<documentid="FEF68D34EAAFC89EC8A4D432E979943F"ID-DOI="10.5852/ejt.2018.399"ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99"ID-ISSN="2118-9773"ID-Zenodo-Dep="1163422"ID-ZooBank="E836E138-D6E2-4F62-B4B3-CE2E073F2B24"IM.bibliography_approvedBy="guilherme"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="guilherme"IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi"IM.taxonomicNames_approvedBy="guilherme"IM.treatments_approvedBy="guilherme"checkinTime="1517404992146"checkinUser="plazi"docAuthor="Zonstein, Sergei, Kunt, Kadir B. & Yağmur, Ersen A."docDate="2018"docId="03B9B44C567B0C29516C84AAFB694D74"docLanguage="en"docName="ejt-399_zonstein_kunt.pdf.imf"docOrigin="European Journal of Taxonomy 399"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Raveniola hyrcanica Dunin 1988"docType="treatment"docVersion="17"lastPageNumber="15"masterDocId="FF80CC3456770C27533C8513FFDA4805"masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia"masterLastPageNumber="93"masterPageNumber="1"pageNumber="13"updateTime="1734611146249"updateUser="guilherme">
<mods:titleid="F8314A8EC2615938B24FC9D6F975234A">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
<bibRefCitationid="EF8178AB567B0C2B506D84A9FC3E49D1"author="Dunin P. M."box="[849,996,442,468]"journalOrPublisher="Zoologicheskii Zhurnal"pageId="12"pageNumber="13"pagination="1245 - 1248"part="67"refId="ref35521"refString="Dunin P. M. 1988. Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan. Zoologicheskii Zhurnal 67 (8): 1245 - 1248. [In Russian with English summary]"title="Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan"type="journal article"year="1988">Dunin, 1988</bibRefCitation>
<figureCitationid="132B19DF567B0C2B52A584CDFDD849FD"box="[409,514,478,504]"captionStart-0="Figs 1–6"captionStart-1="Figs 7–12"captionStartId-0="50.[189,241,1734,1760]"captionStartId-1="51.[189,241,1719,1745]"captionTargetBox-0="[189,1399,265,1680]"captionTargetBox-1="[189,1399,265,1680]"captionTargetId-0="figure@50.[189,1399,265,1680]"captionTargetId-1="figure@51.[189,1399,265,1680]"captionTargetPageId-0="50"captionText-0="Figs 1–6.Raveniola spp., ♀♀ (1–3, 5–6) and ♂ (4). Chelicerae (1), apical edge of maxilla (2–3), palpal organ (4) and tarsus I (5–6); ventroapical (1), ventral (2–4, 6) and dorsal (5) views. — 1–2. R. micropa (Ausserer, 1871). 3–5. R. pontica (Spassky, 1937). 6. R. hyrcanica Dunin, 1988."captionText-1="Figs 7–12. Raveniola spp., ♀♀. Tarsal organ, legs I (7–8, 10, 12) and IV (9, 11), dorsal views. — 7–9. R. hyrcanica Dunin, 1988. 10. R. pontica (Spassky, 1937). 11. R. micropa (Ausserer, 1871). 12. R. zaitzevi (Charitonov, 1948)."httpUri-0="https://zenodo.org/record/1163424/files/figure.png"httpUri-1="https://zenodo.org/record/1163426/files/figure.png"pageId="12"pageNumber="13">Figs 6–9</figureCitation>
,
<figureCitationid="132B19DF567B0C2B513284CCFD8049FC"box="[526,602,479,505]"captionStart="Figs 13–18"captionStartId="52.[189,241,1725,1751]"captionTargetBox="[189,1399,265,1680]"captionTargetId="figure@52.[189,1399,265,1680]"captionText="Figs 13–18. Raveniola spp., ♀♀. Bases of trichobothria, legs I (13, 15–16, 18) and IV (14, 17), dorsal views.—13–14.R.hyrcanicaDunin,1988.15.R.pontica(Spassky,1937).16–17.R.micropa(Ausserer, 1871). 18. R. zaitzevi (Charitonov, 1948)."httpUri="https://zenodo.org/record/1163428/files/figure.png"pageId="12"pageNumber="13">13–14</figureCitation>
,
