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<document id="8D2997205FC04FFDB38A23C0F95ABDE8" ID-DOI="10.1080/00222930802002535" ID-ISSN="1464-5262" ID-Zenodo-Dep="5219397" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629340601867" checkinUser="felipe" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Scholtz, Clarke H." docDate="2008" docId="1F77BC72FFCB305523C9FEC7FC58FD1A" docLanguage="en" docName="JNatHist.42.21-24.1477-1508.pdf" docOrigin="Journal of Natural History 42 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222930802002535" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Coptorhina nitefacta Gillet 1932" docType="treatment" docVersion="3" lastPageNumber="1507" masterDocId="E34EC40AFFD430752211FF81FFF5FF8C" masterDocTitle="Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="1508" masterPageNumber="1477" pageNumber="1506" updateTime="1699068592934" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="631A4E1AA3E9DBCA6A08A1D7F7EEB7DB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EB63F801CA7725D82825F9778DCBA1AD">Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="66D428B7B6D24DCAF007CC65155BB442">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="06911CD2066BB1D6AA0A3B1480A526E1">Akhmetova, Lilia A.</mods:namePart>
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<mods:namePart id="41D3D9E64899CD908EA8E05D06A594FE">Scholtz, Clarke H.</mods:namePart>
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<mods:typeOfResource id="9FAA0D57DBF17F58039BA62551709168">text</mods:typeOfResource>
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<mods:titleInfo id="10A4483D2AAD92370B0478B73F879FA2">
<mods:title id="D34E2E74F80F563953765609CE271128">Journal of Natural History</mods:title>
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<mods:part id="5389171B9A0767BF7794F326FDF06ABE">
<mods:date id="AFC5A5F17E9BDDF6ACCE9D9D0F6E250B">2008</mods:date>
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<mods:number id="56FB5CEF16B072969B924DF42D3BC7F9">2008-06-30</mods:number>
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<mods:detail id="E29B5EF25AB4FD540AADFFCBCF0AE655" type="volume">
<mods:number id="E605FD4168902FC1EE5566252090FADC">42</mods:number>
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<mods:number id="94DE6FB9BE6D43BDE508E42E2DB7040E">21 - 24</mods:number>
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<mods:url id="21E6DC769B67272E76A0FD4C186109BB">http://dx.doi.org/10.1080/00222930802002535</mods:url>
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<mods:classification id="03084FC0CB94DB752681E50A8518AFE5">journal article</mods:classification>
<mods:identifier id="C4374F9BD5C43787D08946E17B0848E5" type="DOI">10.1080/00222930802002535</mods:identifier>
<mods:identifier id="B758EA98A155D84BB66AEAA5BAD88B16" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="8F7AA43956EEB3BEE88D4141EC7A4A4A" type="Zenodo-Dep">5219397</mods:identifier>
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<treatment id="1F77BC72FFCB305523C9FEC7FC58FD1A" LSID="urn:lsid:plazi:treatment:1F77BC72FFCB305523C9FEC7FC58FD1A" httpUri="http://treatment.plazi.org/id/1F77BC72FFCB305523C9FEC7FC58FD1A" lastPageId="32" lastPageNumber="1507" pageId="31" pageNumber="1506">
<subSubSection id="DFC45EEFFFCB306A23C9FEC7FCFDFED2" box="[472,776,326,350]" pageId="31" pageNumber="1506" type="nomenclature">
<paragraph id="97610D64FFCB306A23C9FEC7FCFDFED2" blockId="31.[146,930,326,415]" box="[472,776,326,350]" pageId="31" pageNumber="1506">
<heading id="CC29BA08FFCB306A23C9FEC7FCFDFED2" box="[472,776,326,350]" centered="true" fontSize="36" level="2" pageId="31" pageNumber="1506" reason="7">
<taxonomicName id="50DE76E7FFCB306A23C9FEC7FCFDFED2" ID-CoL="Y885" authority="Gillet" authorityName="Gillet" authorityYear="1932" box="[472,776,326,350]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1506" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFCB306A23C9FEC7FD4AFED2" bold="true" box="[472,703,326,350]" italics="true" pageId="31" pageNumber="1506">Coptorhina nitefacta</emphasis>
Gillet
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFCB306A23EFFEE7FD16FEF2" box="[510,739,358,382]" pageId="31" pageNumber="1506" type="description">
<paragraph id="97610D64FFCB306A23EFFEE7FD16FEF2" blockId="31.[146,930,326,415]" box="[510,739,358,382]" pageId="31" pageNumber="1506">
(
<figureCitation id="0FE511E1FFCB306A2017FEE7FD85FEF2" box="[518,624,358,382]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="31" pageNumber="1506">Figures 5</figureCitation>
