treatments-xml/data/D7/CA/01/D7CA01C826CA5B47A9138A3BE8FE53CB.xml

<document id="2FCA24A1817D86AE0A9BBC2894E7C0F7" ID-DOI="10.3897/zookeys.1209.125897" ID-publisher-id="125897" URI-arpha="6ED9C06E-4DBB-5A79-A98B-FE427E12ED58" URI-zoobank="http://zoobank.org/CEE78803-61EB-40CC-8D63-46142E6383A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1723117912200" checkinUser="pensoft" docAuthor="Harden, Curt W. &amp; Caterino, Michael S." docDate="2024" docId="D7CA01C826CA5B47A9138A3BE8FE53CB" docLanguage="en" docName="ZooKeys 1209: 69-197" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/125897/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Serranillus jeanneli Barr 1995" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="197" masterDocId="6ED9C06E4DBB5A79A98BFE427E12ED58" masterDocTitle="Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" masterLastPageNumber="197" masterPageNumber="69" pageNumber="69" updateTime="1723146111653" updateUser="admin">
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<mods:title id="D9CFF0BEBA2AA950B9BDA4BFD77A6FB1">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
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<mods:namePart id="DC5ED63123410201BB9AAB15DA4F0340">Harden, Curt W.</mods:namePart>
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<mods:affiliation id="0850F2D851BECC727CBF338633CE8EDA">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:namePart id="FCC26354722E2B34DA4C9DA470331B12">Caterino, Michael S.</mods:namePart>
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<mods:affiliation id="EB00EBD677333A674B8E9F68B3C04070">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:date id="23D20EA18154E943429917A5BA9F4F05">2024</mods:date>
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<mods:number id="76737446C2537AA6EC661CE9F919AB3B">1209</mods:number>
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<paragraph id="DDEA02394DF976B0A53EF66EE9E12E36">
<taxonomicName id="60A431A7DE0B0A62353725D1DD0AD330" ID-CoL="6YL78" authority="Barr, 1995" authorityName="Barr" authorityYear="1995" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="95947B2D0ACA07D90A380EEC3C36FE56" italics="true">Serranillus jeanneli</emphasis>
<bibRefCitation id="097414371E89F0CB12C7DF1C1F109C36" author="Barr Jr TC" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239 - 248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr, 1995</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="7C24848CA5E457E8213BF082D0237B72" type="description">
<paragraph id="E2CFCB8DF68B051C4A7EE47646B38311">
<figureCitation id="D84AC93CE5BFF90C0616C9207B654AC2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of eastern Nearctic Anillini. A Serranillus dunavani (South Carolina, Oconee Co.) B Anillinus moseleyae (Tennessee, Sevier Co.) C Anillinus dentatus (South Carolina, Abbeville Co.) D Serranillus jeanneli (South Carolina, Oconee Co.) E Anillinus folkertsioides (Alabama, Blount Co.) F Anillinus pecki (North Carolina, Avery Co.) G Anillinus sp. “ Georgia, Barnes Creek sp. 1 ” H Anillinus montrex (South Carolina, York Co.) I Anillinus sp. “ Tennessee, Kings Saltpeter Cave ” J Anillinus virginiae (Virginia, Bath Co.) K Anillinus erwini (Virginia, Grayson Co.) L Serranillus sp. (North Carolina, Graham Co.) M Anillinus sp. “ South Carolina, Waldrop Stone ” (South Carolina, Pickens Co.) N Anillinus sp. “ Alabama, Horseshoe Cave sp. 1 ” O Anillinus loweae (North Carolina, Haywood Co.) P Anillinus sp. “ Kentucky, Hestand sp. 1 ” Q Anillinus indianae (Indiana, Lawrence Co.) R Anillinus cf. barberi (West Virginia, Greenbrier Co.) S Anillinus hirsutus (Alabama, Madison Co.). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1209.125897.figure2" httpUri="https://binary.pensoft.net/fig/1110341">Figs 2 D</figureCitation>
, 
<figureCitation id="02E5E89818344EF0297575F70A617FB5" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution map of Serranillus species that occur in South Carolina. The locality for Serranillus sp. “ South Carolina, Coon Branch ” is the same as the single South Carolina S. jeanneli occurrence, and is not shown. Data are from Harden (2024)." figureDoi="10.3897/zookeys.1209.125897.figure16" httpUri="https://binary.pensoft.net/fig/1110355">16</figureCitation>
, 
<figureCitation id="02DFC66FFCE3F6426627658E3E654347" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Multiple aspects of median lobes of South Carolina Serranillus species, progressively rotated dorsally from right lateral aspect (upper left) to left lateroventral aspect (lower right) A Serranillus dunavani B Serranillus jeanneli C Serranillus monadnock, holotype (Kings Mountain, SC) D Serranillus monadnock, paratype (Little Mountain, SC). Photographs not to same scale." figureDoi="10.3897/zookeys.1209.125897.figure17" httpUri="https://binary.pensoft.net/fig/1110356">17 B</figureCitation>
, 
<figureCitation id="F6E2168AD672FC6AEEC99476DC63168F" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 A – D</figureCitation>
, 
<figureCitation id="6C138E01DC7CE9D7C705534AB6F0699A" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 T</figureCitation>
, 
<figureCitation id="E1E56D5A9DA1C2C32B3B6D92AD3381D4" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Anillini habitat in South Carolina A Waldrop Stone Falls, Pickens Co. (Anillinus mica, Anillinus micamicus, Anillinus murrayae, Serranillus dunavani) B Lower Whitewater River gorge, Oconee Co. (Anillinus cherokee, A. murrayae, Anillinus sp. “ South Carolina, Coon Branch ”, S. dunavani, Serranillus jeanneli, Serranillus sp. “ South Carolina, Coon Branch ” C Long Cane Creek, Abbeville Co. (Anillinus chandleri, Anillinus dentatus, Anillinus jancae, S. dunavani). The holotypes of A. mica and A. micamicus were collected under embedded rocks uphill of the wooden staircase in A. The holotype of A. jancae was collected in a pipe trap set on a hill just out of view in C." figureDoi="10.3897/zookeys.1209.125897.figure25" httpUri="https://binary.pensoft.net/fig/1110364">25 B</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="FC81242A111E60601BB4378CD3F71A60">
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<treatmentCitation id="8315446E0672C675CE8B6D5FE98DD4B3" author="Barr Jr TC" page="247" year="1995">
<taxonomicName id="FE401D0101851EA6364A5767B54CA8D8" ID-CoL="6YL78" authority="Barr 1995: 247" authorityName="Barr" authorityPageNumber="247" authorityYear="1995" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="ABC2B33D348DC31C9A9578A781327229" italics="true">Serranillus jeanneli</emphasis>
<comment id="A7E424F9EF7AB896F7D485BABF407C52">
<bibRefCitation id="B7E54BE85B2C80492AD5133E358466F5" author="Barr Jr TC" firstAuthor="Barr" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" page="247" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239–248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr 1995: 247</bibRefCitation>
.
