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<document id="8E169B92481AEF56F0CFAC22C771C151" ID-DOI="10.11646/zootaxa.5448.1.10" ID-ISSN="1175-5326" ID-Zenodo-Dep="11231037" ID-ZooBank="9B7FFD06-AEA1-4258-A3E6-BDE2ADE72482" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716278060123" checkinUser="plazi" docAuthor="Bilton, David T. &amp; Mlambo, Musa C." docDate="2024" docId="3E74251DFFC5FFBAFF3DF69CA09D8B30" docLanguage="en" docName="zootaxa.5448.1.10.pdf" docOrigin="Zootaxa 5448 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Relictorygmus riparius Bilton &amp; Mlambo 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="147" masterDocId="C24D5D65FFC4FFBEFFAAF252A0468C71" masterDocTitle="A new, apparently lotic species of Relictorygmus from the Northern Cape Kamiesberg, South Africa (Coleoptera, Hydrophilidae, Cylominae)" masterLastPageNumber="150" masterPageNumber="143" pageNumber="144" updateTime="1716278469310" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="37D8ECE59A540AEC312B5DE0BD8917E4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE0D00AC494C626B6735EDCC7338B3FA">
<mods:title id="9D538A36BC8A1C3F9C105F4236ADF945">A new, apparently lotic species of Relictorygmus from the Northern Cape Kamiesberg, South Africa (Coleoptera, Hydrophilidae, Cylominae)</mods:title>
</mods:titleInfo>
<mods:name id="F49F8DF0287F12AD4CC1F41BFFF90363" type="personal">
<mods:role id="5804CBF87EA5F9A70F0D54286240B1F4">
<mods:roleTerm id="16D8C26BDB2783095798A36EF0F45FC7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2FFB91058682B8534402C501D546EF1">Bilton, David T.</mods:namePart>
<mods:affiliation id="53556B131D7BFA93193A9A6276BFE93E">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth PL 4 8 AA, UK &amp; Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, Johannesburg 2006, South Africa</mods:affiliation>
</mods:name>
<mods:name id="4E53159361AE9C3F66C1D8F031C014DD" type="personal">
<mods:role id="F566EC486CB79F9F7C56466CBADF455C">
<mods:roleTerm id="4C9A3787502A513E05F21459E3A5613C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E57EA95C34439CE6AE945966E9578CB5">Mlambo, Musa C.</mods:namePart>
<mods:affiliation id="8781C98697309E718B0F91BC2E502A80">Department of Freshwater Invertebrates, Albany Museum, and Department of Zoology and Entomology, Rhodes University, Grahamstown 6140, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2B0FB28DCB640DA85F1A07CB0011EBAC">text</mods:typeOfResource>
<mods:relatedItem id="4040BC4456997D4E55E6E60D2D60F7DD" type="host">
<mods:titleInfo id="E4DD1A5E908F79D3E6CA808A81104F82">
<mods:title id="8106E7FC05B7F06165CFBB156F4D69D4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3CC79351AF44EB2F2F264F8C6484B8AC">
<mods:date id="E45C1EBE07629C5573CF1D9E2E422C8F">2024</mods:date>
<mods:detail id="1C01370B5D84E48C6CF99BCEE7A70B6C" type="pubDate">
<mods:number id="377AB0A19412FF8B22F3F7C8AC416CF9">2024-05-09</mods:number>
</mods:detail>
<mods:detail id="A4C597D99DD9C3F903A3722E157177A9" type="volume">
<mods:number id="9C91A2DCA714F1283BDF08822FA7099D">5448</mods:number>
</mods:detail>
<mods:detail id="B6757D97D2EF58FA26278DD02622D117" type="issue">
<mods:number id="12051AC0DBCE50B72890E9285CFCF84B">1</mods:number>
</mods:detail>
<mods:extent id="0132EEFEB1C8B7BF0AE64F2E12A27096" unit="page">
<mods:start id="83A2BD9768190F3F2552F23CEB1B9979">143</mods:start>
<mods:end id="6DE868555F7D926DC70ED2CDF8AA77DC">150</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="CAC20668A0B1FAEBDBEB1052EEAEF275">
<mods:url id="DFCF6088C796A15826BF588D2CD39062">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
</mods:location>
<mods:classification id="4261D29CB4B1F6295DF7DA1339FA074A">journal article</mods:classification>
<mods:identifier id="94597D9D77F7E35116E46CA717657E93" type="DOI">10.11646/zootaxa.5448.1.10</mods:identifier>
<mods:identifier id="D42FA68B8D30835BB37F786D8E05CD7F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A69F19978F5FBEF075BE834B390A2245" type="Zenodo-Dep">11231037</mods:identifier>
<mods:identifier id="A3916AEBFDE8CDFD72B931DF9A087080" type="ZooBank">9B7FFD06-AEA1-4258-A3E6-BDE2ADE72482</mods:identifier>
</mods:mods>
<treatment id="3E74251DFFC5FFBAFF3DF69CA09D8B30" ID-DOI="http://doi.org/10.5281/zenodo.11231039" ID-Zenodo-Dep="11231039" LSID="urn:lsid:plazi:treatment:3E74251DFFC5FFBAFF3DF69CA09D8B30" httpUri="http://treatment.plazi.org/id/3E74251DFFC5FFBAFF3DF69CA09D8B30" lastPageId="4" lastPageNumber="147" pageId="1" pageNumber="144">
<subSubSection id="FEC7C780FFC5FFBFFF3DF69CA2548898" box="[151,530,1230,1257]" pageId="1" pageNumber="144" type="nomenclature">
<paragraph id="B662940BFFC5FFBFFF3DF69CA2548898" blockId="1.[151,530,1230,1293]" box="[151,530,1230,1257]" pageId="1" pageNumber="144">
<heading id="ED2A2367FFC5FFBFFF3DF69CA2548898" bold="true" box="[151,530,1230,1257]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
<emphasis id="84A94819FFC5FFBFFF3DF69CA2548898" bold="true" box="[151,530,1230,1257]" pageId="1" pageNumber="144">
<taxonomicName id="71DDEF88FFC5FFBFFF3DF69CA1EB8898" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[151,429,1230,1257]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC5FFBFFF3DF69CA1EB8898" bold="true" box="[151,429,1230,1257]" italics="true" pageId="1" pageNumber="144">Relictorygmus riparius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9F9AF562FFC5FFBFFE1EF69DA2548898" box="[436,530,1231,1257]" pageId="1" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FEC7C780FFC5FFBFFF3DF6A1A154897C" box="[151,274,1267,1293]" pageId="1" pageNumber="144" type="description">
<paragraph id="B662940BFFC5FFBFFF3DF6A1A154897C" blockId="1.[151,530,1230,1293]" box="[151,274,1267,1293]" pageId="1" pageNumber="144">
(