,
<figureCitation id="0FE511E1FFCB306A206FFEE7FD44FEF2" box="[638,689,358,382]" captionStart="Figures 9–14" captionStartId="7.[146,222,1424,1446]" captionTargetBox="[146,1104,165,1387]" captionTargetId="figure-1@7.[146,1104,165,1387]" captionTargetPageId="7" captionText="Figures 9–14. Coptorhina spp. 9, wing; 10–11, aedeagus in dorsal and lateral view; 12, internal sac of aedeagus; 13, internal sac sclerite arrowed in Figure 12; 14, abdomen in lateral view (14A, female, 14B, male). 9, 10, 12, 13C–E, 14, C. auspicata (13C, lectotype, Enkeldoorn, Zimbabwe, 13D, Cinergy Game farm, RSA, 13E, Shesheke, Zambia); 11, 13I–J, C. nitidipennis (13I, Cinergy Game farm, RSA, 13J, paratype, ‘‘Caffraria’’); 13A–B, C. excavata sp. n. (13A, holotype, 13B, paratype, Mamathes, Lesotho); 13F, C. klugii, Blouberg, RSA; 13G, C. davidi, paratype, Mpika, Congo; 13H, C. nitefacta, holotype, Bura, Kenya. 9, 13 and 14 are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219401" httpUri="https://zenodo.org/record/5219401/files/figure.png" pageId="31" pageNumber="1506">13H</figureCitation>
,
<figureCitation id="0FE511E1FFCB306A20AEFEE7FD2EFEF2" box="[703,731,358,382]" captionStart="Figures 20–22" captionStartId="26.[213,289,1398,1420]" captionTargetBox="[213,1171,165,1362]" captionTargetId="figure-56@26.[213,1171,165,1362]" captionTargetPageId="26" captionText="Figures 20–22. Coptorhina spp., distribution map. 20, C. auspicata; 21, C. excavata sp. n.; 22, C. nitefacta." figureDoi="http://doi.org/10.5281/zenodo.5219409" httpUri="https://zenodo.org/record/5219409/files/figure.png" pageId="31" pageNumber="1506">22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFCB306A2283FE06FC57FE13" box="[146,930,391,415]" pageId="31" pageNumber="1506" type="reference_group">
<paragraph id="97610D64FFCB306A2283FE06FC57FE13" blockId="31.[146,930,326,415]" box="[146,930,391,415]" pageId="31" pageNumber="1506">
<treatmentCitationGroup id="B7CE2A4AFFCB306A2283FE06FC57FE13" box="[146,930,391,415]" pageId="31" pageNumber="1506">
<treatmentCitation id="167F2B75FFCB306A2283FE06FDC7FE13" author="Gillet JJE" box="[146,562,391,415]" page="327" pageId="31" pageNumber="1506" year="1932">
<taxonomicName id="50DE76E7FFCB306A2283FE06FDC7FE13" ID-CoL="Y885" authority="Gillet 1932: 327" authorityName="Gillet" authorityPageNumber="327" authorityYear="1932" box="[146,562,391,415]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1506" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFCB306A2283FE06FE8CFE13" box="[146,377,391,415]" italics="true" pageId="31" pageNumber="1506">Coptorhina nitefacta</emphasis>
<bibRefCitation id="F34F7095FFCB306A2393FE06FDC7FE13" author="Gillet JJE" box="[386,562,391,415]" pageId="31" pageNumber="1506" pagination="323 - 332" refId="ref19299" refString="Gillet JJE. 1932. Lamellicornes coprophages (Coleoptera). Bull Soc Entomol Belg. 72: 323 - 332." type="journal article" year="1932">Gillet 1932:327</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="167F2B75FFCB306A2051FE06FCF9FE13" author="Ferreira M" box="[576,780,391,415]" page="357" pageId="31" pageNumber="1506" year="1972">
<bibRefCitation id="F34F7095FFCB306A2051FE06FCF9FE13" author="Ferreira M" box="[576,780,391,415]" pageId="31" pageNumber="1506" pagination="5 - 1088" refId="ref19177" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Entomol Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972:357</bibRefCitation>
</treatmentCitation>
; 1939:34, 38.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DFC45EEFFFCB30552283FE59FC58FD1A" lastPageId="32" lastPageNumber="1507" pageId="31" pageNumber="1506" type="materials_examined">
<paragraph id="97610D64FFCB306A2283FE59FE57FE7C" blockId="31.[146,418,472,496]" box="[146,418,472,496]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FE59FE57FE7C" box="[146,418,472,496]" italics="true" pageId="31" pageNumber="1506">
<typeStatus id="4865B3C6FFCB306A2283FE59FF3FFE7C" box="[146,202,472,496]" pageId="31" pageNumber="1506">Type</typeStatus>
material examined
</emphasis>
</paragraph>
<paragraph id="97610D64FFCB306A2283FD80FDC2FDD5" blockId="31.[146,1102,513,601]" pageId="31" pageNumber="1506">
<taxonomicName id="50DE76E7FFCB306A2283FD80FE8BFD95" authorityName="Gillet" authorityYear="1932" box="[146,382,513,537]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1506" phylum="Arthropoda" rank="species" species="nitefacta">
<emphasis id="A5AAD176FFCB306A2283FD80FE8BFD95" box="[146,382,513,537]" italics="true" pageId="31" pageNumber="1506">Coptorhina nitefacta</emphasis>
</taxonomicName>
.
