189 lines
19 KiB
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189 lines
19 KiB
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<document id="DA9F3BB43B8FC3C74B2201905BA39E76" ID-DOI="10.5281/zenodo.282224" ID-GBIF-Dataset="3c9db890-c1df-4340-8be1-8e5cd631aeed" ID-ISSN="1175-5326" ID-Zenodo-Dep="282224" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460519716872" checkinUser="plazi" docAuthor="Plisko, Jadwiga Danuta" docDate="2012" docId="03F54361FF9FFFA8E68BD6B8B8EB6DD7" docLanguage="en" docName="zt03458p058.pdf" docOrigin="Zootaxa 3458" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Parachilota wahlbergi Michaelsen 1899" docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FFCC3B19FFB8FF8FE61CD62FBD796E71" masterDocTitle="An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)" masterLastPageNumber="58" masterPageNumber="4" pageNumber="43" updateTime="1698314471110" updateUser="plazi">
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<mods:title id="2E4C64AEA9416680F932677A2CBF11EB">An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)</mods:title>
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<mods:namePart id="53B7D43FD9817B4F11650A2F31C0A37D">Plisko, Jadwiga Danuta</mods:namePart>
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<mods:title id="1C01CAB4B17C9BDE206C2391C1AE1929">Zootaxa</mods:title>
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<mods:date id="29E13EDDFFC0109312DE0CC8524E8945">2012</mods:date>
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<mods:detail id="0B21F3AFE88290E65866648F57B67CA5" type="volume">
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<mods:number id="85CE1B33CA31B45302D969BB3336E3FE">3458</mods:number>
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<mods:identifier id="52FED26344972DD7B5CAD035BA78A7B1" type="DOI">10.5281/zenodo.282224</mods:identifier>
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<mods:identifier id="0E486D2EFE07C7A9B69DEACF1FA5D57F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="22D504DDC6AEB1255AF95871CF50CC78" type="Zenodo-Dep">282224</mods:identifier>
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<treatment id="03F54361FF9FFFA8E68BD6B8B8EB6DD7" ID-DOI="http://doi.org/10.5281/zenodo.6171414" ID-GBIF-Taxon="119525058" ID-Zenodo-Dep="6171414" LSID="urn:lsid:plazi:treatment:03F54361FF9FFFA8E68BD6B8B8EB6DD7" httpUri="http://treatment.plazi.org/id/03F54361FF9FFFA8E68BD6B8B8EB6DD7" lastPageNumber="43" pageId="39" pageNumber="43">
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<subSubSection id="C346A1FCFF9FFFA8E68BD6B8BFEC6EC0" box="[151,661,151,177]" pageId="39" pageNumber="43" type="nomenclature">
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<paragraph id="8BE3F277FF9FFFA8E68BD6B8BFEC6EC0" blockId="39.[151,661,151,177]" box="[151,661,151,177]" pageId="39" pageNumber="43">
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<heading id="D0AB451BFF9FFFA8E68BD6B8BFEC6EC0" bold="true" box="[151,661,151,177]" fontSize="11" level="1" pageId="39" pageNumber="43" reason="1">
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD6B8BFEC6EC0" ID-CoL="f7305698-3f52-4e06-904f-b9e642649536" authority="Michaelsen, 1899" authorityName="Michaelsen" authorityYear="1899" box="[151,661,151,177]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD6B8BFEC6EC0" bold="true" box="[151,661,151,177]" pageId="39" pageNumber="43">
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<emphasis id="B9282E65FF9FFFA8E68BD6B8BCDD6EC0" bold="true" box="[151,420,151,177]" italics="true" pageId="39" pageNumber="43">Parachilota wahlbergi</emphasis>
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(Michaelsen, 1899)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346A1FCFF9FFFA8E68BD6F0BF286FFC" pageId="39" pageNumber="43" type="reference_group">
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<paragraph id="8BE3F277FF9FFFA8E68BD6F0BF296E85" blockId="39.[151,1081,222,398]" box="[151,592,222,245]" pageId="39" pageNumber="43">
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<treatmentCitationGroup id="AB4CD559FF9FFFA8E68BD6F0BF296E85" box="[151,592,222,245]" pageId="39" pageNumber="43">
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD6F0BF356E85" authority="Michaelsen, 1899b: 441" authorityName="Michaelsen" authorityPageNumber="441" authorityYear="1899" box="[151,588,222,245]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD6F0BC366E84" box="[151,335,223,245]" italics="true" pageId="39" pageNumber="43">Chilota wahlbergi</emphasis>
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<treatmentCitation id="0AFDD466FF9FFFA8E749D6F1BF356E85" author="Michaelsen" box="[341,588,222,244]" page="441" pageId="39" pageNumber="43" year="1899">
