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581 lines
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<document id="37D0183058D206D28FDDD3F1208809C0" ID-CLB-Dataset="121554" ID-DOI="10.11646/zootaxa.5174.4.6" ID-GBIF-Dataset="b009d163-c746-470f-a3d7-95dbc8726627" ID-ISSN="1175-5326" ID-Zenodo-Dep="6986469" ID-ZooBank="196884A6-1AF7-4C16-B2C5-D05C7F78E1CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1660327463178" checkinUser="plazi" docAuthor="Kiałka, Agata & Ruta, Rafał" docDate="2022" docId="A0384152FF9A02384DE7FB8DFF5CF98C" docLanguage="en" docName="zootaxa.5174.4.6.pdf" docOrigin="Zootaxa 5174 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atopida White 1846" docType="treatment" docVersion="8" lastPageNumber="412" masterDocId="5C01392AFF9802334D70FFB3FF9BFFAA" masterDocTitle="Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae)" masterLastPageNumber="443" masterPageNumber="401" pageNumber="403" updateTime="1699367011641" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="64E78D9B7C388758547BD07FCC0C8CD0">Revision of Atopida White, 1846 (Coleoptera: Scirtoidea: Scirtidae)</mods:title>
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<mods:name id="5A6D0619948C9BEB03744F297B6BD9BA" type="personal">
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<mods:namePart id="7B3EA50C0CE88346633ACF39A950DF86">Kiałka, Agata</mods:namePart>
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<mods:nameIdentifier id="FE9A98E14B8953C90DF1883743BA019D" type="ORCID">0000-0001-8883-9035</mods:nameIdentifier>
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<mods:affiliation id="7254C3C0B8B500CE38FB369F444AD142">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. & agata. kialka @ gmail. com; https: // orcid. org / 0000 - 0001 - 8883 - 9035</mods:affiliation>
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<mods:nameIdentifier id="D668156A6A814CD55FBA819CF42B313F" type="email">agata.kialka@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EF410CC1FDC7A0303C55F4349D1EA79E">Ruta, Rafał</mods:namePart>
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<mods:affiliation id="5E937ABEC5506E2EC2A0E3AF8DD0A9AF">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. & rafal. ruta @ uwr. edu. pl; https: // orcid. org / 0000 - 0001 - 8515 - 2385</mods:affiliation>
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<mods:nameIdentifier id="161ACC84B431376B3400665040F4F53B" type="email">rafal.ruta@uwr.edu.pl</mods:nameIdentifier>
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<mods:title id="6BA7CFC9C8579165E5EBFAA8A6F029C5">Zootaxa</mods:title>
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<treatment id="A0384152FF9A02384DE7FB8DFF5CF98C" ID-DOI="http://doi.org/10.5281/zenodo.6993092" ID-GBIF-Taxon="199094419" ID-Zenodo-Dep="6993092" LSID="urn:lsid:plazi:treatment:A0384152FF9A02384DE7FB8DFF5CF98C" httpUri="http://treatment.plazi.org/id/A0384152FF9A02384DE7FB8DFF5CF98C" lastPageId="11" lastPageNumber="412" pageId="2" pageNumber="403">
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<subSubSection id="608BA3CFFF9A02314DE7FB8DFE08FBF2" box="[151,403,1086,1113]" pageId="2" pageNumber="403" type="nomenclature">
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<paragraph id="282EF044FF9A02314DE7FB8DFE08FBF2" blockId="2.[151,403,1086,1149]" box="[151,403,1086,1113]" pageId="2" pageNumber="403">
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<heading id="73664728FF9A02314DE7FB8DFE08FBF2" bold="true" box="[151,403,1086,1113]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
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<taxonomicName id="EF918BC7FF9A02314DE7FB8DFE08FBF2" authority="White, 1846" authorityName="White" authorityYear="1846" box="[151,403,1086,1113]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE52C56FF9A02314DE7FB8DFE08FBF2" bold="true" box="[151,403,1086,1113]" pageId="2" pageNumber="403">
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<emphasis id="1AE52C56FF9A02314DE7FB8DFF6DFBF3" bold="true" box="[151,246,1086,1113]" italics="true" pageId="2" pageNumber="403">Atopida</emphasis>
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<bibRefCitation id="4C008DB5FF9A02314D8DFB8DFE08FBF2" author="White, A." box="[253,403,1086,1112]" pageId="2" pageNumber="403" pagination="1 - 24" refId="ref40229" refString="White, A. (1846) Insects. Insects of New Zealand. In: Richardson, J. & Gray, J. E. (Eds.), The Zoology of the Voyage of H. M. S. Erebus and Terror, under the command of Captain Sir James Clark Ross, R. N., F. R. S., during the years 1839 to 1843. 2 (4). Green and Longman, London, pp. 1 - 24, pls. 1 - 6." type="book chapter" year="1846">White, 1846</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="608BA3CFFF9A02314DE7FBD0FE82FBD6" box="[151,281,1122,1149]" pageId="2" pageNumber="403" type="description">
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<paragraph id="282EF044FF9A02314DE7FBD0FE82FBD6" blockId="2.[151,403,1086,1149]" box="[151,281,1122,1149]" pageId="2" pageNumber="403">
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(
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<figureCitation id="B0AAECC1FF9A02314DEFFBD0FE8BFBD7" box="[159,272,1123,1149]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-12="FIGURE 13" captionStart-13="FIGURE 14" captionStart-14="FIGURE 15" captionStart-15="FIGURE 16" captionStart-16="FIGURE 17" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1859,1883]" captionStartId-1="4.[151,250,1894,1918]" captionStartId-10="14.[151,250,783,807]" captionStartId-11="14.[151,250,1955,1979]" captionStartId-12="20.[151,250,1824,1848]" captionStartId-13="22.[151,250,1854,1878]" captionStartId-14="26.[151,250,1934,1958]" captionStartId-15="33.[151,250,1982,2006]" captionStartId-16="39.[151,250,1897,1921]" captionStartId-2="5.[151,250,1792,1816]" captionStartId-3="6.[151,250,1704,1728]" captionStartId-4="7.[151,250,1740,1764]" captionStartId-5="8.[151,250,1562,1586]" captionStartId-6="9.[151,250,1928,1952]" captionStartId-7="10.[151,250,1905,1929]" captionStartId-8="11.[151,250,1333,1357]" captionStartId-9="12.[151,250,1441,1465]" captionTargetBox-0="[223,1364,181,1835]" captionTargetBox-1="[259,1328,181,1871]" captionTargetBox-10="[269,1317,181,759]" captionTargetBox-11="[151,1436,832,1932]" captionTargetBox-12="[151,1436,777,1801]" captionTargetBox-13="[151,1436,633,1830]" captionTargetBox-14="[151,1436,777,1910]" captionTargetBox-15="[165,1421,813,1959]" captionTargetBox-16="[218,1369,181,1873]" captionTargetBox-2="[151,1436,181,1768]" captionTargetBox-3="[151,1436,633,1680]" captionTargetBox-4="[151,1436,181,1716]" captionTargetBox-5="[151,1436,451,1538]" captionTargetBox-6="[151,1436,630,1905]" captionTargetBox-7="[171,1416,489,1881]" captionTargetBox-8="[151,1436,181,1309]" captionTargetBox-9="[151,1436,181,1417]" captionTargetId-0="figure-19@3.[223,1364,181,1835]" captionTargetId-1="figure-19@4.[259,1328,181,1871]" captionTargetId-10="figure-19@14.[269,1317,181,759]" captionTargetId-11="figure-35@14.[151,1436,832,1932]" captionTargetId-12="figure-263@20.[151,1436,777,1801]" captionTargetId-13="figure-197@22.[151,1436,633,1830]" captionTargetId-14="figure-332@26.[151,1436,777,1910]" captionTargetId-15="figure-287@33.[165,1421,813,1959]" captionTargetId-16="figure-19@39.[218,1369,181,1873]" captionTargetId-2="figure-19@5.[151,1436,181,1768]" captionTargetId-3="figure-229@6.[151,1436,633,1680]" captionTargetId-4="figure-19@7.[151,1436,181,1716]" captionTargetId-5="figure-157@8.[151,1436,451,1538]" captionTargetId-6="figure-199@9.[151,1436,630,1905]" captionTargetId-7="figure-162@10.[171,1416,489,1881]" captionTargetId-8="figure-19@11.[151,1436,181,1309]" captionTargetId-9="figure-19@12.[151,1436,181,1417]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="14" captionTargetPageId-11="14" captionTargetPageId-12="20" captionTargetPageId-13="22" captionTargetPageId-14="26" captionTargetPageId-15="33" captionTargetPageId-16="39" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="FIGURE 1. Atopida, dorsal view.A) A. castanea, male; B) same, female; C) same, female with longer setation, D) same, female with not explanate anterolateral angles of pronotum; E) A. hirta, male; F) same, female; G) A. impressa, male, dark form; H) same, male, light form; I) same, female; J) A. insularis sp. nov., male; K) same, female; L) A. lescheni sp. nov., male; M) same, male, light form; N) A. montana, male; O) same, female; P) A. paparoa sp. nov., male; Q) same, male, colour form. Scale bars = 2.0 mm." captionText-1="FIGURE 2. Atopida, dorsal view. A) A. paparoa sp. nov., female; B) A. paringa sp. nov., male; C) A. proba, male; D) same, female; E) A. suffusa, male; F) same, female; G) A. suturalis, male; H) same, female; I) A. tuhua, male; J) s
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<emphasis id="1AE52C56FF9A02314C60FBD1FE82FBD6" bold="true" box="[272,281,1122,1148]" pageId="2" pageNumber="403">)</emphasis>
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</paragraph>
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<paragraph id="282EF044FF9A02314DE7FB14FD9DFB6A" blockId="2.[151,1131,1191,1246]" box="[151,518,1191,1216]" pageId="2" pageNumber="403">
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<treatmentCitation id="A930D655FF9A02314DE7FB14FE1EFB15" author="White, A." box="[151,389,1191,1215]" page="8" pageId="2" pageNumber="403" year="1846">