</comment>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EFPDK" type="Neotype male">
<paragraph id="A1C4425019D206BCE7CE1ECBC9A5E6F2">
<heading id="883FBF6E3C2BDBB478667371EF3DF542" reason="title">
<typeStatus id="218CA986F2594F616C0009A992107877" type="neotype">Neotype</typeStatus>
male
</heading>
</paragraph>
<paragraph id="CE23332311549626A4F41C851081566F">
<materialsCitation id="6C3A8E52A58C11FCA203FF8EE2DF00F2" collectingDate="1969-08-13" collectionCode="CMNH" collectorName="T. Barr" county="Macon Co." determinerName="C. W. Harden" location="Coweeta Exp. Sta. Ball Creek 3700" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<emphasis id="DB5B861175225D2FEBC8A740398DD840" bold="true">
(
<named-content id="D4C360E7176563A11A40F733FEBDA0E0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="A04382AE5986CA9283A8B8FF52AE04FA" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
)
</emphasis>
, 
<emphasis id="81E725E8D1A2D9A7F4A34810AAE60845" bold="true">here designated.</emphasis>
Dissected, with abdomen glued to point and genitalia in glycerin in plastic microvial pinned below labels, labeled: “ NC: 
<collectingCounty id="9BE7116BEDA80802AAFBCD550DC4B247">Macon Co.</collectingCounty>
# 62 
<location id="F3783624BC55C0315C37681106046C9E" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:F3783624BC55C0315C37681106046C9E" county="Macon Co." name="Coweeta Exp. Sta. Ball Creek 3700">Coweeta Exp. Sta. Ball Creek 3700</location>
’ 
<date id="BB859EFF6462482690E81A050FB28C94" value="1969-08-13">
<collectingDate id="DD3B0CEBB36F05CF5F48A83EB97CB7F0" value="1969-08-13">13 Aug 1969</collectingDate>
</date>
<collectorName id="EBAE8DCD8ED848684EFDE5E79D8139E5">T. Barr</collectorName>
” “ THOMAS C. BARR COLLECTION 2011 Acc. No. 38,014 ” “ 
<taxonomicName id="9315B86395C1D6801C0C19CBE9689877" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="9C7008DEDA88337CD5351480507753E0" italics="true">Serranillus jeanneli</emphasis>
</taxonomicName>
<specimenCount id="4A6BE258DCD4CDE8CADD2D4EB968164C" count="1" type="male">♂</specimenCount>
det. 
<determinerName id="CD99A01FD23BB3709DF2DEBE7BB5569B">
C. W. 
<bibRefCitation id="92F1315864D1B5F323ADAD2029AE8009" DOI="10.5281/zenodo.10983000" author="Harden" firstAuthor="Harden" refId="B39" refString="Harden CW (2024) Distribution of Nearctic Anillini (Coleoptera, Carabidae, Trechinae) (Version 01) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.10983000" year="2024">Harden 2024</bibRefCitation>
</determinerName>
” “ 
<typeStatus id="F376C6F5D3A94C491E57C044B480D4B1" type="neotype">NEOTYPE</typeStatus>
<taxonomicName id="94EE30AFB20430CC8FB1F5D592FA12FF" authority="Barr, 1995" authorityName="Barr" authorityYear="1995" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<originalDetermination id="577C434559B186A75D082759AC4DCFD0">
<emphasis id="41EDCE8F32CF517ADEB6B953CF450892" italics="true">Serranillus jeanneli</emphasis>
<bibRefCitation id="41968285A9AD872226807A8ED0CF6D02" author="Barr Jr TC" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239 - 248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr, 1995</bibRefCitation>
</originalDetermination>
</taxonomicName>
des. 
<bibRefCitation id="D08687B6F9E2CF125995E7C59CE008D8" DOI="10.1093/isd/ixae009" author="Caterino MS &amp; Harden CW" refId="B15" refString="Caterino MS, Harden CW (2024) A review of Appalachian Dasycerus Brongniart and the recognition of cryptic diversity within Dasycerus carolinensis (Coleoptera: Staphylinidae: Dasycerinae). Insect Systematics and Diversity 8 (2) [ixae 009]: 1 - 22. https: // doi. org / 10.1093 / isd / ixae 009" year="2024">Harden &amp; Caterino 2024</bibRefCitation>
[red cardstock] ”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EOQDK" type="material">
<paragraph id="F0D995071C212834DA6F6F2563F50CA7">
<heading id="2B59AD805856F364E2FE462D59A0CC92" reason="title">Material examined</heading>
</paragraph>
<paragraph id="1318FF9CAC32FD393001432350A5DCAE">
<emphasis id="AD8975D4F5B7A599F3DB317C61B26EA0" bold="true">
(
<emphasis id="0019D25BB8DE28A0A90C16873D206EDD" bold="true" italics="true">n</emphasis>
= 67). 
<materialsCitation id="FDAE3B3E38C6D72EC2E3A9F7093060A2" collectingDate="2021-05-11" collectionCode="CUAC" collectorName="M. Caterino &amp; A. Haberski" country="United States of America" county="Rabun Co." latitude="34.913" location="Rabun Cliffs" longLatPrecision="55" longitude="-83.2978" municipality="Chattahoochee National Forest" specimenCode="CUAC 000135496" specimenCount="1" specimenCount-male="1" stateProvince="Georgia">
<collectingCountry id="01B5BA2924A7CA1DA149388CD6D2A4CC" name="United States of America">USA</collectingCountry>
• 
<emphasis id="DA8109A9B9E75C8EF69108C683CE4B66" bold="true">
<collectingRegion id="283CF88172254F0FF00AA4A53DD4AE82" country="United States of America" name="Georgia">Georgia</collectingRegion>
</emphasis>
• 
<specimenCount id="F06C7304D3B22486DBF1DC110B7E1726" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="B78B08E72F67C37F38B0DB077A456AD0" bold="true">
<collectingCounty id="7019252DEEF70CB000F2DE4467DEE03E">Rabun Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="33BBDB014864F4BDBE01CC86B76B26EA">Chattahoochee National Forest</collectingMunicipality>
, 
<location id="F6873723A88CF716E08685EAECC2A1D2" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:F6873723A88CF716E08685EAECC2A1D2" country="United States of America" county="Rabun Co." latitude="34.913" longLatPrecision="55" longitude="-83.2978" municipality="Chattahoochee National Forest" name="Rabun Cliffs" stateProvince="Georgia">Rabun Cliffs</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EARDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.297800,34.913000]}">
<geoCoordinate id="B7E5351989DF89AB14810C079663E76A" degrees="34.913" direction="north" orientation="latitude" precision="55" value="34.913">34.913</geoCoordinate>
, 
<geoCoordinate id="1183978818777A172A124650E87DBD20" degrees="83.2978" direction="west" orientation="longitude" precision="5" value="-83.2978">- 83.2978</geoCoordinate>
</named-content>
; 
<date id="A3047BA152ED2B73676D3C6ECF7348F5" value="2021-05-11">
<collectingDate id="3EBA7786DC0F26D070AFA32BC4F80993" value="2021-05-11">11 May 2021</collectingDate>
</date>
; 
<collectorName id="A118E573C0A84C574A3B7EC33212FF66">M. Caterino</collectorName>
and 
<collectorName id="820B2E90A536212DBF79E58036189B3C">A. Haberski</collectorName>
leg.; MSC- 7026, 
<specimenCode id="96411C7AB1C7B748988DCE75295E7735" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000135496</specimenCode>
</materialsCitation>
</emphasis>
; • 
<materialsCitation id="665F2B5D6568AE79D3CBCFE585E55E00" collectingDate="2021-05-11" collectionCode="CUAC" collectorName="M. Caterino &amp; A. Haberski" country="United States of America" county="Rabun Co." latitude="34.913" location="Rabun Cliffs" longLatPrecision="55" longitude="-83.2978" municipality="Chattahoochee National Forest" specimenCode="CUAC 000172330, CUAC 000172322" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Georgia">
<specimenCount id="DBFC5C1BBE9911ED1AA238962EB82CAC" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="B7DFA687F94B5055B307FB90EAEBD874" count="1" type="female">1 ♀</specimenCount>
; same data as previous; 
<specimenCode id="A6969A7AD6BA57376ABEAA2E457122B7" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000172330</specimenCode>
and 
<specimenCode id="B04F2A6791A617D50A68835AF0EDE7C9" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000172322</specimenCode>
</materialsCitation>
; • 
<materialsCitation id="920C994077C3FA9A18D829634893B142" collectingDate="2014-07-09" collectionCode="CWHc" collectorName="T. Lawton" country="United States of America" county="Rabun Co." latitude="34.967" location="Rabun Bald" longLatPrecision="71" longitude="-83.299" specimenCount="2" specimenCount-female="2" stateProvince="Georgia">
<specimenCount id="8D26C2297C0EFD0845CC8A5EE7407CB9" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="FA9B4320BF6F9021CADE58A8C445CC52" bold="true">
