treatments-xml/data/03/BB/11/03BB115251269B68479FFF4CFEFFF85A.xml

<document id="48FD7669BED2F49ECE0F52E849ACE354" ID-CLB-Dataset="32005" ID-DOI="10.11646/zootaxa.4323.4.4" ID-GBIF-Dataset="cafdc21f-367c-4abc-b93c-bf4338483003" ID-ISSN="1175-5326" ID-Zenodo-Dep="995854" ID-ZooBank="5641Cb17-Fb71-4Ebc-B178-0891C31B6Abb" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506321515914" checkinUser="plazi" docAuthor="Reynoso-Velasco, Daniel" docDate="2017" docId="03BB115251269B68479FFF4CFEFFF85A" docLanguage="en" docName="zootaxa.4323.4.4.pdf" docOrigin="Zootaxa 4323 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Carvalhoiella montandoni Reynoso-Velasco 2017, NEW COMBINATION" docType="treatment" docVersion="7" lastPageNumber="506" masterDocId="FF82692A51259B6B4708FFDBFFE4FF9C" masterDocTitle="New combinations, status, and species of Neotropical Ambrysini (Heteroptera: Naucoridae: Cryphocricinae)" masterLastPageNumber="518" masterPageNumber="503" pageNumber="506" updateTime="1698468928599" updateUser="ExternalLinkService">
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<mods:title id="95830ABFCC93A8EDD62D80D7AB473B4E">New combinations, status, and species of Neotropical Ambrysini (Heteroptera: Naucoridae: Cryphocricinae)</mods:title>
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<treatment id="03BB115251269B68479FFF4CFEFFF85A" ID-DOI="http://doi.org/10.5281/zenodo.6015853" ID-GBIF-Taxon="134839750" ID-Zenodo-Dep="6015853" LSID="urn:lsid:plazi:treatment:03BB115251269B68479FFF4CFEFFF85A" httpUri="http://treatment.plazi.org/id/03BB115251269B68479FFF4CFEFFF85A" lastPageNumber="506" pageId="3" pageNumber="506">
<subSubSection id="C308F3CF51269B68479FFF4CFEE9FF4F" pageId="3" pageNumber="506" type="nomenclature">
<paragraph id="8BADA04451269B68479FFF4CFC7DFF2E" blockId="3.[151,921,151,211]" box="[151,921,151,178]" pageId="3" pageNumber="506">
<heading id="D0E5172851269B68479FFF4CFC7DFF2E" bold="true" box="[151,921,151,178]" fontSize="11" level="1" pageId="3" pageNumber="506" reason="1">
<emphasis id="B9667C5651269B68479FFF4CFC7DFF2E" bold="true" box="[151,921,151,178]" pageId="3" pageNumber="506">
<taxonomicName id="4C12DBC751269B68479FFF4CFD8EFF2E" ID-CoL="5XDHB" authority="(La Rivers)" authorityName="Reynoso-Velasco" authorityYear="2017" baseAuthorityName=": Sites &amp; Nichols" baseAuthorityYear="1963" box="[151,618,151,178]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni" status="comb. nov.">
<emphasis id="B9667C5651269B68479FFF4CFE31FF2D" bold="true" box="[151,469,151,177]" italics="true" pageId="3" pageNumber="506">Carvalhoiella montandoni</emphasis>
(La Rivers)
</taxonomicName>
<taxonomicNameLabel id="A255C12D51269B684579FF43FC7DFF2E" box="[625,921,152,178]" pageId="3" pageNumber="506">NEW COMBINATION</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BADA04451269B68479FFF61FEE9FF4F" blockId="3.[151,921,151,211]" box="[151,269,186,211]" pageId="3" pageNumber="506">
<figureCitation id="1329BCC151269B68479FFF61FEE9FF4F" box="[151,269,186,211]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Figs. 8–13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C308F3CF51269B68479FFED9FC0DFEE8" pageId="3" pageNumber="506" type="reference_group">
<paragraph id="8BADA04451269B68479FFED9FC0DFEE8" blockId="3.[151,1288,258,373]" pageId="3" pageNumber="506">
<taxonomicName id="4C12DBC751269B68479FFED9FDC4FE84" ID-CoL="CPX6" authority="La Rivers 1963" authorityName=", La Rivers" authorityYear="1963" box="[151,544,258,280]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9667C5651269B68479FFED9FE9EFE84" box="[151,378,258,280]" italics="true" pageId="3" pageNumber="506">Ambrysus montandoni</emphasis>
<bibRefCitation id="EF83DDB551269B684688FED8FDC4FE84" author="La" box="[384,544,258,280]" pageId="3" pageNumber="506" refString="La Rivers, I. (1963) Two new Ambrysi (Hemiptera: Naucoridae). Biological Society of Nevada Occasional Papers, 1, 1 - 7." type="journal article" year="1963">La Rivers 1963</bibRefCitation>
</taxonomicName>
: Biol. Soc. 