<materialsCitation id="27B60739FFCB306A2383FD80FDC6FDD5" collectingDate="1912-03" collectionCode="MNHN" elevation="1050" pageId="31" pageNumber="1506" specimenCount="1" typeStatus="holotype">
<typeStatus id="4865B3C6FFCB306A2383FD80FE08FD95" box="[402,509,513,537]" pageId="31" pageNumber="1506" type="holotype">Holotype</typeStatus>
<emphasis id="A5AAD176FFCB306A201AFD85FDE8FD94" box="[523,541,516,536]" italics="true" pageId="31" pageNumber="1506">„</emphasis>
with labels ‘‘AFR. OR. ANGL. (WA-TaïTA) BURA Alluaud and Jeannel
<date id="E3602BA4FFCB306A23F5FDA0FDAAFDB5" box="[484,607,545,569]" pageId="31" pageNumber="1506" value="1912-03">
<collectingDate id="F324D24CFFCB306A23F5FDA0FDAAFDB5" box="[484,607,545,569]" pageId="31" pageNumber="1506" value="1912-03">Mars 1912</collectingDate>
</date>
–
<quantity id="5026A081FFCB306A206EFDA0FD3BFDB5" box="[639,718,545,569]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="31" pageNumber="1506" unit="m" value="1050.0">
<elevation id="1CF3EA57FFCB306A206EFDA0FD3BFDB5" box="[639,718,545,569]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="31" pageNumber="1506" unit="m" value="1050.0">1050 m</elevation>
</quantity>
– St. 61’’, ‘‘
<taxonomicName id="50DE76E7FFCB306A214AFDA0FF27FDD5" authority="Gillet" authorityName="Gillet" authorityYear="1932" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1506" phylum="Arthropoda" rank="species" species="nitefacta">Coptorhina nitefacta Gillet</taxonomicName>
type.’’, ‘‘Holotype’’ (
<collectionCode id="F1CF95A1FFCB306A23DBFDC0FDDEFDD5" box="[458,555,577,601]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="31" pageNumber="1506">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="97610D64FFCB306A2283FD0BFE2BFD2E" blockId="31.[146,478,650,674]" box="[146,478,650,674]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FD0BFE2BFD2E" box="[146,478,650,674]" italics="true" pageId="31" pageNumber="1506">Additional material examined</emphasis>
</paragraph>
<paragraph id="97610D64FFCB306A2283FD33FF05FD66" blockId="31.[146,1103,690,746]" pageId="31" pageNumber="1506">
<materialsCitation id="27B60739FFCB306A2283FD33FF19FD66" collectingDate="1999-11-30" collectionCode="JJCM" collectorName="M. Snizek" country="Kenya" location="Garissa" pageId="31" pageNumber="1506" specimenCount="1" stateProvince="Garissa">
<emphasis id="A5AAD176FFCB306A2283FD33FF2CFD46" bold="true" box="[146,217,690,714]" pageId="31" pageNumber="1506">
<collectingCountry id="EFC94DF4FFCB306A2283FD33FF2CFD46" box="[146,217,690,714]" name="Kenya" pageId="31" pageNumber="1506">Kenya</collectingCountry>
</emphasis>
:
<collectingRegion id="551AC386FFCB306A22FBFD33FEB5FD46" box="[234,320,690,714]" country="Kenya" name="Garissa" pageId="31" pageNumber="1506">Garissa</collectingRegion>
env. [00
<emphasis id="A5AAD176FFCB306A23B5FD32FE5BFD47" bold="true" box="[420,430,691,715]" italics="true" pageId="31" pageNumber="1506">°</emphasis>
279 S 39
<emphasis id="A5AAD176FFCB306A2018FD32FDE6FD47" bold="true" box="[521,531,691,715]" italics="true" pageId="31" pageNumber="1506">°</emphasis>
399 E],
<date id="E3602BA4FFCB306A2074FD33FCB6FD46" box="[613,835,690,714]" pageId="31" pageNumber="1506" value="1999-11-30">