<collectingCounty id="6BCB19158893AB7ADBDD3547C61C8A00">Rabun Co.</collectingCounty>
</emphasis>
; 
<location id="9332A730660D165E50367677DCD036C4" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:9332A730660D165E50367677DCD036C4" country="United States of America" county="Rabun Co." latitude="34.967" longLatPrecision="71" longitude="-83.299" name="Rabun Bald" stateProvince="Georgia">Rabun Bald</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQRDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.299000,34.967000]}">
<geoCoordinate id="CC63DE23EAD87F00416AAD25E5A38A8A" degrees="34.967" direction="north" orientation="latitude" precision="55" value="34.967">34.967</geoCoordinate>
, 
<geoCoordinate id="90F7D5C84B783CC8D77B33AAB9AB9CB0" degrees="83.299" direction="west" orientation="longitude" precision="55" value="-83.299">- 83.299</geoCoordinate>
</named-content>
; 
<date id="DC63F78BE8A45EF86B23F3FD6AD989AD" value="2014-07-09">
<collectingDate id="BBD07954F7276A8AB94A36686621F001" value="2014-07-09">9 Jul. 2014</collectingDate>
</date>
; 
<collectorName id="A3923612C237211CC7A5440002FCCDC2">T. Lawton</collectorName>
leg.; 
<abbrev id="ABBRID0EVRDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="BD1C3246537B0660524F06B77569739D">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="CDE9CB97C217B673CD874D433920F5BE" collectingDate="2014-05-24" collectionCode="CWHc" collectorName="T. Lawton" country="United States of America" county="Rabun Co." latitude="34.967" location="Rabun Bald" longLatPrecision="71" longitude="-83.299" specimenCount="1" specimenCount-female="1" stateProvince="Georgia">
<specimenCount id="EC1291E69AB6FE9C1AB6DDFDD8CDFFD2" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="E86A2BC3AFA933FEACE8763B011DAD04" bold="true">
<collectingCounty id="9453EF1D8CDE612C656DDB6C17EEEDEC">Rabun Co.</collectingCounty>
</emphasis>
; 
<location id="FF4D0E78533D9F766FF1B2C40144D157" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:FF4D0E78533D9F766FF1B2C40144D157" country="United States of America" county="Rabun Co." latitude="34.967" longLatPrecision="71" longitude="-83.299" name="Rabun Bald" stateProvince="Georgia">Rabun Bald</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5RDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.299000,34.967000]}">
<geoCoordinate id="4AFE12BAE8A3A68EB82E637BB33DE60A" degrees="34.967" direction="north" orientation="latitude" precision="55" value="34.967">34.967</geoCoordinate>
, 
<geoCoordinate id="135BBEB11CC840B2208AD1E26BF0E6FF" degrees="83.299" direction="west" orientation="longitude" precision="55" value="-83.299">- 83.299</geoCoordinate>
</named-content>
; 
<date id="D7AFFC28DF9A60622D1AB5559091B039" value="2014-05-24">
<collectingDate id="2D90EAA456B33CA698EE18EBCDCA7A49" value="2014-05-24">24 May 2014</collectingDate>
</date>
; 
<collectorName id="EC9ABE1BE7FE4D903A618A42ABD3F4A9">T. Lawton</collectorName>
leg.; 
<abbrev id="ABBRID0EDSDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="9A314B056303E7C76D89CFF3345ECB7A">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="5D255DC8866BBC2880410B331F918BD6" collectingDate="2020-07-02" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Rabun Co." latitude="34.9708" location="Rabun Bald" longLatPrecision="6" longitude="-83.3032" specimenCount="34" specimenCount-female="15" specimenCount-male="19" stateProvince="Georgia">
<specimenCount id="3E0CB42D5ECAD235246571B1C44B8B34" count="19" type="male">19 ♂</specimenCount>
, 
<specimenCount id="FA87879EB8737A2BF953AE4C9365312E" count="15" type="female">15 ♀</specimenCount>
; 
<emphasis id="AA438CE1470263B1E281DF2B26F6B20A" bold="true">
<collectingCounty id="A20974FF2D08C2534BF9EB88881DE9D6">Rabun Co.</collectingCounty>
</emphasis>
; 
<location id="A5B206F4289CF2CF7482BEFCC602CE8C" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:A5B206F4289CF2CF7482BEFCC602CE8C" country="United States of America" county="Rabun Co." latitude="34.9708" longLatPrecision="6" longitude="-83.3032" name="Rabun Bald" stateProvince="Georgia">Rabun Bald</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMSDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.303200,34.970800]}">
<geoCoordinate id="65D735A33570C13ECE13159232376054" degrees="34.9708" direction="north" orientation="latitude" precision="5" value="34.9708">34.9708</geoCoordinate>
, 
<geoCoordinate id="2BA045CBD133BC95DC0FA20380CF3ACF" degrees="83.3032" direction="west" orientation="longitude" precision="5" value="-83.3032">- 83.3032</geoCoordinate>
</named-content>
; 
<date id="8E6704604DC5EBC5BAB9A120D4FC5221" value="2020-07-02">
<collectingDate id="3EE8A534398A8EA023FA51F4A4776DEB" value="2020-07-02">2 Jul. 2020</collectingDate>
</date>
; 
<collectorName id="116D835382D6A087059C65F3D8048935">C. W. Harden</collectorName>
leg.; 
<abbrev id="ABBRID0ERSDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="1282DBAA6B6B97DA2CF995781CC89D23">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="F101F16078F5834588D126FB188B6BC3" collectingDate="2023-06-05" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Rabun Co." latitude="34.9759" location="south of Beegum Gap" longLatPrecision="6" longitude="-83.3041" municipality="Chattahoochee National Forest" specimenCount="1" specimenCount-male="1" stateProvince="Georgia">
<specimenCount id="A9C8F83F207968EDE0535AC0BEB87F88" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="9BAF463BD378CC149F5D434F6EB5053B" bold="true">
<collectingCounty id="3ABA49A1367D440D7CB0864076FF6022">Rabun Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="BEBB80D63A143C0BECE18A5DDDD83DB7">Chattahoochee National Forest</collectingMunicipality>
, 
<location id="50098573AF706B6996FE404E5180A70F" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:50098573AF706B6996FE404E5180A70F" country="United States of America" county="Rabun Co." latitude="34.9759" longLatPrecision="6" longitude="-83.3041" municipality="Chattahoochee National Forest" name="south of Beegum Gap" stateProvince="Georgia">south of Beegum Gap</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1SDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.304100,34.975900]}">
<geoCoordinate id="8A7E0E64806BB81921FE14782993451A" degrees="34.9759" direction="north" orientation="latitude" precision="5" value="34.9759">34.9759</geoCoordinate>
, 
<geoCoordinate id="C9B30C13815ABB1BE4BCF1F6C6AB4A4C" degrees="83.3041" direction="west" orientation="longitude" precision="5" value="-83.3041">- 83.3041</geoCoordinate>
</named-content>
; 
<date id="B7DE205FF4705EA043FF29F8216F5427" value="2023-06-05">
<collectingDate id="5CD56D06D969680B9070EDBDA59A0A2A" value="2023-06-05">5 Jun. 2023</collectingDate>
</date>
; 
<collectorName id="5A5E863E180E81374E5EDF49A8C3CED8">C. W. Harden</collectorName>
leg.; 
<abbrev id="ABBRID0E6SDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="1C0BA1AD8B7EBBBB3EFC9D0AE9832ECD">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="458B04CF672E975762E4DED65E55E724" collectingDate="1965-05-23" collectionCode="CMNH" collectorName="H. R. Steeves" country="United States of America" county="Macon Co." latitude="35.045" location="Highlands" longLatPrecision="71" longitude="-83.451" municipality="Coweeta Hydrological Lab" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="North Carolina">
<emphasis id="B404F09DCB0635F7605D671EC45A38CA" bold="true">
<collectingRegion id="C73D51F68494E63B74C5939AAEF01216" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="0676881585481EAED7B0181FF723B9E1" count="4" type="male">4 ♂</specimenCount>
, 
<specimenCount id="37B97F58195D2FCBCBA1DCC0454F3C1D" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="BFFDDF2C6E94DD9BC6B00F696E9453DA" bold="true">
<collectingCounty id="D383440E7F0D34F15F386EC43803D2C9">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="8FD00D6A66AB81A2EA73BBFFF1CC751E">Coweeta Hydrological Lab</collectingMunicipality>
, ca. 