<collectingRegion id="49D66EA651269B68459CFED9FD05FE84" box="[660,737,258,280]" country="United States of America" name="Nevada" pageId="3" pageNumber="506">Nevada</collectingRegion>
Occas. Pap. 1: 1–5 (original description). 
<taxonomicName id="4C12DBC751269B68479FFEFAFD7BFEAB" ID-CoL="CPX6" authority=": La Rivers 1965" authorityName="La Rivers" authorityYear="1965" box="[151,671,289,311]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni" subGenus="Syncollus">
<emphasis id="B9667C5651269B68479FFEFAFF1FFEAB" box="[151,251,289,311]" italics="true" pageId="3" pageNumber="506">Ambrysus</emphasis>
(
<emphasis id="B9667C5651269B684601FEFAFE8EFEAB" box="[265,362,289,311]" italics="true" pageId="3" pageNumber="506">Syncollus</emphasis>
) 
<emphasis id="B9667C5651269B684671FEFAFE16FEAB" box="[377,498,289,311]" italics="true" pageId="3" pageNumber="506">montandoni</emphasis>
: 
<bibRefCitation id="EF83DDB551269B6846F7FEF9FD7BFEAB" author="La" box="[511,671,289,311]" pageId="3" pageNumber="506" refString="La Rivers, I. (1965) The subgenera of the genus Ambrysus (Hemiptera, Naucoridae). Biological Society of Nevada Occasional Papers, 4, 1 - 7." type="journal article" year="1965">La Rivers 1965</bibRefCitation>
</taxonomicName>
. Biol. Soc. 
<collectingRegion id="49D66EA651269B68441BFEFAFC84FEAB" box="[787,864,289,311]" country="United States of America" name="Nevada" pageId="3" pageNumber="506">Nevada</collectingRegion>
Occas. Pap., 4: 5 (subgenus designation). 
<taxonomicName id="4C12DBC751269B68479FFE9BFD80FECA" ID-CoL="CPX6" authority=": Sites &amp; Nichols 1999" authorityName="Sites &amp; Nichols" authorityYear="1999" box="[151,612,320,342]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9667C5651269B68479FFE9BFE9EFECA" box="[151,378,320,342]" italics="true" pageId="3" pageNumber="506">Ambrysus montandoni</emphasis>
: 
<bibRefCitation id="EF83DDB551269B68468FFE9BFD80FECA" author="Sites" box="[391,612,320,342]" pageId="3" pageNumber="506" refString="Sites, R. W. &amp; Nichols, B. J. (1999) Egg architecture of Naucoridae (Heteroptera): internal and external structure of the chorion and micropyle. Proceedings of the Entomological Society of Washington, 101, 1 - 25." type="journal article" year="1999">Sites &amp; Nichols 1999</bibRefCitation>
</taxonomicName>
. Proc. Entomol. Soc. Wash. 101: 5–6 (egg description). 