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<taxonomicName id="EF918BC7FF9A02314DE7FB14FE1EFB15" authority="White, 1846: 8" authorityName="White" authorityPageNumber="8" authorityYear="1846" box="[151,389,1191,1215]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE52C56FF9A02314DE7FB14FF7CFB15" box="[151,231,1191,1215]" italics="true" pageId="2" pageNumber="403">Atopida</emphasis>
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<bibRefCitation id="4C008DB5FF9A02314D9EFB14FE1EFB15" author="White, A." box="[238,389,1191,1215]" pageId="2" pageNumber="403" pagination="1 - 24" refId="ref40229" refString="White, A. (1846) Insects. Insects of New Zealand. In: Richardson, J. & Gray, J. E. (Eds.), The Zoology of the Voyage of H. M. S. Erebus and Terror, under the command of Captain Sir James Clark Ross, R. N., F. R. S., during the years 1839 to 1843. 2 (4). Green and Longman, London, pp. 1 - 24, pls. 1 - 6." type="book chapter" year="1846">White, 1846: 8</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(new genus,
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</paragraph>
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</subSubSection>
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<subSubSection id="608BA3CFFF9A02314F7CFB1BFBF0FB15" box="[524,1131,1191,1216]" pageId="2" pageNumber="403" type="reference_group">
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<paragraph id="282EF044FF9A02314F7CFB1BFBF0FB15" blockId="2.[151,1131,1191,1246]" box="[524,1131,1191,1216]" pageId="2" pageNumber="403">
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<typeStatus id="F72A4EE6FF9A02314F7CFB1BFDA3FB6A" box="[524,568,1192,1216]" pageId="2" pageNumber="403">type</typeStatus>
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species:
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<taxonomicName id="EF918BC7FF9A02314FE5FB14FC50FB15" authority="White, 1846" authorityName="White" authorityYear="1846" box="[661,971,1191,1215]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="castanea">
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<emphasis id="1AE52C56FF9A02314FE5FB14FCDEFB15" box="[661,837,1191,1215]" italics="true" pageId="2" pageNumber="403">Atopida castanea</emphasis>
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<bibRefCitation id="4C008DB5FF9A02314E3BFB14FC50FB15" author="White, A." box="[843,971,1191,1215]" pageId="2" pageNumber="403" pagination="1 - 24" refId="ref40229" refString="White, A. (1846) Insects. Insects of New Zealand. In: Richardson, J. & Gray, J. E. (Eds.), The Zoology of the Voyage of H. M. S. Erebus and Terror, under the command of Captain Sir James Clark Ross, R. N., F. R. S., during the years 1839 to 1843. 2 (4). Green and Longman, London, pp. 1 - 24, pls. 1 - 6." type="book chapter" year="1846">White, 1846</bibRefCitation>
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</taxonomicName>
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, by monotypy).
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</paragraph>
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</treatmentCitationGroup>
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<paragraph id="282EF044FF9A02314DE7FB75FD1CFB74" blockId="2.[151,1131,1191,1246]" box="[151,647,1222,1246]" pageId="2" pageNumber="403">
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<treatmentCitationGroup id="0881D76AFF9A02314DE7FB75FD1CFB74" box="[151,647,1222,1246]" pageId="2" pageNumber="403">
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<treatmentCitation id="A930D655FF9A02314DE7FB75FD95FB74" ID-DOI="http://doi.org/10.5281/zenodo.6043345" ID-Zenodo-Dep="6043345" author="Kialka, A. & Ruta, R." box="[151,526,1222,1246]" httpUri="http://treatment.plazi.org/id/F54487B2FFD9061960F9FA4BFC672A6A" page="27" pageId="2" pageNumber="403" year="2017">
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<taxonomicName id="EF918BC7FF9A02314DE7FB75FD95FB74" authority=": Kialka & Ruta 2017: 21 - 27" authorityName=": Kialka & Ruta" authorityPageNumber="21 - 27" authorityYear="2017" box="[151,526,1222,1246]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE52C56FF9A02314DE7FB75FF7CFB74" box="[151,231,1222,1246]" italics="true" pageId="2" pageNumber="403">Atopida</emphasis>
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:
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|
|
<bibRefCitation id="4C008DB5FF9A02314D85FB75FD95FB74" author="Kialka, A. & Ruta, R." box="[245,526,1222,1246]" pageId="2" pageNumber="403" pagination="1 - 76" refId="ref39179" refString="Kialka, A. & Ruta, R. (2017) An illustrated catalogue of the New Zealand marsh beetles (Coleoptera: Scirtidae). Zootaxa, 4366 (1), 1 - 76. https: // doi. org / 10.11646 / zootaxa. 4366.1.1" type="journal article" year="2017">Kiałka & Ruta 2017: 21–27</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</treatmentCitation>
|
|||
|
|
(catalogue)
|
|||
|
|
</treatmentCitationGroup>
|
|||
|
|
</paragraph>
|
|||
|
|
<subSubSection id="608BA3CFFF9A02314DE7FABFFD72FAC4" pageId="2" pageNumber="403" type="diagnosis">
|
|||
|
|
<paragraph id="282EF044FF9A02314DE7FABFFD72FAC4" blockId="2.[151,1437,1292,2039]" pageId="2" pageNumber="403">
|
|||
|
|
<emphasis id="1AE52C56FF9A02314DE7FABFFE89FA8C" bold="true" box="[151,274,1292,1318]" pageId="2" pageNumber="403">Diagnosis.</emphasis>
|
|||
|
|
Body elongated, small to large; pronotum usually widest in anterior 1/3, rarely in 1/2 of its length; subantennal fossae deep; apical labial palpomere articulated on the side of the preapical one; trigonium of penis not divided, parameres of tegmen without modifications.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="608BA3CFFF9A02394DB7FACBFE78FE7B" lastPageId="10" lastPageNumber="411" pageId="2" pageNumber="403" type="description">
|
|||
|
|
<paragraph id="282EF044FF9A02314DB7FACBFB91FA55" blockId="2.[151,1437,1292,2039]" pageId="2" pageNumber="403">
|
|||
|
|
<emphasis id="1AE52C56FF9A02314DB7FACBFEEEFA38" bold="true" box="[199,373,1400,1426]" pageId="2" pageNumber="403">Redescription.</emphasis>
|
|||
|
|
Male. Body elongated, small to large (TL
|
|||
|
|
<quantity id="EF695DA1FF9A02314E4FFACBFC53FA38" box="[831,968,1400,1426]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="6.9" metricValueMin="2.4" pageId="2" pageNumber="403" unit="mm" value="4.65" valueMax="6.9" valueMin="2.4">2.4–6.9 mm</quantity>
|
|||
|
|
), 1.9–2.4× longer than wide, sides of elytra subparallel, moderately convex. Body yellowish-brown to dark brown, legs and antenna often lighter. Punctation on head and pronotum usually granulate, strong and dense, punctures often touching each other, punctation on elytra simple, not granulate. Body covered with short to long, adherent to erect setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="282EF044FF9A02354DB7F9BBFC22FDCB" blockId="2.[151,1437,1292,2039]" lastBlockId="6.[151,1437,151,609]" lastPageId="6" lastPageNumber="407" pageId="2" pageNumber="403">
|
|||
|
|
Head rather wide, W/L ca. 1.2, base of head covered with pronotum. Eyes big, oval, slightly protuberant, gently faceted. Distance from posterior margin of eye to occipital ridge distinctly longer than diameter of eye. Distance from the ventral margin of the eye to subgenal ridge (
|
|||
|
|
<figureCitation id="B0AAECC1FF9A02314FAFF9E3FCC2F9C0" box="[735,857,1616,1642]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="2" pageNumber="403">Figs 3A, C</figureCitation>
|
|||
|
|
) varies from 0.4–1.0 diameter of eye. Supraantennal ridges (
|
|||
|
|
<figureCitation id="B0AAECC1FF9A02314D9CF9C7FEDBF924" box="[236,320,1652,1678]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="2" pageNumber="403">Fig. 3E</figureCitation>
|
|||
|
|
) well-marked, slightly to significantly elevated, extending into clypeal margin. Subantennal fossae (
|
|||
|
|
<figureCitation id="B0AAECC1FF9A02314DEFF92BFF63F918" box="[159,248,1688,1714]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="2" pageNumber="403">Fig. 3A</figureCitation>
|
|||
|
|
) deep, wide, reach margin of eye and toward bottom of head, separated from eye with subocular carinae. Frontoclypeal suture absent. Clypeus short, strongly transverse, lateral margins strongly converging anteriorly, anterior edge straight to subtly semicircularly emarginated. Anteclypeus well visible, narrower and slightly longer than clypeus. Ventral side of head between subgenal ridges concave. Gular area convex, square. Submentum wider and slightly longer than mentum, flat, hexagonal, square to transverse, separated from gula. Antennae filiform; antennomere 1 cylindrical, broad, slightly flattened; antennomere 2 globular to cylindrical, the shortest; antennomere 3 narrow, ca. 2.2× longer than antennomere 2, antennomeres 4–11 distinctly wider than antennomere 3, antennomere 4 slightly longer than antennomere 3, antennomeres 4–10 as long as antennomere 3, apical antennomere longer, as long as antennomeres 2 and 3 combined. Labrum (
|
|||
|
|
<figureCitation id="B0AAECC1FF9A02314E7AF80BFCFFF878" box="[778,868,1976,2002]" captionStart="FIGURE 4" captionStartId="6.[151,250,1704,1728]" captionTargetBox="[151,1436,633,1680]" captionTargetId="figure-229@6.[151,1436,633,1680]" captionTargetPageId="6" captionText="FIGURE 4. Atopida suturalis, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6986483" httpUri="https://zenodo.org/record/6986483/files/figure.png" pageId="2" pageNumber="403">Fig. 4C</figureCitation>
|
|||
|
|
) narrower than clypeus, transversely rectangular, anterior margin deeply semicircularly indented, with long setae, front angles rounded. Mandibles (
|
|||
|
|
