170 lines
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170 lines
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<document id="A6B347EBE9006B0E5D9C4B6E3800EFEF" ID-DOI="10.5281/zenodo.211734" ID-GBIF-Dataset="1837d7e6-3418-4081-9525-50f0b78d2125" ID-ISSN="1175-5326" ID-Zenodo-Dep="211734" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460502342988" checkinUser="plazi" docAuthor="Bauer, Aaron M., Jackman, Todd R., Sadlier, Ross A. & Whitaker, Anthony H." docDate="2012" docId="5C4F87E1FFB1FFB6088FF944D5D3B4B0" docLanguage="en" docName="zt03404p052.pdf" docOrigin="Zootaxa 3404" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhacodactylus Fitzinger 1843" docType="treatment" docVersion="10" lastPageNumber="17" masterDocId="A076FF99FFBEFFA60818FF81D705B67C" masterDocTitle="Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="16" updateTime="1698312116073" updateUser="plazi">
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<mods:titleInfo id="171102F5805B8F82B7DF663DF0422E88">
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<mods:title id="482B3963CCC3E6368216830E1CA509F7">Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus)</mods:title>
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<mods:namePart id="EE2740CA9E68216FE0071A6E33386B40">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="AA45F74CBBBB00041AFD7354D26169C0">Jackman, Todd R.</mods:namePart>
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<mods:namePart id="6842A02B4CA17AD6E282BC56DED1723A">Sadlier, Ross A.</mods:namePart>
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<mods:namePart id="A4EDF1532E2FBBE0ADBFB858D6D55966">Whitaker, Anthony H.</mods:namePart>
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<mods:typeOfResource id="7F153E241686013D61CBE518F0888610">text</mods:typeOfResource>
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<mods:title id="E4C2EEEF9BD8E116C1B6813FA42A2AF7">Zootaxa</mods:title>
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<mods:date id="E127F0601702B21E9B02C4ADD9941196">2012</mods:date>
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<mods:detail id="74B07A26EBAB385A6332E365AE8AFDA1" type="volume">
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<mods:number id="7F80F7F5B592D849DC7C8BA21AC0A198">3404</mods:number>
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<mods:identifier id="A9BF8E90C8DBA52D186CEB8608E06863" type="DOI">10.5281/zenodo.211734</mods:identifier>
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<mods:identifier id="A751B276C4CE41438854207812655B68" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5DBA109713BAD9830287D9E89A3A42AA" type="Zenodo-Dep">211734</mods:identifier>
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<treatment id="5C4F87E1FFB1FFB6088FF944D5D3B4B0" ID-DOI="http://doi.org/10.5281/zenodo.6166570" ID-GBIF-Taxon="119521676" ID-Zenodo-Dep="6166570" LSID="urn:lsid:plazi:treatment:5C4F87E1FFB1FFB6088FF944D5D3B4B0" httpUri="http://treatment.plazi.org/id/5C4F87E1FFB1FFB6088FF944D5D3B4B0" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection id="9CFC657CFFB1FFA9088FF944D5E1B13B" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="D45936F7FFB1FFA9088FF944D50DB0A3" blockId="15.[151,520,1733,1759]" box="[151,520,1733,1759]" pageId="15" pageNumber="16">
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<heading id="8F11819BFFB1FFA9088FF944D50DB0A3" bold="true" box="[151,520,1733,1759]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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<taxonomicName id="13E64D74FFB1FFA9088FF944D50DB0A3" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[151,520,1733,1759]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="E692EAE5FFB1FFA9088FF944D50DB0A3" bold="true" box="[151,520,1733,1759]" pageId="15" pageNumber="16">
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<emphasis id="E692EAE5FFB1FFA9088FF944D64CB0A3" bold="true" box="[151,329,1733,1759]" italics="true" pageId="15" pageNumber="16">Rhacodactylus</emphasis>
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<bibRefCitation id="B0774B06FFB1FFA90948F944D50DB0A3" author="Fitzinger" box="[336,520,1733,1759]" pageId="15" pageNumber="16" refString="Fitzinger, L. (1843) Systema Reptilium (Amblyglossae). Braumuller et Seidel, Vindobonae [Vienna]. 106 pp. + vi, 3 pp. index (unnumbered)." type="book" year="1843">Fitzinger, 1843</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D45936F7FFB1FFA9088FF88AD5E1B13B" blockId="15.[151,1436,1803,2008]" pageId="15" pageNumber="16">
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<emphasis id="E692EAE5FFB1FFA9088FF88AD7FAB158" bold="true" box="[151,255,1803,1828]" pageId="15" pageNumber="16">Content.</emphasis>
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<taxonomicName id="13E64D74FFB1FFA90911F88DD5E0B158" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[265,741,1804,1828]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leachianus">