<figureCitation id="2EE6888EFFC5FFBFFF35F6A1A14C897C" box="[159,266,1267,1293]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[152,255,1546,1570]" captionStartId-1="3.[152,255,1359,1383]" captionStartId-2="4.[152,255,1436,1460]" captionStartId-3="5.[152,255,1891,1915]" captionTargetBox-0="[192,1394,189,1502]" captionTargetBox-1="[154,1434,489,1336]" captionTargetBox-2="[154,1434,453,1413]" captionTargetBox-3="[341,1247,181,1866]" captionTargetId-0="figure-27@2.[180,1408,179,1520]" captionTargetId-1="figure-139@3.[154,1434,489,1336]" captionTargetId-2="figure-164@4.[154,1434,453,1413]" captionTargetId-3="figure-17@5.[341,1247,181,1866]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." captionText-1="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." captionText-2="FIGURE 3. Relictorygmus species, detail of pronotal and elytral punctation. A) R. riparius sp. nov.; B) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018." captionText-3="FIGURE 4. Relictorygmus habitat and distribution. A) Type locality of R. riparius sp. nov., South Africa, Northern Cape Province, Kamiesberg (photo Andy Foggo); B) distribution of Relictorygmus species in South Africa. Green symbol = R. riparius sp. nov., red symbol = R. repentinus (Hebauer, 2002), yellow symbol = R. trevornoahi Seidel, Minoshima, Arriaga- Varela &amp; Fikáček, 2018." figureDoi-0="http://doi.org/10.5281/zenodo.11231041" figureDoi-1="http://doi.org/10.5281/zenodo.11231044" figureDoi-2="http://doi.org/10.5281/zenodo.11231046" figureDoi-3="http://doi.org/10.5281/zenodo.11231048" httpUri-0="https://zenodo.org/record/11231041/files/figure.png" httpUri-1="https://zenodo.org/record/11231044/files/figure.png" httpUri-2="https://zenodo.org/record/11231046/files/figure.png" httpUri-3="https://zenodo.org/record/11231048/files/figure.png" pageId="1" pageNumber="144">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FEC7C780FFC5FFBFFF3DF769A4738994" pageId="1" pageNumber="144" type="materials_examined">
<paragraph id="B662940BFFC5FFBFFF3DF769A3C28908" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
<emphasis id="84A94819FFC5FFBFFF3DF769A17C8925" bold="true" box="[151,314,1338,1365]" pageId="1" pageNumber="144">Type locality.</emphasis>
<materialsCitation id="06B59E56FFC5FFBFFEEFF768A3C68908" collectorName="De Kruis" country="South Africa" elevation="1168" latitude="-30.360386" location="Northern Cape Province" longLatPrecision="1" longitude="18.116982" pageId="1" pageNumber="144" specimenCount="1" stateProvince="Northern Cape">
<emphasis id="84A94819FFC5FFBFFEEFF768A1A78925" bold="true" box="[325,481,1338,1364]" pageId="1" pageNumber="144">
<collectingCountry id="CECAD49BFFC5FFBFFEEFF768A1A78925" box="[325,481,1338,1364]" name="South Africa" pageId="1" pageNumber="144">South Africa</collectingCountry>
</emphasis>
,
<collectingRegion id="74195AE9FFC5FFBFFE59F769A3488924" box="[499,782,1339,1365]" country="South Africa" name="Northern Cape" pageId="1" pageNumber="144">Northern Cape Province</collectingRegion>
, Kamiesberg, stream beside dirt road from
<collectorName id="1B28F1DDFFC5FFBFFABBF769A53D8924" box="[1297,1403,1339,1365]" pageId="1" pageNumber="144">De Kruis</collectorName>
to Leliefontein,
<geoCoordinate id="D3E9F2CCFFC5FFBFFE84F70DA1AF8908" box="[302,489,1375,1401]" degrees="30" direction="south" minutes="21" orientation="latitude" pageId="1" pageNumber="144" precision="1" seconds="37.39" value="-30.360386">30° 21’ 37.39” S</geoCoordinate>
<geoCoordinate id="D3E9F2CCFFC5FFBFFE45F70DA2EB8908" box="[495,685,1375,1401]" degrees="18" direction="east" minutes="07" orientation="longitude" pageId="1" pageNumber="144" precision="1" seconds="01.13" value="18.116982">18° 07’ 01.13” E</geoCoordinate>
,
<quantity id="712539EEFFC5FFBFFD13F70DA34B8908" box="[697,781,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="1" pageNumber="144" unit="m" value="1168.0">
<elevation id="3DF07338FFC5FFBFFD13F70DA34B8908" box="[697,781,1375,1401]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="1" pageNumber="144" unit="m" value="1168.0">1168 m</elevation>
</quantity>
(
<figureCitation id="2EE6888EFFC5FFBFFCB6F70DA3318908" box="[796,887,1375,1401]" captionStart="FIGURE 4" captionStartId="5.[152,255,1891,1915]" captionTargetBox="[341,1247,181,1866]" captionTargetId="figure-17@5.[341,1247,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Relictorygmus habitat and distribution. A) Type locality of R. riparius sp. nov., South Africa, Northern Cape Province, Kamiesberg (photo Andy Foggo); B) distribution of Relictorygmus species in South Africa. Green symbol = R. riparius sp. nov., red symbol = R. repentinus (Hebauer, 2002), yellow symbol = R. trevornoahi Seidel, Minoshima, Arriaga- Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231048" httpUri="https://zenodo.org/record/11231048/files/figure.png" pageId="1" pageNumber="144">Fig. 4A</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B662940BFFC5FFBFFF6DF7D1A50E89B0" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
<emphasis id="84A94819FFC5FFBFFF6DF7D1A13089ED" bold="true" box="[199,374,1410,1437]" pageId="1" pageNumber="144">Type material.</emphasis>
<materialsCitation id="06B59E56FFC5FFBFFED4F7D1A50E89B0" collectingDate="2023-09-18" collectionCode="AMG" collectorName="D T Bilton" country="South Africa" county="Leliefon" elevation="1168" location="Kamiesberg" municipality="De Kruis" pageId="1" pageNumber="144" specimenCount="2" typeStatus="holotype">
<emphasis id="84A94819FFC5FFBFFED4F7D1A1AD89EC" bold="true" box="[382,491,1411,1437]" pageId="1" pageNumber="144">
<typeStatus id="69662AA9FFC5FFBFFED4F7D1A1AD89EC" box="[382,491,1411,1437]" pageId="1" pageNumber="144" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<date id="C263B2CBFFC5FFBFFDFCF7D1A29789EC" box="[598,721,1411,1437]" pageId="1" pageNumber="144" value="2023-09-18">
<collectingDate id="D2274B23FFC5FFBFFDFCF7D1A29789EC" box="[598,721,1411,1437]" pageId="1" pageNumber="144" value="2023-09-18">18/ix/2023</collectingDate>
</date>
<collectingCountry id="CECAD49BFFC5FFBFFD73F7D1A32F89EC" box="[729,873,1411,1437]" name="South Africa" pageId="1" pageNumber="144">South Africa</collectingCountry>
