treatments-xml/data/03/6D/87/036D87E7D532FFCDFF6CB7E4FDDBFEA3.xml

<document id="4EE8C4A9C06169BC3F9F35D5AC8D381D" ID-DOI="10.5281/zenodo.181238" ID-GBIF-Dataset="c9731fe4-5bb9-4792-8c6a-929a0bb00e82" ID-ISSN="1175-5326" ID-Zenodo-Dep="181238" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459829043349" checkinUser="plazi" docAuthor="Nieto, Carolina &amp; Richard, Barton" docDate="2008" docId="036D87E7D532FFCDFF6CB7E4FDDBFEA3" docLanguage="en" docName="zt01727p021.pdf" docOrigin="Zootaxa 1727" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Cloeodes espinillo Nieto &amp; Richard, 2008, n. sp." docType="treatment" docVersion="10" lastPageNumber="11" masterDocId="FF54FF9FD535FFC7FFFBB32BFF8DFFD9" masterDocTitle="The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="8" updateTime="1698222712371" updateUser="plazi">
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<mods:title id="A4FAE39B2965303AB35D6D8DBDCDD015">The genus Cloeodes (Ephemeroptera: Baetidae) in Argentina with new generic synonymy and new species</mods:title>
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<mods:namePart id="B04096CBD9477EFF489B425CBA74530B">Nieto, Carolina</mods:namePart>
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<heading id="D033819DD532FFC0FF6CB7E4FE36FB30" bold="true" box="[151,443,1231,1257]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B9B0EAE3D532FFC0FF6CB7E4FE36FB30" bold="true" box="[151,443,1231,1257]" pageId="7" pageNumber="8">
<taxonomicName id="4CC44D72D532FFC0FF6CB7E4FEFFFB30" ID-CoL="5ZC5V" box="[151,370,1231,1257]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="espinillo" status="sp. nov.">
<emphasis id="B9B0EAE3D532FFC0FF6CB7E4FEFFFB30" bold="true" box="[151,370,1231,1257]" italics="true" pageId="7" pageNumber="8">Cloeodes espinillo</emphasis>
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(
<figureCitation id="13FF2A74D532FFC0FF64B7D9FEA1FAD5" box="[159,300,1266,1292]" captionStart-0="FIGURES 15 – 19" captionStart-1="FIGURES 20 – 30" captionStartId-0="8.[151,269,1557,1581]" captionStartId-1="9.[151,269,1933,1957]" captionTargetBox-0="[159,1416,652,1556]" captionTargetBox-1="[185,1375,198,1932]" captionTargetId-0="figure@8.[151,1436,652,1601]" captionTargetId-1="figure@9.[184,1390,170,1935]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" captionText-1="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri-0="https://zenodo.org/record/181241/files/figure.png" httpUri-1="https://zenodo.org/record/181242/files/figure.png" pageId="7" pageNumber="8">Figs. 15–30</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3DE657AD532FFCFFF6CB669FB4CF944" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8B7B36F1D532FFC0FF6CB669FC95F9AD" blockId="7.[151,1437,1346,2012]" pageId="7" pageNumber="8">
Male imago (
<figureCitation id="13FF2A74D532FFC0FEC8B669FE04FA85" box="[307,393,1346,1372]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="7" pageNumber="8">Fig. 15</figureCitation>
). Length: body, 
<quantity id="4C3C9B14D532FFC0FDB3B669FD54FA85" box="[584,729,1346,1372]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="6.5" metricValueMin="6.4" pageId="7" pageNumber="8" unit="mm" value="6.45" valueMax="6.5" valueMin="6.4">6.4–6.5 mm</quantity>
. Fore wings, 5.9–6.0; hind wings 
<quantity id="4C3C9B14D532FFC0FB98B669FB79FA85" box="[1123,1268,1346,1372]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="7" pageNumber="8" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
. Head yellowish brown, compound eyes pale yellow, height of stalk half length of eye diameter. Antennae pale yellow. Thorax yellowish brown, sterna yellowish brown. Legs pale yellow. Leg I: tibia 1.5 times the length of femur, tarsi as long as tibia with 4 segments decreasing in length apically. Fore wings (
<figureCitation id="13FF2A74D532FFC0FBC1B691FB02FA0D" box="[1082,1167,1466,1492]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="7" pageNumber="8">Fig. 16</figureCitation>
) hyaline, except costal and subcostal spaces translucent. 