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<bibRefCitation id="EFCD8F86FF9FFFA8E749D6F1BF636E85" author="Michaelsen" box="[341,538,222,244]" pageId="39" pageNumber="43" refString="Michaelsen, W. (1899 b) Revision der Kinberg'schen Oligochaeten-Typen. Ofversigt af Kongliga Vetenskaps-akademiens Forhandlingar, 56, 413 - 448." type="journal article" year="1899" yearSuffix="b">Michaelsen, 1899b</bibRefCitation>
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: 441
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE3F277FF9FFFA8E68BD6D2BFCA6F40" blockId="39.[151,1081,222,398]" pageId="39" pageNumber="43">
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<treatmentCitationGroup id="AB4CD559FF9FFFA8E68BD6D2BFCA6F40" pageId="39" pageNumber="43">
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD6D2BF396F63" authority="Michaelsen 1900: 147" authorityName="Michaelsen" authorityPageNumber="147" authorityYear="1900" box="[151,576,253,275]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD6D2BC366F62" box="[151,335,253,275]" italics="true" pageId="39" pageNumber="43">Chilota wahlbergi</emphasis>
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:
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<treatmentCitation id="0AFDD466FF9FFFA8E740D6D2BF396F63" author="Michaelsen" box="[348,576,253,275]" page="147" pageId="39" pageNumber="43" year="1900">
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<bibRefCitation id="EFCD8F86FF9FFFA8E740D6D2BF776F63" author="Michaelsen" box="[348,526,253,275]" pageId="39" pageNumber="43" refString="Michaelsen, W. (1900) Oligochaeta. In: Das Tierreich. Lief. 10. Berlin: R. Friedlander & Sohn, 575 pp." type="book" year="1900">Michaelsen 1900</bibRefCitation>
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: 147
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</treatmentCitation>
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</taxonomicName>
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; 1912: 146;
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<treatmentCitation id="0AFDD466FF9FFFA8E4A2D6D2BE9D6F63" author="Reynolds" box="[702,996,253,275]" page="189" pageId="39" pageNumber="43" year="1976">
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<bibRefCitation id="EFCD8F86FF9FFFA8E4A2D6D2BEC86F62" author="Reynolds" box="[702,945,253,275]" pageId="39" pageNumber="43" refString="Reynolds J. W. & Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Ottawa: Runge Press. 217 pp." type="book" year="1976">Reynolds & Cook 1976</bibRefCitation>
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: 189
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</treatmentCitation>
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; NHRS.
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD733BC366F43" box="[151,335,284,306]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD733BC366F43" box="[151,335,284,306]" italics="true" pageId="39" pageNumber="43">Chilota wahlbergi</emphasis>
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</taxonomicName>
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f.
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<emphasis id="B9282E65FF9FFFA8E776D732BCCB6F43" box="[362,434,285,306]" italics="true" pageId="39" pageNumber="43">typicus</emphasis>
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:
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<treatmentCitation id="0AFDD466FF9FFFA8E7A3D734BFD76F40" author="Michaelsen" box="[447,686,283,305]" page="416" pageId="39" pageNumber="43" year="1913">
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<bibRefCitation id="EFCD8F86FF9FFFA8E7A3D734BF026F40" author="Michaelsen" box="[447,635,283,305]" pageId="39" pageNumber="43" refString="Michaelsen, W. (1913 c) The Oligochaeta of Natal and Zululand. Annals of the Natal Museum, 2 (4), 397 - 457." type="journal article" year="1913" yearSuffix="c">Michaelsen 1913c</bibRefCitation>
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: 416
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BE3F277FF9FFFA8E68BD715BF286FFC" blockId="39.[151,1081,222,398]" pageId="39" pageNumber="43">
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<treatmentCitationGroup id="AB4CD559FF9FFFA8E68BD715BF286FFC" pageId="39" pageNumber="43">
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD715BC366F21" box="[151,335,314,336]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD715BC366F21" box="[151,335,314,336]" italics="true" pageId="39" pageNumber="43">Chilota wahlbergi</emphasis>
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</taxonomicName>
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f.