<figureCitation id="B0AAECC1FF9A02314992F86FFAA2F85C" box="[1250,1337,2012,2038]" captionStart="FIGURE 4" captionStartId="6.[151,250,1704,1728]" captionTargetBox="[151,1436,633,1680]" captionTargetId="figure-229@6.[151,1436,633,1680]" captionTargetPageId="6" captionText="FIGURE 4. Atopida suturalis, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6986483" httpUri="https://zenodo.org/record/6986483/files/figure.png" pageId="2" pageNumber="403">Fig. 4A</figureCitation>
|
|||
|
|
) slightly asymmetrical, narrowly triangular, ca. 2× longer than width at bases, apices unidentate, acute; with short carina on the external edge, abruptly curved in apical third, with setae covering 2/3 of its length; internal edge with triangular denticle and carina with 3 smaller denticles, not covered with setae, denticles developed somewhat better on left mandible; prostheca absent; molar region without denticles or setation. Maxillae (
|
|||
|
|
<figureCitation id="B0AAECC1FF9E0235495FFEB0FB13FEB7" box="[1071,1160,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1704,1728]" captionTargetBox="[151,1436,633,1680]" captionTargetId="figure-229@6.[151,1436,633,1680]" captionTargetPageId="6" captionText="FIGURE 4. Atopida suturalis, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6986483" httpUri="https://zenodo.org/record/6986483/files/figure.png" pageId="6" pageNumber="407">Fig. 4C</figureCitation>
|
|||
|
|
): basistipes with distinct setation; maxillary palpi short; palpomere 1 poorly subconical, slightly longer than wide, palpomeres 2 and 3 about 2× longer than palpomere 1, apical maxillary palpomere about same length as palpomere 3, slightly narrower, bulbous at base and pointed at apex, covered with setae at basal 2/3. Galea slightly elongated, apex with
|
|||
|
|
<specimenCount id="3E973BCDFF9E02354840FEDCFA07FE23" box="[1328,1436,367,393]" pageId="6" pageNumber="407" type="generic" typeStatus="types">two types</specimenCount>
|
|||
|
|
of setae—some of setae short and straight, forming fan-shaped structure, other setae much longer, “feathered”, and irregularly arranged. Lacinia elongated, covered with straight, sparse setae in the lower half and with longer, denser, bulkier on apex. Labium (
|
|||
|
|
<figureCitation id="B0AAECC1FF9E02354CB7FE68FDBDFE5F" box="[455,550,475,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1704,1728]" captionTargetBox="[151,1436,633,1680]" captionTargetId="figure-229@6.[151,1436,633,1680]" captionTargetPageId="6" captionText="FIGURE 4. Atopida suturalis, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6986483" httpUri="https://zenodo.org/record/6986483/files/figure.png" pageId="6" pageNumber="407">Fig. 4D</figureCitation>
|
|||
|
|
): mentum subtrapezoidal, as wide as long; praementum strongly sclerotized, narrow at base, then strongly widening, anterior edge semicircular, covered with numerous straight setae. Labial palpomere 1 angular, long, without setae; palpomere 2 broadly oval, larger than palpomere 1; apical palpomere inserted laterally, oval, slightly shorter than palpomere 2, without setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="7CEEA0CCFF9B02304DE7F8F0FE9BF841" ID-DOI="http://doi.org/10.5281/zenodo.6986473" ID-Zenodo-Dep="6986473" httpUri="https://zenodo.org/record/6986473/files/figure.png" pageId="3" pageNumber="404" startId="3.[151,250,1859,1883]" targetBox="[223,1364,181,1835]" targetPageId="3">
|
|||
|
|
<paragraph id="282EF044FF9B02304DE7F8F0FE9BF841" blockId="3.[151,1436,1859,2027]" pageId="3" pageNumber="404">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304DE7F8F0FE89F8F1" bold="true" box="[151,274,1859,1883]" pageId="3" pageNumber="404">FIGURE 1.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304C67F8F0FEFCF8F1" authorityName=": Kialka & Ruta" authorityYear="2017" box="[279,359,1859,1883]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304C67F8F0FEFCF8F1" box="[279,359,1859,1883]" italics="true" pageId="3" pageNumber="404">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, dorsal view. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304F60F8F0FD1EF8F1" authorityName="White" authorityYear="1846" box="[528,645,1859,1883]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="castanea">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304F60F8F0FD1EF8F1" box="[528,645,1859,1883]" italics="true" pageId="3" pageNumber="404">A. castanea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; B) same, female; C) same, female with longer setation, D) same, female with not explanate anterolateral angles of pronotum; E)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304FA7F8D4FCBEF8D5" authorityName="Broun" authorityYear="1880" box="[727,805,1895,1919]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="hirta">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304FA7F8D4FCBEF8D5" box="[727,805,1895,1919]" italics="true" pageId="3" pageNumber="404">A. hirta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; F) same, female; G)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304939F8D4FB5AF8D5" authorityName="Broun" authorityYear="1914" box="[1097,1217,1895,1919]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="impressa">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304939F8D4FB5AF8D5" box="[1097,1217,1895,1919]" italics="true" pageId="3" pageNumber="404">A. impressa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male, dark form; H) same, male, light form; I) same, female; J)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304F38F838FD26F809" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[584,701,1931,1955]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304F38F838FD26F809" box="[584,701,1931,1955]" italics="true" pageId="3" pageNumber="404">A. insularis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9B02304FB2F838FC89F809" bold="true" box="[706,786,1931,1955]" pageId="3" pageNumber="404">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9B02304FB2F838FC89F809" box="[706,786,1931,1955]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; K) same, female; L)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B0230495EF838FB07F809" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[1070,1180,1931,1955]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9B0230495EF838FB07F809" box="[1070,1180,1931,1955]" italics="true" pageId="3" pageNumber="404">A. lescheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9B023049D2F838FB69F809" bold="true" box="[1186,1266,1931,1955]" pageId="3" pageNumber="404">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9B023049D2F838FB69F809" box="[1186,1266,1931,1955]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; M) same, male, light form; N)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304C1AF81CFE44F86D" authorityName=", Broun" authorityYear="1921" box="[362,479,1967,1991]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="montana">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304C1AF81CFE44F86D" box="[362,479,1967,1991]" italics="true" pageId="3" pageNumber="404">A. montana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; O) same, female; P)
|
|||
|
|
<taxonomicName id="EF918BC7FF9B02304F8CF81CFCF6F86D" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[764,877,1967,1991]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9B02304F8CF81CFCF6F86D" box="[764,877,1967,1991]" italics="true" pageId="3" pageNumber="404">A. paparoa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9B02304E03F81CFC5FF86D" bold="true" box="[883,964,1967,1991]" pageId="3" pageNumber="404">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9B02304E03F81CFC5FF86D" box="[883,964,1967,1991]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; Q) same, male, colour form. Scale bars = 2.0 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="7CEEA0CCFF9C02374DE7F8D5FD86F840" ID-DOI="http://doi.org/10.5281/zenodo.6986477" ID-Zenodo-Dep="6986477" httpUri="https://zenodo.org/record/6986477/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1894,1918]" targetBox="[259,1328,181,1871]" targetPageId="4">
|
|||
|
|
<paragraph id="282EF044FF9C02374DE7F8D5FD86F840" blockId="4.[151,1437,1894,2027]" pageId="4" pageNumber="405">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374DE7F8D5FE8FF8D4" bold="true" box="[151,276,1894,1918]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374C6BF8D4FEF0F8D5" authorityName=": Kialka & Ruta" authorityYear="2017" box="[283,363,1895,1919]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374C6BF8D4FEF0F8D5" box="[283,363,1895,1919]" italics="true" pageId="4" pageNumber="405">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, dorsal view. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374F6BF8D4FD16F8D4" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[539,653,1895,1918]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374F6BF8D4FD16F8D4" box="[539,653,1895,1918]" italics="true" pageId="4" pageNumber="405">A. paparoa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9C02374FE4F8D4FD7DF8D5" bold="true" box="[660,742,1895,1919]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9C02374FE4F8D4FD7DF8D5" box="[660,742,1895,1919]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, female; B)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374E17F8D4FC4FF8D4" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[871,980,1895,1918]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="paringa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374E17F8D4FC4FF8D4" box="[871,980,1895,1918]" italics="true" pageId="4" pageNumber="405">A. paringa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9C02374EABF8D4FBB6F8D5" bold="true" box="[987,1069,1895,1919]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9C02374EABF8D4FBB6F8D5" box="[987,1069,1895,1919]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; C)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C023749EBF8D4FB6FF8D5" authorityName="Sharp" authorityYear="1878" box="[1179,1268,1895,1919]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="proba">
