<documentid="7D312C11844DC2C82A64C93A2FDA833D"ID-CLB-Dataset="8134"ID-DOI="10.11646/zootaxa.4927.1.2"ID-GBIF-Dataset="f8e774a8-36c4-482c-8d53-adc4612ea281"ID-ISSN="1175-5326"ID-Zenodo-Dep="4533849"ID-ZooBank="E4C28E67-3072-4023-BD47-C2FD9CECAAE5"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_approvedBy="admin"IM.taxonomicNames_approvedBy="admin"checkinTime="1613034223463"checkinUser="plazi"docAuthor="Lyu, Zhi-Tong, Zeng, Zhao-Chi, Wang, Jian, Liu, Zu-Yao, Huang, Ya-Qiong, Li, Wen-Zhou & Wang, Ying-Yong"docDate="2021"docId="03B087AFFFACFFB3628EFF28FD44D5E2"docLanguage="en"docName="zootaxa.4927.1.2.pdf"docOrigin="Zootaxa 4927 (1)"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Panophrys daoji Lyu, Zeng, Wang & Wang 2021, sp. nov."docType="treatment"docVersion="10"lastPageNumber="26"masterDocId="FF89FFD7FFBCFFA26219FFBFFF8DD303"masterDocTitle="Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus"masterLastPageNumber="40"masterPageNumber="9"pageNumber="25"updateTime="1720383473109"updateUser="admin"zenodo-license-document="CLOSED">
<mods:titleid="D6AF6F1199111731CAE3C1F1B91FD088">Four new species of Panophrys (Anura, Megophryidae) from eastern China, with discussion on the recognition of Panophrys as a distinct genus</mods:title>
<mods:affiliationid="CE481BC3BF6C4CB657AE9C3F542FE3DB">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859</mods:affiliation>
<mods:affiliationid="95282611B575D17291310591F8B9DE2B">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399</mods:affiliation>
<mods:affiliationid="4485ED28D9389529DA383AF7650542E4">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Shenzhen Shuanghuamu Biological Technology Co., Ltd, Shenzhen 51800, PR China & wangj 397 @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 4249 - 7767</mods:affiliation>
<mods:affiliationid="9099EB1A9E96201B34BE2182EFF55A32">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China & Institute of Ecology and Evolution, University of Bern, Bern 3012, Switzerland.</mods:affiliation>
<mods:affiliationid="6A8B72A472ACEF2C077C8A778230E3B1">Fujian Daiyun Mountain National Nature Reserve, Quanzhou 362500, PR China & Fujian Wildlife Conservation Center, Fujian Provincial Department of Forestry, Fuzhou 350003, PR China</mods:affiliation>
<mods:affiliationid="40B1ED658BB6895EADCC18E8983528BC">State Key Laboratory of Biocontrol / The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, PR China</mods:affiliation>
<taxonomicNameid="4C194D3AFFACFFB2628EFF28FD26D3B2"authority="Lyu, Zeng, Wang & Wang"authorityName="Lyu, Zeng, Wang & Wang"authorityYear="2021"box="[151,683,151,178]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB26335FF63FE13D3F7"ID-CoL="5N6Z"authorityName="Kuhl & Van Hasselt"authorityYear="1822"box="[300,414,220,244]"class="Amphibia"family="Megophryidae"genus="Megophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="genus">
<bibRefCitationid="EF884B48FFACFFB263F4FF63FD0ED3F7"author="Liu, Z. Y. & Chen, G. L. & Zhu, T. Q. & Zeng, Z. C. & Lyu, Z. T. & Wang, J. & Messenger, K. & Greenberg, A. J. & Guo, Z. X. & Yang, Z. H. & Shi, S. H. & Wang, Y. Y."box="[493,643,220,244]"pageId="16"pageNumber="25"pagination="723 - 731"refId="ref18211"refString="Liu, Z. Y., Chen, G. L., Zhu, T. Q., Zeng, Z. C., Lyu, Z. T., Wang, J., Messenger, K., Greenberg, A. J., Guo, Z. X., Yang, Z. H., Shi, S. H. & Wang, Y. Y. (2018) Prevalence of cryptic species in morphologically uniform taxa-Fast speciation and evolutionary radiation in Asian frogs. Molecular Phylogenetics and Evolution, 127, 723 - 731. https: // doi. org / 10.1016 / j. ympev. 2018.06.020"type="journal article"year="2018">