<taxonomicName id="4C12DBC751269B68479FFE85FD80FEE8" ID-CoL="CPX6" authority=": Rodrigues et al. 2012" authorityName="Rodrigues et al." authorityYear="2012" box="[151,612,350,373]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9667C5651269B68479FFE85FE9EFEE8" box="[151,378,350,372]" italics="true" pageId="3" pageNumber="506">Ambrysus montandoni</emphasis>
: 
<bibRefCitation id="EF83DDB551269B68468FFE85FD80FEE8" author="Rodrigues" box="[391,612,350,373]" pageId="3" pageNumber="506" refString="Rodrigues, H. D. D., de Melo, A. L. &amp; Sites, R. W. (2012) Redescription of Ambrysus montandoni La Rivers, 1963 (Hemiptera: Heteroptera: Naucoridae), with the first records for Brazil. Zootaxa, 3357, 63 - 68." type="journal article" year="2012">
Rodrigues 
<emphasis id="B9667C5651269B6846FDFEBBFDC6FEE8" box="[501,546,350,373]" italics="true" pageId="3" pageNumber="506">et al</emphasis>
. 2012
</bibRefCitation>
</taxonomicName>
, Zootaxa 3357: 63–68 (redescription).
</paragraph>
</subSubSection>
<subSubSection id="C308F3CF51269B68479FFE7AFC04FD62" pageId="3" pageNumber="506" type="diagnosis">
<paragraph id="8BADA04451269B68479FFE7AFC04FD62" blockId="3.[151,1437,417,1990]" pageId="3" pageNumber="506">
<emphasis id="B9667C5651269B68479FFE7AFEF5FE26" bold="true" box="[151,273,417,442]" pageId="3" pageNumber="506">Diagnosis.</emphasis>
Body length 
<quantity id="4CEA0DA151269B6846AFFE7AFD8CFE25" box="[423,616,417,442]" metricMagnitude="-2" metricUnit="m" metricValue="1.208" metricValueMax="1.296" metricValueMin="1.1199999999999999" pageId="3" pageNumber="506" unit="mm" value="12.08" valueMax="12.96" valueMin="11.2">11.20–12.96 mm</quantity>
. The dorsum is tuberculate, dull, and irregularly mottled (
<figureCitation id="1329BCC151269B6843E4FE7AFAD4FE25" box="[1260,1328,417,442]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 8</figureCitation>
). A sharp sulcus marks the anterior margin of the posterior transverse band on the pronotum. Tergum VI of the male lacks an accessory genitalic process (
<figureCitation id="1329BCC151269B6846EBFE32FDC8FD9D" box="[483,556,489,514]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 9</figureCitation>
). Laterosternites are absent on III–V (males) and III–IV (females) and the abdominal venter is lightly colored with a wide, glabrous, yellow marginal band. The midline of sterna V–VIII (males) and V–VII (females) are mostly glabrous and impubis. The pseudoparameres are large, broad, sharply angled posterolaterad, with angulate apices, and overlapping and extending beyond the lateral lobes of tergum VIII (
<figureCitation id="1329BCC151269B684797FDA2FF10FD0E" box="[159,244,633,658]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 10</figureCitation>
). The proctiger is setose in the lateral thirds and nearly devoid of setae in the middle third (
<figureCitation id="1329BCC151269B684200FDA2FABEFD0E" box="[1288,1370,633,658]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 11</figureCitation>
). The parameres have an apical tuft of elongate setae (
<figureCitation id="1329BCC151269B6845E9FD46FCDFFD2A" box="[737,827,669,694]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 12</figureCitation>
). The ventral margins of the phallosoma have two hemispherical, lamellate, sclerotized lobes near the base of the endosoma (
<figureCitation id="1329BCC151269B6844DFFD1AFBCEFD46" box="[983,1066,705,730]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 13</figureCitation>