|
|||
|
|
<emphasis id="1AE52C56FF9C023749EBF8D4FB6FF8D5" box="[1179,1268,1895,1919]" italics="true" pageId="4" pageNumber="405">A. proba</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; D) same, female; E)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374C78F838FEF6F809" authorityName="Broun" authorityYear="1910" box="[264,365,1931,1955]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="suffusa">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374C78F838FEF6F809" box="[264,365,1931,1955]" italics="true" pageId="4" pageNumber="405">A. suffusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; F) same, female; G)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374FFAF838FD64F809" authorityName="White" authorityYear="1846" box="[650,767,1931,1955]" class="Insecta" family="Ptinidae" genus="Ptinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374FFAF838FD64F809" box="[650,767,1931,1955]" italics="true" pageId="4" pageNumber="405">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; H) same, female; I)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374967F838FBF6F809" authorityName="Kiałka & Ruta" authorityYear="2022" box="[1047,1133,1931,1955]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="tuhua">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374967F838FBF6F809" box="[1047,1133,1931,1955]" italics="true" pageId="4" pageNumber="405">A. tuhua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; J) same, female; K)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C023748F6F838FF40F86D" authorityName="Broun" authorityYear="1921" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="villosa">
|
|||
|
|
<emphasis id="1AE52C56FF9C023748F6F838FF40F86D" italics="true" pageId="4" pageNumber="405">A. villosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; L) same, female; M)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374C86F81CFDF3F86C" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[502,616,1967,1990]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="waipoua" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374C86F81CFDF3F86C" box="[502,616,1967,1990]" italics="true" pageId="4" pageNumber="405">A. waipoua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9C02374F1DF81CFD26F86D" bold="true" box="[621,701,1967,1991]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9C02374F1DF81CFD26F86D" box="[621,701,1967,1991]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; N)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C02374E56F81CFC05F86D" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[806,926,1967,1991]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="walkerae" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9C02374E56F81CFC05F86D" box="[806,926,1967,1991]" italics="true" pageId="4" pageNumber="405">A. walkerae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9C02374ED3F81CFC69F86D" bold="true" box="[931,1010,1967,1991]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9C02374ED3F81CFC69F86D" box="[931,1010,1967,1991]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; O)
|
|||
|
|
<taxonomicName id="EF918BC7FF9C0237492CF81CFA9FF86D" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[1116,1284,1967,1991]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="westlandensis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9C0237492CF81CFA9FF86D" box="[1116,1284,1967,1991]" italics="true" pageId="4" pageNumber="405">A. westlandensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9C02374879F81CFAC3F86D" bold="true" box="[1289,1368,1967,1991]" pageId="4" pageNumber="405">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9C02374879F81CFAC3F86D" box="[1289,1368,1967,1991]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; P) same, female. Scale bars = 2.0 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="7CEEA0CCFF9D02364DE7F8B3FDC1F802" ID-DOI="http://doi.org/10.5281/zenodo.6986479" ID-Zenodo-Dep="6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1792,1816]" targetBox="[151,1436,181,1768]" targetPageId="5">
|
|||
|
|
<paragraph id="282EF044FF9D02364DE7F8B3FDC1F802" blockId="5.[151,1437,1792,1960]" pageId="5" pageNumber="406">
|
|||
|
|
<emphasis id="1AE52C56FF9D02364DE7F8B3FE8FF8B2" bold="true" box="[151,276,1792,1816]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9D02364C6CF8B3FEF7F8B2" authorityName=": Kialka & Ruta" authorityYear="2017" box="[284,364,1792,1816]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9D02364C6CF8B3FEF7F8B2" box="[284,364,1792,1816]" italics="true" pageId="5" pageNumber="406">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, morphological details. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF9D02364FF6F8B2FD66F8B2" authorityName="White" authorityYear="1846" box="[646,765,1793,1816]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="castanea">
|
|||
|
|
<emphasis id="1AE52C56FF9D02364FF6F8B2FD66F8B2" box="[646,765,1793,1816]" italics="true" pageId="5" pageNumber="406">A. castanea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C)
|
|||
|
|
<taxonomicName id="EF918BC7FF9D02364C3CF896FE58F896" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[332,451,1828,1852]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9D02364C3CF896FE58F896" box="[332,451,1828,1852]" italics="true" pageId="5" pageNumber="406">A. insularis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9D02364CBCF896FD85F897" bold="true" box="[460,542,1829,1853]" pageId="5" pageNumber="406">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9D02364CBCF896FD85F897" box="[460,542,1829,1853]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E)
|
|||
|
|
<taxonomicName id="EF918BC7FF9D02364C4AF8FAFE32F8CA" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[314,425,1864,1888]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9D02364C4AF8FAFE32F8CA" box="[314,425,1864,1888]" italics="true" pageId="5" pageNumber="406">A. lescheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9D02364CC0F8FAFD9AF8CB" bold="true" box="[432,513,1865,1889]" pageId="5" pageNumber="406">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9D02364CC0F8FAFD9AF8CB" box="[432,513,1865,1889]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male, head, projection of supraantennal ridge marked with arrow; F)
|
|||
|
|
<taxonomicName id="EF918BC7FF9D023649BFF8FAFADFF8CA" authorityName="White" authorityYear="1846" box="[1231,1348,1864,1888]" class="Insecta" family="Ptinidae" genus="Ptinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF9D023649BFF8FAFADFF8CA" box="[1231,1348,1864,1888]" italics="true" pageId="5" pageNumber="406">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="7CEEA0CCFF9E02354DE7F91BFA97F915" ID-DOI="http://doi.org/10.5281/zenodo.6986483" ID-Zenodo-Dep="6986483" box="[151,1292,1704,1728]" httpUri="https://zenodo.org/record/6986483/files/figure.png" pageId="6" pageNumber="407" startId="6.[151,250,1704,1728]" targetBox="[151,1436,633,1680]" targetPageId="6">
|
|||
|
|
<paragraph id="282EF044FF9E02354DE7F91BFA97F915" blockId="6.[151,1292,1704,1728]" box="[151,1292,1704,1728]" pageId="6" pageNumber="407">
|
|||
|
|
<emphasis id="1AE52C56FF9E02354DE7F91BFE88F96A" bold="true" box="[151,275,1704,1728]" pageId="6" pageNumber="407">FIGURE 4.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9E02354C69F91BFE52F96A" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[281,457,1704,1728]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF9E02354C69F91BFE52F96A" box="[281,457,1704,1728]" italics="true" pageId="6" pageNumber="407">Atopida suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, mouthparts. A) mandible, B) maxilla, C) labrum, D) labium. Scale bar = 0.2 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="282EF044FF9E02354DB7F95CFEB3F84B" blockId="6.[151,1437,1774,2017]" pageId="6" pageNumber="407">
|
|||
|
|
Pronotum (
|
|||
|
|
<figureCitation id="B0AAECC1FF9E02354C36F95DFE12F8A3" box="[326,393,1774,1801]" captionStart="FIGURE 5" captionStartId="7.[151,250,1740,1764]" captionTargetBox="[151,1436,181,1716]" captionTargetId="figure-19@7.[151,1436,181,1716]" captionTargetPageId="7" captionText="FIGURE 5. Atopida, pronotum. A) A. castanea, male; B) same, female; C) A. hirta, male; D) same, female; E) A. impressa, male; F) same, female; G) A. insularis sp. nov., male; H) same, female; I) A. lescheni sp. nov., male; J) A. montana, male; K) same, female; L) A. paparoa sp. nov., male; M) same, female; N) A. paringa sp. nov., male; O) A. proba, male; P) same, female; Q) A. suffusa, male; R) same, female; S) A. suturalis, male; T) same, female; U) A. tuhua sp. nov., male; V) same, female; W) A. villosa, male; X) same, female; Y) A. waipoua sp. nov., male; Z) A. walkerae sp. nov., male; AA) A. westlandensis sp. nov., male; BB) same, female. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986489" httpUri="https://zenodo.org/record/6986489/files/figure.png" pageId="6" pageNumber="407">Fig. 5</figureCitation>
|
|||
|
|