<locationDeviation id="C0015378182BD31E0691104E6420DB15">
<quantity id="4DD8CDDCB965EC11DC7DD962502F024F" metricMagnitude="4" metricUnit="m" metricValue="2.0921472" unit="mi" value="13.0">13 miles</quantity>
west of
</locationDeviation>
<location id="461B1C0F950B6F27BE7627EBDC6639A6" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:461B1C0F950B6F27BE7627EBDC6639A6" country="United States of America" county="Macon Co." latitude="35.045" longLatPrecision="71" longitude="-83.451" municipality="Coweeta Hydrological Lab" name="Highlands" stateProvince="North Carolina">Highlands</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKTDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.451000,35.045000]}">
<geoCoordinate id="92C67B5FA9E00433F8FC0682E7E7CE8B" degrees="35.045" direction="north" orientation="latitude" precision="55" value="35.045">35.045</geoCoordinate>
, 
<geoCoordinate id="1BA557BC4C856F34AC305553A5338040" degrees="83.451" direction="west" orientation="longitude" precision="55" value="-83.451">- 83.451</geoCoordinate>
</named-content>
; 
<date id="55AD45575D2A519B7459859EF5D44B81" value="1965-05-23">
<collectingDate id="4BAEF6192B10DA9293ED61376F78969D" value="1965-05-23">23 May 1965</collectingDate>
</date>
; 
<collectorName id="D53C236085E3C6823CD45CF7E5AC2E1E">H. R. Steeves</collectorName>
leg.; 
<named-content id="7FEAD93FAA3A7E958E90FB0F647FEB87" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="751063D677BCF478AEB17DE5C24E5BAE" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="23B21CC12D9F1F9B910A14D5090E2B1C" collectingDate="1969-08-13" collectionCode="CMNH" collectorName="T. C. Barr" country="United States of America" county="Macon Co." latitude="35.0339" location="Ball Creek" longLatPrecision="6" longitude="-83.4505" municipality="Coweeta Experimental Station" specimenCount="1" specimenCount-female="1" stateProvince="North Carolina">
<specimenCount id="D612148EE0379BEB3C834DBD603D12FD" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="1814A148FE7DA6E7D634BA554E6B47D7" bold="true">
<collectingCounty id="2B83B3A44A2514C380243205DD6708C3">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="7913E5DC37A6EC59C494E6B730CAA3C4">Coweeta Experimental Station</collectingMunicipality>
[sic.], 
<location id="B4B46C58B0C5ABB1305008D6180DC8A9" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:B4B46C58B0C5ABB1305008D6180DC8A9" country="United States of America" county="Macon Co." latitude="35.0339" longLatPrecision="6" longitude="-83.4505" municipality="Coweeta Experimental Station" name="Ball Creek" stateProvince="North Carolina">Ball Creek</location>
, # 62, 3700 ’; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYTDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.450500,35.033900]}">
<geoCoordinate id="296E66CCB7478745129115B5BD86469F" degrees="35.0339" direction="north" orientation="latitude" precision="5" value="35.0339">35.0339</geoCoordinate>
, 
<geoCoordinate id="774F4DD728CCDDB5FA1F71F1EF2D1C2A" degrees="83.4505" direction="west" orientation="longitude" precision="5" value="-83.4505">- 83.4505</geoCoordinate>
</named-content>
; 
<date id="6AA198E418FBAEA0FE1440A86E6F3D54" value="1969-08-13">
<collectingDate id="7DA4E498C80A6EDAD44F9F3E1C338785" value="1969-08-13">13 Aug. 1969</collectingDate>
</date>
; 
<collectorName id="69B894746044F4819838148D5C35FA4A">T. C. Barr</collectorName>
leg.; 
<named-content id="94D6487B0106BA27ABCDB57FFD872579" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="2A1AFA74CFAE694E50A6003B0988FB9F" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="93D0D805F6689E8A6A22697E855FD6DF" collectingDate="1969-08-13" collectionCode="CMNH" collectorName="T. C. Barr" country="United States of America" county="Macon Co." latitude="35.0432" location="Ball Creek" longLatPrecision="6" longitude="-83.4535" municipality="Coweeta Experimental Station" specimenCount="2" specimenCount-female="2" stateProvince="North Carolina">
<specimenCount id="AF39DD1A1548A138ADE40CF7AEF63AF9" count="2" type="female">2 ♀</specimenCount>
; 
<emphasis id="4EAF955B35EC89A98CBFFD01F89949EE" bold="true">
<collectingCounty id="4C3E90B57ACD045512A17F4BFE71FA88">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="18E727B2F625D9DCCA97F6BF02E24856">Coweeta Experimental Station</collectingMunicipality>
[sic.], 
<location id="A73CDA318FBF41637793C983FF696397" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:A73CDA318FBF41637793C983FF696397" country="United States of America" county="Macon Co." latitude="35.0432" longLatPrecision="6" longitude="-83.4535" municipality="Coweeta Experimental Station" name="Ball Creek" stateProvince="North Carolina">Ball Creek</location>
# 42, 3100 ’; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EGUDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.453500,35.043200]}">
<geoCoordinate id="8A9B547AF765CB2AE5B23E1AF59BB498" degrees="35.0432" direction="north" orientation="latitude" precision="5" value="35.0432">35.0432</geoCoordinate>
, 
<geoCoordinate id="897E7A045961EB0D760592BCC61130FB" degrees="83.4535" direction="west" orientation="longitude" precision="5" value="-83.4535">- 83.4535</geoCoordinate>
</named-content>
; 
<date id="277EBA0E936ADCFD89BB8294B23DD1BC" value="1969-08-13">
<collectingDate id="C8C6278CA37E33C1D9EA2787B1C4332F" value="1969-08-13">13 Aug. 1969</collectingDate>
</date>
; 
<collectorName id="6EA206DC6AE2FBB70847D261F31F05DA">T. C. Barr</collectorName>
leg.; 
<named-content id="B5AE6277B9359D4B4C6271E46F543E40" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="43DEE0E13018559C85C86482EF5D70FF" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="E63ECB6915696CB188D3AF329B239753" collectingDate="1970-08-08" collectionCode="CMNH" collectorName="T. C. Barr" country="United States of America" county="Macon Co." location="Highlands" municipality="Turtle Pond Creek" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<specimenCount id="1DBDF6E3EDE757C1574172F2D61FF6B7" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="47D0F50CB358173B98B4A01552DB5D99" bold="true">
<collectingCounty id="EBB2DC9BA7710A8C6250153B6F94C185">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="12740E65BD901C412DF2A9841D822CD7">Turtle Pond Creek</collectingMunicipality>
, ca. 