). The distal margin of the female subgenital plate is deeply cleft medially (see 
<bibRefCitation id="EF83DDB551269B684587FD3EFC60FD62" author="Rodrigues" box="[655,900,741,766]" pageId="3" pageNumber="506" refString="Rodrigues, H. D. D., de Melo, A. L. &amp; Sites, R. W. (2012) Redescription of Ambrysus montandoni La Rivers, 1963 (Hemiptera: Heteroptera: Naucoridae), with the first records for Brazil. Zootaxa, 3357, 63 - 68." type="journal article" year="2012">
Rodrigues 
<emphasis id="B9667C5651269B684402FD3DFCDEFD62" box="[778,826,741,766]" italics="true" pageId="3" pageNumber="506">et al</emphasis>
. 2012
</bibRefCitation>
, 
<figureCitation id="1329BCC151269B684486FD3EFC36FD62" box="[910,978,741,766]" captionStart="FIGURES 8 – 13" captionStartId="5.[151,264,1929,1952]" captionTargetBox="[151,1436,213,1884]" captionTargetId="figure@5.[151,1436,213,1884]" captionTargetPageId="5" captionText="FIGURES 8 – 13. Carvalhoiella montandoni. (8) Dorsal habitus, (9) abdominal terga VI and VII of male, (10) abdominal tergum VIII of male with pseudoparameres, (11) male genital capsule, (12) male genital capsule with terga IX and X (proctiger) removed, (13) ventral surface of phallosoma." httpUri="https://zenodo.org/record/995858/files/figure.png" pageId="3" pageNumber="506">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C308F3CF51269B6847CFFCD2FC0DFAFE" pageId="3" pageNumber="506" type="discussion">
<paragraph id="8BADA04451269B6847CFFCD2FA9CFB15" blockId="3.[151,1437,417,1990]" pageId="3" pageNumber="506">
<emphasis id="B9667C5651269B6847CFFCD2FEA8FCBE" bold="true" box="[199,332,777,802]" pageId="3" pageNumber="506">Discussion.</emphasis>
The expression of laterosternites has been partially suppressed in this species. Laterosternites III– V in males and III–IV in females are absent, and the mesal margin of V in females is incomplete; the laterosternites are complete thereafter, although difficult to discern beneath the dense hair pile. Although this large species is superficially reminiscent of 
<taxonomicName id="4C12DBC751269B6846DAFCADFD9FFC11" authorityName="Usinger" authorityYear="1941" box="[466,635,886,909]" class="Insecta" family="Naucoridae" genus="Cataractocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B9667C5651269B6846DAFCADFD9FFC11" box="[466,635,886,909]" italics="true" pageId="3" pageNumber="506">Cataractocoris</emphasis>
</taxonomicName>
, it shares diagnostic attributes consistent with 
<taxonomicName id="4C12DBC751269B684380FCAEFAC0FC12" authorityName="De Carlo" authorityYear="1963" box="[1160,1316,885,910]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B9667C5651269B684380FCAEFAC0FC12" box="[1160,1316,885,910]" italics="true" pageId="3" pageNumber="506">Carvalhoiella</emphasis>
</taxonomicName>
, including the loss of setae on the abdominal mediosternites, the head is not produced anteriorly, the anterior margin of the pronotum is only slightly concave to receive the head, the mesal margins of the laterosternites are diagonally offset from one another, the ventral abdominal lateral band is wide, and the pseudoparameres are large and posterolaterally directed (
<tableCitation id="C69095FF51269B6846B0FBDEFDEAFB82" box="[440,526,1029,1054]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Comparison of character states among genera of Ambrysini." pageId="3" pageNumber="506">Table 1</tableCitation>
