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<document id="243AD82290ECCAD8FF99E2FEB02E9FC7" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFE9E409FEFDFF0FFBF7FCC3" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Trubatsa ganensis Merle, Pacaud, Ledon &amp; Goret, 2024, n. sp." docType="treatment" docUuid="0EBD7F4D-8646-456B-BD5C-10FA4A8625DD" docUuidSource="ZooBank" docVersion="8" lastPageNumber="508" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="508" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="E2CA7A2E61BE97FFA360B79330C85411">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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</mods:titleInfo>
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<mods:namePart id="B47DAE845587F233F508F74288A669F9">Merle, Didier</mods:namePart>
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</mods:name>
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<mods:namePart id="3253ED375A0AC3D55A763EF05AE90730">Pacaud, Jean-Michel</mods:namePart>
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</mods:name>
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<mods:namePart id="DBA407CAD94BAAE718A4B4E917DD71B2">Ledon, Daniel</mods:namePart>
<mods:affiliation id="D7AB0432E64AFA19412E7E63A402E1C8">Muséum national d’histoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="A4DA3B8D34E0BA1C9E7879EADE499C0B" type="email">didier.merle@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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<mods:namePart id="EC41384010994DA2C72CB22045AE4CEE">Goret, Bernard</mods:namePart>
<mods:affiliation id="9E46FF8BB5911360E74E1D2B52B4029C">Raoul Ponchon, F- 31500 Toulouse (France) bernardgoret @ hotmail. com</mods:affiliation>
<mods:nameIdentifier id="441E89F73D1FC690A242D794E7FB2F7D" type="email">bernardgoret@hotmail.com</mods:nameIdentifier>
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<mods:title id="7B96B2D277548D5EADBB9E3DE7957BA8">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:date id="F80A5D728DFCCB8107236BF9AF398AE9">2024</mods:date>
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<mods:url id="557448DD8468C85411EF60471A0A1915">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf</mods:url>
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<mods:classification id="CA8258EF2B8AF5A68489DCB624045537">journal article</mods:classification>
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<treatment id="038E87DAFFE9E409FEFDFF0FFBF7FCC3" ID-DOI="http://doi.org/10.5281/zenodo.13985812" ID-GBIF-Taxon="244735073" ID-Zenodo-Dep="13985812" LSID="urn:lsid:zoobank.org:act:0EBD7F4D-8646-456B-BD5C-10FA4A8625DD" httpUri="http://treatment.plazi.org/id/038E87DAFFE9E409FEFDFF0FFBF7FCC3" lastPageNumber="508" pageId="15" pageNumber="508" scope_class="Gastropoda" scope_family="Muricidae" scope_phylum="Mollusca" scope_subFamily="Typhinae">
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<subSubSection id="C33D6547FFE9E409FEFDFF0FFDC6FF41" box="[325,583,215,243]" pageId="15" pageNumber="508" type="nomenclature">
<paragraph id="8B9836CCFFE9E409FEFDFF0FFDC6FF41" blockId="15.[325,583,215,274]" box="[325,583,215,243]" pageId="15" pageNumber="508">
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<taxonomicName id="4C274D4FFFE9E409FEFDFF0FFD84FF42" ID-CoL="CN5MK" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[325,517,215,243]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="ganensis" status="sp. nov.">
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<emphasis id="B953EADEFFE9E409FEFDFF0FFD84FF42" bold="true" box="[325,517,215,243]" italics="true" pageId="15" pageNumber="508">Trubatsa ganensis</emphasis>
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<taxonomicNameLabel id="A26057A5FFE9E409FDB4FF0EFDC6FF41" box="[524,583,216,242]" pageId="15" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C33D6547FFE9E409FEE5FF21FC86FEFE" pageId="15" pageNumber="508" type="description">
<paragraph id="8B9836CCFFE9E409FEE5FF21FDAFFEA2" blockId="15.[325,583,215,274]" box="[349,558,247,274]" pageId="15" pageNumber="508">
(
<figureCitation id="131C2A49FFE9E409FEDEFF21FE27FEA1" box="[358,422,247,274]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5</figureCitation>
A-D; 25I-K)
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CFEEAFC86FEFE" blockId="15.[132,775,316,333]" box="[132,775,316,333]" pageId="15" pageNumber="508">
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urn:lsid:zoobank.org:act:
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<paragraph id="8B9836CCFFE9E409FF3CFEABFE2CFE1C" blockId="15.[132,774,381,432]" pageId="15" pageNumber="508">
<treatmentCitationGroup id="AB3711E2FFE9E409FF3CFEABFE2CFE1C" pageId="15" pageNumber="508">
<taxonomicName id="4C274D4FFFE9E409FF3CFEABFEE6FE26" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[132,359,381,405]" class="Gastropoda" family="Muricidae" genus="Trubasta" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FF3CFEABFF56FE26" box="[132,215,381,405]" italics="true" pageId="15" pageNumber="508">Trubasta</emphasis>
[
<emphasis id="B953EADEFFE9E409FF5DFEA8FF7AFE26" box="[229,251,382,405]" italics="true" pageId="15" pageNumber="508">sic</emphasis>
]
<emphasis id="B953EADEFFE9E409FEB1FEA8FEE6FE26" box="[265,359,382,405]" italics="true" pageId="15" pageNumber="508">parisiensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A8610DDFFE9E409FEC7FEABFDA4FE26" author="MERLE D." box="[383,549,381,405]" page="169" pageId="15" pageNumber="508" year="1990">
<bibRefCitation id="EFB64B3DFFE9E409FEC7FEABFDA4FE26" author="MERLE D." box="[383,549,381,405]" pageId="15" pageNumber="508" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">Merle 1990: 169</bibRefCitation>
</treatmentCitation>
, pl. 3, figs 13-15, pl. 4, fig. 12 [
<emphasis id="B953EADEFFE9E409FF69FE4EFF74FE1C" box="[209,245,408,431]" italics="true" pageId="15" pageNumber="508">non</emphasis>
d’Orbigny, 1850].