<collectingCountry id="F3D37661D532FFC0FDE6B6C9FDD5FA25" box="[541,600,1506,1532]" name="India" pageId="7" pageNumber="8">Hind</collectingCountry>
wings (
<figureCitation id="13FF2A74D532FFC0FD4FB6C9FC9EFA25" box="[692,787,1506,1532]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="7" pageNumber="8">Figs. 17</figureCitation>
a and b) present, 0.2 times the length of fore wing; costal projection pointed and located in the basal third of anterior margin. Posterior margin of the metanotum strongly projected dorsoventrally (
<figureCitation id="13FF2A74D532FFC0FDCAB519FD07F995" box="[561,650,1586,1612]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="7" pageNumber="8">Fig. 18</figureCitation>
). Abdomen pale yellow. Genitalia (
<figureCitation id="13FF2A74D532FFC0FBCEB519FB01F995" box="[1077,1164,1586,1612]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="7" pageNumber="8">Fig. 19</figureCitation>
) with bases of forceps close together, segment III rounded. Cerci pale yellow.
</paragraph>
<paragraph id="8B7B36F1D532FFC0FF3DB5A9FC69F91D" blockId="7.[151,1437,1346,2012]" pageId="7" pageNumber="8">
Female imago. Length: body, 
<quantity id="4C3C9B14D532FFC0FDC9B5A9FD4BF945" box="[562,710,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="7" pageNumber="8" unit="mm" value="6.65" valueMax="6.7" valueMin="6.6">6.6–6.7 mm</quantity>
. Fore wings, 
<quantity id="4C3C9B14D532FFC0FC97B5A9FC40F945" box="[876,973,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="7" pageNumber="8" unit="mm" value="7.2">7.2 mm</quantity>
; hind wings, 
<quantity id="4C3C9B14D532FFC0FB89B5A9FB5EF945" box="[1138,1235,1666,1692]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="8" unit="mm" value="1.2">1.2 mm</quantity>
. Similar to male imago except thorax yellow. Abdomen yellow with tracheae pigmented.
</paragraph>
<paragraph id="8B7B36F1D532FFCFFF3DB5F9FECCFF03" blockId="7.[151,1437,1346,2012]" lastBlockId="8.[151,1437,152,618]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Nymph (
<figureCitation id="13FF2A74D532FFC0FED5B5F9FE08F935" box="[302,389,1746,1772]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="7" pageNumber="8">Fig. 20</figureCitation>
). Length: body, 5.5–6.0 mm; cerci, 2.8–3.0 mm; terminal filament, 
<quantity id="4C3C9B14D532FFC0FB6CB5F9FB7EF935" box="[1175,1267,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="8" unit="mm" value="2.5">2.5 mm</quantity>
. Head yellowish brown, longer than wide; compound eyes of the male yellowish brown. Antennae pale yellow, 3 times the head capsule; pedicel 1.5 times the length of scape. Mouthpart: labrum (
<figureCitation id="13FF2A74D532FFC0FC11B409FBB2F8E5" box="[1002,1087,1826,1852]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="7" pageNumber="8">Fig. 21</figureCitation>
) with one subapical seta centrally and two setae near lateral margin, anterior margin with basally bifid setae near midline and apically bifid setae near lateral margin. Left mandible (
<figureCitation id="13FF2A74D532FFC0FD39B459FC94F855" box="[706,793,1906,1932]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="7" pageNumber="8">Fig. 22</figureCitation>
) without setae between prostheca and mola, thumb of molar area transverse to anterior margin. Right mandible without setae between prostheca and mola (similar to 
<figureCitation id="13FF2A74D532FFC0FF4DB4E9FE82F805" box="[182,271,1986,2012]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="7" pageNumber="8">Fig. 58</figureCitation>
). Hypopharynx (
<figureCitation id="13FF2A74D532FFC0FE23B4E9FDBCF805" box="[472,561,1986,2012]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="7" pageNumber="8">Fig. 23</figureCitation>
): lingua with a rounded projection. Maxillae (similar to 