) transverse, subrectangular, narrower at base than in anterior part, in most species widest in anterior 1/3. Lateral edges rounded, even or crenulate. Anterior margin slightly rounded, not projecting beyond anterolateral angles, covering base of head. Anterolateral angles subtriangular, often acute and projecting. Posterolateral angles obtuse to acute, well-marked. Basal portion of pronotum with well-marked groove, basal margin of pronotum bisinuate (
|
|||
|
|
<figureCitation id="B0AAECC1FF9E02354F73F8CDFDFAF832" box="[515,609,1918,1944]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="6" pageNumber="407">Fig. 3D</figureCitation>
|
|||
|
|
). Prosternal process elongate (
|
|||
|
|
<figureCitation id="B0AAECC1FF9E02354EB5F8CDFBD2F832" box="[965,1097,1918,1944]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="6" pageNumber="407">Figs 3B, C</figureCitation>
|
|||
|
|
), ca. 3–4× longer than wide, lanceolate to fusiform, apex narrow, bent ventrally, covered with setae. Procoxae long, subconical, procoxal cavities widely open.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="7CEEA0CCFF9F02344DE7F97FFDE2F832" ID-DOI="http://doi.org/10.5281/zenodo.6986489" ID-Zenodo-Dep="6986489" httpUri="https://zenodo.org/record/6986489/files/figure.png" pageId="7" pageNumber="408" startId="7.[151,250,1740,1764]" targetBox="[151,1436,181,1716]" targetPageId="7">
|
|||
|
|
<paragraph id="282EF044FF9F02344DE7F97FFDE2F832" blockId="7.[151,1437,1740,1944]" pageId="7" pageNumber="408">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344DE7F97FFE8FF94E" bold="true" box="[151,276,1740,1764]" pageId="7" pageNumber="408">FIGURE 5.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344C6CF97FFEF7F94E" authorityName=": Kialka & Ruta" authorityYear="2017" box="[284,364,1740,1764]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344C6CF97FFEF7F94E" box="[284,364,1740,1764]" italics="true" pageId="7" pageNumber="408">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, pronotum. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344F7CF97FFD18F94E" authorityName="White" authorityYear="1846" box="[524,643,1740,1764]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="castanea">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344F7CF97FFD18F94E" box="[524,643,1740,1764]" italics="true" pageId="7" pageNumber="408">A. castanea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; B) same, female; C)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344ED9F97FFC63F94E" authorityName="Broun" authorityYear="1880" box="[937,1016,1740,1764]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="hirta">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344ED9F97FFC63F94E" box="[937,1016,1740,1764]" italics="true" pageId="7" pageNumber="408">A. hirta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; D) same, female; E)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F0234486EF97FFA0DF94E" authorityName="Broun" authorityYear="1914" box="[1310,1430,1740,1764]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="impressa">
|
|||
|
|
<emphasis id="1AE52C56FF9F0234486EF97FFA0DF94E" box="[1310,1430,1740,1764]" italics="true" pageId="7" pageNumber="408">A. impressa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; F) same, female; G)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344CDAF943FD84F8A2" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[426,543,1776,1800]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344CDAF943FD84F8A2" box="[426,543,1776,1800]" italics="true" pageId="7" pageNumber="408">A. insularis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344F56F943FDECF8A2" bold="true" box="[550,631,1776,1800]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344F56F943FDECF8A2" box="[550,631,1776,1800]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; H) same, female; I)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344EE2F943FB9AF8A2" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[914,1025,1776,1800]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344EE2F943FB9AF8A2" box="[914,1025,1776,1800]" italics="true" pageId="7" pageNumber="408">A. lescheni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344978F943FBC2F8A2" bold="true" box="[1032,1113,1776,1800]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344978F943FBC2F8A2" box="[1032,1113,1776,1800]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; J)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F023449B0F943FAAEF8A2" authorityName=", Broun" authorityYear="1921" box="[1216,1333,1776,1800]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="montana">
|
|||
|
|
<emphasis id="1AE52C56FF9F023449B0F943FAAEF8A2" box="[1216,1333,1776,1800]" italics="true" pageId="7" pageNumber="408">A. montana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; K) same, female; L)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344C36F8A7FE2DF886" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[326,438,1812,1836]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344C36F8A7FE2DF886" box="[326,438,1812,1836]" italics="true" pageId="7" pageNumber="408">A. paparoa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344CCBF8A7FD90F886" bold="true" box="[443,523,1812,1836]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344CCBF8A7FD90F886" box="[443,523,1812,1836]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; M) same, female; N)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344E5AF8A7FC0EF886" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[810,917,1812,1836]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="paringa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344E5AF8A7FC0EF886" box="[810,917,1812,1836]" italics="true" pageId="7" pageNumber="408">A. paringa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344EEBF8A7FC71F886" bold="true" box="[923,1002,1812,1836]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344EEBF8A7FC71F886" box="[923,1002,1812,1836]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; O)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344924F8A7FB37F886" authorityName="Sharp" authorityYear="1878" box="[1108,1196,1812,1836]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="proba">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344924F8A7FB37F886" box="[1108,1196,1812,1836]" italics="true" pageId="7" pageNumber="408">A. proba</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; P) same, female; Q)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344DC9F88BFE86F8FA" authorityName="Broun" authorityYear="1910" box="[185,285,1848,1872]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="suffusa">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344DC9F88BFE86F8FA" box="[185,285,1848,1872]" italics="true" pageId="7" pageNumber="408">A. suffusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; R) same, female; S)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344F4AF88BFD34F8FA" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[570,687,1848,1872]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344F4AF88BFD34F8FA" box="[570,687,1848,1872]" italics="true" pageId="7" pageNumber="408">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; T) same, female; U)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344EBEF88BFBBFF8FA" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[974,1060,1848,1872]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tuhua" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344EBEF88BFBBFF8FA" box="[974,1060,1848,1872]" italics="true" pageId="7" pageNumber="408">A. tuhua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F0234495BF88BFBE0F8FA" bold="true" box="[1067,1147,1848,1872]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F0234495BF88BFBE0F8FA" box="[1067,1147,1848,1872]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; V) same, female; W)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344DE7F8EFFF6CF8DE" authorityName="Broun" authorityYear="1921" box="[151,247,1884,1908]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="villosa">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344DE7F8EFFF6CF8DE" box="[151,247,1884,1908]" italics="true" pageId="7" pageNumber="408">A. villosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; X) same, female; Y)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344F66F8EFFD12F8DE" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[534,649,1884,1908]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="waipoua" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344F66F8EFFD12F8DE" box="[534,649,1884,1908]" italics="true" pageId="7" pageNumber="408">A. waipoua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344FFFF8EFFD44F8DE" bold="true" box="[655,735,1884,1908]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344FFFF8EFFD44F8DE" box="[655,735,1884,1908]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; Z)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F02344E39F8EFFC59F8DE" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[841,962,1884,1908]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="walkerae" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F02344E39F8EFFC59F8DE" box="[841,962,1884,1908]" italics="true" pageId="7" pageNumber="408">A. walkerae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344EB8F8EFFB82F8DE" bold="true" box="[968,1049,1884,1908]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344EB8F8EFFB82F8DE" box="[968,1049,1884,1908]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; AA)
|
|||
|
|