NC//
<location id="B302C2D0FFC5FFBFFC05F7D1A47089EC" LSID="urn:lsid:plazi:treatment:3E74251DFFC5FFBAFF3DF69CA09D8B30:B302C2D0FFC5FFBFFC05F7D1A47089EC" box="[943,1078,1411,1437]" country="South Africa" county="Leliefon" municipality="De Kruis" name="Kamiesberg" pageId="1" pageNumber="144">Kamiesberg</location>
—stream beside// road from
<collectingMunicipality id="56060E71FFC5FFBFFAD6F7D1A09389B0" pageId="1" pageNumber="144">De Kruis</collectingMunicipality>
to
<collectingCounty id="5F03EC87FFC5FFBFFF53F7F5A11189B0" box="[249,343,1447,1473]" pageId="1" pageNumber="144">Leliefon</collectingCounty>
// -tein 302137.39S 180701.13E//
<quantity id="712539EEFFC5FFBFFD7FF7F5A36F89B0" box="[725,809,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="1" pageNumber="144" unit="m" value="1168.0">
<elevation id="3DF07338FFC5FFBFFD7FF7F5A36F89B0" box="[725,809,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.168" pageId="1" pageNumber="144" unit="m" value="1168.0">1168 m</elevation>
</quantity>
<collectorName id="1B28F1DDFFC5FFBFFC9AF7F5A3EE89B0" box="[816,936,1447,1473]" pageId="1" pageNumber="144">D T Bilton</collectorName>
leg.” and red
<typeStatus id="69662AA9FFC5FFBFFBECF7F5A4EE89B0" box="[1094,1192,1447,1473]" pageId="1" pageNumber="144" type="holotype">holotype</typeStatus>
label (
<collectionCode id="D0CC0CCEFFC5FFBFFB5EF7F5A57A89B0" box="[1268,1340,1447,1473]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="1" pageNumber="144">AMG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B662940BFFC5FFBFFF6DF799A4738994" blockId="1.[151,1437,1338,2013]" box="[199,1077,1482,1509]" pageId="1" pageNumber="144">
<materialsCitation id="06B59E56FFC5FFBFFF6DF799A4778994" box="[199,1073,1482,1509]" collectingDate="2023-09-18" collectionCode="AMG, CDTB, NMPC, NMW" collectorName="D T Bilton" country="South Africa" county="Leliefon" elevation="1168" location="Kamiesberg" municipality="De Kruis" pageId="1" pageNumber="144" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<emphasis id="84A94819FFC5FFBFFF6DF799A1798994" bold="true" box="[199,319,1483,1509]" pageId="1" pageNumber="144">
<typeStatus id="69662AA9FFC5FFBFFF6DF799A1798994" box="[199,319,1483,1509]" pageId="1" pageNumber="144" type="paratype">Paratypes</typeStatus>
</emphasis>
(5):
<specimenCount id="A0DB5F82FFC5FFBFFEDFF799A1DD8994" box="[373,411,1483,1509]" count="2" pageId="1" pageNumber="144" type="male">2♂</specimenCount>
,
<specimenCount id="A0DB5F82FFC5FFBFFE0CF799A18F8994" box="[422,457,1483,1509]" count="3" pageId="1" pageNumber="144" type="female">3♀</specimenCount>
same data as holotype (
<collectionCode id="D0CC0CCEFFC5FFBFFD7CF799A35A8994" box="[726,796,1483,1509]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="1" pageNumber="144">AMG</collectionCode>
,
<collectionCode id="D0CC0CCEFFC5FFBFFC8DF799A3328994" box="[807,884,1483,1509]" pageId="1" pageNumber="144">CDTB</collectionCode>
,
<collectionCode id="D0CC0CCEFFC5FFBFFCD5F799A3978994" box="[895,977,1483,1509]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="144">NMPC</collectionCode>
,
<collectionCode id="D0CC0CCEFFC5FFBFFC77F799A46C8994" box="[989,1066,1483,1509]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="144">NMW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEC7C780FFC5FFBDFF6DF7BCA3758A82" lastPageId="3" lastPageNumber="146" pageId="1" pageNumber="144" type="description">
<paragraph id="B662940BFFC5FFBFFF6DF7BCA50E8A5C" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
<emphasis id="84A94819FFC5FFBFFF6DF7BCA11F8A79" bold="true" box="[199,345,1518,1544]" pageId="1" pageNumber="144">Description.</emphasis>
Size:
<typeStatus id="69662AA9FFC5FFBFFE0AF7BDA24B8A78" box="[416,525,1519,1545]" pageId="1" pageNumber="144" type="holotype">Holotype</typeStatus>
: BL
<quantity id="712539EEFFC5FFBFFDE9F7BCA2ED8A78" box="[579,683,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="144" unit="mm" value="5.8">5.80 mm</quantity>
; EL
<quantity id="712539EEFFC5FFBFFD4AF7BDA30E8A78" box="[736,840,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="144" unit="mm" value="3.8">3.80 mm</quantity>
; EW 03.20 mm.
<typeStatus id="69662AA9FFC5FFBFFBA2F7BDA43F8A78" box="[1032,1145,1519,1545]" pageId="1" pageNumber="144" type="paratype">Paratypes</typeStatus>
: Male BL
<quantity id="712539EEFFC5FFBFFB5BF7BCA5DE8A78" box="[1265,1432,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.3" metricValueMin="5.2" pageId="1" pageNumber="144" unit="mm" value="5.25" valueMax="5.3" valueMin="5.2">5.20–5.30 mm</quantity>
; EL
<quantity id="712539EEFFC5FFBFFF15F440A1238A5C" box="[191,357,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="3.375" metricValueMax="3.45" metricValueMin="3.3" pageId="1" pageNumber="144" unit="mm" value="3.375" valueMax="3.45" valueMin="3.3">3.30–3.45 mm</quantity>
; EW
<quantity id="712539EEFFC5FFBFFE09F440A20F8A5C" box="[419,585,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.15" metricValueMin="3.05" pageId="1" pageNumber="144" unit="mm" value="3.0999999999999996" valueMax="3.15" valueMin="3.05">3.05–3.15 mm</quantity>
. Females BL
<quantity id="712539EEFFC5FFBFFD4BF441A3C18A5C" box="[737,903,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.4" metricValueMin="6.3" pageId="1" pageNumber="144" unit="mm" value="6.35" valueMax="6.4" valueMin="6.3">6.30–6.40 mm</quantity>
; EL
<quantity id="712539EEFFC5FFBFFC11F440A4278A5C" box="[955,1121,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.25" metricValueMin="4.05" pageId="1" pageNumber="144" unit="mm" value="4.15" valueMax="4.25" valueMin="4.05">4.05–4.25 mm</quantity>
; EW
<quantity id="712539EEFFC5FFBFFB34F440A5028A5C" box="[1182,1348,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="3.375" metricValueMax="3.45" metricValueMin="3.3" pageId="1" pageNumber="144" unit="mm" value="3.375" valueMax="3.45" valueMin="3.3">3.30–3.45 mm</quantity>
.