<figureCitation id="13FF2A74D532FFC0FB29B4E9FAA7F805" box="[1234,1322,1986,2012]" captionStart="FIGURES 55 – 67" captionStartId="15.[151,269,1707,1731]" captionTargetBox="[285,1294,214,1652]" captionTargetId="figure@15.[254,1351,194,1683]" captionTargetPageId="15" captionText="FIGURES 55 – 67. Cloeodes penai. Nymph. 55, antennae. Mouthparts (Figs. 56 – 61): 56, labrum, left d. v., right v. v.; 57, left mandible v. v.; 58, right mandible v. v.; 59, hypopharynx v. v.; 60, maxilla v. v.; 61, labium, left d. v., right v. v. 62, leg I; 63, femur I apical margin; 64, tarsal claw I. 65, posterior margin of tergum IV. 66, gill IV. 67, paraproct." httpUri="https://zenodo.org/record/181245/files/figure.png" pageId="7" pageNumber="8">Fig. 60</figureCitation>
), palpi as long as galea-lacinia, two segmented, segment II with a constriction. Labium (
<figureCitation id="13FF2A74D53DFFCFFBB2B3B3FB2CFF6B" box="[1097,1185,152,178]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 24</figureCitation>
) with segment III of palpi rounded.
</paragraph>
<paragraph id="8B7B36F1D53DFFCFFF3DB3C3FEBDFEA3" blockId="8.[151,1437,152,618]" pageId="8" pageNumber="9">
Thorax yellowish brown (
<figureCitation id="13FF2A74D53DFFCFFE0DB3C3FDC3FEDB" box="[502,590,232,258]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 20</figureCitation>
). Fore wing pads pale yellow. Sterna pale yellow. Legs (
<figureCitation id="13FF2A74D53DFFCFFB1AB3C3FABBFEDB" box="[1249,1334,232,258]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 25</figureCitation>
) yellow. Dorsal edge of femora with a row of pointed spines, apically with a subquadrangular projection with a pair of blunt spines (
<figureCitation id="13FF2A74D53DFFCFFECDB213FE02FE8B" box="[310,399,312,338]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 26</figureCitation>
). Tarsi with a row of short spines. Tarsal claws 0.4 times the length of tarsi. 
<collectingCountry id="F3D37661D53DFFCFFAE5B213FAD4FE8B" box="[1310,1369,312,338]" name="India" pageId="8" pageNumber="9">Hind</collectingCountry>
wing pads present.
</paragraph>
<paragraph id="8B7B36F1D53DFFCFFF3DB2A3FB4CF944" blockId="8.[151,1437,152,618]" lastBlockId="8.[151,1436,1666,2013]" pageId="8" pageNumber="9">
Abdomen yellowish brown, segments II–III and VI with brown spots as in 
<figureCitation id="13FF2A74D53DFFCFFBCEB2A3FB06FE7B" box="[1077,1163,392,418]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 20</figureCitation>
, segment IX–X brown, posterior margin of terga with spines (
<figureCitation id="13FF2A74D53DFFCFFDA4B29BFD34FE13" box="[607,697,432,458]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 27</figureCitation>
). Sterna pale yellow, segments II–VI with a small tuft of long fine setae laterally on each side (
<figureCitation id="13FF2A74D53DFFCFFDE8B2F3FDE6FE2B" box="[531,619,472,498]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 28</figureCitation>
). Paraprocts with spines apically (similar to 
<figureCitation id="13FF2A74D53DFFCFFB8AB2F3FB4AFE2B" box="[1137,1223,472,498]" captionStart="FIGURES 43 – 51" captionStartId="13.[151,269,1887,1911]" captionTargetBox="[219,1274,190,1838]" captionTargetId="figure@13.[151,1436,162,1867]" captionTargetPageId="13" captionText="FIGURES 43 – 51. Cloeodes opacus n. sp. Nymph. 43, general view, dorsal. Mouthparts (Figs. 44 – 46): 44, labrum, left d. v., right v. v.; 45, labrum: anterior margin; 46, labium, left d. v., right v. v. 47, leg I; 48, dorsal margin of femur I. 49, posterior margin of tergum IV. 50, gill IV. 51, paraproct. Figs. 52 – 54. Cloeodes penai. Male imago: 52, fore wing; 53 a, hind wing; 53 b, hind wing detail; 54, genitalia v. v." httpUri="https://zenodo.org/record/181244/files/figure.png" pageId="8" pageNumber="9">Fig. 51</figureCitation>
). Gills translucent white, 2 times the length of each tergum, main trachea pigmented (
<figureCitation id="13FF2A74D53DFFCFFC4CB12BFB9DFDC3" box="[951,1040,512,538]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 29</figureCitation>
). Caudal filaments yellow, apical third yellowish brown, with flattened setae basally sclerotized (