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<emphasis id="B9282E65FF9FFFA8E776D715BCB36F21" box="[362,458,314,336]" italics="true" pageId="39" pageNumber="43">pulchrior</emphasis>
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:
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<treatmentCitation id="0AFDD466FF9FFFA8E7CBD715BFBF6F21" author="Michaelsen" box="[471,710,314,336]" page="416" pageId="39" pageNumber="43" year="1913">
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<bibRefCitation id="EFCD8F86FF9FFFA8E7CBD715BFEA6F21" author="Michaelsen" box="[471,659,314,336]" pageId="39" pageNumber="43" refString="Michaelsen, W. (1913 c) The Oligochaeta of Natal and Zululand. Annals of the Natal Museum, 2 (4), 397 - 457." type="journal article" year="1913" yearSuffix="c">Michaelsen 1913c</bibRefCitation>
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: 416
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</treatmentCitation>
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;
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<treatmentCitation id="0AFDD466FF9FFFA8E4CDD715BEEE6F21" author="Pickford" box="[721,919,314,336]" page="461" pageId="39" pageNumber="43" year="1927">
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<bibRefCitation id="EFCD8F86FF9FFFA8E4CDD715BE1D6F21" author="Pickford" box="[721,868,314,336]" pageId="39" pageNumber="43" refString="Pickford, G. E. (1927) The Acanthodriline earthworms of South Africa east of the 25 ° meridian. Albany Museum, 3, 441 - 465." type="journal article" year="1927">Pickford 1927</bibRefCitation>
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: 461
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</treatmentCitation>
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; ZMH.
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD776BF296F1E" ID-CoL="f7305698-3f52-4e06-904f-b9e642649536" authority="Pickford 1937: 336" authorityName="Pickford" authorityPageNumber="336" authorityYear="1937" box="[151,592,345,367]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD776BC056F1E" box="[151,380,345,367]" italics="true" pageId="39" pageNumber="43">Parachilota wahlbergi</emphasis>
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:
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<treatmentCitation id="0AFDD466FF9FFFA8E795D776BF296F1E" author="Pickford" box="[393,592,345,367]" page="336" pageId="39" pageNumber="43" year="1937">
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<bibRefCitation id="EFCD8F86FF9FFFA8E795D776BF646F1E" author="Pickford" box="[393,541,345,367]" pageId="39" pageNumber="43" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer & Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
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: 336
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0AFDD466FF9FFFA8E446D776BF856F1E" author="Plisko" box="[602,764,345,367]" page="30" pageId="39" pageNumber="43" year="2008">
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<bibRefCitation id="EFCD8F86FF9FFFA8E446D776BFAF6F1E" author="Plisko" box="[602,726,345,367]" pageId="39" pageNumber="43" refString="Plisko, J. D. (2008) New South African Acanthodrilinae earthworm species, with new data for some earlier known members of the genus Parachilota (Oligochaeta: Acanthodrilidae). African Invertebrates, 49 (2), 21 - 36." type="journal article" year="2008">Plisko 2008</bibRefCitation>
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: 30
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</treatmentCitation>
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; BMNH; NMSAD.
