EFA58FE50" ID-CoL="6SF56" authorityName="Gardner and Shelley" authorityYear="1989" box="[1281,1433,398,424]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="heterogona">
<emphasis id="78D4F954FFE18B5E3041FE2EFA58FE50" box="[1281,1433,398,424]" italics="true" pageId="9" pageNumber="8">O. heterogona</emphasis>
</taxonomicName>
, is roughly similar in form to the telopodite (
<emphasis id="78D4F954FFE18B5E37FCFE0FFD25FE31" bold="true" box="[700,740,431,457]" pageId="9" pageNumber="8">pac</emphasis>
,
<figureCitation id="D29B39C3FFE18B5E37B3FE0FFCAFFE31" box="[755,878,431,457]" captionStart="Figures 21–24" captionStartId="13.[192,271,1764,1788]" captionTargetBox="[192,1440,229,1734]" captionTargetId="figure-60@13.[192,1440,229,1734]" captionTargetPageId="13" captionText="Figures 21–24. Male Ochrogramma bentona (Chamberlin, 1952). 21) Left gonopod, mesal view. 22) Left gonopod, posterior view. 23) Tip of left gonopod anterior angiocoxite, posterior view. 24) Arborescent processes of gonopod, mesoanterior view. Abbreviations: aac, anterior angiocoxite; c, coxa, cc, colpocoxite; f, flagellocoxite; pac, posterior angiocoxite; t, gonopod telopodite." figureDoi="http://doi.org/10.5281/zenodo.14662388" httpUri="https://zenodo.org/record/14662388/files/figure.png" pageId="9" pageNumber="8">Fig. 21, 22</figureCitation>
). Flagellocoxites are present in all species and are filamentous and branched (
<emphasis id="78D4F954FFE18B5E34B3FE71FE3DFE13" bold="true" box="[499,508,465,491]" pageId="9" pageNumber="8">f</emphasis>
,
<figureCitation id="D29B39C3FFE18B5E3749FE71FD96FE13" box="[521,599,465,491]" captionStart="Figures 21–24" captionStartId="13.[192,271,1764,1788]" captionTargetBox="[192,1440,229,1734]" captionTargetId="figure-60@13.[192,1440,229,1734]" captionTargetPageId="13" captionText="Figures 21–24. Male Ochrogramma bentona (Chamberlin, 1952). 21) Left gonopod, mesal view. 22) Left gonopod, posterior view. 23) Tip of left gonopod anterior angiocoxite, posterior view. 24) Arborescent processes of gonopod, mesoanterior view. Abbreviations: aac, anterior angiocoxite; c, coxa, cc, colpocoxite; f, flagellocoxite; pac, posterior angiocoxite; t, gonopod telopodite." figureDoi="http://doi.org/10.5281/zenodo.14662388" httpUri="https://zenodo.org/record/14662388/files/figure.png" pageId="9" pageNumber="8">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="4A1F2546FFE18B5E3440FE52FCC5FD6B" blockId="9.[191,1442,230,793]" pageId="9" pageNumber="8">
<taxonomicName id="8DA05EC5FFE18B5E3440FE52FE5EFDF4" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[256,415,498,524]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE18B5E3440FE52FE5EFDF4" box="[256,415,498,524]" italics="true" pageId="9" pageNumber="8">Ochrogramma</emphasis>
</taxonomicName>
was considered by
<bibRefCitation id="2E3158B7FFE18B5E3739FE52FC6EFDF5" author="Gardner MR &amp; Shelley RM" box="[633,943,498,525]" pageId="9" pageNumber="8" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
to be distributed entirely to the south of the
<taxonomicName id="8DA05EC5FFE18B5E3581FDB4FEB2FDD6" authority="River" authorityName="River" box="[193,371,532,558]" class="Actinopterygii" family="Percopsidae" genus="Columbia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Percopsiformes" pageId="9" pageNumber="8" phylum="Chordata" rank="genus">Columbia River</taxonomicName>
, but this was an artifact of insufficient collecting in
<collectingRegion id="8864EBA4FFE18B5E36F0FDB4FBFBFDD6" box="[944,1082,532,558]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
.