<figureCitation id="13FF2A74D53DFFCFFC65B103FC74FD9B" box="[926,1017,552,578]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 30</figureCitation>
). Cerci (
<figureCitation id="13FF2A74D53DFFCFFB9DB103FB4DFD9B" box="[1126,1216,552,578]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 30</figureCitation>
) with long spines toward external margin every two segments, terminal filament with long spines toward midline dorsally and 
<emphasis id="B9B0EAE3D53DFFCFFF3DB5A9FEDEF945" bold="true" box="[198,339,1666,1692]" pageId="8" pageNumber="9">Etymology.</emphasis>
<taxonomicName id="4CC44D72D53DFFCFFEA1B5A8FE49F944" box="[346,452,1667,1693]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="espinillo">Espinillo</taxonomicName>
is the name of one of the rivers where this species was collected.
</paragraph>
</subSubSection>
<caption id="DFBB6679D53DFFCFFF6CB53EFA11F9F5" box="[151,1436,1557,1581]" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="8" pageNumber="9" targetBox="[159,1416,652,1556]" targetPageId="8">
<paragraph id="8B7B36F1D53DFFCFFF6CB53EFA11F9F5" blockId="8.[151,1436,1557,1581]" box="[151,1436,1557,1581]" pageId="8" pageNumber="9">
<emphasis id="B9B0EAE3D53DFFCFFF6CB53EFED2F9F4" bold="true" box="[151,351,1557,1581]" pageId="8" pageNumber="9">FIGURES 15–19.</emphasis>
<taxonomicName id="4CC44D72D53DFFCFFE93B53DFDBDF9F4" box="[360,560,1558,1581]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="espinillo" status="sp. nov.">
<emphasis id="B9B0EAE3D53DFFCFFE93B53DFDBDF9F4" box="[360,560,1558,1581]" italics="true" pageId="8" pageNumber="9">Cloeodes espinillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2835798D53DFFCFFDC2B53EFDFAF9F4" box="[569,631,1557,1581]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9B0EAE3D53DFFCFFDC2B53EFDFAF9F4" bold="true" box="[569,631,1557,1581]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
Male imago: 15, head and thorax, dorsal; 16, fore wing; 17a, hind wing;
</paragraph>
</caption>
<subSubSection id="C3DE657AD53DFFCDFF3DB581FDE6FF03" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8B7B36F1D53DFFCDFF3DB581FDE6FF03" blockId="8.[151,1436,1666,2013]" lastBlockId="10.[151,1436,152,378]" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<emphasis id="B9B0EAE3D53DFFCFFF3DB581FECBF91D" bold="true" box="[198,326,1706,1732]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="4CC44D72D53DFFCFFEB5B580FDA4F91D" box="[334,553,1707,1732]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="espinillo" status="sp. nov.">
<emphasis id="B9B0EAE3D53DFFCFFEB5B580FDA4F91D" box="[334,553,1707,1732]" italics="true" pageId="8" pageNumber="9">Cloeodes espinillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2835798D53DFFCFFDCAB581FDF8F91D" box="[561,629,1706,1732]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B9B0EAE3D53DFFCFFDCAB581FDF8F91D" bold="true" box="[561,629,1706,1732]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished from the other species of the genus by the following combination of characters. In the male imago, 1) hind wings 0.2 times the length of the fore wings (
<figureCitation id="13FF2A74D53DFFCFFF64B5D0FEA0F8CC" box="[159,301,1787,1813]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="8" pageNumber="9">Figs. 16, 17</figureCitation>
a); 2) hind wings (
<figureCitation id="13FF2A74D53DFFCFFDF9B5D0FDDBF8CC" box="[514,598,1787,1813]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
b) with costal projection located in the first third of the anterior margin; 3) genitalia (
<figureCitation id="13FF2A74D53DFFCFFED4B408FE08F8E4" box="[303,389,1827,1853]" captionStart="FIGURES 15 – 19" captionStartId="8.[151,269,1557,1581]" captionTargetBox="[159,1416,652,1556]" captionTargetId="figure@8.[151,1436,652,1601]" captionTargetPageId="8" captionText="FIGURES 15 – 19. Cloeodes espinillo n. sp. Male imago: 15, head and thorax, dorsal; 16, fore wing; 17 a, hind wing;" httpUri="https://zenodo.org/record/181241/files/figure.png" pageId="8" pageNumber="9">Fig. 19</figureCitation>