). It lacks many of the distinguishing features of the genus 
<taxonomicName id="4C12DBC751269B6843BCFBDEFAC6FB82" box="[1204,1314,1029,1054]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B9667C5651269B6843BCFBDEFAC6FB82" box="[1204,1314,1029,1054]" italics="true" pageId="3" pageNumber="506">Ambrysus</emphasis>
</taxonomicName>
, including deep invagination of the head into the pronotum, continuous ventral abdominal hair pile, continuous mesal margins of the laterosternites, narrow ventral abdominal lateral band, small male pseudoparameres, and male accessory genitalic process of tergum VI (
<tableCitation id="C69095FF51269B6846F1FBAAFDA9FB16" box="[505,589,1137,1162]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionText="TABLE 1. Comparison of character states among genera of Ambrysini." pageId="3" pageNumber="506">Table 1</tableCitation>
). Thus, 
<taxonomicName id="4C12DBC751269B6845A3FBAAFC43FB16" box="[683,935,1137,1162]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9667C5651269B6845A3FBAAFC43FB16" box="[683,935,1137,1162]" italics="true" pageId="3" pageNumber="506">Ambrysus montandoni</emphasis>
</taxonomicName>
is transferred to the genus 
<taxonomicName id="4C12DBC751269B6843DDFBAAFA94FB16" authorityName="De Carlo" authorityYear="1963" box="[1237,1392,1137,1162]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="genus">
<emphasis id="B9667C5651269B6843DDFBAAFA94FB16" box="[1237,1392,1137,1162]" italics="true" pageId="3" pageNumber="506">Carvalhoiella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BADA04451269B6847CFFB4EFC0DFAFE" blockId="3.[151,1437,417,1990]" pageId="3" pageNumber="506">
This species is known only from the 
<collectingCountry id="F305E0D451269B684589FB4EFD36FB32" box="[641,722,1173,1198]" name="Guyana" pageId="3" pageNumber="506">Guiana</collectingCountry>
Shield in 
<collectingCountry id="F305E0D451269B684459FB4EFC7CFB32" box="[849,920,1173,1198]" name="Brazil" pageId="3" pageNumber="506">Brazil</collectingCountry>
, 
<collectingCountry id="F305E0D451269B6844AFFB4DFBE5FB32" box="[935,1025,1174,1198]" name="Guyana" pageId="3" pageNumber="506">Guyana</collectingCountry>
, and 
<collectingCountry id="F305E0D451269B68434CFB4EFB5FFB32" box="[1092,1211,1173,1198]" name="Venezuela" pageId="3" pageNumber="506">Venezuela</collectingCountry>
. It is perhaps most famously known from El Tobogan de la Selva in 
<collectingCountry id="F305E0D451269B6845CEFB62FCD8FB4E" box="[710,828,1209,1234]" name="Venezuela" pageId="3" pageNumber="506">Venezuela</collectingCountry>
, which is a large, smooth, rolling granitic slide where water flows in a laminar fashion. 
<taxonomicName id="4C12DBC751269B684505FB06FCD1FB6A" authorityName="Reynoso-Velasco" authorityYear="2017" baseAuthorityName=": Sites &amp; Nichols" baseAuthorityYear="1963" box="[525,821,1245,1270]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">
<emphasis id="B9667C5651269B684505FB06FCD1FB6A" box="[525,821,1245,1270]" italics="true" pageId="3" pageNumber="506">Carvalhoiella montandoni</emphasis>
</taxonomicName>
can be found here along with 
<taxonomicName id="4C12DBC751269B684380FB05FAD5FB6A" authorityName="La Rivers" authorityYear="1954" box="[1160,1329,1245,1270]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="maldonadus">
<emphasis id="B9667C5651269B684380FB05FAD5FB6A" box="[1160,1329,1245,1270]" italics="true" pageId="3" pageNumber="506">C. maldonadus</emphasis>