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<paragraph id="8B9836CCFFE9E409FF3CFE1EFD6EFD9C" blockId="15.[130,776,455,744]" pageId="15" pageNumber="508">
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<materialsCitation id="3B4F3C91FFE9E409FF3CFE1EFD6EFD9C" ID-GBIF-Occurrence="4958671424" collectionCode="MNHN" collectorName="de Gan &amp; Varone" country="France" county="Basin" location="Gan" municipality="Pyrenees-Atlantiques" pageId="15" pageNumber="508" specimenCount="1" stateProvince="Aquitaine" typeStatus="holotype">
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. —
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.
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•
<collectingRegion id="49E3F82EFFE9E409FDBBFE11FDE2FE6C" box="[515,611,455,479]" country="France" name="Aquitaine" pageId="15" pageNumber="508">Aquitaine</collectingRegion>
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,
<collectingMunicipality id="6BFCACB6FFE9E409FD11FE11FF77FE49" pageId="15" pageNumber="508">Pyrénées-Atlantiques</collectingMunicipality>
,
<location id="8EF86017FFE9E409FF47FE34FEA8FE49" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FF47FE34FEA8FE49" box="[255,297,482,506]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Gan" pageId="15" pageNumber="508" stateProvince="Aquitaine">Gan</location>
(
<location id="8EF86017FFE9E409FE8DFE34FE04FE49" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FE8DFE34FE04FE49" box="[309,389,482,506]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Tuilerie" pageId="15" pageNumber="508" stateProvince="Aquitaine">Tuilerie</location>
);
<location id="8EF86017FFE9E409FE2DFE34FD82FE4A" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FE2DFE34FD82FE4A" box="[405,515,482,505]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Uppermost" pageId="15" pageNumber="508" stateProvince="Aquitaine">Uppermost</location>
part of the
<location id="8EF86017FFE9E409FDCAFE34FC86FE49" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FDCAFE34FC86FE49" box="[626,775,482,506]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Marnes de Gan" pageId="15" pageNumber="508" stateProvince="Aquitaine">
Marnes
<collectorName id="26D2531AFFE9E409FD78FE34FC86FE49" box="[704,775,482,506]" pageId="15" pageNumber="508">de Gan</collectorName>
</location>
only (top biozone N12/base biozone NP13);
<location id="8EF86017FFE9E409FD9BFE2AFD1DFDA7" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FD9BFE2AFD1DFDA7" box="[547,668,508,532]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Early Eocene" pageId="15" pageNumber="508" stateProvince="Aquitaine">Early Eocene</location>
(
<location id="8EF86017FFE9E409FD1EFE2AFD7CFDA7" LSID="urn:lsid:plazi:treatment:038E87DAFFE9E409FEFDFF0FFBF7FCC3:8EF86017FFE9E409FD1EFE2AFD7CFDA7" box="[678,765,508,532]" country="France" county="Basin" municipality="Pyrenees-Atlantiques" name="Ypresian" pageId="15" pageNumber="508" stateProvince="Aquitaine">Ypresian</location>
);
<collectionCode id="ED36AE09FFE9E409FF3AFDC1FF56FD9D" box="[130,215,535,558]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A90520 (
<collectorName id="26D2531AFFE9E409FEF2FDC1FE10FD9C" box="[330,401,535,559]" pageId="15" pageNumber="508">Varone</collectorName>
coll.), H:
<quantity id="4CDF9B29FFE9E409FE4FFDC1FDBFFD9D" box="[503,574,535,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="508" unit="mm" value="17.0">17 mm</quantity>
(
<figureCitation id="131C2A49FFE9E409FDF5FDC0FD35FD9D" box="[589,692,534,559]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5A, B</figureCitation>
;
<figureCitation id="131C2A49FFE9E409FD07FDC0FD63FD9C" box="[703,738,534,559]" captionStart="FIG" captionStartId="44.[132,143,1530,1547]" captionTargetBox="[132,1455,215,1484]" captionTargetId="figure-374@44.[700,1257,764,1304]" captionTargetPageId="44" captionText="FIG. 25. — New species of Muricopsis Bucquoy &amp; Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13985823" httpUri="https://zenodo.org/record/13985823/files/figure.png" pageId="15" pageNumber="508">25I</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CFDE7FE90FD5B" blockId="15.[130,776,455,744]" pageId="15" pageNumber="508">
Add updates up until 2024-11-29 04:23:56
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<materialsCitation id="3B4F3C91FFE9E409FF3CFDE7FD34FDD7" ID-GBIF-Occurrence="4958671348" pageId="15" pageNumber="508">
Add updates up until 2024-10-24 21:38:35
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<emphasis id="B953EADEFFE9E409FF3CFDE7FEA1FDFA" bold="true" box="[132,288,561,585]" pageId="15" pageNumber="508">
<typeStatus id="549C886EFFE9E409FF3CFDE7FF58FDFA" box="[132,217,561,585]" pageId="15" pageNumber="508" type="paratype">Paratype</typeStatus>
.