) with bases of forceps close together, segment III rounded. In the nymph, 1) hind wing pads present; 2) lingua with a rounded projection (
<figureCitation id="13FF2A74D53DFFCFFD0DB460FCDDF8BC" box="[758,848,1867,1893]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 23</figureCitation>
); 3) dorsal edge of femora with a row of pointed spines, apically with a subquadrangular projection with a pair of blunt spines (
<figureCitation id="13FF2A74D53DFFCFFBBDB458FB2DF854" box="[1094,1184,1907,1933]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 26</figureCitation>
); 4) abdominal color pattern yellowish brown with segments II–III and VI with brown spots as in 
<figureCitation id="13FF2A74D53DFFCFFBDEB4B0FBF6F86C" box="[1061,1147,1947,1973]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 20</figureCitation>
, segments IX–X brown; 5) caudal filaments with flattened setae basally sclerotized (
<figureCitation id="13FF2A74D53DFFCFFCA8B4E8FC26F804" box="[851,939,1987,2013]" captionStart="FIGURES 20 – 30" captionStartId="9.[151,269,1933,1957]" captionTargetBox="[185,1375,198,1932]" captionTargetId="figure@9.[184,1390,170,1935]" captionTargetPageId="9" captionText="FIGURES 20 – 30. Cloeodes espinillo n. sp. Nymph. 20, general view. Mouthparts (Figs. 21 – 24): 21, labrum, left d. v., right v. v.; 22, left mandible v. v.; 23, hypopharynx v. v.; 24, labium, left d. v., right v. v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus." httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="8" pageNumber="9">Fig. 30</figureCitation>
), cerci with long spines toward external margin every two segments and terminal filament with long spines toward midline dorsally and ventrally every two segments (similar to 
<figureCitation id="13FF2A74D53FFFCDFDFCB3EBFDD3FF03" box="[519,606,192,218]" captionStart="FIGURES 5 – 14" captionStartId="6.[151,269,1938,1962]" captionTargetBox="[267,1354,190,1909]" captionTargetId="figure@6.[203,1390,162,1984]" captionTargetPageId="6" captionText="FIGURES 5 – 14. Cloeodes barituensis n. sp. Nymph. 5, general view, dorsal. 6, antennae. Mouthparts (Figs. 7 – 10): 7, labrum, left d. v., right v. v.; 8, right mandible v. v.; 9, hypopharynx v. v.; 10, maxilla v. v. 11, leg I. 12, posterior margin of tergum IV. 13, gill IV. 14, terminal filament, left cercus, right caudal filament." httpUri="https://zenodo.org/record/181240/files/figure.png" pageId="10" pageNumber="11">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DFBB6679D53CFFCEFF6CB4A6FC34F833" httpUri="https://zenodo.org/record/181242/files/figure.png" pageId="9" pageNumber="10" targetBox="[185,1375,198,1932]" targetPageId="9">
<paragraph id="8B7B36F1D53CFFCEFF6CB4A6FC34F833" blockId="9.[151,1436,1933,2026]" pageId="9" pageNumber="10">
<emphasis id="B9B0EAE3D53CFFCEFF6CB4A6FED2F87C" bold="true" box="[151,351,1933,1957]" pageId="9" pageNumber="10">FIGURES 20–30.</emphasis>
<taxonomicName id="4CC44D72D53CFFCEFE9CB4A6FDA3F87D" box="[359,558,1933,1956]" class="Insecta" family="Baetidae" genus="Cloeodes" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="espinillo" status="sp. nov.">
<emphasis id="B9B0EAE3D53CFFCEFE9CB4A6FDA3F87D" box="[359,558,1933,1956]" italics="true" pageId="9" pageNumber="10">Cloeodes espinillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2835798D53CFFCEFDCDB4A6FDF9F87C" box="[566,628,1933,1957]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B9B0EAE3D53CFFCEFDCDB4A6FDF9F87C" bold="true" box="[566,628,1933,1957]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
Nymph. 20, general view. Mouthparts (Figs. 21–24): 21, labrum, left d.v., right v.v.; 22, left mandible v.v.; 23, hypopharynx v.v.; 24, labium, left d.v., right v.v. 25, leg I; 26, dorsal margin of femur I. 27, posterior margin of tergum IV; 28, sternum IV. 29, gill IV. 30, cercus.