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<emphasis id="E692EAE5FFB1FFA90911F88DD537B158" box="[265,562,1804,1828]" italics="true" pageId="15" pageNumber="16">Rhacodactylus leachianus</emphasis>
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(
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<bibRefCitation id="B0774B06FFB1FFA90A5DF88DD5D9B158" author="Cuvier" box="[581,732,1804,1828]" pageId="15" pageNumber="16" refString="Cuvier, G. (1829) Le Regne Animal Distribue d'apres son Organisation, pour Servir de Base a l'Histoire Naturelle des Animaux et d'Introduction a l'Anatomie Comparee, Nouvelle Edition, Revue et Augmentee. Tome 2. Deterville et Crochard, Paris, xv + 406 pp., pls. 5 - 13." type="book chapter" year="1829">Cuvier, 1829</bibRefCitation>
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)
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</taxonomicName>
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(
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<figureCitation id="4CDD2A72FFB1FFA90AE0F88DD43BB158" box="[760,830,1804,1828]" captionStart="FIGURE 3" captionStartId="13.[151,250,1852,1875]" captionTargetBox="[151,1435,259,1799]" captionTargetId="figure@13.[151,1436,259,1801]" captionTargetPageId="13" captionText="FIGURE 3. Representatives of each of the eight genera of endemic New Caledonian diplodactylid geckos. A. Oedodera marmorata. B. Dierogekko nehoueensis. C. Bavayia pulchella. D. Paniegekko madjo. E. Eurydactylodes occidentalis. F. Rhacodactylus leachianus. G. Correlophus belepensis sp. nov. H. Mniarogekko jalu sp. nov. Photos A, E, G – H by A. H. Whitaker, photo B by A. M. Bauer, photos C – D by R. A. Sadlier. Photo F courtesy of Mark O’Shea." httpUri="https://zenodo.org/record/211737/files/figure.png" pageId="15" pageNumber="16">Fig. 3</figureCitation>
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F),
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<taxonomicName id="13E64D74FFB1FFA90B71F88CD3B9B158" authority="Bavay, 1869" authorityName="Bavay" authorityYear="1869" box="[873,1212,1804,1828]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="auriculatus">
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<emphasis id="E692EAE5FFB1FFA90B71F88CD309B158" box="[873,1036,1804,1828]" italics="true" pageId="15" pageNumber="16">R. auriculatus</emphasis>
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(
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<bibRefCitation id="B0774B06FFB1FFA90C07F88DD3B1B158" author="Bavay" box="[1055,1204,1804,1828]" pageId="15" pageNumber="16" refString="Bavay, A. (1869) Catalogue des reptiles de la Nouvelle-Caledonie et description d'especes nouvelles. Memoires de la Societe Linneenne de Normandie, 15, 1 - 37. Bocage, J. V. Barboza du (1873 a) Note sur quelques geckotiens nouveaux ou peu connus de la Nouvelle Caledonie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 201 - 207." type="journal article" year="1869">Bavay, 1869</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicName id="13E64D74FFB1FFA90CD3F88CD631B13B" authority="Bocage, 1873" authorityName="Bocage" authorityYear="1873" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="trachyrhynchus">
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<emphasis id="E692EAE5FFB1FFA90CD3F88CD299B158" box="[1227,1436,1804,1828]" italics="true" pageId="15" pageNumber="16">R. trachyrhynchus</emphasis>
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<bibRefCitation id="B0774B06FFB1FFA9088FF8B1D631B13B" author="Bocage" box="[151,308,1839,1864]" pageId="15" pageNumber="16" refString="Bocage, J. V. Barboza du. (1873) Sur quelques sauriens nouveaux de la Nouvelle Caledonie et de l'Australie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 228 - 232." type="journal article" year="1873">Bocage, 1873</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="13E64D74FFB1FFA90927F8B0D5E5B13B" authority="Boulenger, 1878" authorityName="Boulenger" authorityYear="1878" box="[319,736,1839,1864]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="trachycephalus">
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<emphasis id="E692EAE5FFB1FFA90927F8B0D50FB134" box="[319,522,1840,1864]" italics="true" pageId="15" pageNumber="16">R. trachycephalus</emphasis>
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(
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<bibRefCitation id="B0774B06FFB1FFA90A00F8AED5D2B13B" author="Boulenger" box="[536,727,1839,1863]" pageId="15" pageNumber="16" refString="Boulenger, G. A. (1878) Description d'un genre nouveau et d'une espece nouvelle de la famille des geckotides. Bulletin de la Societe Zoologique de France, 3, 68 - 70, pl. 2." type="journal article" year="1878">Boulenger, 1878</bibRefCitation>
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)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CFC657CFFB1FFA908DFF8D5D31CB110" box="[199,1049,1875,1900]" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="D45936F7FFB1FFA908DFF8D5D31CB110" blockId="15.[151,1436,1803,2008]" box="[199,1049,1875,1900]" pageId="15" pageNumber="16">