</paragraph>
<paragraph id="B662940BFFC5FFBFFF6DF465A3058AE8" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
Colour: Dorsum (
<figureCitation id="2EE6888EFFC5FFBFFE3EF465A1AB8A20" box="[404,493,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="1" pageNumber="144">Fig. 1A</figureCitation>
) dark brown to black; pronotal margins paler, reddish brown. Antennal stem and maxillary and labial palpi and yellowish brown, antennal club darker; legs reddish brown, tarsi paler. Venter (
<figureCitation id="2EE6888EFFC5FFBFFAC5F409A0E08AE8" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="1" pageNumber="144">Fig. 2</figureCitation>
) reddish brown; underside of head and metaventrite darker.
</paragraph>
<paragraph id="B662940BFFC5FFBFFF6DF4F1A50B8B3C" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
Head: Labrum (
<figureCitation id="2EE6888EFFC5FFBFFED4F4F1A2558ACC" box="[382,531,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="1" pageNumber="144">Fig. 1A &amp; C</figureCitation>
) strongly transverse; anterior margin with broad, shallow emargination; dorsal surface shining, microreticulation absent, with sparse, fine punctation, punctures bearing long, fine adpressed setae in basal half. Clypeus, frons and vertex shining; without microreticulation, with sparse, shallow, medium punctures bearing short, peg-like sensillae.Compound eyes moderately large; inter-ocular distance approximately 5x transverse diameter of eye. Maxillary palpi moderately long, last palpomere cylindrical, penultimate widened apically.
</paragraph>
<paragraph id="B662940BFFC5FFBFFF6DF505A2BF8BAC" blockId="1.[151,1437,1338,2013]" pageId="1" pageNumber="144">
Pronotum: Transverse, widest at base (
<figureCitation id="2EE6888EFFC5FFBFFDDCF505A2898B00" box="[630,719,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="1" pageNumber="144">Fig. 1A</figureCitation>
). Anterior margin strongly emarginated, emargination sinuate in centre; posterior margin strongly bisinuate around centre; lateral margins strongly arcuately rounded. Anterior and posterior angles obtusely rounded. Surface strongly shining; microreticulation absent, with sparse, shallow, medium punctures bearing short, peg-like sensillae as on frons.
</paragraph>
<caption id="E2A2C483FFC6FFBCFF32F458A31E8AFE" ID-DOI="http://doi.org/10.5281/zenodo.11231041" ID-Zenodo-Dep="11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="2" pageNumber="145" startId="2.[152,255,1546,1570]" targetBox="[192,1394,189,1502]" targetPageId="2" targetType="figure">
<paragraph id="B662940BFFC6FFBCFF32F458A31E8AFE" blockId="2.[151,1437,1546,1679]" pageId="2" pageNumber="145">
<emphasis id="84A94819FFC6FFBCFF32F458A15C8A53" bold="true" box="[152,282,1546,1570]" pageId="2" pageNumber="145">FIGURE 1.</emphasis>
<taxonomicName id="71DDEF88FFC6FFBCFE8BF458A2658A52" box="[289,547,1546,1571]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="84A94819FFC6FFBCFE8BF458A1F58A53" box="[289,435,1546,1570]" italics="true" pageId="2" pageNumber="145">Relictorygmus</emphasis>
species. A
</taxonomicName>
)
<taxonomicName id="71DDEF88FFC6FFBCFD98F459A2E48A53" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[562,674,1547,1570]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC6FFBCFD98F459A2E48A53" box="[562,674,1547,1570]" italics="true" pageId="2" pageNumber="145">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC6FFBCFD03F459A2BC8A52" bold="true" box="[681,762,1547,1571]" pageId="2" pageNumber="145">
<taxonomicNameLabel id="9F9AF562FFC6FFBCFD03F459A2BC8A52" box="[681,762,1547,1571]" pageId="2" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, dorsal habitus; B)
<taxonomicName id="71DDEF88FFC6FFBCFB8DF459A4D08A53" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[1063,1174,1547,1570]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC6FFBCFB8DF459A4D08A53" box="[1063,1174,1547,1570]" italics="true" pageId="2" pageNumber="145">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC6FFBCFB37F459A4A98A52" bold="true" box="[1181,1263,1547,1571]" pageId="2" pageNumber="145">
<taxonomicNameLabel id="9F9AF562FFC6FFBCFB37F459A4A98A52" box="[1181,1263,1547,1571]" pageId="2" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, lateral habitus; C)
<taxonomicName id="71DDEF88FFC6FFBCFEBAF47DA1C68A37" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[272,384,1583,1606]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC6FFBCFEBAF47DA1C68A37" box="[272,384,1583,1606]" italics="true" pageId="2" pageNumber="145">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC6FFBCFE2DF47DA19F8A36" bold="true" box="[391,473,1583,1607]" pageId="2" pageNumber="145">
<taxonomicNameLabel id="9F9AF562FFC6FFBCFE2DF47DA19F8A36" box="[391,473,1583,1607]" pageId="2" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype frontal view of head; D)
<taxonomicName id="71DDEF88FFC6FFBCFCEEF47DA3F28A37" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[836,948,1583,1606]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC6FFBCFCEEF47DA3F28A37" box="[836,948,1583,1606]" italics="true" pageId="2" pageNumber="145">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC6FFBCFC16F47DA4488A36" bold="true" box="[956,1038,1583,1607]" pageId="2" pageNumber="145">
<taxonomicNameLabel id="9F9AF562FFC6FFBCFC16F47DA4488A36" box="[956,1038,1583,1607]" pageId="2" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype, aedeagus; E)
<taxonomicName id="71DDEF88FFC6FFBCFAA2F47DA2EA8A1A" authority="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek, 2018" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC6FFBCFAA2F47DA5DB8A37" box="[1288,1437,1582,1606]" italics="true" pageId="2" pageNumber="145">R. trevornoahi</emphasis>
<bibRefCitation id="D24CE9FAFFC6FFBCFF3DF400A2EA8A1A" author="Seidel, M. &amp; Minoshima, Y. N. &amp; Arriaga-Varela, E. &amp; Fikacek, M." box="[151,684,1618,1643]" pageId="2" pageNumber="145" pagination="375 - 402" refId="ref3959" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. &amp; Fikacek, M. (2018) Breaking a disjunct distribution: a review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375 - 402. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.2.011" type="journal article" year="2018">Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018</bibRefCitation>
</taxonomicName>
paratype, aedeagus; F)
<taxonomicName id="71DDEF88FFC6FFBCFC35F401A4958A1A" authority="(Hebauer, 2002)" baseAuthorityName="Hebauer" baseAuthorityYear="2002" box="[927,1235,1618,1643]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="repentinus">
<emphasis id="84A94819FFC6FFBCFC35F401A4608A1B" box="[927,1062,1619,1642]" italics="true" pageId="2" pageNumber="145">R. repentinus</emphasis>
(
<bibRefCitation id="D24CE9FAFFC6FFBCFB9EF400A48D8A1A" author="Hebauer, F." box="[1076,1227,1618,1643]" pageId="2" pageNumber="145" pagination="3 - 24" refId="ref3735" refString="Hebauer, F. (2002) New Hydrophilidae of the Old World (Coleoptera, Hydrophilidae). Acta Coleopterologica, 18, 3 - 24." type="journal article" year="2002">Hebauer, 2002</bibRefCitation>
)
</taxonomicName>
holotype, aedeagus (after
<bibRefCitation id="D24CE9FAFFC6FFBCFF7EF424A1E48AFE" author="Seidel, M. &amp; Minoshima, Y. N. &amp; Arriaga-Varela, E. &amp; Fikacek, M." box="[212,418,1654,1679]" pageId="2" pageNumber="145" pagination="375 - 402" refId="ref3959" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. &amp; Fikacek, M. (2018) Breaking a disjunct distribution: a review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375 - 402. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.2.011" type="journal article" year="2018">
Seidel
<emphasis id="84A94819FFC6FFBCFEB0F425A1018AFF" box="[282,327,1654,1678]" italics="true" pageId="2" pageNumber="145">et al</emphasis>
. (2018))
</bibRefCitation>
. Scale bars A–C) = 1 mm; D–F) = 100 μm.