</taxonomicName>
in cracks in the rock and under dead leaves. At El Tobogan, this species was collected with 
<taxonomicName id="4C12DBC751269B68439BFADAFA73FA86" authority="Sites" authorityName="Sites" authorityYear="2015" box="[1171,1431,1281,1306]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="shorti">
<emphasis id="B9667C5651269B68439BFADAFAB4FA86" box="[1171,1360,1281,1306]" italics="true" pageId="3" pageNumber="506">Ambrysus shorti</emphasis>
Sites
</taxonomicName>
, 
<taxonomicName id="4C12DBC751269B68479FFAFEFCE2FAA2" authority="(La Rivers)" authorityName="Reynoso-Velasco" authorityYear="2017" baseAuthorityName="Reynoso-Velasco" baseAuthorityYear="1965" box="[151,774,1317,1342]" class="Insecta" family="Naucoridae" genus="Carvalhoiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="maldonadus">
<emphasis id="B9667C5651269B68479FFAFEFE2EFAA2" box="[151,458,1317,1342]" italics="true" pageId="3" pageNumber="506">Carvalhoiella maldonadus</emphasis>
(La Rivers) (Naucoridae)
</taxonomicName>
, 
<taxonomicName id="4C12DBC751269B684410FAFEFAD4FAA2" authority="Miller" authorityName="Miller" box="[792,1328,1317,1342]" class="Insecta" family="Dytiscidae" genus="Fontidessus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="toboganensis">
<emphasis id="B9667C5651269B684410FAFEFBA4FAA2" box="[792,1088,1317,1342]" italics="true" pageId="3" pageNumber="506">Fontidessus toboganensis</emphasis>
Miller (Dytiscidae)
</taxonomicName>
, 
<taxonomicName id="4C12DBC751269B68424BFAFDFE6BFAFE" authority="Oliva &amp; Short" authorityName="Oliva &amp; Short" authorityYear="2012" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="garciai">
<emphasis id="B9667C5651269B68424BFAFDFF03FAFD" italics="true" pageId="3" pageNumber="506">Berosus garciai</emphasis>
Oliva &amp; Short
</taxonomicName>
, and 
<taxonomicName id="4C12DBC751269B6846C2FA92FC01FAFE" authority="Short &amp; Garcia" authorityName="Short &amp; Garcia" authorityYear="2010" box="[458,997,1353,1378]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="petra">
<emphasis id="B9667C5651269B6846C2FA92FD90FAFD" box="[458,628,1353,1378]" italics="true" pageId="3" pageNumber="506">Oocyclus petra</emphasis>
Short &amp; García (Hydrophilidae)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C308F3CF51269B6847CFFAB6FEFFF85A" pageId="3" pageNumber="506" type="materials_examined">
<paragraph id="8BADA04451269B6847CFFAB6FB0BFA1A" blockId="3.[151,1437,417,1990]" box="[199,1263,1389,1414]" pageId="3" pageNumber="506">
<materialsCitation id="3B7AAA1951269B6847CFFAB6FB0BFA1A" ID-GBIF-Occurrence="1655995275" box="[199,1263,1389,1414]" country="Venezuela" location="Upper Cunucunuma River." municipality="Mount Marahuaca" pageId="3" pageNumber="506" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="B9667C5651269B6847CFFAB6FEBBFA1A" bold="true" box="[199,351,1389,1414]" pageId="3" pageNumber="506">
<typeStatus id="54A91EE651269B6847CFFAB6FEE6FA1A" box="[199,258,1389,1414]" pageId="3" pageNumber="506">Type</typeStatus>
locality
</emphasis>
: 
<collectingCountry id="F305E0D451269B684666FAB6FE00FA1A" box="[366,484,1389,1414]" name="Venezuela" pageId="3" pageNumber="506">Venezuela</collectingCountry>
, Territorio 
<collectingRegion id="49D66EA651269B684569FAB5FD3EFA1A" box="[609,730,1390,1414]" country="Venezuela" name="Amazonas" pageId="3" pageNumber="506">Amazonas</collectingRegion>
, 