<materialsCitationid="3B713CE4FFACFFB2628EFE9DFE86D287"ID-GBIF-Occurrence="3038228302"collectingDate="2017-07-31"collectorName="Jian Wang & Zhao-Chi Zeng "country="China"county="Tiantai County"elevation="680"latitude="29.2781"location="Mt Tiantai"longLatPrecision="6"longitude="121.0996"municipality="Taizhou City"pageId="16"pageNumber="25"specimenCount="1"stateProvince="Zhejiang"typeStatus="holotype">
<figureCitationid="13222A3CFFACFFB263A0FE9DFE76D23F"box="[441,507,290,316]"captionStart="FIGURE 4"captionStartId="18.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-25@18.[151,1436,181,1842]"captionTargetPageId="18"captionText="FIGURE 4. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a006209 of Panophrys daoji sp. nov. in life. (E)–(F) dorsolateral views of adult male paratype SYS a006211 and adult female paratype SYS a006214."figureDoi="http://doi.org/10.5281/zenodo.4533858"httpUri="https://zenodo.org/record/4533858/files/figure.png"pageId="16"pageNumber="25">Fig. 4</figureCitation>
<collectingDateid="EFE3E991FFACFFB266C8FE9DFAEDD23F"box="[1233,1376,290,316]"pageId="16"pageNumber="25"value="2017-07-31">31 July 2017</collectingDate>
<materialsCitationid="3B713CE4FFACFFB262DEFE31FBA2D2CF"ID-GBIF-Occurrence="3038228301"collectingDate="2017-07-31"collectionCode="CIB"collectorName="Jian Wang & Zhao-Chi Zeng"country="China"county="Tiantai County"elevation="680"latitude="29.2781"location="Mt Tiantai"longLatPrecision="6"longitude="121.0996"municipality="Taizhou City"pageId="16"pageNumber="25"specimenCode="CIB 116076"specimenCount="1"stateProvince="Zhejiang"typeStatus="paratype">
<specimenCodeid="DBBF9EC2FFACFFB261DBFE31FBC1D2AA"box="[962,1100,398,425]"collectionCode="CIB"country="China"httpUri="http://grbio.org/cool/vxkr-andy"name="Chengdu Institute of Biology"pageId="16"pageNumber="25">CIB 116076</specimenCode>
) and two adult females (SYS a006213–6214), collected at the same time from the same locality as the holotype
<taxonomicNameid="4C194D3AFFACFFB26022FE69FDF8D2F3"authorityName="Lyu, Zeng, Wang & Wang"authorityYear="2021"box="[571,629,470,496]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji">
is used as a noun in apposition, and refers to the Master Daoji (ḋằñ ®), also known as Ji Gong (ằẌ). He purportedly possessed supernatural powers which he used to help the poor and stand up to injustice, and became a famous legend in Chinese culture and a well-known deity in Chinese folk religion. Master Daoji was born in Yongning Village situated at the foot of Mt Tiantai where is the
(n = 2) in adult females; (2) tympanum edge distinct and raised, upper margin in contact with supratympanic fold; (3) vomerine teeth absent; (4) margin of tongue not notched; (5) hindlimbs short, heels not meeting; (6) tibio-tarsal articulation reaching at center of tympanum; (7) TIB/SVL 0.39–0.42, FTL/SVL 0.57–0.64; (8) narrow lateral fringes on fingers present, one subarticular tubercle present at the base of each finger, relative finger lengths II <I <IV <III; (9) rudimentary web and narrow lateral fringes on toes present, one subarticular tubercle present at the base of each toe; (10) dorsal body’s skin texture rough, with densely-distributed granules and tubercles, discontinuous “X” or “)(”-shaped skin ridge on center of dorsum, short transverse skin ridges present on dorsal shank and thigh; (11) flank with densely-distributed tubercles; (12) a small horn-like tubercle present at edge of upper eyelid; (13) supratympanic fold distinct with raised conical tubercles; (14) coloration of dorsal surfaces variable, a dark brown triangular marking with light edge between eyes, a dark “X” or “)(”-shaped marking, with light edge, present on the center of dorsum; (15) single subgular vocal sac present in males; (16) black nuptial spines present on the finger I in breeding males.