<collectingDate id="F324D24CFFCB306A2074FD33FCB6FD46" box="[613,835,690,714]" pageId="31" pageNumber="1506" value="1999-11-30">30 November 1999</collectingDate>
</date>
,
<collectorName id="3A2B68B2FFCB306A2140FD33FC32FD46" box="[849,967,690,714]" pageId="31" pageNumber="1506">M. Snižek</collectorName>
leg. 1 spm. (
<collectionCode id="F1CF95A1FFCB306A228BFD53FF16FD66" box="[154,227,722,746]" pageId="31" pageNumber="1506">JJCM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="97610D64FFCB306A2283FC9FFF0AFCBA" blockId="31.[146,255,798,822]" box="[146,255,798,822]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FC9FFF0AFCBA" box="[146,255,798,822]" italics="true" pageId="31" pageNumber="1506">Diagnosis</emphasis>
</paragraph>
<paragraph id="97610D64FFCB306A2283FCC6FDB7FCF3" blockId="31.[146,1102,839,895]" pageId="31" pageNumber="1506">
This species differs from other similar
<taxonomicName id="50DE76E7FFCB306A207AFCC6FCB7FCD3" box="[619,834,839,863]" class="Insecta" family="Scarabaeidae" genus="Coptorhina" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="1506" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A5AAD176FFCB306A207AFCC6FD1DFCD3" box="[619,744,839,863]" italics="true" pageId="31" pageNumber="1506">Coptorhina</emphasis>
species
</taxonomicName>
in having shiny, very sparsely punctate upper side of body.
</paragraph>
<paragraph id="97610D64FFCB306A2283FC39FEE4FC5C" blockId="31.[146,273,952,976]" box="[146,273,952,976]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FC39FEE4FC5C" box="[146,273,952,976]" italics="true" pageId="31" pageNumber="1506">Description</emphasis>
</paragraph>
<paragraph id="97610D64FFCB306A2283FC61FDC6FB94" blockId="31.[146,1103,992,1048]" pageId="31" pageNumber="1506">Strongly convex, black, shiny beetle. Body length 9.5–13.5 mm, width 6.5–9.5 mm. Dorsal surface without visible setae.</paragraph>
<paragraph id="97610D64FFCB306A2283FBD0FD66FAA5" blockId="31.[146,1103,1105,1321]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FBD0FF3BFBE5" box="[146,206,1105,1129]" italics="true" pageId="31" pageNumber="1506">Head</emphasis>
. Clypeus with protruding anterior angles separated by deep, right-angled sinuation (
<figureCitation id="0FE511E1FFCB306A2301FBF0FE9AFB05" box="[272,367,1137,1161]" captionStart="Figures 1–8" captionStartId="5.[146,222,1554,1576]" captionTargetBox="[146,1104,167,1518]" captionTargetId="figure-1@5.[146,1104,165,1518]" captionTargetPageId="5" captionText="Figures 1–8. Coptorhina spp. 1, lateral view of body; 2–6, head and pronotum in dorsal view; 7–8, pronotum in dorsal view. 1–2, C. auspicata; 3, C. nitidipennis; 4, C. excavata sp. n; 5, C. nitefacta; 6, C. klugii; 7, C. angolensis; 8, C. davidi. Figures are not to scale." figureDoi="http://doi.org/10.5281/zenodo.5219399" httpUri="https://zenodo.org/record/5219399/files/figure.png" pageId="31" pageNumber="1506">Figure 5</figureCitation>
). Protruding angles acute to narrowly rounded apically, with longitudinal carinae on the upper side. Genae obtusely rounded. Genal sutures feebly distinct. Lateral margins of clypeus sinuate near genal sutures in some individuals. Frontal suture feebly visible, broadly interrupted in the middle. Head very densely and evenly punctuate, punctures sometimes adjacent and sculpture appears rugose. Antennal clubs dark-brown.