<taxonomicName id="EF918BC7FF9F023449E6F8EFFAA4F8DE" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[1174,1343,1884,1908]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="westlandensis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9F023449E6F8EFFAA4F8DE" box="[1174,1343,1884,1908]" italics="true" pageId="7" pageNumber="408">A. westlandensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9F02344835F8EFFA0EF8DE" bold="true" box="[1349,1429,1884,1908]" pageId="7" pageNumber="408">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9F02344835F8EFFA0EF8DE" box="[1349,1429,1884,1908]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male; BB) same, female. Not to the same scale.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="282EF044FF90023B4DB7FF24FB1AFEEA" blockId="8.[151,1437,151,433]" pageId="8" pageNumber="409">Scutellar shield triangular, subequilateral (slightly longer than it is wide), flat, with slightly rounded edges and acute apex. Elytra elongated, sides subparallel in proximal 1/2, angle between elytra and pronotum well-marked in dorsal view. Elytra distinctly wider than pronotum. Humeri pronounced, apices regularly rounded. Elytra in most species without longitudinal carinae. Epipleuron flat, narrow, slightly narrower than metepisternum at base, then regularly tapering towards apex of elytra, complete to apex. Sides of elytra not explanate.</paragraph>
|
|||
|
|
<paragraph id="282EF044FF90023B4DB7FEFAFA06FE01" blockId="8.[151,1437,151,433]" pageId="8" pageNumber="409">
|
|||
|
|
Hind wings (
|
|||
|
|
<figureCitation id="B0AAECC1FF90023B4C2BFEFAFE3BFEC9" box="[347,416,329,355]" captionStart="FIGURE 6" captionStartId="8.[151,250,1562,1586]" captionTargetBox="[151,1436,451,1538]" captionTargetId="figure-157@8.[151,1436,451,1538]" captionTargetPageId="8" captionText="FIGURE 6. Atopida, hind wings. A) A. suturalis, macropterous wing; B) A. impressa brachypterous wing." figureDoi="http://doi.org/10.5281/zenodo.6986491" httpUri="https://zenodo.org/record/6986491/files/figure.png" pageId="8" pageNumber="409">Fig. 6</figureCitation>
|
|||
|
|
) ca. 2.2× longer than its width; radial cell subtriangular; apical portion of r3 well visible, straight; RML arcuate, forming acute angle with MP
|
|||
|
|
<subScript id="B415F201FF90023B4FAFFE33FD63FE24" attach="right" box="[735,760,384,398]" fontSize="6" pageId="8" pageNumber="409">1+2</subScript>
|
|||
|
|
; CuA
|
|||
|
|
<subScript id="B415F201FF90023B4E4BFE33FCD8FE24" attach="left" box="[827,835,384,398]" fontSize="6" pageId="8" pageNumber="409">1</subScript>
|
|||
|
|
long, oblique, joining MP
|
|||
|
|
<subScript id="B415F201FF90023B4919FE33FB19FE24" attach="left" box="[1129,1154,384,398]" fontSize="6" pageId="8" pageNumber="409">3+4</subScript>
|
|||
|
|
and CAS; anal field with well-markedAP
|
|||
|
|
<subScript id="B415F201FF90023B4C3AFE10FEFFFE1B" attach="left" box="[330,356,419,433]" fontSize="6" pageId="8" pageNumber="409">3+4</subScript>
|
|||
|
|
.Medial field with3free veins;pigmented areas in radial cell, medial,radial,central and apical fields.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="7CEEA0CCFF90023B4DE7F9A9FB4AF998" ID-DOI="http://doi.org/10.5281/zenodo.6986491" ID-Zenodo-Dep="6986491" box="[151,1233,1562,1586]" httpUri="https://zenodo.org/record/6986491/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,250,1562,1586]" targetBox="[151,1436,451,1538]" targetPageId="8">
|
|||
|
|
<paragraph id="282EF044FF90023B4DE7F9A9FB4AF998" blockId="8.[151,1233,1562,1586]" box="[151,1233,1562,1586]" pageId="8" pageNumber="409">
|
|||
|
|
<emphasis id="1AE52C56FF90023B4DE7F9A9FE88F998" bold="true" box="[151,275,1562,1586]" pageId="8" pageNumber="409">FIGURE 6.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF90023B4C69F9A9FEF2F998" authorityName=": Kialka & Ruta" authorityYear="2017" box="[281,361,1562,1586]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF90023B4C69F9A9FEF2F998" box="[281,361,1562,1586]" italics="true" pageId="8" pageNumber="409">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, hind wings. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF90023B4F62F9A9FD13F998" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[530,648,1562,1586]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF90023B4F62F9A9FD13F998" box="[530,648,1562,1586]" italics="true" pageId="8" pageNumber="409">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, macropterous wing; B)
|
|||
|
|
<taxonomicName id="EF918BC7FF90023B4EF4F9A9FC60F998" authorityName="Broun" authorityYear="1914" box="[900,1019,1562,1586]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="impressa">
|
|||
|
|
<emphasis id="1AE52C56FF90023B4EF4F9A9FC60F998" box="[900,1019,1562,1586]" italics="true" pageId="8" pageNumber="409">A. impressa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
brachypterous wing.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="282EF044FF90023B4DB7F9EAFD89F8E2" blockId="8.[151,1437,1625,2042]" pageId="8" pageNumber="409">
|
|||
|
|
Mesoventrite short, anterior edge with pair of oblique procoxal rests, suture present throughout mesoventral process. Prosternal process rest (
|
|||
|
|
<figureCitation id="B0AAECC1FF90023B4F74F9CFFDC1F93D" box="[516,602,1660,1687]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="8" pageNumber="409">Fig. 3B</figureCitation>
|
|||
|
|
) triangular, narrow to wide, with slightly rounded edges, slightly concave. Mesoventral process (
|
|||
|
|
<figureCitation id="B0AAECC1FF90023B4CE3F913FE70F910" box="[403,491,1696,1722]" captionStart="FIGURE 3" captionStartId="5.[151,250,1792,1816]" captionTargetBox="[151,1436,181,1768]" captionTargetId="figure-19@5.[151,1436,181,1768]" captionTargetPageId="5" captionText="FIGURE 3. Atopida, morphological details. A) A. castanea, head, anterior view; B) same, ventral portion of prosternum and mesoventrite; C) A. insularis sp. nov., ventral portion of head, prosternum and mesoventrite; D) same, head and pronotum, dorsal view; E) A. lescheni sp. nov., male, head, projection of supraantennal ridge marked with arrow; F) A. suturalis, female, head, projection of supraantennal ridge marked with arrow. Abbreviations: mp—mesoventral process, pp—prosternal process, sgr—subgenal ridge, soc—subocular carina." figureDoi="http://doi.org/10.5281/zenodo.6986479" httpUri="https://zenodo.org/record/6986479/files/figure.png" pageId="8" pageNumber="409">Fig. 3B</figureCitation>
|
|||
|
|
) triangular to subrectangular, ca. 1.5× longer than wide, apex bilobed. Mesocoxal cavities widely separated. Mesepisterna moderately wide, rhomboid, middle part with depression for procoxae. Mesepimeron narrower than mesepisternum, triangular. Mesocoxal cavities closed laterally by narrow processes of mesepimeron and metasternum. Mesocoxae weakly elongated, ovally transverse (ca. 1.6× longer than wide), slightly protruding and triangular.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="282EF044FF90023B4DB7F8E2FAE8F825" blockId="8.[151,1437,1625,2042]" pageId="8" pageNumber="409">Metaventrite long, convex. Metaventral discrimen present in posterior 1/4–1/2. Metepisternum wide, rhomboid. Metacoxae big, ca. 3.5× longer than wide. Metacoxal plates touching each other mesally over the entire length.</paragraph>
|
|||
|
|
<paragraph id="282EF044FF90023B4DB7F82AFB12F853" blockId="8.[151,1437,1625,2042]" pageId="8" pageNumber="409">Legs long, femora not enlarged, not saltatorial, extending beyond elytral margin; tibiae slightly longer than femora, slender and, especially metatibiae, subtly S-shaped, slightly extended at apex; with well-marked paired longitudinal carinae; tibial spurs well developed, dorsal spurs ca. 1.5× longer than ventral.</paragraph>
|
|||
|
|
<paragraph id="282EF044FF91023A4DB7FF24FD53FF7E" blockId="9.[151,1437,150,607]" pageId="9" pageNumber="410">Abdomen longer than wide, moderately convex. Ventrites 1–5 of subequal length, regularly covered with setae; ventrite 5 rounded to subtriangular, apex rounded.</paragraph>
|
|||
|
|
<paragraph id="282EF044FF91023A4DB7FF6DFE4CFDF5" blockId="9.[151,1437,150,607]" pageId="9" pageNumber="410">
|
|||
|
|
Morphology of terminal sclerites of males uniform (
|
|||
|
|
<figureCitation id="B0AAECC1FF91023A4E57FF6DFC37FF52" box="[807,940,222,248]" captionStart="FIGURE 7" captionStartId="9.[151,250,1928,1952]" captionTargetBox="[151,1436,630,1905]" captionTargetId="figure-199@9.[151,1436,630,1905]" captionTargetPageId="9" captionText="FIGURE 7. Atopida, male terminal segments and genitalia (A, C) and gut contents (B). A) A. suturalis, B) same, gut contents, C) A. paparoa sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6986493" httpUri="https://zenodo.org/record/6986493/files/figure.png" pageId="9" pageNumber="410">Figs 7A, C</figureCitation>
|
|||
|
|
). Sternite VIII transversely rectangular, Ushaped with very few setae and punctures concentrated on apical margin, distinctly wider than longer; sternite IX rectangular, consisting of subtriangular hemisternites, apical part covered with medium length setae, apodemes short and barely visible. Tergite VIII rectangular with rather distinct apodemes, numerous short setae in apical part; tergite IX elongated, apical part membranous, almost devoid of setae, with long apodemes. Aedeagus symmetrical. Tegmen with small styli, parameres widely rectangular, with rounded apical portions to narrowly triangular with pointed apices, with scarce punctures and numerous rows of delicate setae in apical part. Trigonium of penis rather narrow, apex rounded or pointed, shorter than parameroids or as long (rarely slightly longer) as parameroids, rarely trigonium widely oval, bulbous. Pala triangular, slightly longer than parameroids, in some species basal portion emarginated. Parameroids elongated, subtriangular to subrectangular, apices rounded, tempered or pointed, apical part covered with punctures.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="7CEEA0CCFF91023A4DE7F83BFEE4F86F" ID-DOI="http://doi.org/10.5281/zenodo.6986493" ID-Zenodo-Dep="6986493" httpUri="https://zenodo.org/record/6986493/files/figure.png" pageId="9" pageNumber="410" startId="9.[151,250,1928,1952]" targetBox="[151,1436,630,1905]" targetPageId="9">