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<emphasis id="E692EAE5FFB1FFA908DFF8D5D667B110" bold="true" box="[199,354,1875,1900]" pageId="15" pageNumber="16">
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<typeStatus id="0B5D8855FFB1FFA908DFF8D5D607B110" box="[199,258,1876,1900]" pageId="15" pageNumber="16">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="13E64D74FFB1FFA90971F8D5D40CB110" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[361,777,1876,1900]" class="Squamata" family="Gekkonidae" genus="Ascalabotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leachianus">
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<emphasis id="E692EAE5FFB1FFA90971F8D5D575B110" box="[361,624,1876,1900]" italics="true" pageId="15" pageNumber="16">Ascalabotes leachianus</emphasis>
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<bibRefCitation id="B0774B06FFB1FFA90A6FF8D5D40CB110" author="Cuvier" box="[631,777,1876,1900]" pageId="15" pageNumber="16" refString="Cuvier, G. (1829) Le Regne Animal Distribue d'apres son Organisation, pour Servir de Base a l'Histoire Naturelle des Animaux et d'Introduction a l'Anatomie Comparee, Nouvelle Edition, Revue et Augmentee. Tome 2. Deterville et Crochard, Paris, xv + 406 pp., pls. 5 - 13." type="book chapter" year="1829">Cuvier, 1829</bibRefCitation>
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</taxonomicName>
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by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CFC657CFFB1FFB608DFF8F6D241B743" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="diagnosis">
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<paragraph id="D45936F7FFB1FFB608DFF8F6D241B743" blockId="15.[151,1436,1803,2008]" lastBlockId="16.[151,1436,151,716]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<emphasis id="E692EAE5FFB1FFA908DFF8F6D644B1EC" bold="true" box="[199,321,1911,1936]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
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<taxonomicName id="13E64D74FFB1FFA90953F8F9D6F5B1EC" box="[331,496,1912,1936]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="E692EAE5FFB1FFA90953F8F9D6F5B1EC" box="[331,496,1912,1936]" italics="true" pageId="15" pageNumber="16">Rhacodactylus</emphasis>
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</taxonomicName>
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may be distinguished from all other New Caledonian diplodactylid geckos by the following combination of character states: body large to very large (maximum
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<quantity id="131E9B12FFB1FFA90C35F81DD3C0B1C8" box="[1069,1221,1948,1972]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.56" metricValueMin="1.25" pageId="15" pageNumber="16" unit="mm" value="190.5" valueMax="256.0" valueMin="125.0">125–256 mm</quantity>
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SVL); head large, skull usually ornamented with bumps, ridges or rugosities; tail variable across species, 30–100% of SVL; dorsal scalation granular, homogeneous; extensive skin folds present or absent; expanded undivided subdigital lamellae under all toes; webbing between digits weakly to strongly developed; claw of digit I of manus and pes positioned lateral to a single, undivided apical lamella; precloacal pores in three to six rows (occasionally up to eight rows, but posteriormost one or two with only scattered pores) in males (49–130 pores in total), longest anterior rows extending on to base of thighs or not; dorsal color pattern highly variable both within and between species.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CFC657CFFAEFFB608DFFECAD216B7B3" pageId="16" pageNumber="17" type="distribution">
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<paragraph id="D45936F7FFAEFFB608DFFECAD216B7B3" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="17">
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<emphasis id="E692EAE5FFAEFFB608DFFECAD665B718" bold="true" box="[199,352,331,356]" pageId="16" pageNumber="17">Distribution.</emphasis>
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<taxonomicName id="13E64D74FFAEFFB60970FECDD508B718" box="[360,525,332,356]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
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<emphasis id="E692EAE5FFAEFFB60970FECDD508B718" box="[360,525,332,356]" italics="true" pageId="16" pageNumber="17">Rhacodactylus</emphasis>
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</taxonomicName>
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spp. occur throughout most of the Grande Terre — as far north as the Dôme de Tiébaghi in the west and the Panié massif in the east — but they have not been recorded in the far north of Grande Terre and among its smaller satellite islands they have only been recorded on one (Île Némou). They are also present on the Île des Pins and its surrounding satellite islands but are absent from the Loyalty Islands.