<collectingCountry id="F330765CFFE9E409FF59FDE7FEA1FDFA" box="[225,288,561,585]" name="France" pageId="15" pageNumber="508">France</collectingCountry>
</emphasis>
Add updates up until 2024-11-29 04:23:56
2024-11-29 05:29:04 +01:00
• 1 spm; same as for the holotype;
Add updates up until 2024-10-24 21:38:35
2024-10-24 23:44:41 +02:00
<collectionCode id="ED36AE09FFE9E409FDEBFDE7FD25FDFB" box="[595,676,561,584]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.R61441 (
<taxonomicName id="4C274D4FFFE9E409FF33FD9DFF44FDD0" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[139,197,587,611]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll., figured specimen of
<bibRefCitation id="EFB64B3DFFE9E409FE01FD9DFDAEFDD7" author="MERLE D." box="[441,559,587,612]" pageId="15" pageNumber="508" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFE9E409FE01FD9DFE70FDD0" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[441,497,587,611]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1990
</bibRefCitation>
: pl. 3, fig. 13)
</materialsCitation>
Add updates up until 2024-11-29 04:23:56
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<materialsCitation id="3B4F3C91FFE9E409FD01FD9AFEB2FD2A" ID-GBIF-Occurrence="4958671397" pageId="15" pageNumber="508">
Add updates up until 2024-10-24 21:38:35
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• 1 spm; idem;
<collectionCode id="ED36AE09FFE9E409FF7AFDB0FE96FDCE" box="[194,279,614,637]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A90518 (
<taxonomicName id="4C274D4FFFE9E409FE30FDB0FE42FDCD" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[392,451,614,638]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll., with complete siphonal canal,
<figureCitation id="131C2A49FFE9E409FF08FD56FF7CFD2B" box="[176,253,640,664]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5C</figureCitation>
;
<figureCitation id="131C2A49FFE9E409FEB1FD56FEACFD2A" box="[265,301,640,665]" captionStart="FIG" captionStartId="44.[132,143,1530,1547]" captionTargetBox="[132,1455,215,1484]" captionTargetId="figure-374@44.[700,1257,764,1304]" captionTargetPageId="44" captionText="FIG. 25. — New species of Muricopsis Bucquoy &amp; Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13985823" httpUri="https://zenodo.org/record/13985823/files/figure.png" pageId="15" pageNumber="508">25J</figureCitation>
)
</materialsCitation>
Add updates up until 2024-11-29 04:23:56
2024-11-29 05:29:04 +01:00
<materialsCitation id="3B4F3C91FFE9E409FE83FD57FDDDFD00" ID-GBIF-Occurrence="4958671413" pageId="15" pageNumber="508">
Add updates up until 2024-10-24 21:38:35
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• 1 spm; idem;
<collectionCode id="ED36AE09FFE9E409FE6EFD57FDABFD2B" box="[470,554,641,664]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A90517 (
<taxonomicName id="4C274D4FFFE9E409FD23FD56FD57FD2B" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[667,726,640,664]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll. with incomplete siphonal canal;
<figureCitation id="131C2A49FFE9E409FE74FD4DFD9CFD01" box="[460,541,667,691]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Figs 5D</figureCitation>
;
<figureCitation id="131C2A49FFE9E409FD91FD4DFDD5FD00" box="[553,596,667,691]" captionStart="FIG" captionStartId="44.[132,143,1530,1547]" captionTargetBox="[132,1455,215,1484]" captionTargetId="figure-374@44.[700,1257,764,1304]" captionTargetPageId="44" captionText="FIG. 25. — New species of Muricopsis Bucquoy &amp; Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13985823" httpUri="https://zenodo.org/record/13985823/files/figure.png" pageId="15" pageNumber="508">25K</figureCitation>
)
</materialsCitation>
Add updates up until 2024-11-29 04:23:56
2024-11-29 05:29:04 +01:00
<materialsCitation id="3B4F3C91FFE9E409FDDCFD4DFE32FD7E" ID-GBIF-Occurrence="4958671336" pageId="15" pageNumber="508">
Add updates up until 2024-10-24 21:38:35
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• 12 spm; idem;
<collectionCode id="ED36AE09FFE9E409FF3EFD60FF59FD7E" box="[134,216,694,717]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A90519 (
<taxonomicName id="4C274D4FFFE9E409FEFDFD63FEFEFD7E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[325,383,693,717]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
coll.)