<taxonomicName id="8DA05EC5FFE18B5E3103FDB4FE92FDA9" ID-CoL="74F57" authority="(Chamberlin, 1952)" baseAuthorityName="Chamberlin" baseAuthorityYear="1952" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="bentona">
<emphasis id="78D4F954FFE18B5E3103FDB4FA83FDD6" box="[1091,1346,532,558]" italics="true" pageId="9" pageNumber="8">Ochrogramma bentona</emphasis>
(Chamberlin, 1952)
</taxonomicName>
, extends north of the river to the Puget Sound region and
<taxonomicName id="8DA05EC5FFE18B5E36B0FD96FAC0FDA8" authorityName="Shear and Marek" authorityYear="2024" box="[1008,1281,566,592]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="skamania" status="sp. nov.">
<emphasis id="78D4F954FFE18B5E36B0FD96FAC0FDA8" box="[1008,1281,566,592]" italics="true" pageId="9" pageNumber="8">Ochrogramma skamania</emphasis>
</taxonomicName>
<emphasis id="78D4F954FFE18B5E3048FD96FA86FDA8" bold="true" box="[1288,1351,566,592]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="63E7442FFFE18B5E3048FD96FA86FDA8" box="[1288,1351,566,592]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is from west central
<collectingRegion id="8864EBA4FFE18B5E340EFDF7FE19FD89" box="[334,472,599,625]" country="United States of America" name="Washington" pageId="9" pageNumber="8">Washington</collectingRegion>
. The other three
<taxonomicName id="8DA05EC5FFE18B5E37D4FDF7FC49FD89" box="[660,904,599,625]" pageId="9" pageNumber="8">
<emphasis id="78D4F954FFE18B5E37D4FDF7FCF2FD89" box="[660,819,599,625]" italics="true" pageId="9" pageNumber="8">Ochrogramma</emphasis>
species
</taxonomicName>
are limited to small areas of
<collectingRegion id="8864EBA4FFE18B5E318AFDF7FADEFD89" box="[1226,1311,599,625]" country="United States of America" name="Oregon" pageId="9" pageNumber="8">Oregon</collectingRegion>
and northern
<collectingRegion id="8864EBA4FFE18B5E35ACFDD9FE9FFD6B" box="[236,350,633,659]" country="United States of America" name="California" pageId="9" pageNumber="8">California</collectingRegion>
(
<bibRefCitation id="2E3158B7FFE18B5E342CFDD9FD51FD6B" author="Gardner MR &amp; Shelley RM" box="[364,656,633,659]" pageId="9" pageNumber="8" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley 1989</bibRefCitation>
, Fig. 220).
</paragraph>
<paragraph id="4A1F2546FFE18B5E3440FD3AFE95FCE1" blockId="9.[191,1442,230,793]" pageId="9" pageNumber="8">
The meaning of the name
<taxonomicName id="8DA05EC5FFE18B5E375CFD3AFD7AFD4C" ID-CoL="7PMVS" authorityName="Gardener and Shelley" authorityYear="1989" box="[540,699,666,692]" class="Diplopoda" family="Caseyidae" genus="Ochrogramma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="78D4F954FFE18B5E375CFD3AFD7AFD4C" box="[540,699,666,692]" italics="true" pageId="9" pageNumber="8">Ochrogramma</emphasis>
</taxonomicName>
was not discussed by
<bibRefCitation id="2E3158B7FFE18B5E36ECFD3AFB1EFD4D" author="Gardner MR &amp; Shelley RM" box="[940,1247,666,693]" pageId="9" pageNumber="8" pagination="177 - 268" refId="ref11180" refString="Gardner MR, Shelley RM. 1989. New records, species and genera of caseyid millipeds from the Pacific Coast of North America (Diplopoda: Chordeumatida: Caseyidae). Pan-Pacific Entomologist 65: 177 - 268." type="journal article" year="1989">Gardner and Shelley (1989)</bibRefCitation>
. Both parts of the name come from Greek;
<emphasis id="78D4F954FFE18B5E3498FD1CFDDCFD2E" box="[472,541,700,726]" italics="true" pageId="9" pageNumber="8">ochros</emphasis>
a prefix meaning “yellow” and
<emphasis id="78D4F954FFE18B5E363DFD1DFC1EFD2E" box="[893,991,701,726]" italics="true" pageId="9" pageNumber="8">gramma,</emphasis>
referring to a “line, writing or drawing” (the root of English words such as grammar, graphics, telegraph, etc.). The combination likely refers to the dorsal yellow stripe.
</paragraph>
</subSubSection>
</treatment>
</document>
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