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</paragraph>
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</subSubSection>
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<subSubSection id="9CFC657CFFAEFFB608DFFE5DD5D3B4B0" pageId="16" pageNumber="17" type="discussion">
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<paragraph id="D45936F7FFAEFFB608DFFE5DD5D3B4B0" blockId="16.[151,1436,151,716]" pageId="16" pageNumber="17">
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<emphasis id="E692EAE5FFAEFFB608DFFE5DD63EB788" bold="true" box="[199,315,476,500]" pageId="16" pageNumber="17">Remarks.</emphasis>
|
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|
The four species here retained in a redefined
|
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<taxonomicName id="13E64D74FFAEFFB60B6EFE5DD31EB788" box="[886,1051,476,500]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
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|
|
<emphasis id="E692EAE5FFAEFFB60B6EFE5DD31EB788" box="[886,1051,476,500]" italics="true" pageId="16" pageNumber="17">Rhacodactylus</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
represent three morphologically distinct units. Although we retrieve a monophyletic
|
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|
|
<taxonomicName id="13E64D74FFAEFFB60AC4FD81D484B464" box="[732,897,512,536]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60AC4FD81D484B464" box="[732,897,512,536]" italics="true" pageId="16" pageNumber="17">Rhacodactylus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
under maximum likelihood in the ND2 tree, the low level of support for this arrangement does not exclude the possibility that each of these units represents an independent lineage with closer affinities to other New Caledonian genera than to one another. Were this the case, the name
|
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|
|
<taxonomicName id="13E64D74FFAEFFB6091CFDEDD6ACB4F8" box="[260,425,620,644]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB6091CFDEDD6ACB4F8" box="[260,425,620,644]" italics="true" pageId="16" pageNumber="17">Rhacodactylus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is linked to
|
|||
|
|
<taxonomicName id="13E64D74FFAEFFB60A2CFDECD5C9B4F8" box="[564,716,620,644]" class="Squamata" family="Gekkonidae" genus="Ascalabotes" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="leachianus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60A2CFDECD5C9B4F8" box="[564,716,620,644]" italics="true" pageId="16" pageNumber="17">R. leachianus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the names
|
|||
|
|
<taxonomicName id="13E64D74FFAEFFB60B62FDEDD3B1B4F8" authority="Bocage, 1873" authorityName="Bocage" authorityYear="1873" box="[890,1204,620,645]" class="Squamata" family="Diplodactylidae" genus="Ceratolophus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60B62FDEDD317B4F8" box="[890,1042,620,644]" italics="true" pageId="16" pageNumber="17">Ceratolophus</emphasis>
|
|||
|
|
<bibRefCitation id="B0774B06FFAEFFB60C02FDECD3B1B4F8" author="Bocage" box="[1050,1204,620,645]" pageId="16" pageNumber="17" refString="Bocage, J. V. Barboza du. (1873) Sur quelques sauriens nouveaux de la Nouvelle Caledonie et de l'Australie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 228 - 232." type="journal article" year="1873">Bocage, 1873</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="13E64D74FFAEFFB60CF3FDEDD652B4DB" authority="Boulenger, 1878" authorityName="Boulenger" authorityYear="1878" class="Squamata" family="Diplodactylidae" genus="Chameleonurus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60CF3FDEDD299B4F8" box="[1259,1436,620,644]" italics="true" pageId="16" pageNumber="17">Chameleonurus</emphasis>
|
|||
|
|
<bibRefCitation id="B0774B06FFAEFFB6088FFD0ED652B4DB" author="Boulenger" box="[151,343,655,679]" pageId="16" pageNumber="17" refString="Boulenger, G. A. (1878) Description d'un genre nouveau et d'une espece nouvelle de la famille des geckotides. Bulletin de la Societe Zoologique de France, 3, 68 - 70, pl. 2." type="journal article" year="1878">Boulenger, 1878</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are available for
|
|||
|
|
<taxonomicName id="13E64D74FFAEFFB60A2FFD10D5D8B4D4" box="[567,733,656,680]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="auriculatus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60A2FFD10D5D8B4D4" box="[567,733,656,680]" italics="true" pageId="16" pageNumber="17">R. auriculatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the live-bearing forms, respectively. See below for a discussion of the revalidation of
|
|||
|
|
<taxonomicName id="13E64D74FFAEFFB60A1DFD34D5CAB4B0" box="[517,719,692,716]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="trachycephalus">
|
|||
|
|
<emphasis id="E692EAE5FFAEFFB60A1DFD34D5CAB4B0" box="[517,719,692,716]" italics="true" pageId="16" pageNumber="17">R. trachycephalus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|