</materialsCitation>
Add updates up until 2024-11-29 04:23:56
2024-11-29 05:29:04 +01:00
<materialsCitation id="3B4F3C91FFE9E409FE0FFD60FE90FD5B" ID-GBIF-Occurrence="4958671405" pageId="15" pageNumber="508">
Add updates up until 2024-10-24 21:38:35
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• 17 spm; idem;
<collectionCode id="ED36AE09FFE9E409FDF7FD60FD23FD7E" box="[591,674,694,717]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="15" pageNumber="508">MNHN</collectionCode>
.F.A72383 (Varone coll.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FF3CFCD6FF74FC81" pageId="15" pageNumber="508" type="description">
<paragraph id="8B9836CCFFE9E409FF3CFCD6FF74FC81" blockId="15.[132,775,767,818]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFCD6FF7FFCA5" baselines="786,786" box="[132,254,768,792]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Additional" pageId="15" pageNumber="508">ADDITIONAL</smallCapsWord>
<smallCapsWord id="8D7EA010FFE9E409FEBBFCD3FEE1FCA5" baselines="786" box="[259,352,773,790]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="material" pageId="15" pageNumber="508">MATERIAL</smallCapsWord>
. —
Add updates up until 2024-11-29 04:23:56
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<materialsCitation id="3B4F3C91FFE9E409FE31FD29FF74FC81" ID-GBIF-Occurrence="4958671352" pageId="15" pageNumber="508">4 spm; idem;MNHN.F.A90521 (Combault coll.).</materialsCitation>
Add updates up until 2024-10-24 21:38:35
2024-10-24 23:44:41 +02:00
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FF3CFC9CFC89FCD2" box="[132,776,841,866]" pageId="15" pageNumber="508" type="etymology">
<paragraph id="8B9836CCFFE9E409FF3CFC9CFC89FCD2" blockId="15.[132,776,841,866]" box="[132,776,841,866]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFC9CFF7DFCD3" baselines="859,860" box="[132,252,842,865]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="15" pageNumber="508">ETYMOLOGY</smallCapsWord>
. — From the
<typeStatus id="549C886EFFE9E409FE38FC9CFE28FCD1" box="[384,425,842,866]" pageId="15" pageNumber="508">type</typeStatus>
locality, Gan (Pyrénées-Atlantiques).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FF3CFCACFD44FBA3" pageId="15" pageNumber="508" type="materials_examined">
<paragraph id="8B9836CCFFE9E409FF3CFCACFE3DFC75" blockId="15.[132,775,889,966]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFCACFF37FC3C" baselines="907,907" box="[132,182,890,913]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="15" pageNumber="508">TYPE</smallCapsWord>
<smallCapsWord id="8D7EA010FFE9E409FF06FCA9FE9BFC23" baselines="908" box="[190,282,895,912]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="15" pageNumber="508">HORIZON</smallCapsWord>
. — Uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13), see
<bibRefCitation id="EFB64B3DFFE9E409FD95FC45FC83FC1F" author="NOLF D. &amp; STEURBAUT E. &amp; CAHUZAC B." box="[557,770,915,940]" pageId="15" pageNumber="508" pagination="164 - 194" refId="ref47746" refString="NOLF D., STEURBAUT E. &amp; CAHUZAC B. 2002. - Progres recents dans la connaissance des gisements cenozoiques en Aquitaine meridionale (Chalosse, Bearn et Bas-Adour: SW France. Revue de Micropaleontologie 45 (3): 164 - 194. https: // doi. org / 10.1016 / S 0035 - 1598 (02) 90017 - 6" type="journal article" year="2002">
Nolf
<emphasis id="B953EADEFFE9E409FDD8FC42FD0EFC18" box="[608,655,915,939]" italics="true" pageId="15" pageNumber="508">et al.</emphasis>
(2002: 173
</bibRefCitation>
, fig. 4), Early Eocene (Ypresian).