</paragraph>
</caption>
<subSubSection id="C3DE657AD53FFFCDFF3DB3CCFDDBFEA3" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8B7B36F1D53FFFCDFF3DB3CCFDDBFEA3" blockId="10.[151,1436,152,378]" pageId="10" pageNumber="11">
<emphasis id="B9B0EAE3D53FFFCDFF3DB3CCFEB6FED8" bold="true" box="[198,315,231,257]" pageId="10" pageNumber="11">Material.</emphasis>
<typeStatus id="547F8853D53FFFCDFEBAB3C3FE3EFEDB" box="[321,435,232,258]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
: male nymph: 
<collectingCountry id="F3D37661D53FFFCDFDA6B3C3FC86FEDB" box="[605,779,232,258]" name="Argentina" pageId="10" pageNumber="11">ARGENTINA</collectingCountry>
, Córdoba, Río Los Espinillos, 
<date id="FF7A1031D53FFFCDFB94B3C3FABFFEDB" box="[1135,1330,232,258]" pageId="10" pageNumber="11" value="1997-09-13" valueMax="1997-09-16" valueMin="1997-09-13">13–16/ IX/ 1997</date>
, Nieto &amp; Molineri colls. 
<typeStatus id="547F8853D53FFFCDFEB7B23BFE4EFEF3" box="[332,451,272,298]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
: 4 nymphs, 
<specimenCount id="9DC2FD78D53FFFCDFDA9B23BFD2FFEF3" box="[594,674,272,298]" pageId="10" pageNumber="11" type="male">5 male</specimenCount>
imagos (reared) and 
<specimenCount id="9DC2FD78D53FFFCDFC64B23BFC73FEF3" box="[927,1022,272,298]" pageId="10" pageNumber="11" type="male">16 male</specimenCount>
imagos, same data as 
<typeStatus id="547F8853D53FFFCDFAF3B23BFAFEFEF3" box="[1288,1395,272,298]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
. 18 nymphs: Depto. Pocho, arroyito afluente Río Pocho, 
<geoCoordinate id="EEF05036D53FFFCDFCFBB213FC00FE8B" box="[768,909,312,338]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="15" value="-31.22139">S 31°13'17''</geoCoordinate>
, 
<geoCoordinate id="EEF05036D53FFFCDFC62B213FBBCFE8B" box="[921,1073,312,338]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="15" value="-65.14056">W 65°08'26''</geoCoordinate>
, 
<quantity id="4C3C9B14D53FFFCDFBC6B213FB15FE8B" box="[1085,1176,312,338]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="10" pageNumber="11" unit="m" value="1030.0">1030 m</quantity>
, 
<date id="FF7A1031D53FFFCDFB58B213FAB7FE8B" box="[1187,1338,312,338]" pageId="10" pageNumber="11" value="2001-11-14">14/ XI/ 2001</date>
, Orce &amp; Nieto colls. Material housed in 
<collectionCode id="EDD5AE34D53FFFCDFDF3B24BFDDFFEA3" box="[520,594,352,378]" pageId="10" pageNumber="11">IFML</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>