</paragraph>
</caption>
<paragraph id="B662940BFFC6FFBCFF6DF4B3A1938BFA" blockId="2.[151,1436,1761,2039]" pageId="2" pageNumber="145">
Elytra: Slightly elongate (
<figureCitation id="2EE6888EFFC6FFBCFE4AF4B3A2708A8A" box="[480,566,1761,1787]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="2" pageNumber="145">Fig.1A</figureCitation>
); sides strongly arcuate, from shoulders to posterior angles.Ten seriate puncture rows, punctures slightly wider than punctures in intervals. Sutural puncture row striate impressed, particularly in posterior half (
<figureCitation id="2EE6888EFFC6FFBCFEC8F57BA1F88B32" box="[354,446,1833,1859]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="2" pageNumber="145">Figs 1A</figureCitation>
&amp;
<figureCitation id="2EE6888EFFC6FFBCFE43F57BA2488B32" box="[489,526,1833,1859]" captionStart="FIGURE 3" captionStartId="4.[152,255,1436,1460]" captionTargetBox="[154,1434,453,1413]" captionTargetId="figure-164@4.[154,1434,453,1413]" captionTargetPageId="4" captionText="FIGURE 3. Relictorygmus species, detail of pronotal and elytral punctation. A) R. riparius sp. nov.; B) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231046" httpUri="https://zenodo.org/record/11231046/files/figure.png" pageId="2" pageNumber="145">3A</figureCitation>
). Serial and interval punctures mostly relatively small and shallow; systematic punctures markedly larger and distinct (
<figureCitation id="2EE6888EFFC6FFBCFDF1F51FA2F18B16" box="[603,695,1869,1895]" captionStart="FIGURE 3" captionStartId="4.[152,255,1436,1460]" captionTargetBox="[154,1434,453,1413]" captionTargetId="figure-164@4.[154,1434,453,1413]" captionTargetPageId="4" captionText="FIGURE 3. Relictorygmus species, detail of pronotal and elytral punctation. A) R. riparius sp. nov.; B) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231046" httpUri="https://zenodo.org/record/11231046/files/figure.png" pageId="2" pageNumber="145">Fig. 3A</figureCitation>
). Highest towards rear of elytra; dorsal profile following a single arc in lateral view (
<figureCitation id="2EE6888EFFC6FFBCFEDAF523A1818BFA" box="[368,455,1905,1931]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="2" pageNumber="145">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="B662940BFFC6FFBDFF6DF5C7A4718DA0" blockId="2.[151,1436,1761,2039]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="146" pageId="2" pageNumber="145">
Venter: Mentum (
<figureCitation id="2EE6888EFFC6FFBCFE3BF5C7A1AF8BDE" box="[401,489,1941,1967]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
) with broadly truncate, dorsally-directed apicomedian projection anteriorly; broad, transverse depression posterior to projection, deepest anteriorly, boundary between the two areas appearing as a transverse ridge; surface shining, with sparse, shallow, medium punctures basolaterally. Submentum (
<figureCitation id="2EE6888EFFC6FFBCFA91F58FA5D28B86" box="[1339,1428,2013,2039]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="2" pageNumber="145">Fig. 2B</figureCitation>
) shining; smooth anteriorly, somewhat rugose posteriorly, with medium, shallow punctures throughout, each bearing a long, yellow-white arcuate seta. Genae shining, microreticulation absent. Gula shining, rugose; punctate and setose as submentum; median ridge present posteriorly. Prosternum (
<figureCitation id="2EE6888EFFC7FFBDFC3FF28DA3AC8C88" box="[917,1002,223,249]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Fig. 2C</figureCitation>
) coarsely microreticulate and rugosely punctate, punctures bearing long, arcuate yellow-white setae; posterior marginal bead broader than anterior, microreticulate; anterior and posterior marginal beads lacking punctures and setae. Mesoventrite (
<figureCitation id="2EE6888EFFC7FFBDFAB2F375A5D38D30" box="[1304,1429,295,321]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Fig. 2D–E</figureCitation>
) angulate, reaching coxal cavities laterally; elevated anterior part subtriangular with weakly sinuate posterior margin; main part rugosely punctate and setose medially; lacking transverse groove posteromesally. Raised median area of metaventrite (
<figureCitation id="2EE6888EFFC7FFBDFEF9F3C1A1EE8DDC" box="[339,424,403,429]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Fig. 2F</figureCitation>
) setose anteriorly and smooth and impunctate posteriorly, with clear longitudinal furrow. Anterior metaventral process wide, triangular, ca. 1.3× as long as maximum width.