<taxonomicNameid="4C194D3AFFACFFB26079FB3DFC97D79F"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[608,794,1154,1180]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB261A3FB3CFBD4D79F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[954,1113,1154,1180]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daiyunensis">
<taxonomicNameid="4C194D3AFFACFFB26725FB3DFF34D7C3"authorityName="Rao & Yang"authorityYear="1997"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="genus">
<tableCitationid="C69B0302FFACFFB263DDFB19FD95D7C3"box="[452,536,1190,1216]"captionStart="TABLE 3"captionStartId="8.[151,239,468,494]"captionTargetPageId="8"captionText="TABLE 3. Diagnostic characters separating all 51 species of the genus Panophrys."httpUri="http://table.plazi.org/id/DF666631FFB4FFAA628EFE6BFB81D2ED"pageId="16"pageNumber="25"tableUuid="DF666631FFB4FFAA628EFE6BFB81D2ED">Table 3</tableCitation>
<taxonomicNameid="4C194D3AFFACFFB262DEFB75FE09D7E7"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[199,388,1226,1252]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB2609DFB74FCA8D7E7"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[644,805,1226,1252]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daiyunensis">
in females), vomerine teeth absent (vs present), heels not meeting (vs overlapping or meeting), and nuptial spines present on the finger I in breeding males (vs absent).
<taxonomicNameid="4C194D3AFFACFFB263DEFAE5FDCDD677"authorityName="Rao & Yang"authorityYear="1997"box="[455,576,1370,1396]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="genus">
<taxonomicNameid="4C194D3AFFACFFB26679FAE4FB38D677"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1120,1205,1370,1396]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB2611BFA1DFC31D6BF"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[770,956,1442,1468]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB266B5FA1CFAF7D6BF"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="2005"box="[1196,1402,1442,1468]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="huangshanensis">
in female), tongue not notched (vs notched), narrow lateral fringes and rudimentary web on toes present (vs both absent), and round light patches on the shoulder absent (vs present).
<taxonomicNameid="4C194D3AFFACFFB26361F9B1FDBFD52B"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[376,562,1550,1576]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB26725F98DFF76D573"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB263EBF9E8FD09D573"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu & Jiang"baseAuthorityYear="2017"box="[498,644,1622,1648]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="lishuiensis">
<taxonomicNameid="4C194D3AFFACFFB26336F9C5FE67D597"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[303,490,1658,1684]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB260F9F9C4FCF3D597"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Messenger & Dahn"baseAuthorityYear="2019"box="[736,894,1658,1684]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="ombrophila">
<taxonomicNameid="4C194D3AFFACFFB266FBF921FA10D5BB"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1250,1437,1694,1720]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB2639CF97CFD93D5DF"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Wu, Peng, Shi, Lu & Wu"baseAuthorityYear="2020"box="[389,542,1731,1756]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="xianjuensis">
<taxonomicNameid="4C194D3AFFACFFB266AEF959FAFED403"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[1207,1395,1766,1792]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
<taxonomicNameid="4C194D3AFFACFFB263BAF890FE70D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Jin"baseAuthorityYear="2014"box="[419,509,1839,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="acuta">
<taxonomicNameid="4C194D3AFFACFFB26013F890FD34D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye, Fei & Xie"baseAuthorityYear="2007"box="[522,697,1838,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="baolongensis">
<taxonomicNameid="4C194D3AFFACFFB260DEF890FCE4D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Fei & Ye"baseAuthorityYear="2009"box="[711,873,1838,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="binlingensis">
<taxonomicNameid="4C194D3AFFACFFB2616EF890FB93D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen"baseAuthorityYear="1994"box="[887,1054,1838,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="caudoprocta">
<taxonomicNameid="4C194D3AFFACFFB26635F890FB09D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Liu"baseAuthorityYear="2014"box="[1068,1156,1838,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="cheni">
<taxonomicNameid="4C194D3AFFACFFB26688F890FAD9D44B"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[1169,1364,1838,1864]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="hoanglienensis">