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<taxonomicName id="4C5C89F4FF9FFFA8E68BD757BF346FFC" authority="Ljungstrom 1972b: 108" authorityName="Ljungstrom" authorityPageNumber="108" authorityYear="1972" box="[151,589,376,398]" class="Clitellata" family="Acanthodrilidae" genus="Udeina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="39" pageNumber="43" phylum="Annelida" rank="species" species="wahlbergi">
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<emphasis id="B9282E65FF9FFFA8E68BD757BC376FFF" box="[151,334,376,398]" italics="true" pageId="39" pageNumber="43">Udeina wahlbergi</emphasis>
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:
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<treatmentCitation id="0AFDD466FF9FFFA8E747D757BF346FFC" author="Ljungstrom" box="[347,589,376,398]" page="108" pageId="39" pageNumber="43" year="1972">
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<bibRefCitation id="EFCD8F86FF9FFFA8E747D757BF636FFF" author="Ljungstrom" box="[347,538,376,398]" pageId="39" pageNumber="43" refString="Ljungstrom, P. O. (1972 b) Taxonomical and ecological notes on the earthworm genus Udeina and a requiem for the South African acanthodrilines. Pedobiologia, 12 (2), 100 - 110." type="journal article" year="1972" yearSuffix="b">Ljungström 1972b</bibRefCitation>
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: 108
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C346A1FCFF9FFFA8E68BD794BE0E6C6B" pageId="39" pageNumber="43" type="materials_examined">
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<paragraph id="8BE3F277FF9FFFA8E68BD794BE0E6C6B" blockId="39.[151,1437,442,935]" pageId="39" pageNumber="43">
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<emphasis id="B9282E65FF9FFFA8E68BD794BC4A6FA2" bold="true" box="[151,307,442,467]" pageId="39" pageNumber="43">
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<typeStatus id="54E74CD5FF9FFFA8E68BD794BDA96FA2" box="[151,208,443,467]" pageId="39" pageNumber="43">Type</typeStatus>
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locality.
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</emphasis>
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Unknown, in
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<collectingCountry id="F34BB2E7FF9FFFA8E7CFD795BF746FA3" box="[467,525,442,466]" name="South Africa" pageId="39" pageNumber="43">
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<collectionCode id="ED4D6AB2FF9FFFA8E7CFD795BF746FA3" box="[467,525,442,466]" pageId="39" pageNumber="43">RSA</collectionCode>
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</collectingCountry>
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. The name ‘Caffraria’ given on original label, translated by Michaelsen (1899) as ‘Kafferlandet’ refers to a name usually understood as ‘South East Africa’, although it may also refers to some other parts of
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<collectingCountry id="F34BB2E7FF9FFFA8E6EFD42DBCFF6C6B" box="[243,390,514,538]" name="South Africa" pageId="39" pageNumber="43">South Africa</collectingCountry>
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. (
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<emphasis id="B9282E65FF9FFFA8E786D42CBFC86C6A" box="[410,689,515,539]" italics="true" pageId="39" pageNumber="43">Encyclopaedia Britanica</emphasis>
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(11th ed., 1911).
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|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C346A1FCFF9FFFA8E6DBD409BEEF6C83" pageId="39" pageNumber="43" type="distribution">
|
|||
|
|
<paragraph id="8BE3F277FF9FFFA8E6DBD409BEEF6C83" blockId="39.[151,1437,442,935]" pageId="39" pageNumber="43">
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E6DBD409BC206C4E" bold="true" box="[199,345,550,575]" pageId="39" pageNumber="43">Distribution</emphasis>
|
|||
|
|
. Known from diverse biotopes, distandly separated sites in three provinces:
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E2A0D409B99A6C4E" bold="true" box="[1212,1251,550,575]" pageId="39" pageNumber="43">GT</emphasis>
|
|||
|
|
: Witpoort Falls, Krokodile river (for f.
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E78BD464BC8C6C12" box="[407,501,587,611]" italics="true" pageId="39" pageNumber="43">pulchior</emphasis>
|
|||
|
|
, in
|
|||
|
|
<bibRefCitation id="EFCD8F86FF9FFFA8E43DD465BF9F6C13" author="Michaelsen" box="[545,742,586,610]" pageId="39" pageNumber="43" refString="Michaelsen, W. (1912) Die terrestrischen Oligochaeten des tropischen Afrikas und ihre geographischen Beziehungen. 3. Zoologie 1. In: Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika Expedition 1907 - 1908. Leipzig: Klinkhardt & Biermann, 3, 1 - 90." type="journal article" year="1912">Michaelsen 1912</bibRefCitation>
|
|||
|
|
). Witpoortje Falls, in wet soil among roots of grass and moss growing behind the spray of the waterfall, refers to Pickford material (1937: 338).