<locationDeviation id="709D5F69E99768FCF89CF4515900215A">
<quantity id="4331CD997DFC4279BA8C07EE299DE3EA" metricMagnitude="3" metricUnit="m" metricValue="6.437376" unit="mi" value="4.0">4 miles</quantity>
west-northwest of
</locationDeviation>
<location id="39E9F48B089BF27D0D4B41698189CC7E" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:39E9F48B089BF27D0D4B41698189CC7E" country="United States of America" county="Macon Co." municipality="Turtle Pond Creek" name="Highlands" stateProvince="North Carolina">Highlands</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EUUDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.260000,35.060000]}">35.06, - 83.26</named-content>
; 
<date id="D39FB4D6C907A2D9DA84CD3BDE6F5AB8" value="1970-08-08">
<collectingDate id="B44C8DA4AA261B14606BA428A7517F54" value="1970-08-08">8 Aug. 1970</collectingDate>
</date>
; 
<collectorName id="C9D0BF5E388935FB7A1E18977AA7D43E">T. C. Barr</collectorName>
leg.; 
<named-content id="941E202254EF26372AA9B5C4DEBC66F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="A2B1D9FCDC058991CE9B0AA911D16DD0" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="D556EC5020605100292805D927B252C6" collectingDate="1971-05-14" collectionCode="CMNH" collectorName="T. C. Barr" country="United States of America" county="Macon Co." location="California Ridge" municipality="Goldmine" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="California">
<specimenCount id="EBFA206A515CFC6BCB4B203943FBC3FA" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="1706E34BDAD8401E155D2011F2D88F38" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="8671DCD4D5C1D810252F55F99D85EA11" bold="true">
<collectingCounty id="D5CE5FE83B7A2B18960BAB4433E1B7BE">Macon Co.</collectingCounty>
</emphasis>
; 
<locationDeviation id="61B7F428512CD84F0C98A20DA62EBDFA">
<quantity id="913172F850BD9A17EE9CF7096F359ADD" metricMagnitude="2" metricUnit="m" metricValue="9.656064" unit="mi" value="0.6">0.6 miles</quantity>
northeast of Goldmine
</locationDeviation>
, 
<location id="37DC8D0E0FA709A8C191119B34E6C18E" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:37DC8D0E0FA709A8C191119B34E6C18E" country="United States of America" county="Macon Co." municipality="Goldmine" name="California Ridge" stateProvince="California">California Ridge</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECVDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.280000,35.100000]}">35.10, - 83.28</named-content>
; 
<date id="6E0EA0264497001B540FE4FC2AB88191" value="1971-05-14">
<collectingDate id="0971C5B9F0CEC52479379E39BED5A1EA" value="1971-05-14">14 May 1971</collectingDate>
</date>
; 
<collectorName id="9FFC4D27AF795C5022C2E952348329E0">T. C. Barr</collectorName>
leg.; 
<named-content id="974DA5438FDFC5AB33BF5013074291A2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="D6DB30FB62F6F78BAAD577156084F3B3" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="C0F0C75FD73A96CE2AD4500407EE3048" collectingDate="2020-08-03" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Macon Co." latitude="35.1554" location="Wayah Road" longLatPrecision="6" longitude="-83.5584" municipality="Nantahala National Forest" specimenCount="5" specimenCount-male="5" stateProvince="North Carolina">
<specimenCount id="EF712E9D4600176FEA125C4A146A03B5" count="5" type="male">5 ♂</specimenCount>
; 
<emphasis id="FB9CB9B685FA6BEEE8C623D1B74B9CA8" bold="true">
<collectingCounty id="6B09C4B54205F0573B3E5A39C0636B18">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="C19B0588417C601622CAC83F85E794C3">Nantahala National Forest</collectingMunicipality>
, off 
<location id="EEACB1001437EBD4DB540EB4FBD85AF5" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:EEACB1001437EBD4DB540EB4FBD85AF5" country="United States of America" county="Macon Co." latitude="35.1554" longLatPrecision="6" longitude="-83.5584" municipality="Nantahala National Forest" name="Wayah Road" stateProvince="North Carolina">Wayah Road</location>
ca. 
<locationDeviation id="9166A4D7636315503F02FEBC2281A4B1">
<quantity id="69292C93D955DA3F9953E4EB488EC352" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
west from Route 64
</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQVDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.558400,35.155400]}">
<geoCoordinate id="2FDC0D5DBEAEF30DB9792202F7B8B583" degrees="35.1554" direction="north" orientation="latitude" precision="5" value="35.1554">35.1554</geoCoordinate>
, 
<geoCoordinate id="A21EDF0F9B575C501F70CC29858EE8EA" degrees="83.5584" direction="west" orientation="longitude" precision="5" value="-83.5584">- 83.5584</geoCoordinate>
</named-content>
; 
<date id="43E63022E5AFFB606E9E240D45CD8CAD" value="2020-08-03">
<collectingDate id="FBC49F0A688907103E9D9A51020907CB" value="2020-08-03">3 Aug. 2020</collectingDate>
</date>
; 
<collectorName id="FAC830B94302297B21F1FF97F610276C">C. W. Harden</collectorName>
leg.; 
<abbrev id="ABBRID0EVVDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="6675979B152B87FE4EBD56782695135A">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="FFEAC948023389C863B57D3049EE20CD" collectingDate="2021-06-04" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Macon Co." latitude="35.1554" location="Wayah Road" longLatPrecision="6" longitude="-83.5584" municipality="Nantahala National Forest" specimenCount="1" specimenCount-female="1" stateProvince="North Carolina">
<specimenCount id="79F564796F7D326A369C64F001980716" count="1" type="female">1 ♀</specimenCount>
; same data as previous; 
<date id="6C6B6DC4D67C787B1F24BCDAEB45A241" value="2021-06-04">
<collectingDate id="F2E729123E40C265E751FA18CBA75310" value="2021-06-04">4 Jun. 2021</collectingDate>
</date>
</materialsCitation>
; • 
<materialsCitation id="A3CFF416D741307954DE5F1D0E4EE263" collectingDate="2019-10-20" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Macon Co." latitude="35.1557" location="Wayah Road" longLatPrecision="6" longitude="-83.5583" municipality="Nantahala National Forest" specimenCount="1" specimenCount-female="1" stateProvince="North Carolina">
<specimenCount id="B580EE4C345645FFBB9BB6C3E74369AF" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="3FE0B8E53E92E592E05F929FB7AAAD57" bold="true">
<collectingCounty id="B7F9B4D331E7F1F6B562F24C4E1C58DE">Macon Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="3BBE153E95B382123E19EF6018779EAF">Nantahala National Forest</collectingMunicipality>
, off 
<location id="C3B8FFC43ACE82556EB08164A67728BF" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:C3B8FFC43ACE82556EB08164A67728BF" country="United States of America" county="Macon Co." latitude="35.1557" longLatPrecision="6" longitude="-83.5583" municipality="Nantahala National Forest" name="Wayah Road" stateProvince="North Carolina">Wayah Road</location>
ca. 