<taxonomicNameid="4C194D3AFFACFFB2677BF890FF6ED46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Liu, Lyu, Zeng & Wang"baseAuthorityYear="2017"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="insularis">
<taxonomicNameid="4C194D3AFFACFFB262E8F8ECFE24D46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Fei & Ye"baseAuthorityYear="1983"box="[241,425,1874,1900]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="jingdongensis">
<taxonomicNameid="4C194D3AFFACFFB263AFF8ECFDE0D46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang"baseAuthorityYear="2012"box="[438,621,1875,1900]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="jinggangensis">
<taxonomicNameid="4C194D3AFFACFFB26062F8ECFD77D46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou"baseAuthorityYear="2017"box="[635,762,1874,1900]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="liboensis">
<taxonomicNameid="4C194D3AFFACFFB26111F8ECFC05D46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang & Zhao"baseAuthorityYear="2020"box="[776,904,1874,1900]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="mirabilis">
<taxonomicNameid="4C194D3AFFACFFB2618FF8ECFC7FD46F"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Li & Zhao"baseAuthorityYear="2014"box="[918,1010,1874,1900]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="obesa">
<taxonomicNameid="4C194D3AFFACFFB2628EF8C8FECCD493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Jiang, Ye & Fei"baseAuthorityYear="2008"box="[151,321,1910,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="sangzhiensis">
<taxonomicNameid="4C194D3AFFACFFB26357F8C8FD99D493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tian & Sun"baseAuthorityYear="1995"box="[334,532,1910,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="shuichengensis">
<taxonomicNameid="4C194D3AFFACFFB26038F8C8FD02D493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu & Hu"baseAuthorityYear="1973"box="[545,655,1911,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="spinata">
<taxonomicNameid="4C194D3AFFACFFB26084F8C8FCF7D493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Shen, Mo & Li"baseAuthorityYear="2010"box="[669,890,1910,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="tuberogranulatus">
<taxonomicNameid="4C194D3AFFACFFB26191F8C8FBC9D493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[904,1092,1911,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="xiangnanensis">
<taxonomicNameid="4C194D3AFFACFFB26698F8C8FACED493"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Zeng & Wang"baseAuthorityYear="2020"box="[1153,1347,1911,1936]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="yangmingensis">
<taxonomicNameid="4C194D3AFFACFFB262DEF825FE0ED4B7"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[199,387,1946,1972]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
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<taxonomicNameid="4C194D3AFFACFFB26159F800FB88D4DB"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[832,1029,1982,2008]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFACFFB2660CF800FB47D4DB"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[1045,1226,1982,2008]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="fansipanensis">
<taxonomicNameid="4C194D3AFFACFFB266C3F800FAF5D4DB"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[1242,1400,1982,2008]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFACFFB36791F800FEACD3B2"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng & Wang"baseAuthorityYear="2019"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"lastPageId="17"lastPageNumber="26"order="Anura"pageId="16"pageNumber="25"phylum="Chordata"rank="species"species="nankunensis">
<taxonomicNameid="4C194D3AFFADFFB36329FF28FE57D3B2"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[304,474,151,177]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="nanlingensis">
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<taxonomicNameid="4C194D3AFFADFFB366E5FF28FF38D3D7"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="fansipanensis">
<taxonomicNameid="4C194D3AFFADFFB362DAFF04FED2D3D6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[195,351,187,213]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFADFFB363C3FF04FD09D3D6"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[474,644,187,213]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="nanlingensis">
<taxonomicNameid="4C194D3AFFADFFB36791FF04FF56D3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che"baseAuthorityYear="2019"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="angka">
<taxonomicNameid="4C194D3AFFADFFB362F0FF60FE2ED3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[233,419,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="binchuanensis">
<taxonomicNameid="4C194D3AFFADFFB363A8FF60FD81D3FB"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang & Wu"baseAuthorityYear="2020"box="[433,524,223,248]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="jiangi">