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CFC08FD44FBA3" blockId="15.[132,775,990,1040]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFC08FF34FC40" baselines="1007,1007" box="[132,181,990,1013]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="15" pageNumber="508">TYPE</smallCapsWord>
<smallCapsWord id="8D7EA010FFE9E409FF03FC35FE97FC47" baselines="1008" box="[187,278,995,1012]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="15" pageNumber="508">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFE9E409FEF9FC08FE04FC46" box="[321,389,990,1013]" name="France" pageId="15" pageNumber="508">France</collectingCountry>
,
<collectingRegion id="49E3F82EFFE9E409FE36FC08FE6FFC45" box="[398,494,990,1014]" country="France" name="Aquitaine" pageId="15" pageNumber="508">Aquitaine</collectingRegion>
Basin, Pyrénées-Atlantiques, Gan (Tuilerie) (
<bibRefCitation id="EFB64B3DFFE9E409FEA6FC2EFE2AFBA3" author="MERLE D." box="[286,427,1016,1040]" pageId="15" pageNumber="508" refId="ref46520" refString="MERLE D. 1986. - Contribution a l'etude paleontologique du gisement cuisien de Gan (Pyrenees Atlantiques): systematique, evolution et paleoecologie. Unpublished Memoir of the Ecole pratique des Hautes Etudes (EPHE), 2 vols, Section Sciences de la Vie et de la Terre, Paris, 394 p." type="book" year="1986">
<taxonomicName id="4C274D4FFFE9E409FEA6FC2EFED6FBA3" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[286,343,1016,1040]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
1986: 8
</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFE9E409FE0EFC2EFD04FBA3" author="NOLF D. &amp; STEURBAUT E. &amp; CAHUZAC B." box="[438,645,1016,1040]" pageId="15" pageNumber="508" pagination="164 - 194" refId="ref47746" refString="NOLF D., STEURBAUT E. &amp; CAHUZAC B. 2002. - Progres recents dans la connaissance des gisements cenozoiques en Aquitaine meridionale (Chalosse, Bearn et Bas-Adour: SW France. Revue de Micropaleontologie 45 (3): 164 - 194. https: // doi. org / 10.1016 / S 0035 - 1598 (02) 90017 - 6" type="journal article" year="2002">
Nolf
<emphasis id="B953EADEFFE9E409FE52FC2FFD98FBA3" box="[490,537,1016,1040]" italics="true" pageId="15" pageNumber="508">et al.</emphasis>
2002: 185
</bibRefCitation>
, 186).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FF3CFBFEFD19FBF3" box="[132,664,1064,1088]" pageId="15" pageNumber="508" type="distribution">
<paragraph id="8B9836CCFFE9E409FF3CFBFEFD19FBF3" blockId="15.[132,664,1064,1088]" box="[132,664,1064,1088]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFBFEFE93FB8D" baselines="1081,1082" box="[132,274,1064,1087]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="15" pageNumber="508">DISTRIBUTION</smallCapsWord>
. — Only known from the
<typeStatus id="549C886EFFE9E409FDA4FBFEFDC7FBF3" box="[540,582,1064,1088]" pageId="15" pageNumber="508">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFE9E409FF3CFBA3FBF7FCC3" pageId="15" pageNumber="508" type="description">
<paragraph id="8B9836CCFFE9E409FF3CFBA3FE94FB3E" blockId="15.[130,777,1141,1741]" box="[132,277,1141,1167]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CFBA3FE94FB3E" baselines="1160,1160" box="[132,277,1141,1167]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="15" pageNumber="508">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CFB42FEDDF97F" blockId="15.[130,777,1141,1741]" pageId="15" pageNumber="508">
Bulbous protoconch of 1.5-2 whorls (see
<bibRefCitation id="EFB64B3DFFE9E409FDFBFB42FD40FB1D" author="MERLE D." box="[579,705,1172,1198]" pageId="15" pageNumber="508" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFE9E409FDFBFB42FD03FB1D" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[579,642,1172,1198]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
1990
</bibRefCitation>
: pl. 4, fig. 12). Teleoconch up to
<quantity id="4CDF9B29FFE9E409FE1AFB62FE73FB7D" box="[418,498,1204,1230]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="508" unit="mm" value="17.0">17 mm</quantity>
(canal siphonal not complete) in height, up to
<quantity id="4CDF9B29FFE9E409FEC7FB02FE4EFB5D" box="[383,463,1236,1262]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="508" unit="mm" value="11.0">11 mm</quantity>
in width, biconic in profile, composed of five whorls. Moderately high spire. Last whorl representing 72% of total height. Spiral angle 56°. Suture weakly impressed. Axial sculpture consisting in four slightly foliate varices per whorl. On spire, anal tube (P1) and very short spine (P2). Anal tube forming angle of approximately 60-65° with axis of shell. Anal tube (P1) totally integrated within varices, eliminating shoulder. Small callus at base of anal tube. On convex part of last whorl and abapically to anal tube four spinelets (P2, s2, P3, P4 and P5). Intervarical spaces usually smooth, but with P3-P5 cords on
<typeStatus id="549C886EFFE9E409FD18F9C4FC82F99F" box="[672,771,1554,1580]" pageId="15" pageNumber="508" type="holotype">holotype</typeStatus>
. Rounded aperture up to 25% of diameter, up to 74% of height of last whorl (including siphonal canal). Edge, erect, smooth, peristome complete. Siphonal canal closed, usually broken, occupying 60% of apertural lenght (complete siphonal canal longer, see