<locationDeviation id="ED7CBBC1734F9BAB570797CD0323E4D4">
<quantity id="55A9930F97E11ABFD516B58D1E2C3A4B" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
west from Route 64
</locationDeviation>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E5VDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.558300,35.155700]}">
<geoCoordinate id="AF651E6EE52E7DBA08EC35779DFB7FC3" degrees="35.1557" direction="north" orientation="latitude" precision="5" value="35.1557">35.1557</geoCoordinate>
, 
<geoCoordinate id="4D346D07C315EA77CFD307544BDAB95F" degrees="83.5583" direction="west" orientation="longitude" precision="5" value="-83.5583">- 83.5583</geoCoordinate>
</named-content>
; 
<date id="F97BB81A8BCEE14C4D59EB38D1D3EFD1" value="2019-10-20">
<collectingDate id="9A4EE8BE6AC5A1484F5A8C955C2D8390" value="2019-10-20">20 Oct. 2019</collectingDate>
</date>
; 
<collectorName id="AB5FC06EEC80679366A524B18DE290F8">C. W. Harden</collectorName>
leg.; 
<abbrev id="ABBRID0EDWDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="61010DFE5925B0FC0378103781016FA0">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="0722715DEF8BEB6923E3DCDB64BC62C6" collectingDate="2020-08-03" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Macon Co." latitude="35.1557" location="Wayah Road" longLatPrecision="6" longitude="-83.5583" municipality="Nantahala National Forest" specimenCount="6" specimenCount-female="6" stateProvince="North Carolina">
<specimenCount id="63228D648212BDB5B317763AC6581359" count="6" type="female">6 ♀</specimenCount>
; same data as previous; 
<date id="ABDD8F33354ECD4BDFAC63F37430513C" value="2020-08-03">
<collectingDate id="64B3AF7C57EF89812F5B4B88213042E4" value="2020-08-03">3 Aug. 2020</collectingDate>
</date>
; 
<abbrev id="ABBRID0EHWDK" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="2BC3D88A628946D1D95CE57B45B65967">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="9A72D27FB5C4ED80C84AFF36E1C40D46" collectingDate="1976-03-18" collectionCode="OSUC" country="United States of America" county="Macon Co." latitude="35.239" location="four miles north of Franklin" longLatPrecision="71" longitude="-83.374" specimenCode="OSUC 442503, OSUC 442504" specimenCount="2" specimenCount-male="2" stateProvince="North Carolina">
<specimenCount id="44203506EF73986415F567E555D5D636" count="2" type="male">2 ♂</specimenCount>
; 
<emphasis id="43F30FFB248120C11BB2951E37FF0424" bold="true">
<collectingCounty id="6CE61781D77A7DAC307FF9921D2776D2">Macon Co.</collectingCounty>
</emphasis>
; 
<location id="DF8C0790A3036A9D4EB90DE09AEFDFAA" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:DF8C0790A3036A9D4EB90DE09AEFDFAA" country="United States of America" county="Macon Co." latitude="35.239" longLatPrecision="71" longitude="-83.374" name="four miles north of Franklin" stateProvince="North Carolina">four miles north of Franklin</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EQWDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.374000,35.239000]}">
<geoCoordinate id="155D0BBACDE8DE4C981E75445AF5F0CF" degrees="35.239" direction="north" orientation="latitude" precision="55" value="35.239">35.239</geoCoordinate>
, 
<geoCoordinate id="69F4D41C86F34DD55EFA1161B507F9C5" degrees="83.374" direction="west" orientation="longitude" precision="55" value="-83.374">- 83.374</geoCoordinate>
</named-content>
; 
<date id="DD9BA5CA2DE47CCE994C134231954B1C" value="1976-03-18">
<collectingDate id="F33A7A729962739C0A34F7DF3C5E877D" value="1976-03-18">18 Mar. 1976</collectingDate>
</date>
; 
<specimenCode id="DE1E0E6DBFA56AD8F33EB9C3D09C9D1E" collectionCode="OSUC" country="USA" httpUri="http://grbio.org/cool/x820-3v0f" name="Oregon State University" type="">OSUC 442503</specimenCode>
and 
<specimenCode id="097F91BCE652E43093D45CA413F297C4" collectionCode="OSUC" country="USA" httpUri="http://grbio.org/cool/x820-3v0f" name="Oregon State University" type="">OSUC 442504</specimenCode>
; 
<named-content id="19AAA51D7C069422FAC8793043C504ED" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/ohio-state-university" xlink_title="Ohio State University Collection">
<collectionCode id="1D9FF334E20383E28C82A5875D4225B1" country="USA" httpUri="http://grbio.org/cool/x820-3v0f" name="Oregon State University" type="">OSUC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="00B7F0F7B5ABDF9CC8898CFBFC289BB0" collectingDate="2022-08-23" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Oconee Co." latitude="35.023" location="Whitewater River" longLatPrecision="71" longitude="-83.004" municipality="Coon Branch Natural Area" specimenCode="CUAC 000185794" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina">
<emphasis id="7CDDC607CBE0DD178FFDDAE6B4E1F0C0" bold="true">
<collectingRegion id="1FF2A0E8F86A35E4EBE6B559C9AF1BD5" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="4242D20DE33146532DDA2F4A90C344E7" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="8BAF4BE88930A8D03B3375E627F3E16A" bold="true">
<collectingCounty id="C58883A03084677D4C9F5D4966DBF210">Oconee Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="793C32CA98728BDA7BD77EA9714DDF85">Coon Branch Natural Area</collectingMunicipality>
, near 
<location id="9F99C933E991092D3E0EB4E640F6424D" LSID="urn:lsid:plazi:treatment:D7CA01C826CA5B47A9138A3BE8FE53CB:9F99C933E991092D3E0EB4E640F6424D" country="United States of America" county="Oconee Co." latitude="35.023" longLatPrecision="71" longitude="-83.004" municipality="Coon Branch Natural Area" name="Whitewater River" stateProvince="South Carolina">Whitewater River</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAXDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.004000,35.023000]}">
<geoCoordinate id="8AC24BEC6E69034D477C93C9F10C5905" degrees="35.023" direction="north" orientation="latitude" precision="55" value="35.023">35.023</geoCoordinate>
, 
<geoCoordinate id="D2EC940482CA209AD23AED8CA5AE7A2D" degrees="83.004" direction="west" orientation="longitude" precision="55" value="-83.004">- 83.004</geoCoordinate>
</named-content>
; 
<date id="44EC04EFEFB00BF8E035404146187D92" value="2022-08-23">
<collectingDate id="163667E59F3D4AE7112A5B498C133170" value="2022-08-23">23 Aug. 2022</collectingDate>
</date>
; 
<collectorName id="966321F7DC7CD25D20D4391494B15B8A">C. W. Harden</collectorName>
leg.; CWH- 454, 
<specimenCode id="236CE34EEA51B0179DA3D4D6DDA2DD7A" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000185794</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="A96009ACD3798E5B87CC42F4998A6946">
GenBank: 
<ext-link id="297A0B604DE4014F334DB22E31E6AFCB" ext-link-type="gen" xlink_href="OR853398" xlink_type="simple">OR 853398</ext-link>
, 
<ext-link id="3EE1840DAA7335432362BC3351082D1B" ext-link-type="gen" xlink_href="OR839364" xlink_type="simple">OR 839364</ext-link>
, 
<ext-link id="A4FF54FBD03B33EACB8EC9E8D8CDC9F9" ext-link-type="gen" xlink_href="OR839681" xlink_type="simple">OR 839681</ext-link>
, 
<ext-link id="FA9739AC8A69F2FDDAD0115182326EDE" ext-link-type="gen" xlink_href="OR837924" xlink_type="simple">OR 837924</ext-link>
, 
<ext-link id="EE9AE37AB3A241CB8814A001706FADEF" ext-link-type="gen" xlink_href="OR838086" xlink_type="simple">OR 838086</ext-link>
, 
<ext-link id="E28D78112DF0906AA460ED9874623781" ext-link-type="gen" xlink_href="OR838255" xlink_type="simple">OR 838255</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EFYDK" type="Literature records">
<paragraph id="EC19B9E81C82325B20DE7382334C3852">
<heading id="A02235A6C85FDDC4291FC713D6DB4433" reason="title">Literature records.</heading>
</paragraph>
<paragraph id="8E0427E873EBF6B6EDEC6600D94CF8A4">
<bibRefCitation id="26D5859F5CEE06007F88703673D9FC13" author="Barr Jr TC" firstAuthor="Barr" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239–248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr (1995)</bibRefCitation>
stated that the type locality was “ along Ball Creek, elevation approximately 
<quantity id="AB901B0943E65873FC0A5A39DDB48492" metricMagnitude="2" metricUnit="m" metricValue="9.5" unit="m" value="950.0">950 m</quantity>
, Coweeta Hydrologic Laboratory, 
<collectingCountry id="A7B15D956AE584175DA89681112A17A0" name="United States of America">U. S.</collectingCountry>
Forest Service, Macon Co., 
<collectingRegion id="05AB0CEA6AA2DD6663B30C5440BBFF24" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
. ” Field notes for the date of collection (
<date id="A618A64E73BAD79D21F4AA59514551DC" value="1969-08-13">13 August 1969</date>
) state that the site was below the first switchback of Ball Creek Road, approximate coordinates 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYYDK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-83.453500,35.043200]}">
<geoCoordinate id="99EA88F048BF16C049858882996E6AFE" degrees="35.0432" direction="north" orientation="latitude" precision="5" value="35.0432">35.0432</geoCoordinate>
, 
<geoCoordinate id="697DCA4D849048ED53CC2996AC0E9679" degrees="83.4535" direction="west" orientation="longitude" precision="5" value="-83.4535">- 83.4535</geoCoordinate>
</named-content>
. Barr also wrote that the species occurred in the Great Balsam Mountains in 
<collectingRegion id="D02672EED7D8E207FD3D2AFD7D8448F0" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
and Towns Co., 
<collectingRegion id="A2386F3745507127813983B1868738A6" country="United States of America" name="Georgia">Georgia</collectingRegion>
. 