</paragraph>
<caption id="E2A2C483FFC7FFBDFF32F71DA47089FA" ID-DOI="http://doi.org/10.5281/zenodo.11231044" ID-Zenodo-Dep="11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146" startId="3.[152,255,1359,1383]" targetBox="[154,1434,489,1336]" targetPageId="3" targetType="figure">
<paragraph id="B662940BFFC7FFBDFF32F71DA47089FA" blockId="3.[151,1436,1359,1419]" pageId="3" pageNumber="146">
<emphasis id="84A94819FFC7FFBDFF32F71DA15E8916" bold="true" box="[152,280,1359,1383]" pageId="3" pageNumber="146">FIGURE 2.</emphasis>
<taxonomicName id="71DDEF88FFC7FFBDFEB4F71DA24E8916" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[286,520,1359,1383]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC7FFBDFEB4F71DA24E8916" box="[286,520,1359,1383]" italics="true" pageId="3" pageNumber="146">Relictorygmus riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC7FFBDFDA4F71DA2188916" bold="true" box="[526,606,1359,1383]" pageId="3" pageNumber="146">
<taxonomicNameLabel id="9F9AF562FFC7FFBDFDA4F71DA2188916" box="[526,606,1359,1383]" pageId="3" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur.
</paragraph>
</caption>
<paragraph id="B662940BFFC7FFBDFF6DF78CA4A08A6A" blockId="3.[151,1437,1501,2032]" pageId="3" pageNumber="146">
Legs: Ventral surface of profemora (
<figureCitation id="2EE6888EFFC7FFBDFDCBF78FA2FF8986" box="[609,697,1501,1527]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Fig. 2G</figureCitation>
) moderately pubescent at base, particularly posteriorly, with very sparse, scattered setae to apex. Meso- and metafemora (
<figureCitation id="2EE6888EFFC7FFBDFCADF453A33C8A6A" box="[775,890,1537,1563]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Figs 2H–I</figureCitation>
) sparsely pubescent throughout.
</paragraph>
<paragraph id="B662940BFFC7FFBDFF6DF477A19B8ABE" blockId="3.[151,1437,1501,2032]" pageId="3" pageNumber="146">
Aedeagus: Elongate (
<figureCitation id="2EE6888EFFC7FFBDFE16F477A2578A4E" box="[444,529,1573,1599]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="3" pageNumber="146">Fig 1D</figureCitation>
). Phallobase rather weakly constricted basally; narrow basal part much shorter than broader anterior portion. Parameres shorter than phallobase, with outer faces arcuate basally, then straight and converging to truncately rounded apices; inner margins weakly arcuate. Median lobe relatively broad, not reaching paramere apices; tapering to truncately rounded apex over distal half; median strut relatively broad. Gonopore apex expanded, slightly subapical.
</paragraph>
<paragraph id="B662940BFFC7FFBDFF6DF48BA3758A82" blockId="3.[151,1437,1501,2032]" box="[199,819,1753,1780]" pageId="3" pageNumber="146">Female: Without obvious external differences to males.</paragraph>
</subSubSection>
<subSubSection id="FEC7C780FFC7FFBAFF6DF4AFA34D8DF8" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146" type="diagnosis">
<paragraph id="B662940BFFC7FFBDFF6DF4AFA1888B66" blockId="3.[151,1437,1501,2032]" box="[199,462,1789,1815]" pageId="3" pageNumber="146">
<emphasis id="84A94819FFC7FFBDFF6DF4AFA1888B66" bold="true" box="[199,462,1789,1815]" pageId="3" pageNumber="146">Differential diagnosis.</emphasis>
</paragraph>
<paragraph id="B662940BFFC7FFBAFF6DF570A34D8DF8" blockId="3.[151,1437,1501,2032]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146">
In overall habitus and structure of the genitalia and most external characters, the new species most closely resembles
<taxonomicName id="71DDEF88FFC7FFBDFEA5F514A1F78B2E" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" box="[271,433,1861,1887]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC7FFBDFEA5F514A1F78B2E" box="[271,433,1861,1887]" italics="true" pageId="3" pageNumber="146">R. trevornoahi</emphasis>
</taxonomicName>
. In common with this species, the ventral surface of the profemora is sparsely pubescent (
<figureCitation id="2EE6888EFFC7FFBDFF35F53BA0BD8BF2" box="[159,251,1897,1923]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="3" pageNumber="146">Fig. 2G</figureCitation>
), rather than densely pubescent at base as in
<taxonomicName id="71DDEF88FFC7FFBDFD52F538A3CB8BF2" baseAuthorityName="Hebauer" baseAuthorityYear="2002" box="[760,909,1898,1923]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="repentinus">
<emphasis id="84A94819FFC7FFBDFD52F538A3CB8BF2" box="[760,909,1898,1923]" italics="true" pageId="3" pageNumber="146">R. repentinus</emphasis>
</taxonomicName>
. It does, however, differ from this species in a number of ways, which are summarised here. The dorsal punctation of
<taxonomicName id="71DDEF88FFC7FFBDFC6BF5DCA47D8BD6" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[961,1083,1934,1959]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC7FFBDFC6BF5DCA47D8BD6" box="[961,1083,1934,1959]" italics="true" pageId="3" pageNumber="146">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC7FFBDFBEEF5DFA4D98BD6" bold="true" box="[1092,1183,1933,1959]" pageId="3" pageNumber="146">
<taxonomicNameLabel id="9F9AF562FFC7FFBDFBEEF5DFA4D98BD6" box="[1092,1183,1933,1959]" pageId="3" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is finer and shallower than that of
<taxonomicName id="71DDEF88FFC7FFBDFEB5F5E0A1848BBA" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" box="[287,450,1969,1995]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC7FFBDFEB5F5E0A1848BBA" box="[287,450,1969,1995]" italics="true" pageId="3" pageNumber="146">R. trevornoahi</emphasis>
</taxonomicName>
. This means that the systematic punctures of the elytra are more clearly distinguishable from background punctation in