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E230D440B9216CF6" bold="true" box="[1068,1112,623,647]" pageId="39" pageNumber="43">MP</emphasis>
|
|||
|
|
: Waterval Boven, behind the waterfall in very wet soil with the grass roots and liverworts. Lake Chrissie, Ermelo area, damp turf near the spring.
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E68BD498BDAD6CBE" bold="true" box="[151,212,695,719]" pageId="39" pageNumber="43">KZN</emphasis>
|
|||
|
|
: Farm between Pietermaritzburg and Greytown, at the roots of ferns. Farm Linwood (Vicinity of Midmar Dam), in the temporary pool during dry phase, under the moist rock.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C346A1FCFF9FFFA8E6DBD4D0B8EB6DD7" pageId="39" pageNumber="43" type="discussion">
|
|||
|
|
<paragraph id="8BE3F277FF9FFFA8E6DBD4D0B8EB6DD7" blockId="39.[151,1437,442,935]" pageId="39" pageNumber="43">
|
|||
|
|
<emphasis id="B9282E65FF9FFFA8E6DBD4D0BC4D6D66" bold="true" box="[199,308,767,791]" pageId="39" pageNumber="43">Remarks</emphasis>
|
|||
|
|
.
|
|||
|
|
<typeStatus id="54E74CD5FF9FFFA8E75FD4D1BCD26D67" box="[323,427,766,790]" pageId="39" pageNumber="43" type="holotype">Holotype</typeStatus>
|
|||
|
|
NHRS 164.
|
|||
|
|
<typeStatus id="54E74CD5FF9FFFA8E422D4D1BFE36D67" box="[574,666,766,790]" pageId="39" pageNumber="43" type="syntype">Syntype</typeStatus>
|
|||
|
|
ZMH V-7494. Other material: SAMC A20615. NMSA/Olig.03650. It may be noted that the collector of the
|
|||
|
|
<typeStatus id="54E74CD5FF9FFFA8E470D50CBFE56D4A" box="[620,668,803,827]" pageId="39" pageNumber="43">type</typeStatus>
|
|||
|
|
material of this species, J.A.Wahlberg, spent some years in Africa travelling and collecting animals and plants. His travels included repeated transits through Cape Town, Durban, large part of
|
|||
|
|
<collectingCountry id="F34BB2E7FF9FFFA8E72FD545BC096DF3" box="[307,368,874,898]" name="South Africa" pageId="39" pageNumber="43">Natal</collectingCountry>
|
|||
|
|
including coast to both Umfolozi rivers, Drakensberg, highveld in Transvaal, Megalisberg, Springbok Flats, across Limpopo river and
|
|||
|
|
<collectingCountry id="F34BB2E7FF9FFFA8E461D5A1BF876DD6" box="[637,766,910,935]" name="Namibia" pageId="39" pageNumber="43">Walvis Bay</collectingCountry>
|
|||
|
|
going to
|
|||
|
|
<collectingCountry id="F34BB2E7FF9FFFA8E574D5A1BEB16DD7" box="[872,968,910,934]" name="Namibia" pageId="39" pageNumber="43">Namibia</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F34BB2E7FF9FFFA8E5E3D5A0B9146DD7" box="[1023,1133,911,934]" name="Botswana" pageId="39" pageNumber="43">Botswana</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFCD8F86FF9FFFA8E260D5A1B8FC6DD7" author="Craig" box="[1148,1413,910,934]" pageId="39" pageNumber="43" refString="Craig, A. & Hummel, C. (1994) Johan August Wahlberg. Travels journals (and some letters). South Africa and Namibia / Botswana, 1838 - 1856. Translated from the Swedish by Michael Roberts. Cape Town, Van Riebeck Society, 249 pp." type="book chapter" year="1994">Craig & Hummel 1994</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|