</paragraph>
<paragraph id="97610D64FFCB306A2283FAE3FE69F9B5" blockId="31.[146,1103,1378,1593]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283FAE3FEF4FAF6" box="[146,257,1378,1402]" italics="true" pageId="31" pageNumber="1506">Pronotum</emphasis>
. Trapezoidal, two times wider than long. Anterior margin and base finely bordered. Sides more or less evenly rounded. Lateral border relatively narrow, evenly tapering basally and apically. Pronotum divided into larger anterior part and smaller posterior part by a transversal carina. The carina is almost as wide as pronotum, narrowly interrupted medially, not serrate. Anterior part of pronotum punctate, excavated laterally in larger specimens. Posterior part sparsely punctate with minute punctures.
</paragraph>
<paragraph id="97610D64FFCB306A2283F9F3FC3BF926" blockId="31.[146,1102,1650,1706]" pageId="31" pageNumber="1506">
<emphasis id="A5AAD176FFCB306A2283F9F3FF2CF906" box="[146,217,1650,1674]" italics="true" pageId="31" pageNumber="1506">Elytra</emphasis>
. Striae fine but distinct, punctate (punctures separated by 3–4 puncture diameters). Intervals flat, very sparsely punctate with minute punctures.
</paragraph>
<paragraph id="97610D64FFF4305522C4FF28FD58FE8D" blockId="32.[213,1169,168,257]" pageId="32" pageNumber="1507">
<emphasis id="A5AAD176FFF4305522C4FF28FEB1FF4D" box="[213,324,169,193]" italics="true" pageId="32" pageNumber="1507">Underside</emphasis>
. Pygidium with fine border, its disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of metasternum which is very sparsely to almost indistinctly punctuate.
</paragraph>
<paragraph id="97610D64FFF4305522C4FEAEFC4EFECB" blockId="32.[213,955,303,327]" box="[213,955,303,327]" pageId="32" pageNumber="1507">
<emphasis id="A5AAD176FFF4305522C4FEAEFECAFECB" box="[213,319,303,327]" italics="true" pageId="32" pageNumber="1507">Aedeagus</emphasis>
. Parameres with widely rounded apices in lateral view.
</paragraph>
<paragraph id="97610D64FFF4305522C4FE01FEB8FE14" blockId="32.[213,333,384,408]" box="[213,333,384,408]" pageId="32" pageNumber="1507">
<emphasis id="A5AAD176FFF4305522C4FE01FEB8FE14" box="[213,333,384,408]" italics="true" pageId="32" pageNumber="1507">Variability</emphasis>
</paragraph>
<paragraph id="97610D64FFF4305522C4FE28FCD7FDAE" blockId="32.[213,1169,425,546]" pageId="32" pageNumber="1507">
The description above is based mostly on the
<typeStatus id="4865B3C6FFF4305520F7FE28FCBBFE4D" box="[742,846,425,449]" pageId="32" pageNumber="1507" type="holotype">holotype</typeStatus>
. The other specimen known to us is smaller and has less developed transversal pronotal carina and slightly or not excavated anterolateral sides of pronotum. It is however impossible to appreciate the interspecific variability from the material available.
</paragraph>
<paragraph id="97610D64FFF4305522C4FDD7FEAEFDE2" blockId="32.[213,347,598,622]" box="[213,347,598,622]" pageId="32" pageNumber="1507">
<emphasis id="A5AAD176FFF4305522C4FDD7FEAEFDE2" box="[213,347,598,622]" italics="true" pageId="32" pageNumber="1507">Distribution</emphasis>
</paragraph>
<paragraph id="97610D64FFF4305522C4FDFFFC58FD1A" blockId="32.[213,941,638,662]" box="[213,941,638,662]" pageId="32" pageNumber="1507">
This species is known from two localities in
<collectingCountry id="EFC94DF4FFF4305520C7FDFFFCD4FD1A" box="[726,801,638,662]" name="Kenya" pageId="32" pageNumber="1507">Kenya</collectingCountry>
(
<figureCitation id="0FE511E1FFF430552120FDFFFC54FD1A" box="[817,929,638,662]" captionStart="Figures 20–22" captionStartId="26.[213,289,1398,1420]" captionTargetBox="[213,1171,165,1362]" captionTargetId="figure-56@26.[213,1171,165,1362]" captionTargetPageId="26" captionText="Figures 20–22. Coptorhina spp., distribution map. 20, C. auspicata; 21, C. excavata sp. n.; 22, C. nitefacta." figureDoi="http://doi.org/10.5281/zenodo.5219409" httpUri="https://zenodo.org/record/5219409/files/figure.png" pageId="32" pageNumber="1507">Figure 22</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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