<figureCitation id="131C2A49FFE9E409FF43F964FECEF97F" box="[251,335,1714,1741]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CF924FE9EF8B9" blockId="15.[132,775,1778,2028]" box="[132,287,1778,1804]" pageId="15" pageNumber="508">
<smallCapsWord id="8D7EA010FFE9E409FF3CF924FE9EF8B9" baselines="1797,1797" box="[132,287,1778,1804]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="15" pageNumber="508">COMPARISONS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFE9E409FF3CF8C4FBF7FCC3" blockId="15.[132,775,1778,2028]" lastBlockId="15.[811,1457,215,880]" pageId="15" pageNumber="508">
<bibRefCitation id="EFB64B3DFFE9E409FF3CF8C4FECCF89F" author="MERLE D." box="[132,333,1810,1836]" pageId="15" pageNumber="508" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFE9E409FF3CF8C4FF42F89F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[132,195,1810,1836]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1990: 169)
</bibRefCitation>
attributed this material collected at Gan to
<taxonomicName id="4C274D4FFFE9E409FF1BF8E7FD90F8FF" ID-CoL="8Z9N2" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[163,529,1841,1868]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FF1BF8E7FED5F8F8" box="[163,340,1841,1867]" italics="true" pageId="15" pageNumber="508">Typhis parisiensis</emphasis>
d’Orbigny, 1850
</taxonomicName>
and to the genus
<taxonomicName id="4C274D4FFFE9E409FD61F8E4FEBBF8DF" ID-CoL="8LT6R" authority="Dall, 1889" authorityName="Dall" authorityYear="1889" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFE9E409FD61F8E4FF38F8D8" italics="true" pageId="15" pageNumber="508">Trubatsa</emphasis>
Dall, 1889
</taxonomicName>
. The generic attribution is correct and the distinction between
<taxonomicName id="4C274D4FFFE9E409FED5F8A7FD55F838" ID-CoL="7PGWL" authority="Jousseaume, 1880" authorityName="Jousseaume" authorityYear="1880" box="[365,724,1905,1932]" class="Gastropoda" family="Muricidae" genus="Siphonochelus" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFE9E409FED5F8A7FD83F838" box="[365,514,1905,1931]" italics="true" pageId="15" pageNumber="508">Siphonochelus</emphasis>
<bibRefCitation id="EFB64B3DFFE9E409FDB1F8A4FD55F838" author="JOUSSEAUME F. P." box="[521,724,1905,1932]" pageId="15" pageNumber="508" pagination="335 - 338" refId="ref45378" refString="JOUSSEAUME F. P. 1880. - Division methodique de la famille des Purpurides. Le Naturaliste 2 (42): 335 - 338. http: // biodiversitylibrary. org / page / 33928744" type="journal article" year="1880">Jousseaume, 1880</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C274D4FFFE9E409FF3EF847FF63F818" ID-CoL="8LT6R" authorityName="Dall" authorityYear="1889" box="[134,226,1937,1963]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFE9E409FF3EF847FF63F818" box="[134,226,1937,1963]" italics="true" pageId="15" pageNumber="508">Trubatsa</emphasis>
</taxonomicName>
is now validated by molecular data (
<bibRefCitation id="EFB64B3DFFE9E409FDC1F844FF3CF878" author="HOUART R. &amp; BUGE B. &amp; ZUCCON D." pageId="15" pageNumber="508" pagination="103 - 147" refId="ref45035" refString="HOUART R., BUGE B. &amp; ZUCCON D. 2021. - A taxonomic update of the Typhinae (Gastropoda: Muricidae) with a review of New Caledonia species and the description of new species from New Caledonia, the South China sea and Western Australia. Journal of Conchology 44 (2): 103 - 147." type="journal article" year="2021">
Houart
<emphasis id="B953EADEFFE9E409FD6AF844FC86F818" box="[722,775,1937,1963]" italics="true" pageId="15" pageNumber="508">et al.</emphasis>
2021
</bibRefCitation>
). Nevertheless, after further comparison, specific attribution of the Gan material to this species incorrect. Firstly, the protoconchs are different, that of
<taxonomicName id="4C274D4FFFE9E409FB0DFF0EFAA2FF42" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1205,1315,216,241]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="ganensis" status="n. sp.">
<emphasis id="B953EADEFFE9E409FB0DFF0EFAA2FF42" box="[1205,1315,216,241]" italics="true" pageId="15" pageNumber="508">T. ganensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFE9E409FA92FF0EFAE5FF41" box="[1322,1380,216,242]" pageId="15" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
is paucispiral (
<bibRefCitation id="EFB64B3DFFE9E409FC3FFF21FB85FEA2" author="MERLE D." box="[903,1028,247,273]" pageId="15" pageNumber="508" pagination="145 - 185" refId="ref46577" refString="MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185." type="journal article" year="1990">
<taxonomicName id="4C274D4FFFE9E409FC3FFF21FC47FEA2" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[903,966,247,273]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
1990
</bibRefCitation>
: pl. 4, fig. 12) and that of
<taxonomicName id="4C274D4FFFE9E409FAADFF2EFA19FEA2" ID-CoL="8Z9N2" authorityName="d'Orbigny" authorityYear="1850" box="[1301,1432,248,273]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FAADFF2EFA19FEA2" box="[1301,1432,248,273]" italics="true" pageId="15" pageNumber="508">T. parisiensis</emphasis>
</taxonomicName>
is multispiral (see juvenile specimen from Barton illustrated by A.