|
|||
|
|
<paragraph id="282EF044FF91023A4DE7F83BFEE4F86F" blockId="9.[151,1436,1928,2025]" pageId="9" pageNumber="410">
|
|||
|
|
<emphasis id="1AE52C56FF91023A4DE7F83BFE88F80B" bold="true" box="[151,275,1928,1953]" pageId="9" pageNumber="410">FIGURE 7.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF91023A4C6AF83AFEF1F80B" authorityName=": Kialka & Ruta" authorityYear="2017" box="[282,362,1929,1953]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF91023A4C6AF83AFEF1F80B" box="[282,362,1929,1953]" italics="true" pageId="9" pageNumber="410">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male terminal segments and genitalia (A, C) and gut contents (B). A)
|
|||
|
|
<taxonomicName id="EF918BC7FF91023A4948F83AFB35F80B" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[1080,1198,1929,1953]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF91023A4948F83AFB35F80B" box="[1080,1198,1929,1953]" italics="true" pageId="9" pageNumber="410">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, B) same, gut contents, C)
|
|||
|
|
<taxonomicName id="EF918BC7FF91023A4DC7F81EFEB3F86E" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[183,296,1965,1988]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF91023A4DC7F81EFEB3F86E" box="[183,296,1965,1988]" italics="true" pageId="9" pageNumber="410">A. paparoa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF91023A4C5EF81EFEE4F86F" box="[302,383,1965,1989]" pageId="9" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="282EF044FF9102394DB7F87CFE78FE7B" blockId="9.[151,1436,1928,2025]" lastBlockId="10.[151,1437,151,465]" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410">
|
|||
|
|
Female. Many species with significant sexual dimorphism. Pronotum often more transverse than in males, punctation of pronotum often sparser than in males, with shining interspaces. Punctation of elytra often coarser than in males. Hind wings often reduced—brachypterous (
|
|||
|
|
<figureCitation id="B0AAECC1FF9202394F89FF08FCC9FF7F" box="[761,850,187,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1562,1586]" captionTargetBox="[151,1436,451,1538]" captionTargetId="figure-157@8.[151,1436,451,1538]" captionTargetPageId="8" captionText="FIGURE 6. Atopida, hind wings. A) A. suturalis, macropterous wing; B) A. impressa brachypterous wing." figureDoi="http://doi.org/10.5281/zenodo.6986491" httpUri="https://zenodo.org/record/6986491/files/figure.png" pageId="10" pageNumber="411">Fig. 6B</figureCitation>
|
|||
|
|
), rarely apterous, no infraspecific variation in wing reduction was observed. Base of elytra in some species closely fitting pronotal base, as a result, humeral portion may project anteriorly. Elytra of species exhibiting wing reduction with curved lateral sides and faintly marked humeri. Abdominal ventrite 4 (in some species also ventrite 3, and occasionally also 5) with paired oval or transverse patches of long, dense setae (
|
|||
|
|
<figureCitation id="B0AAECC1FF9202394CABFEF8FD85FECF" box="[475,542,331,357]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="10" pageNumber="411">Fig. 8</figureCitation>
|
|||
|
|
). Apex of ventrite 5 usually deeply emarginated at apex (e.g.
|
|||
|
|
<figureCitation id="B0AAECC1FF92023949B8FEF8FAA1FECF" box="[1224,1338,331,357]" captionStart="FIGURE 8" captionStartId="10.[151,250,1905,1929]" captionTargetBox="[171,1416,489,1881]" captionTargetId="figure-162@10.[171,1416,489,1881]" captionTargetPageId="10" captionText="FIGURE 8. Atopida, abdominal segments of females. A) A. castanea; B) A. hirta; C) A. impressa; D) same, tergite VII; E) A. insularis sp. nov.; F) A. montana, male; G) A. paparoa sp. nov.; H) A. proba; I) same, tergite VII; J) A. suffusa; K) A. suturalis; L) same, tergite VII; M) A. tuhua; N) A. villosa; O) A. westlandensis sp. nov. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="10" pageNumber="411">Figs 8C, J</figureCitation>
|
|||
|
|
). Tergite VII modified in some species, its posterior portion forming rectangular flap or concavity. Ovipositor (
|
|||
|
|
<figureCitation id="B0AAECC1FF9202394860FEDCFACFFE23" box="[1296,1364,367,393]" captionStart="FIGURE 9" captionStartId="11.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@11.[151,1436,181,1309]" captionTargetPageId="11" captionText="FIGURE 9. Atopida, female genital tract. A) A. insularis sp. nov., B) A. villosa. Abbreviations: bscl—bursal sclerite, gla— gland, sVIII—sternite VIII, tVIII—tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.6986505" httpUri="https://zenodo.org/record/6986505/files/figure.png" pageId="10" pageNumber="411">Fig. 9</figureCitation>
|
|||
|
|
) long, branchlets absent; coxites narrow, membranous, styli apical; bursal sclerites present, either small, rod-like (
|
|||
|
|
<figureCitation id="B0AAECC1FF920239484FFE20FA0FFE07" box="[1343,1428,403,429]" captionStart="FIGURE 9" captionStartId="11.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@11.[151,1436,181,1309]" captionTargetPageId="11" captionText="FIGURE 9. Atopida, female genital tract. A) A. insularis sp. nov., B) A. villosa. Abbreviations: bscl—bursal sclerite, gla— gland, sVIII—sternite VIII, tVIII—tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.6986505" httpUri="https://zenodo.org/record/6986505/files/figure.png" pageId="10" pageNumber="411">Fig. 9B</figureCitation>
|
|||
|
|
) or larger, plate-like (
|
|||
|
|
<figureCitation id="B0AAECC1FF9202394C0CFE04FE4DFE7B" box="[380,470,439,465]" captionStart="FIGURE 9" captionStartId="11.[151,250,1333,1357]" captionTargetBox="[151,1436,181,1309]" captionTargetId="figure-19@11.[151,1436,181,1309]" captionTargetPageId="11" captionText="FIGURE 9. Atopida, female genital tract. A) A. insularis sp. nov., B) A. villosa. Abbreviations: bscl—bursal sclerite, gla— gland, sVIII—sternite VIII, tVIII—tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.6986505" httpUri="https://zenodo.org/record/6986505/files/figure.png" pageId="10" pageNumber="411">Fig. 9A</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="7CEEA0CCFF9202394DE7F8C2FB10F87B" ID-DOI="http://doi.org/10.5281/zenodo.6986501" ID-Zenodo-Dep="6986501" httpUri="https://zenodo.org/record/6986501/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,250,1905,1929]" targetBox="[171,1416,489,1881]" targetPageId="10">
|
|||
|
|
<paragraph id="282EF044FF9202394DE7F8C2FB10F87B" blockId="10.[151,1437,1905,2001]" pageId="10" pageNumber="411">
|
|||
|
|
<emphasis id="1AE52C56FF9202394DE7F8C2FE88F823" bold="true" box="[151,275,1905,1929]" pageId="10" pageNumber="411">FIGURE 8.</emphasis>
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394C6AF8C2FEF1F823" authorityName=": Kialka & Ruta" authorityYear="2017" box="[282,362,1905,1929]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9202394C6AF8C2FEF1F823" box="[282,362,1905,1929]" italics="true" pageId="10" pageNumber="411">Atopida</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, abdominal segments of females. A)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394F97F8C1FCC5F823" authorityName="White" authorityYear="1846" box="[743,862,1906,1929]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="castanea">
|
|||
|
|
<emphasis id="1AE52C56FF9202394F97F8C1FCC5F823" box="[743,862,1906,1929]" italics="true" pageId="10" pageNumber="411">A. castanea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; B)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394EFCF8C1FC41F823" authorityName="Broun" authorityYear="1880" box="[908,986,1905,1929]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="hirta">
|
|||
|
|
<emphasis id="1AE52C56FF9202394EFCF8C1FC41F823" box="[908,986,1905,1929]" italics="true" pageId="10" pageNumber="411">A. hirta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; C)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394978F8C1FB1BF823" authorityName="Broun" authorityYear="1914" box="[1032,1152,1906,1929]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="impressa">
|
|||
|
|
<emphasis id="1AE52C56FF9202394978F8C1FB1BF823" box="[1032,1152,1906,1929]" italics="true" pageId="10" pageNumber="411">A. impressa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; D) same, tergite VII; E)
|
|||
|
|
<taxonomicName id="EF918BC7FF92023948F6F8C1FF6BF807" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF92023948F6F8C1FF6BF807" italics="true" pageId="10" pageNumber="411">A. insularis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9202394D86F826FEDDF807" bold="true" box="[246,326,1941,1965]" pageId="10" pageNumber="411">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9202394D86F826FEDDF807" box="[246,326,1941,1965]" pageId="10" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
; F)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394C00F825FE7EF807" authorityName=", Broun" authorityYear="1921" box="[368,485,1942,1965]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="montana">
|
|||
|
|