<bibRefCitation id="16DE4292D25260AF7B5373B573A8DDE0" DOI="10.11646/zootaxa.1740.1.4" author="Sokolov IM &amp; Carlton CE" firstAuthor="Sokolov" issue="1" journalOrPublisher="Zootaxa" pagination="37" refId="B85" refString="Sokolov IM, Carlton CE (2008) Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A. Zootaxa 1740 (1): 37. https://doi.org/10.11646/zootaxa.1740.1.4" title="Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A." volume="1740" year="2008">Sokolov and Carlton (2008)</bibRefCitation>
state they studied specimens of 
<taxonomicName id="623741BDB12FCA94FA2A70B4E8FB7AEE" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="35F45D3B41A1F5220DBF2AB1FD161520" italics="true">S. jeanneli</emphasis>
</taxonomicName>
from “ White County, 
<collectingRegion id="4B6EABDE3CA2FFABF67220DB2AA2A661" country="United States of America" name="Georgia">Georgia</collectingRegion>
” without further data. The 
<taxonomicName id="F1F4EBDCE4A02940ADF4B8620B8DA658" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="EA120E9F5611C710B6054E204931AA40" italics="true">Serranillus</emphasis>
</taxonomicName>
that we have studied from Towns Co. and White Co., 
<collectingRegion id="B92B820F1BB123FF036783440D2C9A8C" country="United States of America" name="Georgia">Georgia</collectingRegion>
are not 
<taxonomicName id="56E4FC6C7CB13CE84093AE17423138FC" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="56F73B20C5A81B2CC9E139391D79DA58" italics="true">S. jeanneli</emphasis>
</taxonomicName>
but two undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E5ZDK" type="diagnosis">
<paragraph id="B2E035676E48334AD5927F5C336C14FA">
<heading id="DE14E88BB9DFD8A6A079F60D7BA24779" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="DCB6AC91A17BEAA2515FC95D76313BBC">
From other species of 
<taxonomicName id="6EC3364C715ADD90AC0B04BCB7DF9695" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="697927642F744513B8FEC6D2002C79DD" italics="true">Serranillus</emphasis>
</taxonomicName>
, 
<taxonomicName id="E4D7D59DAD69ACF2F9BF798F318F1F13" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="F18013BAB6B91EA079BCF9C46F16E7BD" italics="true">S. jeanneli</emphasis>
</taxonomicName>
is best distinguished by the male median lobe of the aedeagus, which has a distinctive carinate shelf on the ventral surface, causing a preapical notch in the ventral margin in right lateral aspect (Fig. 
<figureCitation id="63C08F868D408868027BCE621CCE94BB" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 D</figureCitation>
). Externally, members of 
<taxonomicName id="6F973AF7EA3EB4E861E444677DFA9F55" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="576C10F42A958EE53B63DBA0C015A591" italics="true">S. jeanneli</emphasis>
</taxonomicName>
are moderately sized for 
<taxonomicName id="62673C4B7240CB3C000CC8C552449DD7" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="E84E8538C934641BD6A323F332EAFD66" italics="true">Serranillus</emphasis>
</taxonomicName>
, with male 
<abbrev id="ABBRID0EM2DK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 
<quantity id="E85D88D8C4857ACC613FD6F1FE5BF50D" metricMagnitude="-3" metricUnit="m" metricValue="2.24" metricValueMax="2.35" metricValueMin="2.13" unit="mm" value="2.24" valueMax="2.35" valueMin="2.13">2.13–2.35 mm</quantity>
and tentatively assigned females 
<abbrev id="ABBRID0EQ2DK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 2.07–2.51. The denticles on the last abdominal ventrite in males differ from other 
<collectingRegion id="FD2D85863FB6C02326B8FB78D26740F1" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
species by having the inner denticle broader and less pronounced than the outer two (Fig. 
<figureCitation id="E8F325F1965C79C0EDA42E76878F9097" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 B</figureCitation>
).
</paragraph>
<caption id="0AF7DEFB32C4A9EF326CCBCF67517C3B" ID-DOI="10.3897/zookeys.1209.125897.figure18" ID-arpha="F98C5F78-80F0-56F0-9EA2-ACFA71C7FE56" httpUri="https://binary.pensoft.net/fig/1110357" startId="F18">
<paragraph id="74D6BFD312E94D408C79CCC4A41D9D5A">
<label id="D589F3B5EBC998EC771DA9D29F2EA89D">Figure 18.</label>
</paragraph>
<paragraph id="C49AB51EB3752EA3BC5E59962402C1AA">
<taxonomicName id="4C8565DA09C31E166D375C475D431F66" authority="Barr." authorityName="Barr." class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="BC6D6DC7AE5EA32985B6F0B71B830398" italics="true">Serranillus jeanneli</emphasis>
Barr.
</taxonomicName>
<emphasis id="51DE9EE0D20131C090702BBAAC4C02F2" bold="true">A</emphasis>
dorsal habitus, abdomen removed for DNA extraction 
<emphasis id="5B34A947914E7B39EAF0242BC99B037B" bold="true">B</emphasis>
last abdominal ventrite, ventral aspect 
<emphasis id="8C9F201A199EA1E238250EE7B17BF8D2" bold="true">C</emphasis>
left paramere, left lateral aspect 
<emphasis id="F44E9A87BB2F815500036E247F18827C" bold="true">D</emphasis>
median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (
<emphasis id="FFB2C20DB8FEF4D2DA4F98CF631D89F8" bold="true">A</emphasis>
); 0.1 mm (
<emphasis id="5625721BFE274FEFDF8954D8F8DD51FC" bold="true">B – D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E33DK" type="redescription">
<paragraph id="00C297897A3CDAE849B39D47E0B15860">
<heading id="0C16ABA35D397F795E0E416421060058" reason="title">Redescription.</heading>
</paragraph>
<paragraph id="726F33D7471424323AF9AF6B7025E313">
<emphasis id="24107FA9CB21A2EC9643F9F632A91575" bold="true" italics="true">Habitus</emphasis>
Robust and convex (Fig. 
<figureCitation id="309ABD3155D94830A167F2BCA0AD40F9" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 A</figureCitation>
), moderately sized for genus, 
<abbrev id="ABBRID0EL4DK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 2.07–2.51. 
<emphasis id="21CD6DF48069338228928F2404890D69" bold="true" italics="true">Integument</emphasis>
Dorsal microsculpture absent from most of head and pronotum, present at center of vertex. Small, irregular patches of weak microsculpture present on disc of pronotum in some individuals. 
<emphasis id="1D7B560B9EB94BD9B0F0C11F4AA4D7AB" bold="true" italics="true">Head</emphasis>
Relatively narrow (male 
<abbrev id="ABBRID0EV4DK" xlink_title="Maximum head width">HW</abbrev>
/ 
<abbrev id="ABBRID0EZ4DK" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.70–0.71), frontoclypeal horn present and well developed. Ocular area behind antennal insertion with a dorsoventral linear tubercle at midpoint of carina. 
<emphasis id="9C5B776E882EE3EFCB19947176A66ED5" bold="true" italics="true">Pronotum</emphasis>
Cordate, with sides broadly rounded and evenly converging posteriorly to constricted base, 
<abbrev id="ABBRID0EA5DK" xlink_title="Pronotal basal width, measured at posterior angles and not including external denticles">PbW</abbrev>
/ 
<abbrev id="ABBRID0EE5DK" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.69–0.73. Approximately 1 / 4 body length (
<abbrev id="ABBRID0EI5DK" xlink_title="Pronotal length, measured along midline">PL</abbrev>
/ 
<abbrev id="ABBRID0EM5DK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.23–0.26). 
<emphasis id="2E59BC317D5875F5CCF05BE598AE8DAA" bold="true" italics="true">Elytra</emphasis>
Ovoid and convex, with rounded humeri, weak traces of three striae present. 
<emphasis id="74BDA9E726EB6F490F046E09647B7BEF" bold="true" italics="true">Legs</emphasis>
Male protarsomeres 1 and 2 expanded and bearing ventral adhesive setae. Male metatrochanters with coarse microsculpture, metafemora unmodified in either sex. 
<emphasis id="010B90FE640C0C874A3593153FBFE518" bold="true" italics="true">Abdominal ventrites</emphasis>
Males with last abdominal ventrite bearing three denticles on posterior margin, the outer two narrower and more prominent than the inner one. 