<figureCitationid="132B19DF567B0C2B515A84CCFD6849FC"box="[614,690,479,505]"captionStart="Figs 19–24"captionStartId="53.[189,241,1725,1751]"captionTargetBox="[189,1399,265,1678]"captionTargetId="figure@53.[189,1399,265,1680]"captionText="Figs 19–24. Raveniola spp., ♀♀. Palpal (19) and tarsal (20–24) claws; apical (19, 21, 23) and lateral (20, 22, 24) views. — 19–20. R. pontica (Spassky, 1937). 21–22. R. hyrcanica Dunin, 1988, legs I and IV. 23–24. R. micropa (Ausserer, 1871), legs I and IV."httpUri="https://zenodo.org/record/1163430/files/figure.png"pageId="12"pageNumber="13">21–22</figureCitation>
<figureCitationid="132B19DF567B0C2B50D28710FBC04A18"box="[1006,1050,515,541]"captionStart="Figure 220"captionStartId="86.[189,272,1534,1560]"captionTargetBox="[190,1398,265,1471]"captionTargetId="figure@86.[189,1398,265,1471]"captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey."httpUri="https://zenodo.org/record/1163496/files/figure.png"pageId="12"pageNumber="13">220</figureCitation>
<bibRefCitationid="EF8178AB567B0C2B528B8755FD4C4A64"author="Dunin P. M."box="[439,662,582,609]"journalOrPublisher="Zoologicheskii Zhurnal"pageId="12"pageNumber="13"pagination="1245 - 1248"part="67"refId="ref35521"refString="Dunin P. M. 1988. Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan. Zoologicheskii Zhurnal 67 (8): 1245 - 1248. [In Russian with English summary]"title="Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan"type="journal article"year="1988">Dunin, 1988: 1245</bibRefCitation>
<bibRefCitationid="EF8178AB567B0C2B52F1879AFD454AA1"author="Platnick N. I."box="[461,671,649,676]"journalOrPublisher="The New York Entomological Society, New York"pageId="12"pageNumber="13"pagination="1940 - 1980"refId="ref36412"refString="Platnick N. I. 1993. Advances in Spider Taxonomy 1988 - 1991, with Synonymies and Transfers 1940 - 1980. The New York Entomological Society, New York."title="Advances in Spider Taxonomy 1988 - 1991, with Synonymies and Transfers"type="book"year="1993">Platnick 1993: 95</bibRefCitation>
. —
<bibRefCitationid="EF8178AB567B0C2B51F3879AFC664AA1"author="Mikhailov K. G."box="[719,956,649,676]"journalOrPublisher="Zoological Museum of the Moscow State University, Moscow"pageId="12"pageNumber="13"refId="ref36266"refString="Mikhailov K. G. 1997. Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei). Zoological Museum of the Moscow State University, Moscow."title="Catalogue of the Spiders of the Territories of the Former Soviet Union (Arachnida, Aranei)"type="book"year="1997">Mikhailov 1997: 20</bibRefCitation>
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<paragraphid="8BAF055A567B0C2A53818271FD084949"blockId="12.[189,1399,1890,2027]"lastBlockId="13.[189,1399,267,332]"lastPageId="13"lastPageNumber="14"pageId="12"pageNumber="13">COLOUR. Carapace reddish chestnut with dense brownish black pilosity, eye tubercle not darkened but with partially fused blackish brown spots bordering eyes; chelicerae, almost whole leg I (except lighter tarsus), femora of palps and legs II–IV dorsally dark reddish chestnut; other parts of legs dorsally with lighter reddish chestnut coloration; sternum, labium, maxillae, palps and legs ventrally light brownish orange; abdomen dorsally light chestnut, with numerous, small lighter brownish spots, ventral abdominal surface and spinnerets light yellowish brown.</paragraph>
<figureCitationid="132B19DF567A0C2A503E8464FC864994"box="[770,860,375,401]"captionStart="Figs 68–73"captionStartId="60.[189,241,1882,1908]"captionTargetBox="[190,1398,265,1816]"captionTargetId="figure@60.[189,1398,265,1816]"captionText="Figs 68–73. Raveniola spp., ♂♂. Carapace and chelicerae, dorsal views. — 68. R. adjarica sp. nov. (holotype). 69. R. dunini sp. nov. (holotype). 70. R. hyrcanica Dunin, 1988 (holotype). 71. R. pontica (Spassky, 1937) (Lazarevskoe). 72. R. birecikensis sp. nov. (holotype). 73. Same species as preceding (paratype, Birecik). Scale bars: 1.0 mm."httpUri="https://zenodo.org/record/1163444/files/figure.png"pageId="13"pageNumber="14">Fig. 70</figureCitation>