<collectingMunicipality id="6BC93A3E51269B6845EEFAB5FC52FA1A" box="[742,950,1389,1414]" pageId="3" pageNumber="506">Mount Marahuaca</collectingMunicipality>
, 
<location id="8ECDF69F51269B6844C9FAB5FB0BFA1A" LSID="urn:lsid:plazi:treatment:03BB115251269B68479FFF4CFEFFF85A:8ECDF69F51269B6844C9FAB5FB0BFA1A" box="[961,1263,1389,1414]" country="Venezuela" municipality="Mount Marahuaca" name="Upper Cunucunuma River." pageId="3" pageNumber="506" stateProvince="Amazonas">Upper Cunucunuma River.</location>
</materialsCitation>
</paragraph>
<paragraph id="8BADA04451269B6847CFFA4AFB41FA36" blockId="3.[151,1437,417,1990]" box="[199,1189,1425,1450]" pageId="3" pageNumber="506">
<materialsCitation id="3B7AAA1951269B6847CFFA4AFB41FA36" ID-GBIF-Occurrence="1655995288" box="[199,1189,1425,1450]" location="Type" pageId="3" pageNumber="506" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<emphasis id="B9667C5651269B6847CFFA4AFE66FA36" bold="true" box="[199,386,1425,1450]" pageId="3" pageNumber="506">
<location id="8ECDF69F51269B6847CFFA4AFEE6FA36" LSID="urn:lsid:plazi:treatment:03BB115251269B68479FFF4CFEFFF85A:8ECDF69F51269B6847CFFA4AFEE6FA36" box="[199,258,1425,1450]" name="Type" pageId="3" pageNumber="506" stateProvince="California">
<typeStatus id="54A91EE651269B6847CFFA4AFEE6FA36" box="[199,258,1425,1450]" pageId="3" pageNumber="506">Type</typeStatus>
</location>
repository
</emphasis>
: 
<collectingRegion id="49D66EA651269B684699FA4AFDE6FA36" box="[401,514,1425,1450]" country="United States of America" name="California" pageId="3" pageNumber="506">California</collectingRegion>
Academy of Sciences (
<typeStatus id="54A91EE651269B684403FA4AFC8BFA36" box="[779,879,1425,1450]" pageId="3" pageNumber="506" type="holotype">holotype</typeStatus>
<specimenCount id="9D146BCD51269B68447EFA49FC6FFA35" box="[886,907,1426,1449]" pageId="3" pageNumber="506" type="male">♂</specimenCount>
by original designation).
</materialsCitation>
</paragraph>
<paragraph id="8BADA04451269B6847CFFA6EFEFFF85A" blockId="3.[151,1437,417,1990]" pageId="3" pageNumber="506">
<emphasis id="B9667C5651269B6847CFFA6EFDA2FA52" bold="true" box="[199,582,1461,1486]" pageId="3" pageNumber="506">Material examined. BRAZIL</emphasis>
: Roraima State, Yanomami indigenous area, Serra de Surucucu (02°49’53.78”N, 63°38’19.78”W), stream without name, 
<date id="FFAC868451269B68443DFA01FC5CFA6E" box="[821,952,1498,1522]" pageId="3" pageNumber="506" value="1991-11-24">24.XI.1991</date>
, V. Py-Daniel and U. Barbosa coll., (1♂ INPA); same data, except collected in river of the waterfall, (1♂ INPA; 1♂, 1♀ DPIC); Yanomami indigenous area, Xitei/Xidéa indigenous park (02°36’20.54”N, 63°52’27.76”W), Igarapé das Irmãs, 
<date id="FFAC868451269B684342F9F9FB28F9A6" box="[1098,1228,1570,1594]" pageId="3" pageNumber="506" value="1993-11-17">17.XI.1993</date>
, V. Py-Daniel and U. Barbosa coll., (1♂, 1♀ INPA). 
<emphasis id="B9667C5651269B68451AF99EFD6EF9C2" bold="true" box="[530,650,1605,1630]" pageId="3" pageNumber="506">GUYANA</emphasis>
: Lethem (
<quantity id="4CEA0DA151269B6845F5F99EFCA1F9C2" box="[765,837,1605,1630]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="506" unit="km" value="25.0">25 km</quantity>
SE), Kumu Falls, 3°15.9'N, 59°43.6'W, 
<date id="FFAC868451269B68420FF99DFA7CF9C2" box="[1287,1432,1605,1630]" pageId="3" pageNumber="506" value="1994-04-04">4 April 1994</date>
, PJ Spangler / colln. #14, side pool at base of falls (2♂, 3 immatures USNM). 