<emphasis id="1AE52C56FF9202394C00F825FE7EF807" box="[368,485,1942,1965]" italics="true" pageId="10" pageNumber="411">A. montana</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male; G)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394F20F825FD5AF807" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[592,705,1942,1965]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="paparoa" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9202394F20F825FD5AF807" box="[592,705,1942,1965]" italics="true" pageId="10" pageNumber="411">A. paparoa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9202394FB7F826FC8CF807" bold="true" box="[711,791,1941,1965]" pageId="10" pageNumber="411">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9202394FB7F826FC8CF807" box="[711,791,1941,1965]" pageId="10" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
; H)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394E35F825FC06F807" authorityName="Sharp" authorityYear="1878" box="[837,925,1941,1965]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="proba">
|
|||
|
|
<emphasis id="1AE52C56FF9202394E35F825FC06F807" box="[837,925,1941,1965]" italics="true" pageId="10" pageNumber="411">A. proba</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; I) same, tergite VII; J)
|
|||
|
|
<taxonomicName id="EF918BC7FF92023949FFF825FB68F807" authorityName="Broun" authorityYear="1910" box="[1167,1267,1941,1965]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="suffusa">
|
|||
|
|
<emphasis id="1AE52C56FF92023949FFF825FB68F807" box="[1167,1267,1941,1965]" italics="true" pageId="10" pageNumber="411">A. suffusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; K)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394851F825FA0EF807" authorityName="" baseAuthorityName="White" baseAuthorityYear="1846" box="[1313,1429,1941,1965]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="1AE52C56FF9202394851F825FA0EF807" box="[1313,1429,1941,1965]" italics="true" pageId="10" pageNumber="411">A. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; L) same, tergite VII; M)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394CE2F809FE73F87B" authorityName="Kiałka & Ruta" authorityYear="2022" box="[402,488,1977,2001]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="tuhua">
|
|||
|
|
<emphasis id="1AE52C56FF9202394CE2F809FE73F87B" box="[402,488,1977,2001]" italics="true" pageId="10" pageNumber="411">A. tuhua</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; N)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394F66F809FDECF87B" authorityName="Broun" authorityYear="1921" box="[534,631,1977,2001]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="villosa">
|
|||
|
|
<emphasis id="1AE52C56FF9202394F66F809FDECF87B" box="[534,631,1977,2001]" italics="true" pageId="10" pageNumber="411">A. villosa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; O)
|
|||
|
|
<taxonomicName id="EF918BC7FF9202394FD5F809FCD5F87B" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[677,846,1977,2001]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="westlandensis" status="sp. nov.">
|
|||
|
|
<emphasis id="1AE52C56FF9202394FD5F809FCD5F87B" box="[677,846,1977,2001]" italics="true" pageId="10" pageNumber="411">A. westlandensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="1AE52C56FF9202394E24F80AFC3FF87B" bold="true" box="[852,932,1977,2001]" pageId="10" pageNumber="411">
|
|||
|
|
<taxonomicNameLabel id="01D6912DFF9202394E24F80AFC3FF87B" box="[852,932,1977,2001]" pageId="10" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Not to the same scale.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="7CEEA0CCFF9302384DE7FA86FDE2FADB" ID-DOI="http://doi.org/10.5281/zenodo.6986505" ID-Zenodo-Dep="6986505" httpUri="https://zenodo.org/record/6986505/files/figure.png" pageId="11" pageNumber="412" startId="11.[151,250,1333,1357]" targetBox="[151,1436,181,1309]" targetPageId="11">
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<paragraph id="282EF044FF9302384DE7FA86FDE2FADB" blockId="11.[151,1436,1333,1393]" pageId="11" pageNumber="412">
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<emphasis id="1AE52C56FF9302384DE7FA86FE8EFAE7" bold="true" box="[151,277,1333,1357]" pageId="11" pageNumber="412">FIGURE 9.</emphasis>
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<taxonomicName id="EF918BC7FF9302384C6DFA86FEF6FAE7" authorityName=": Kialka & Ruta" authorityYear="2017" box="[285,365,1333,1357]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
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<emphasis id="1AE52C56FF9302384C6DFA86FEF6FAE7" box="[285,365,1333,1357]" italics="true" pageId="11" pageNumber="412">Atopida</emphasis>
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</taxonomicName>
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, female genital tract. A)
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<taxonomicName id="EF918BC7FF9302384F04FA86FD70FAE7" authority="Kiałka & Ruta, 2022" authorityName="Kiałka & Ruta" authorityYear="2022" box="[628,747,1333,1357]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">
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<emphasis id="1AE52C56FF9302384F04FA86FD70FAE7" box="[628,747,1333,1357]" italics="true" pageId="11" pageNumber="412">A. insularis</emphasis>
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</taxonomicName>
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<emphasis id="1AE52C56FF9302384F83FA86FCDEFAE7" bold="true" box="[755,837,1333,1357]" pageId="11" pageNumber="412">
|
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<taxonomicNameLabel id="01D6912DFF9302384F83FA86FCDEFAE7" box="[755,837,1333,1357]" pageId="11" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
|
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</emphasis>
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, B)
|
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<taxonomicName id="EF918BC7FF9302384E06FA86FC43FAE7" authorityName="Broun" authorityYear="1921" box="[886,984,1333,1357]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="villosa">
|
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|
<emphasis id="1AE52C56FF9302384E06FA86FC43FAE7" box="[886,984,1333,1357]" italics="true" pageId="11" pageNumber="412">A. villosa</emphasis>
|
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</taxonomicName>
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. Abbreviations: bscl—bursal sclerite, gla— gland, sVIII—sternite VIII, tVIII—tergite VIII.
|
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</paragraph>
|
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</caption>
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<subSubSection id="608BA3CFFF9302384DB7FA2CFF5CF98C" pageId="11" pageNumber="412" type="discussion">
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<paragraph id="282EF044FF9302384DB7FA2CFF5CF98C" blockId="11.[151,1436,1439,1574]" pageId="11" pageNumber="412">
|
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<emphasis id="1AE52C56FF9302384DB7FA2CFEA0FA13" bold="true" box="[199,315,1439,1465]" pageId="11" pageNumber="412">Remarks.</emphasis>
|
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<bibRefCitation id="4C008DB5FF9302384C31FA2CFE48FA13" author="Sharp, D." box="[321,467,1439,1465]" pageId="11" pageNumber="412" pagination="40 - 59" refId="ref40097" refString="Sharp, D. (1878) On the Dascillidae of New Zealand. The Annals and Magazine of Natural History, 2, 40 - 59. https: // doi. org / 10.1080 / 00222937808682380" type="journal article" year="1878">Sharp (1878)</bibRefCitation>
|
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|
erroneously described labial palpi of
|
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<taxonomicName id="EF918BC7FF9302384E03FA2CFC57FA13" authorityName=": Kialka & Ruta" authorityYear="2017" box="[883,972,1439,1465]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="1AE52C56FF9302384E03FA2CFC57FA13" box="[883,972,1439,1465]" italics="true" pageId="11" pageNumber="412">Atopida</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
as “not forked”, i. e. the apical palpomere arising from the top of the penultimate one. This mistake was corrected by
|
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|
|
<bibRefCitation id="4C008DB5FF9302384E8BFA77FB0FFA77" author="Broun, T." box="[1019,1172,1475,1502]" pageId="11" pageNumber="412" pagination="975 - 1504" refId="ref38944" refString="Broun, T. (1893) Manual of the New Zealand Coleoptera. Parts V - VII. Government Printer, Wellington, XVII + 530 pp. [pp. I - XVII + pp. 975 - 1504]" type="book chapter" year="1893">Broun (1893)</bibRefCitation>
|
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|
|
who examined several species of
|
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|
<taxonomicName id="EF918BC7FF9302384C7EFA54FEFCF9AB" authorityName=": Kialka & Ruta" authorityYear="2017" box="[270,359,1511,1537]" class="Insecta" family="Scirtidae" genus="Atopida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis id="1AE52C56FF9302384C7EFA54FEFCF9AB" box="[270,359,1511,1537]" italics="true" pageId="11" pageNumber="412">Atopida</emphasis>
|
|||
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|
</taxonomicName>
|
|||
|
|
, all with furcate labial palpi, i. e. with apical palpomere arising from the side of the penultimate one.
|
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</paragraph>
|
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</subSubSection>
|
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</treatment>
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</document>
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