<taxonomicName id="71DDEF88FFC7FFBDFDACF584A2C48B9E" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[518,642,2006,2031]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC7FFBDFDACF584A2C48B9E" box="[518,642,2006,2031]" italics="true" pageId="3" pageNumber="146">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC7FFBDFD26F587A2AF8B9E" bold="true" box="[652,745,2005,2031]" pageId="3" pageNumber="146">
<taxonomicNameLabel id="9F9AF562FFC7FFBDFD26F587A2AF8B9E" box="[652,745,2005,2031]" pageId="3" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
In addition, the serial punctures are more markedly striateimpressed in
<taxonomicName id="71DDEF88FFC0FFBAFE9BF2C5A1938CC0" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" box="[305,469,151,177]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC0FFBAFE9BF2C5A1938CC0" box="[305,469,151,177]" italics="true" pageId="4" pageNumber="147">R. trevornoahi</emphasis>
</taxonomicName>
compared to the new species (
<figureCitation id="2EE6888EFFC0FFBAFC90F2C5A3C78CC0" box="[826,897,151,177]" captionStart="FIGURE 3" captionStartId="4.[152,255,1436,1460]" captionTargetBox="[154,1434,453,1413]" captionTargetId="figure-164@4.[154,1434,453,1413]" captionTargetPageId="4" captionText="FIGURE 3. Relictorygmus species, detail of pronotal and elytral punctation. A) R. riparius sp. nov.; B) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231046" httpUri="https://zenodo.org/record/11231046/files/figure.png" pageId="4" pageNumber="147">Fig. 3</figureCitation>
). The excavation on the mentum is deeper and narrower in
<taxonomicName id="71DDEF88FFC0FFBAFE8BF2E9A1DC8CA5" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[289,410,187,212]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC0FFBAFE8BF2E9A1DC8CA5" box="[289,410,187,212]" italics="true" pageId="4" pageNumber="147">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC0FFBAFE0BF2E9A1BD8CA4" bold="true" box="[417,507,187,213]" pageId="4" pageNumber="147">
<taxonomicNameLabel id="9F9AF562FFC0FFBAFE0BF2E9A1BD8CA4" box="[417,507,187,213]" pageId="4" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
when compared with
<taxonomicName id="71DDEF88FFC0FFBAFD5FF2E9A3DE8CA4" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" box="[757,920,187,213]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC0FFBAFD5FF2E9A3DE8CA4" box="[757,920,187,213]" italics="true" pageId="4" pageNumber="147">R. trevornoahi</emphasis>
</taxonomicName>
. The prosternum of the new species (
<figureCitation id="2EE6888EFFC0FFBAFA94F2E9A5D28CA4" box="[1342,1428,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="4" pageNumber="147">Fig. 2C</figureCitation>
) is also more setose and rugose, the raised anterior portion of its mesoventrite is triangular (
<figureCitation id="2EE6888EFFC0FFBAFB33F28DA5578C88" box="[1177,1297,223,249]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="4" pageNumber="147">Fig. 2D–E</figureCitation>
) as opposed to heart-shaped and its mesoventrite (
<figureCitation id="2EE6888EFFC0FFBAFDEAF351A2DE8D6C" box="[576,664,259,285]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="4" pageNumber="147">Fig. 2D</figureCitation>
) lacks an obvious transverse groove posteromesally. The punctation on the ventral face of the femora of
<taxonomicName id="71DDEF88FFC0FFBAFD8EF375A2808D30" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" box="[548,710,295,321]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC0FFBAFD8EF375A2808D30" box="[548,710,295,321]" italics="true" pageId="4" pageNumber="147">R. trevornoahi</emphasis>
</taxonomicName>
is denser, with coarser setae, than in the new species (
<figureCitation id="2EE6888EFFC0FFBAFAB5F375A5D78D30" box="[1311,1425,295,321]" captionStart="FIGURE 2" captionStartId="3.[152,255,1359,1383]" captionTargetBox="[154,1434,489,1336]" captionTargetId="figure-139@3.[154,1434,489,1336]" captionTargetPageId="3" captionText="FIGURE 2. Relictorygmus riparius sp. nov., paratype. A) venter; B) ventral view of head; C) prosetrnum; D) mesoventrite; E) detail of anterior mesoventrite; F) metaventrite; G) profemur; H) mesofemur; I) metafemur." figureDoi="http://doi.org/10.5281/zenodo.11231044" httpUri="https://zenodo.org/record/11231044/files/figure.png" pageId="4" pageNumber="147">Fig. 2G–I</figureCitation>
). Lastly, the aedeagus differs from both previously described species in a number of details, including the relative lengths and shape of the different elements (
<figureCitation id="2EE6888EFFC0FFBAFD2DF33DA2B88DF8" box="[647,766,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1546,1570]" captionTargetBox="[192,1394,189,1502]" captionTargetId="figure-27@2.[180,1408,179,1520]" captionTargetPageId="2" captionText="FIGURE 1. Relictorygmus species. A) R. riparius sp. nov. holotype, dorsal habitus; B) R. riparius sp. nov. holotype, lateral habitus; C) R. riparius sp. nov. holotype frontal view of head; D) R. riparius sp. nov. holotype, aedeagus; E) R. trevornoahi Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018 paratype, aedeagus; F) R. repentinus (Hebauer, 2002) holotype, aedeagus (after Seidel et al. (2018)). Scale bars A–C) = 1 mm; D–F) = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.11231041" httpUri="https://zenodo.org/record/11231041/files/figure.png" pageId="4" pageNumber="147">Fig. 1D–F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FEC7C780FFC0FFBAFF6DF3C1A4C18DDC" box="[199,1159,403,429]" pageId="4" pageNumber="147" type="etymology">
<paragraph id="B662940BFFC0FFBAFF6DF3C1A4C18DDC" blockId="4.[151,1437,151,429]" box="[199,1159,403,429]" pageId="4" pageNumber="147">
<emphasis id="84A94819FFC0FFBAFF6DF3C1A1088DDC" bold="true" box="[199,334,403,429]" pageId="4" pageNumber="147">Etymology.</emphasis>
Named in reference to the habitat, at the banks of a small mountain river.