<bibRefCitation id="EFB64B3DFFE9E409FCF2FEE1FC64FEE2" author="MORTON A." box="[842,997,311,337]" pageId="15" pageNumber="508" refId="ref47587" refString="MORTON A. 2023. -- A Collection of Eocene and Oligocene Fossils. Available at: www. dmap. co. uk / fossils / (last access on november 2023)." type="journal volume" year="2023">Morton (2023</bibRefCitation>
; www.dmap.co.uk/fossils). In addition, the Bartonian specimens are more sculptured than the specimens from Gan (
<figureCitation id="131C2A49FFE9E409FC12FEA1FB9BFE22" box="[938,1050,374,401]" captionStart="FIG" captionStartId="14.[132,143,1183,1200]" captionTargetBox="[132,1455,215,1138]" captionTargetId="figure-499@14.[698,1267,213,769]" captionTargetPageId="14" captionText="FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton." figureDoi="http://doi.org/10.5281/zenodo.13985770" httpUri="https://zenodo.org/record/13985770/files/figure.png" pageId="15" pageNumber="508">Fig. 5E, F</figureCitation>
). They display stronger spines (P2 to P5) and P6, spiral cords (P2 to P5) in the intervarical spaces and a partition recalling that of
<taxonomicName id="4C274D4FFFE9E409FB2CFE61FA8DFE63" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1172,1292,439,464]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="gaasensis">
<emphasis id="B953EADEFFE9E409FB2CFE61FA8DFE63" box="[1172,1292,439,464]" italics="true" pageId="15" pageNumber="508">T. gaasensis</emphasis>
</taxonomicName>
(Tournouër in Benoist, 1880) from the Rupelian of Aquitaine Basin.
<taxonomicName id="4C274D4FFFE9E409FA39FE01FC51FDA3" ID-CoL="8YPH6" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FA39FE01FC51FDA3" italics="true" pageId="15" pageNumber="508">Trubatsa parisiensis</emphasis>
</taxonomicName>
occur in the Bartonian from
<collectingCountry id="F330765CFFE9E409FAB2FE20FAE4FDA3" box="[1290,1381,502,528]" name="United Kingdom" pageId="15" pageNumber="508">England</collectingCountry>
and in the Priabonian from
<collectingCountry id="F330765CFFE9E409FBAEFDC0FBFDFD83" box="[1046,1148,534,560]" name="Germany" pageId="15" pageNumber="508">Germany</collectingCountry>
(
<bibRefCitation id="EFB64B3DFFE9E409FB37FDC1FAD9FD83" author="VON KOENEN A." box="[1167,1368,534,561]" pageId="15" pageNumber="508" pagination="1 - 280" refId="ref49077" refString="VON KOENEN A. 1889. - Das Norddeutsche Unter-Oligocan und seine Mollusken-Fauna. Abhandlungen zur geologischen Specialkarte von Preussen und den Thuringischen Staaten 10 (1): 1 - 280." type="journal article" year="1889">von Koenen 1889</bibRefCitation>
).
<bibRefCitation id="EFB64B3DFFE9E409FAD7FDC0FCF7FDE2" author="MERLE D." pageId="15" pageNumber="508" pagination="257 - 262" refId="ref46465" refString="MERLE D. 1985. - Les communautes fossiles en tant qu'indicateurs paleobiologiques de milieux: exemples pris sur le Cuisien de Gan (Pyrenees-Atlantiques) et l'Egerien de Hongrie. Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser. 7, sect. C, 24: 257 - 262." type="journal article" year="1985">
<taxonomicName id="4C274D4FFFE9E409FAD7FDC0FA2EFD83" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1391,1455,534,560]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1985)
</bibRefCitation>
also suggested that
<taxonomicName id="4C274D4FFFE9E409FBEAFDE0FA2EFDE3" authority="Cossmann, 1902" authorityName="Cossmann" authorityYear="1902" box="[1106,1455,566,592]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="sinuosus">
<emphasis id="B953EADEFFE9E409FBEAFDE0FB6EFDE3" box="[1106,1263,566,592]" italics="true" pageId="15" pageNumber="508">Typhis sinuosus</emphasis>
<bibRefCitation id="EFB64B3DFFE9E409FB4EFDE0FA2EFDE3" author="COSSMANN M." box="[1270,1455,566,592]" pageId="15" pageNumber="508" pagination="55 - 209" refId="ref43634" refString="COSSMANN M. 1902. - Mollusques eoceniques de la Loire-inferieure - Fascicule 1. Bulletin de la Societe des Sciences naturelles de l'Ouest de la France 2 (2): 55 - 209." type="journal article" year="1902">Cossmann, 1902</bibRefCitation>
</taxonomicName>
from the Bartonian of Bois-Gouët (