<emphasis id="B9667C5651269B68432CF9B2FB36F91E" bold="true" box="[1060,1234,1641,1666]" pageId="3" pageNumber="506">VENEZUELA</emphasis>
: Venezuela Exp., Territ. Amazonas, Culebra, N. Duida, 
<date id="FFAC868451269B68455EF955FD09F93A" box="[598,749,1677,1702]" pageId="3" pageNumber="506" value="1950-06-09">June 9, 1950</date>
/ J. Maldonado Capriles Coll. / 
<taxonomicName id="4C12DBC751269B684378F956FF04F956" authority="La Rivers" authorityName=", La Rivers" authorityYear="1963" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">Ambrysus montandoni La Rivers</taxonomicName>
HOLOTYPE / 
<taxonomicName id="4C12DBC751269B68469FF96AFCF0F956" authority="La Rivers" authorityName=", La Rivers" authorityYear="1963" box="[407,788,1713,1738]" class="Insecta" family="Naucoridae" genus="Ambrysus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="montandoni">Ambrysus montandoni La Rivers</taxonomicName>
, determined by Ira La Rivers / Ira La Rivers Collection, Bequeathed to the California Academy of Sciences—1978 / California Academy of Sciences Type No. 13411 (Holotype ♂ CAS); Amazonas State, N Puerto Ayachucho, nr Iboruwa, &quot;Tobogancito&quot;, 
<date id="FFAC868451269B6843B6F921FA62F88E" box="[1214,1414,1786,1810]" pageId="3" pageNumber="506" value="2009-01-13">13 January 2009</date>
/ 5°48.414'N, 67°26.313'W, 
<quantity id="4CEA0DA151269B6846C0F8C5FDE1F8A9" box="[456,517,1822,1846]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="506" unit="m" value="80.0">80 m</quantity>
, VZ-20090113-02, coll: J. Camacho (1♂, 1♀ UMC); Amazonas State, S Puerto Ayachucho, El Tobogan de la Selva, 
<date id="FFAC868451269B68453AF899FD0FF8C6" box="[562,747,1858,1882]" pageId="3" pageNumber="506" value="2009-01-14">14 January 2009</date>
/ 5°23.207'N, 67°36.922'W, 
<quantity id="4CEA0DA151269B684324F89AFB91F8C5" box="[1068,1141,1857,1882]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="506" unit="m" value="125.0">125 m</quantity>
, VZ-20090114-01, coll: J. Camacho (4♂, 4♀ UMC); Amazonas, 
<quantity id="4CEA0DA151269B68455FF8BDFD45F8E2" box="[599,673,1893,1918]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="506" unit="km" value="42.0">42 km</quantity>
S of Puerto Ayacucho Tobogan, 
<date id="FFAC868451269B68432DF8BDFB03F8E2" box="[1061,1255,1894,1918]" pageId="3" pageNumber="506" value="1989-01-19">19 January 1989</date>
, J.T. Polhemus, CL2371, (2♂, 2♀ UMC); Bolivar, 
<quantity id="4CEA0DA151269B684517F851FD9CF83E" box="[543,632,1929,1954]" metricMagnitude="5" metricUnit="m" metricValue="1.08" pageId="3" pageNumber="506" unit="km" value="108.0">108 km</quantity>
. SW Sta. Elena de Uairen, 
<date id="FFAC868451269B6844A1F851FBCBF83E" box="[937,1071,1929,1954]" pageId="3" pageNumber="506" value="1978-07-04">4 July 1978</date>
, R.S. Miller / blackwater stream (1♂ UMC).
</paragraph>
</subSubSection>
</treatment>
</document>