. Clypeus and eye group as in
<figureCitationid="132B19DF567A0C2A57EE8464FAE04994"box="[1234,1338,375,401]"captionStart="Figs 102–116"captionStartId="66.[189,241,1755,1781]"captionTargetBox="[204,1382,265,1739]"captionTargetId="figure@66.[187,1396,250,1752]"captionText="Figs 102–116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt & Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani & Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm."httpUri="https://zenodo.org/record/1163456/files/figure.png"pageId="13"pageNumber="14">Fig. 104</figureCitation>
. Eye diameters and interdistances: AME 0.22(0.29), ALE 0.28, PLE 0.22, PME 0.16, AME–AME 0.17(0.10), ALE–AME 0.15(0.11), ALE–PLE 0.20, PLE–PME 0.10, PME–PME 0.56. Chelicerae: each furrow with 10–11 promarginal teeth and 0–1 mesobasal denticles. Maxillae with 8 large cuspules each. Sternal sigilla oval and submarginal, as shown in
<figureCitationid="132B19DF567A0C2A51958701FCD94A2E"box="[681,771,529,556]"captionStart="Figs 59–67"captionStartId="59.[189,241,1662,1688]"captionTargetBox="[189,1398,265,1617]"captionTargetId="figure@59.[189,1398,265,1617]"captionText="Figs 59–67. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (60–62, 64–65, 67). Left chelicera (59–61; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (62–64), and spinnerets (65– 67); prolateral (59–61,65, 67)and ventral (62–64,66) views.— 59–60. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm."httpUri="https://zenodo.org/record/1163442/files/figure.png"pageId="13"pageNumber="14">Fig. 62.</figureCitation>
. Scopula (dense, scopuliform hairs moderately long): distal on metatarsi I–II, entire on tarsi I–II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–12 each on tibiae, 13–18 on metatarsi, 15–19 on tarsi, 10 on cymbium. Paired tarsal claws: inner and outer margins with 7–8 teeth each.
<paragraphid="8BAF055A567A0C2A538183E2FBDA4F0E"blockId="13.[189,1024,1777,1804]"box="[189,1024,1777,1804]"pageId="13"pageNumber="14">COLOUR. As in male, but legs I–IV dorsally uniformly reddish-chestnut.</paragraph>
<figureCitationid="132B19DF567A0C2A51D28225FC9E4F55"box="[750,836,1846,1872]"captionStart="Figs 84–89"captionStartId="63.[189,241,1892,1918]"captionTargetBox="[189,1398,266,1873]"captionTargetId="figure@63.[189,1398,265,1878]"captionText="Figs 84–89. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm."httpUri="https://zenodo.org/record/1163450/files/figure.png"pageId="13"pageNumber="14">Fig. 86</figureCitation>
, cephalic part noticeably elevated over thoracic part. Clypeus and eye group as in
<figureCitationid="132B19DF567A0C2A5160824EFD194F72"box="[604,707,1885,1911]"captionStart="Figs 117–128"captionStartId="67.[189,241,1621,1647]"captionTargetBox="[193,1382,278,1581]"captionTargetId="figure@67.[189,1398,265,1581]"captionText="Figs 117–128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm."httpUri="https://zenodo.org/record/1163458/files/figure.png"pageId="13"pageNumber="14">Fig. 119</figureCitation>
. Eye diameters and interdistances: AME 0.20(0.28), ALE 0.32, PLE 0.25, PME 0.16, AME–AME 0.22(0.15), ALE–AME 0.23(0.19), ALE–PLE 0.22, PLE–PME 0.08, PME–PME 0.71. Each cheliceral furrow with 10 promarginal teeth and 3–5 mesobasal denticles. Maxillae trapezoidal, each with 7–8 cuspules.