<taxonomicNameid="4C194D3AFFADFFB36003FF60FD38D3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Zeng, Lyu & Wang"baseAuthorityYear="2019"box="[538,693,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="jiulianensis">
<taxonomicNameid="4C194D3AFFADFFB360DAFF60FCEBD3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Li, Xu, Liu, Jiang, Wei & Wang"baseAuthorityYear="2018"box="[707,870,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="leishanensis">
<taxonomicNameid="4C194D3AFFADFFB3616AFF60FC3ED3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Yang"baseAuthorityYear="2014"box="[883,947,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="lini">
<taxonomicNameid="4C194D3AFFADFFB36632FF60FB6DD3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Lyu & Wang"baseAuthorityYear="2019"box="[1067,1248,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="mufumontana">
<taxonomicNameid="4C194D3AFFADFFB366F4FF60FA1BD3FA"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Wang, Liu & Wang"baseAuthorityYear="2019"box="[1261,1430,223,249]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="nanlingensis">
<taxonomicNameid="4C194D3AFFADFFB3628EFEBCFEB2D21E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Lyu, Liu & Wang"baseAuthorityYear="2020"box="[151,319,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="shimentaina">
<taxonomicNameid="4C194D3AFFADFFB36349FEBCFDAFD21E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang, Deng, Liu, Wu & Liu"baseAuthorityYear="2019"box="[336,546,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="shunhuangensis">
<taxonomicNameid="4C194D3AFFADFFB3602AFEBCFC95D21E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[563,792,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="wuliangshanensis">
<taxonomicNameid="4C194D3AFFADFFB36145FEBCFB84D21E"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[860,1033,259,285]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="wushanensis">
<taxonomicNameid="4C194D3AFFADFFB3637FFE98FE4AD242"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che"baseAuthorityYear="2019"box="[358,455,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="angka">
<taxonomicNameid="4C194D3AFFADFFB363CEFE98FDF0D242"baseAuthorityName="Inger & Romer"baseAuthorityYear="1961"box="[471,637,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="brachykolos">
<taxonomicNameid="4C194D3AFFADFFB36095FE98FCCCD242"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Nguyen, Pham, Nguyen, Luong & Ziegler"baseAuthorityYear="2020"box="[652,833,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="caobangensis">
<taxonomicNameid="4C194D3AFFADFFB36148FE98FC71D242"baseAuthorityName="Rao & Yang"baseAuthorityYear="1997"box="[849,1020,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="daweimontis">
<taxonomicNameid="4C194D3AFFADFFB36612FE98FB5DD242"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Wang & Wang"baseAuthorityYear="2019"box="[1035,1232,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="dongguanensis">
<taxonomicNameid="4C194D3AFFADFFB366F9FE98FA18D242"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley"baseAuthorityYear="2018"box="[1248,1429,295,321]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="fansipanensis">
<taxonomicNameid="4C194D3AFFADFFB3628EFEF4FF79D267"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Liu, Li, Wei, Xu, Cheng, Wang & Wu"baseAuthorityYear="2020"box="[151,244,331,356]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="jiangi">
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<taxonomicNameid="4C194D3AFFADFFB363B6FEF4FDD9D266"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Li, Xu, Liu, Jiang, Wei & Wang"baseAuthorityYear="2018"box="[431,596,331,357]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="leishanensis">
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<taxonomicNameid="4C194D3AFFADFFB36757FEF4FEBBD28A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="wuliangshanensis">
<taxonomicNameid="4C194D3AFFADFFB363A5FED0FDFAD28A"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"baseAuthorityName="Ye & Fei"baseAuthorityYear="1995"box="[444,631,367,393]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="binchuanensis">
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; head width shorter than head length, HDW/HDL 0.99; snout rounded in dorsal view, projecting, sloping posteriorly to mouth in profile, protruding well beyond margin of lower jaw; dorsal surface of head flat; eye small, ED/HDL 0.39; nostril obliquely ovoid; pupil vertical; canthus rostralis well developed, curved above nostril; loreal region sloping; internasal distance larger than interorbital distance; tympanum edge distinct and raised, upper margin in contact with supratympanic fold; choanae large ovoid, situated at base of maxilla; vomerine teeth absent; margin of tongue not notched.