<collectingCountry id="F330765CFFE9E409FB19FD80FB6CFDC3" box="[1185,1261,598,624]" name="France" pageId="15" pageNumber="508">France</collectingCountry>
, Loire-Atlantique) was a junior subjective synonym of
<taxonomicName id="4C274D4FFFE9E409FB16FDA1FAB3FD23" ID-CoL="8YPH6" baseAuthorityName="d'Orbigny" baseAuthorityYear="1850" box="[1198,1330,631,656]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="parisiensis">
<emphasis id="B953EADEFFE9E409FB16FDA1FAB3FD23" box="[1198,1330,631,656]" italics="true" pageId="15" pageNumber="508">T. parisiensis</emphasis>
</taxonomicName>
. According to
<bibRefCitation id="EFB64B3DFFE9E409FCF0FD40FC54FD03" author="MERLE D." box="[840,981,661,688]" pageId="15" pageNumber="508" pagination="257 - 262" refId="ref46465" refString="MERLE D. 1985. - Les communautes fossiles en tant qu'indicateurs paleobiologiques de milieux: exemples pris sur le Cuisien de Gan (Pyrenees-Atlantiques) et l'Egerien de Hongrie. Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser. 7, sect. C, 24: 257 - 262." type="journal article" year="1985">
<taxonomicName id="4C274D4FFFE9E409FCF0FD40FC06FD03" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[840,903,662,688]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
(1985)
</bibRefCitation>
, the shape of the varices of the
<typeStatus id="549C886EFFE9E409FAA0FD40FAC7FD03" box="[1304,1350,662,688]" pageId="15" pageNumber="508">type</typeStatus>
specimen (Dumas coll.) of
<taxonomicName id="4C274D4FFFE9E409FC65FD60FBCAFD7C" authorityName="Cossmann" authorityYear="1902" box="[989,1099,694,719]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="sinuosus">
<emphasis id="B953EADEFFE9E409FC65FD60FBCAFD7C" box="[989,1099,694,719]" italics="true" pageId="15" pageNumber="508">T. sinuosus</emphasis>
</taxonomicName>
recalls that of
<taxonomicName id="4C274D4FFFE9E409FB58FD63FABAFD7C" ID-CoL="8LT6R" authorityName="Dall" authorityYear="1889" box="[1248,1339,693,719]" class="Gastropoda" family="Muricidae" genus="Trubatsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFE9E409FB58FD63FABAFD7C" box="[1248,1339,693,719]" italics="true" pageId="15" pageNumber="508">Trubatsa</emphasis>
</taxonomicName>
. However, it is a juvenile (H:
<quantity id="4CDF9B29FFE9E409FC4DFD00FBB6FD43" box="[1013,1079,726,752]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="508" unit="mm" value="8.0">8 mm</quantity>
) and the varices of the juveniles of
<taxonomicName id="4C274D4FFFE9E409FC95FD23FB09FCBC" ID-CoL="8YPJH" authority="Cossmann, 1882" authorityName="Cossmann" authorityYear="1882" box="[813,1160,757,783]" class="Gastropoda" family="Muricidae" genus="Typhina" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="rutoti">
<emphasis id="B953EADEFFE9E409FC95FD23FC47FCBC" box="[813,966,757,783]" italics="true" pageId="15" pageNumber="508">Typhina rutoti</emphasis>
Cossmann, 1882
</taxonomicName>
are also similar. Moreover, the
<typeStatus id="549C886EFFE9E409FCEFFCC0FC04FC83" box="[855,901,790,816]" pageId="15" pageNumber="508">type</typeStatus>
specimen of
<taxonomicName id="4C274D4FFFE9E409FBABFCC0FB03FC9C" authorityName="Cossmann" authorityYear="1902" box="[1043,1154,790,815]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="sinuosus">
<emphasis id="B953EADEFFE9E409FBABFCC0FB03FC9C" box="[1043,1154,790,815]" italics="true" pageId="15" pageNumber="508">T. sinuosus</emphasis>
</taxonomicName>
is worn, lacks a protoconch and finally lacks diagnostic characters. Thus, we consider
<taxonomicName id="4C274D4FFFE9E409FC97FC80FC1FFCDC" authorityName="Cossmann" authorityYear="1902" box="[815,926,854,879]" class="Gastropoda" family="Muricidae" genus="Typhis" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="508" phylum="Mollusca" rank="species" species="sinuosus">
<emphasis id="B953EADEFFE9E409FC97FC80FC1FFCDC" box="[815,926,854,879]" italics="true" pageId="15" pageNumber="508">T. sinuosus</emphasis>
</taxonomicName>
as a
<emphasis id="B953EADEFFE9E409FC69FC80FBEEFCDC" box="[977,1135,853,879]" italics="true" pageId="15" pageNumber="508">nomen dubium</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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