</paragraph>
</subSubSection>
<caption id="E2A2C483FFC0FFBAFF32F7CEA22589A8" ID-DOI="http://doi.org/10.5281/zenodo.11231046" ID-Zenodo-Dep="11231046" httpUri="https://zenodo.org/record/11231046/files/figure.png" pageId="4" pageNumber="147" startId="4.[152,255,1436,1460]" targetBox="[154,1434,453,1413]" targetPageId="4" targetType="figure">
<paragraph id="B662940BFFC0FFBAFF32F7CEA22589A8" blockId="4.[151,1436,1436,1497]" pageId="4" pageNumber="147">
<emphasis id="84A94819FFC0FFBAFF32F7CEA15E89C5" bold="true" box="[152,280,1436,1460]" pageId="4" pageNumber="147">FIGURE 3.</emphasis>
<taxonomicName id="71DDEF88FFC0FFBAFEB5F7CEA24589C5" box="[287,515,1436,1460]" pageId="4" pageNumber="147">
<emphasis id="84A94819FFC0FFBAFEB5F7CEA1F789C5" box="[287,433,1436,1460]" italics="true" pageId="4" pageNumber="147">Relictorygmus</emphasis>
species
</taxonomicName>
, detail of pronotal and elytral punctation. A)
<taxonomicName id="71DDEF88FFC0FFBAFC61F7CFA47F89C5" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[971,1081,1437,1460]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC0FFBAFC61F7CFA47F89C5" box="[971,1081,1437,1460]" italics="true" pageId="4" pageNumber="147">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC0FFBAFB95F7CFA4D689C4" bold="true" box="[1087,1168,1437,1461]" pageId="4" pageNumber="147">
<taxonomicNameLabel id="9F9AF562FFC0FFBAFB95F7CFA4D689C4" box="[1087,1168,1437,1461]" pageId="4" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B)
<taxonomicName id="71DDEF88FFC0FFBAFB17F7CFA22689A8" authority="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek, 2018" authorityName="Seidel, Minoshima, Arriaga-Varela &amp; Fikacek" authorityYear="2018" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="trevornoahi">
<emphasis id="84A94819FFC0FFBAFB17F7CFA51789C5" box="[1213,1361,1436,1460]" italics="true" pageId="4" pageNumber="147">R. trevornoahi</emphasis>
<bibRefCitation id="D24CE9FAFFC0FFBAFAFDF7CEA22689A8" author="Seidel, M. &amp; Minoshima, Y. N. &amp; Arriaga-Varela, E. &amp; Fikacek, M." pageId="4" pageNumber="147" pagination="375 - 402" refId="ref3959" refString="Seidel, M., Minoshima, Y. N., Arriaga-Varela, E. &amp; Fikacek, M. (2018) Breaking a disjunct distribution: a review of the Southern Hemisphere genera Cylorygmus and Relictorygmus gen. nov. (Hydrophilidae: Cylominae). Annales Zoologici, 68, 375 - 402. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.2.011" type="journal article" year="2018">Seidel, Minoshima, Arriaga-Varela &amp; Fikáček, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="FEC7C780FFC0FFBAFF6DF478A09D8B30" pageId="4" pageNumber="147" type="distribution">
<paragraph id="B662940BFFC0FFBAFF6DF478A09D8B30" blockId="4.[151,1437,1578,1857]" pageId="4" pageNumber="147">
<emphasis id="84A94819FFC0FFBAFF6DF478A1B18A34" bold="true" box="[199,503,1578,1605]" pageId="4" pageNumber="147">Distribution and ecology.</emphasis>
Known only from the
<typeStatus id="69662AA9FFC0FFBAFCABF479A3778A34" box="[769,817,1579,1605]" pageId="4" pageNumber="147">type</typeStatus>
locality (
<figureCitation id="2EE6888EFFC0FFBAFC0AF479A3A18A34" box="[928,999,1579,1605]" captionStart="FIGURE 4" captionStartId="5.[152,255,1891,1915]" captionTargetBox="[341,1247,181,1866]" captionTargetId="figure-17@5.[341,1247,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Relictorygmus habitat and distribution. A) Type locality of R. riparius sp. nov., South Africa, Northern Cape Province, Kamiesberg (photo Andy Foggo); B) distribution of Relictorygmus species in South Africa. Green symbol = R. riparius sp. nov., red symbol = R. repentinus (Hebauer, 2002), yellow symbol = R. trevornoahi Seidel, Minoshima, Arriaga- Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231048" httpUri="https://zenodo.org/record/11231048/files/figure.png" pageId="4" pageNumber="147">Fig. 4</figureCitation>
), beside a small mountain river in the Kamiesberg,
<collectingRegion id="74195AE9FFC0FFBAFE87F41DA2058A18" box="[301,579,1615,1641]" country="South Africa" name="Northern Cape" pageId="4" pageNumber="147">Northern Cape Province</collectingRegion>
,
<collectingCountry id="CECAD49BFFC0FFBAFDE5F41DA2A58A18" box="[591,739,1615,1641]" name="South Africa" pageId="4" pageNumber="147">South Africa</collectingCountry>
. Beetles were found living in mats of wet moss, immediately beside the stream at the time of sampling, and within areas of seasonal seepage on peaty soil (
<figureCitation id="2EE6888EFFC0FFBAFB44F421A50A8AFC" box="[1262,1356,1651,1677]" captionStart="FIGURE 4" captionStartId="5.[152,255,1891,1915]" captionTargetBox="[341,1247,181,1866]" captionTargetId="figure-17@5.[341,1247,181,1866]" captionTargetPageId="5" captionText="FIGURE 4. Relictorygmus habitat and distribution. A) Type locality of R. riparius sp. nov., South Africa, Northern Cape Province, Kamiesberg (photo Andy Foggo); B) distribution of Relictorygmus species in South Africa. Green symbol = R. riparius sp. nov., red symbol = R. repentinus (Hebauer, 2002), yellow symbol = R. trevornoahi Seidel, Minoshima, Arriaga- Varela &amp; Fikáček, 2018." figureDoi="http://doi.org/10.5281/zenodo.11231048" httpUri="https://zenodo.org/record/11231048/files/figure.png" pageId="4" pageNumber="147">Fig. 4A</figureCitation>
). They were revealed when moss was removed and wetted, allowing them to float free of the microhabitat. At the time of collection it was assumed that this was a species of
<taxonomicName id="71DDEF88FFC0FFBAFD4AF4E9A3208AA4" authorityName="Brulle" authorityYear="1835" box="[736,870,1723,1749]" class="Insecta" family="Hydrophilidae" genus="Coelostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="84A94819FFC0FFBAFD4AF4E9A3208AA4" box="[736,870,1723,1749]" italics="true" pageId="4" pageNumber="147">Coelostoma</emphasis>
</taxonomicName>
, meaning many additional specimens (at least 10) were released. The only other beetle present in this microhabitat was
<taxonomicName id="71DDEF88FFC0FFBAFC0EF48DA5218A88" authority="Komarek, 2004" authorityName="Komarek" authorityYear="2004" box="[932,1383,1759,1785]" class="Insecta" family="Hydrophilidae" genus="Anacaena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="glabriventris">
<emphasis id="84A94819FFC0FFBAFC0EF48DA4EB8A88" box="[932,1197,1759,1785]" italics="true" pageId="4" pageNumber="147">Anacaena glabriventris</emphasis>
Komarek, 2004
</taxonomicName>
. The limited hydrofuge pubescence on the femora suggests that the respiratory bubble of
<taxonomicName id="71DDEF88FFC0FFBAFB96F551A4F28B6D" authority="Bilton &amp; Mlambo, 2024" authorityName="Bilton &amp; Mlambo" authorityYear="2024" box="[1084,1204,1795,1820]" class="Insecta" family="Hydrophilidae" genus="Relictorygmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="riparius" status="sp. nov.">
<emphasis id="84A94819FFC0FFBAFB96F551A4F28B6D" box="[1084,1204,1795,1820]" italics="true" pageId="4" pageNumber="147">R. riparius</emphasis>
</taxonomicName>
<emphasis id="84A94819FFC0FFBAFB10F551A5558B6C" bold="true" box="[1210,1299,1795,1821]" pageId="4" pageNumber="147">
<taxonomicNameLabel id="9F9AF562FFC0FFBAFB10F551A5558B6C" box="[1210,1299,1795,1821]" pageId="4" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is relatively small.
</paragraph>
</subSubSection>
</treatment>
</document>
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