</paragraph>
<paragraphid="8BA636B9FFADFFB362DEFD0CFF72D03A"blockId="17.[151,1437,151,1761]"pageId="17"pageNumber="26">Lower arm length 0.20 of SVL and hand length 0.23 of SVL; relative finger lengths II <I <IV <III; tip of finger rounded, slightly dilated; one subarticular tubercle present at base of each finger; fingers without webs, narrow lateral fringes present on fingers; outer and inner metacarpal tubercles distinct, inner metacarpal tubercle observably enlarged.</paragraph>
<paragraphid="8BA636B9FFADFFB362DEFCFCFDBFD0EE"blockId="17.[151,1437,151,1761]"pageId="17"pageNumber="26">Shank length 0.42 of SVL and foot 0.64 of SVL; tibio-tarsal articulation reaches at center of tympanum when hindlimb stretched alongside body; heels not meeting when hindlimbs held at right angles to body; relative toe lengths I <II <V <III <IV; tips of toes rounded, slightly dilated; one subarticular tubercle present at base of each toe; rudimentary web and lateral fringes present between toes; tarsal folds absent; inner metatarsal tubercle rounded, and outer metatarsal tubercle absent.</paragraph>
<paragraphid="8BA636B9FFADFFB362DEFC48FDCBD7C6"blockId="17.[151,1437,151,1761]"pageId="17"pageNumber="26">Dorsal body’s skin texture rough with densely-distributed granules and tubercles; discontinuous “X” shaped skin ridge on center of dorsum; short transverse skin ridges present on dorsal shank and thigh; a small horn-like tubercle present at edge of upper eyelid; supratympanic fold distinct, with distinct raised tubercles, curving from posterior corner of eye, posteroventrally to above insertion of arm; flank with densely-distributed tubercles. Ventral surface smooth; densely-distributed tubercles on thigh; pectoral gland relatively small, closer to axilla; single femoral gland on posterior surface of thigh.</paragraph>
Dorsal surface dark brown; scarlet granules on the dorsal head, supratympanic fold, dorsum, and flanks; a dark brown triangular marking with light edge between eyes; a dark brown “X” shaped marking with light edge on center of dorsum; dark stripes below eyes; iris red. Ventral surface grayish brown, with a large white patch and tiny scarlet spots on the belly; three dark longitudinal stripes on the throat; palms pale, soles grayish brown, tips of digits greyish white, metacarpal and metatarsal tubercles pale; pectoral glands and femoral glands white.
<tableCitationid="C69B0302FFADFFB3613CFA18FCF1D6C2"box="[805,892,1446,1473]"captionStart="TABLE 5"captionStartId="19.[151,239,151,175]"captionTargetPageId="19"captionText="TABLE 5. Measurements (in mm) of the type series of Panophrys daoji sp. nov., * for the holotype."httpUri="http://table.plazi.org/id/DF666631FFAFFFB1628EFF28FB47D3AC"pageId="17"pageNumber="26"tableUuid="DF666631FFAFFFB1628EFF28FB47D3AC">Table 5</tableCitation>
. All specimens were similar in morphology but their colorations were variable. Females are larger than males. SYS a006210 is dark gray on dorsum with “)(”- shaped skin ridge and “)(”-shaped marking on center of dorsum. SYS a006211 (
<figureCitationid="13222A3CFFADFFB36614FA50FBECD50A"box="[1037,1121,1519,1545]"captionStart="FIGURE 4"captionStartId="18.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-25@18.[151,1436,181,1842]"captionTargetPageId="18"captionText="FIGURE 4. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a006209 of Panophrys daoji sp. nov. in life. (E)–(F) dorsolateral views of adult male paratype SYS a006211 and adult female paratype SYS a006214."figureDoi="http://doi.org/10.5281/zenodo.4533858"httpUri="https://zenodo.org/record/4533858/files/figure.png"pageId="17"pageNumber="26">Fig. 4E</figureCitation>
) is pale brown dorsally with underdeveloped “)(”-shaped skin ridge on center of dorsum. SYS a006212 is yellowish brown on dorsum. SYS a006214 (
<figureCitationid="13222A3CFFADFFB3631EF988FED6D552"box="[263,347,1591,1617]"captionStart="FIGURE 4"captionStartId="18.[151,250,1866,1890]"captionTargetBox="[151,1436,181,1842]"captionTargetId="figure-25@18.[151,1436,181,1842]"captionTargetPageId="18"captionText="FIGURE 4. (A)–(D) dorsolateral view, ventral view, right hand, and right foot of adult male holotype SYS a006209 of Panophrys daoji sp. nov. in life. (E)–(F) dorsolateral views of adult male paratype SYS a006211 and adult female paratype SYS a006214."figureDoi="http://doi.org/10.5281/zenodo.4533858"httpUri="https://zenodo.org/record/4533858/files/figure.png"pageId="17"pageNumber="26">Fig. 4F</figureCitation>
) is reddish brown on dorsum with light patches on the shoulder.
<taxonomicNameid="4C194D3AFFADFFB36075F9E4FCABD576"authority="Lyu & Zeng & Wang & Liu & Huang & Li & Wang, 2021"authorityName="Lyu & Zeng & Wang & Liu & Huang & Li & Wang"authorityYear="2021"box="[620,806,1627,1653]"class="Amphibia"family="Megophryidae"genus="Panophrys"kingdom="Animalia"order="Anura"pageId="17"pageNumber="26"phylum="Chordata"rank="species"species="daoji"status="sp. nov.">