</paragraph>
<paragraphid="8BAF055A56790C295381841FFC7B496A"blockId="14.[189,1398,267,368]"pageId="14"pageNumber="15">LEGS. Scopula: dense, scopuliform hairs moderately long. Trichobothria: 2 rows of 10–12 each on tibiae, 12–17 on metatarsi, 13–16 on tarsi, 11 on palpal tarsus. Paired tarsal claws: outer margin with 5–6 teeth, inner one with 4–5 teeth. Palpal claw with 4 promarginal teeth.</paragraph>
<paragraphid="8BAF055A56790C295381848BFD804A6F"blockId="14.[189,1399,408,618]"pageId="14"pageNumber="15">SPINATION. Femora I–II with one basodorsal spine and 2–3 dorsal bristles; patellae I–II and palpal patella aspinose. Palp: femur d1, pd1(0); tibia v2–2–2(3); tarsus d7(8). Leg I: femur d1, pd1; tibia v2–1–2– 2(3); metatarsus v2–2–2(3). Leg II: femur d1; pd1; tibia p1(0)–1, v2–2–3; metatarsus v2–2–2(3). Leg III: femur d1(0)–1, pd1, rd1(0)–1; patella p1, r1; tibia p1–1, r1–1, v2–2–3; metatarsus p1–1–1, r1–1–1, v2–1(0)–2–3. Leg IV: femur d1–1–1(0)–1(0), pd1, rd1; patella r1; tibia p1–1, r1–1, v2–2–3; metatarsus p1–1–1–1, r1–1–1–1, v2–2–1–2–2.</paragraph>
Carapace length in males varies from 7.25 to 8.57, in females from 6.88 to 10.13. In some females, the dorsal abdominal pattern is almost indistinct. Spermathecae show a quite insignificant variation (cf.
<taxonomicNameid="4C107ED956790C2952848142FCED4C6E"authority="C. A. Mey"authorityName="C.A. Mey"box="[440,823,1104,1131]"class="Magnoliopsida"family="Fagaceae"genus="Quercus"higherTaxonomySource="GBIF"kingdom="Plantae"order="Fagales"pageId="14"pageNumber="15"phylum="Tracheophyta"rank="species"species="castaneifolia">
<figureCitationid="132B19DF56790C29504A8143FBC64C6F"box="[886,1052,1104,1130]"captionStart="Figs 223–229"captionStartId="88.[189,241,1652,1678]"captionTargetBox="[189,1399,306,1600]"captionTargetId="figure@88.[189,1399,265,1600]"captionText="Figs 223–229. Habitats of Raveniola spp. — 223–224. Azerbaijan, southern macroslope of Caucasus Major,forested midlands inBelokany area, betweenLagodekhi,Georgia [thetypelocality of R. zaitzevi (Charitonov, 1948)] and Zakataly, Azerbaijan. 225–227. Abkhazia, canyon of the Bzyb River, forested biotopes inhabited by R.pontica (Spassky, 1937). 228–229.Azerbaijan, foothillsof TalyshMts,Hyrcan Reserve, humid localities typical for R. hyrcanica Dunin, 1988. Images courtesy of Natasha Snegovaya (223–224, 228–229) and Mykola Kovblyuk (225–227)."httpUri="https://zenodo.org/record/1163502/files/figure.png"pageId="14"pageNumber="15">Figs 228–229</figureCitation>
). Unlike other congeners, the representatives of this species do not exclusively use natural hideouts but build simple open burrows in the forest floor. Burrow length reaches 30–35 cm, with diameter varying from 2.5 to 3.0 cm. It comprises a fairly straight vertical hole which then curves and ends in an expanded sub-horizontal dwelling chamber; no silk-lining was found (
<bibRefCitationid="EF8178AB56790C29515D81F7FD364CFB"author="Dunin P. M."box="[609,748,1252,1278]"journalOrPublisher="Zoologicheskii Zhurnal"pageId="14"pageNumber="15"pagination="1245 - 1248"part="67"refId="ref35521"refString="Dunin P. M. 1988. Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan. Zoologicheskii Zhurnal 67 (8): 1245 - 1248. [In Russian with English summary]"title="Mygalomorphic spiders (Aranei, Mygalomorphae) of Azerbaijan"type="journal article"year="1988">Dunin 1988</bibRefCitation>
South Azerbaijan: Talysh Mts (the northwestern continuation of Alborz). See
<figureCitationid="132B19DF56790C2957768044FB6A4D74"box="[1098,1200,1367,1393]"captionStart="Figure 220"captionStartId="86.[189,272,1534,1560]"captionTargetBox="[190,1398,265,1471]"captionTargetId="figure@86.[189,1398,265,1471]"captionText="Figure 220. Localities of Raveniola spp. in the eastern part of the Caucasus: R. dunini sp. nov. (), R. hyrcanica Dunin, 1988 () and R. zaitzevi (Charitonov, 1948) ().Abbreviation:TU = Turkey."httpUri="https://zenodo.org/record/1163496/files/figure.png"pageId="14"pageNumber="15">Fig. 220</figureCitation>