<emphasis id="6F0EDF11D759E4FF0D98CDC0A1758A77" bold="true" italics="true">Male genitalia</emphasis>
Ring sclerite long (
<abbrev id="ABBRID0E35DK" xlink_title="Ring sclerite length">RL</abbrev>
/ 
<abbrev id="ABBRID0EA6DK" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.32), similar in form to that of 
<taxonomicName id="A5E7849A49D714E372013E338AA4B0E1" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="AA1929E74F4F0EB75CDCBBF2C71F7071" italics="true">S. dunavani</emphasis>
</taxonomicName>
. Median lobe strongly asymmetrical (Fig. 
<figureCitation id="94112C254FD632476DDAD9270E7C3377" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Multiple aspects of median lobes of South Carolina Serranillus species, progressively rotated dorsally from right lateral aspect (upper left) to left lateroventral aspect (lower right) A Serranillus dunavani B Serranillus jeanneli C Serranillus monadnock, holotype (Kings Mountain, SC) D Serranillus monadnock, paratype (Little Mountain, SC). Photographs not to same scale." figureDoi="10.3897/zookeys.1209.125897.figure17" httpUri="https://binary.pensoft.net/fig/1110356">17 B</figureCitation>
), abruptly enlarged beyond base, outline a blunt-topped broad triangle in lateral aspects. Ventral face with a carinate shelf projecting below ventral margin; in right lateral aspect causing the ventral margin to appear notched (Fig. 
<figureCitation id="9C0802961A447044520AC0759FA8934E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 D</figureCitation>
). Apex of median lobe abruptly narrowed to a small curved hooklike apex visible in dorsal or ventral aspects. Internal sac of median lobe with two long sclerites. The right sclerite ribbonlike and corkscrewed along right side of internal sac, appearing as a dark curved shape in right lateral aspect (Fig. 
<figureCitation id="704C67C57516BC3ABAA00020DB063E01" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 D</figureCitation>
), narrowing distally, where it protrudes from the ostium as a long evenly curved spine. The left sclerite is stouter, gradually narrowing along its sinuate length until it protrudes from the ostium beside the right sclerite as a bluntly hooked and curved spine. The left side of the ostium is dominated by a large rolled sclerite that curves dorsally from the left face of the median lobe into the internal sac, where it is rolled over itself twice; in right dorsolateral aspect, the scrolled sclerite appears to be a complex sclerotized structure resembling several stacked plates or a group of blunt spines (Fig. 
<figureCitation id="79FE02B7C5884A320AD0C4243BEA42CF" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 D</figureCitation>
); the true rolled shape is visible in posterior or anterior aspects. Flagellum not observed, possibly represented by a narrow, lightly sclerotized structure located in a similar position as the prominent flagellum in 
<taxonomicName id="1C950D035BBA5245189AFDA34EE9055D" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="5F66A58D46E56EE27C9C7B1FBFB04074" italics="true">S. dunavani</emphasis>
</taxonomicName>
(cf. Fig. 
<figureCitation id="15DB1BEFB2EF872B5A923B374D290982" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Serranillus dunavani (Jeannel) A habitus B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D right paramere, right lateral aspect E median lobe, right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – E)." figureDoi="10.3897/zookeys.1209.125897.figure15" httpUri="https://binary.pensoft.net/fig/1110354">15 E</figureCitation>
). The parameres are as in 
<taxonomicName id="F0EA541CCE5238CD1E5105F6E7EAD035" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="D53D04EFED0D64C5CC68635776FBB250" italics="true">S. dunavani</emphasis>
</taxonomicName>
, thus the right paramere is minute and asetose and the left paramere is large and conchoid, with a large base (Fig. 
<figureCitation id="F542B6100A1227C9AAB04F4F7DB0B44F" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Serranillus jeanneli Barr. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C left paramere, left lateral aspect D median lobe (with right paramere attached, black arrow), right dorsolateral aspect. Scale bars: 1 mm (A); 0.1 mm (B – D)." figureDoi="10.3897/zookeys.1209.125897.figure18" httpUri="https://binary.pensoft.net/fig/1110357">18 C, D</figureCitation>
). The basal lobes of the median lobe are similarly asymmetrical, with the right lobe reduced to a thin strap and the left lobe larger and cup-shaped. 
<emphasis id="3F0A83D1F12188FF35711042D4CDA08B" bold="true" italics="true">Female genitalia</emphasis>
Spermatheca small, less curved than in 
<taxonomicName id="9362C6792775ED8E800621278C7B4F84" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="28FF1BE05867AAE3AF73488616915257" italics="true">S. dunavani</emphasis>
</taxonomicName>
and with base less swollen (Fig. 
<figureCitation id="BE345DDD8EF405D37BBE1E4377DB2F1D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 T</figureCitation>
). Spermathecal duct long and coiled.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPBEK" type="distribution">
<paragraph id="67AE59BDC54E3C2084E178508310B9CE">
<heading id="5252E4E317162806E70789D0FE3F9055" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="75394D73FAA22B6A61C3886A736BBB5C">
Notwithstanding the comments of 
<bibRefCitation id="333F99A88EAE9A8E44E8A3261A8D5604" author="Barr Jr TC" firstAuthor="Barr" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239–248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr (1995)</bibRefCitation>
and 
<bibRefCitation id="B436EC9F3CDAEA7951060E531433EB37" DOI="10.11646/zootaxa.1740.1.4" author="Sokolov IM &amp; Carlton CE" firstAuthor="Sokolov" issue="1" journalOrPublisher="Zootaxa" pagination="37" refId="B85" refString="Sokolov IM, Carlton CE (2008) Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A. Zootaxa 1740 (1): 37. https://doi.org/10.11646/zootaxa.1740.1.4" title="Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A." volume="1740" year="2008">Sokolov and Carlton (2008)</bibRefCitation>
, we have seen specimens only from a small area centered around the North Carolina-South Carolina-Georgia corner (Fig. 
<figureCitation id="AC4F8C5FD534DD7C72C5E8DD4EDB8BBC" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution map of Serranillus species that occur in South Carolina. The locality for Serranillus sp. “ South Carolina, Coon Branch ” is the same as the single South Carolina S. jeanneli occurrence, and is not shown. Data are from Harden (2024)." figureDoi="10.3897/zookeys.1209.125897.figure16" httpUri="https://binary.pensoft.net/fig/1110355">16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBCEK" type="Sympatry">
<paragraph id="45A7619FEE5B93666C95A5B4A1235EA2">
<heading id="6FA3260056DB04A0F1925DE82BDFDD64" reason="title">Sympatry.</heading>
</paragraph>
<paragraph id="3AE43445F80B1C8511462DCA22CEA799">
At Coon Branch in Oconee Co., 
<collectingRegion id="1A06797925E8EF2AE80831E326576DE5" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, this species co-occurs with 
<taxonomicName id="B5362C84EEC570F4FCBA8DC64BDAE3D7" baseAuthorityName="Jeannel" baseAuthorityYear="1963" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="0DCF526C40F30D4FAC7251AA5B01DC64" italics="true">S. dunavani</emphasis>
</taxonomicName>
, 
<emphasis id="E6F54E7A571CB6EEA039B661B60E118A" italics="true">S.</emphasis>
sp. “ 
<collectingRegion id="7E2C526C4FA8D2B23269877E0FF4F6A2" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, Coon Branch ”, 
<taxonomicName id="0FDDE27181E290F57B695C8FE2530076" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="murrayae">
<emphasis id="E6DA1BC3578EFAA3F422A406F9825E61" italics="true">Anillinus murrayae</emphasis>
</taxonomicName>
, 
<taxonomicName id="C6B8DFD2D8F8BD247FEBE837FBFBB87D" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cherokee">
<emphasis id="2E9168586ECF124CA86A151A17078C84" italics="true">Anillinus cherokee</emphasis>
</taxonomicName>
, and 
<emphasis id="903AAA5871AE23AF5A5BEA5041F7857F" italics="true">A.</emphasis>
sp. “ 
<collectingRegion id="DD8D24E7F95671D3C23AA7358FD00047" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, Coon Branch ”.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWDEK" type="natural history">
<paragraph id="07CCA4824F9C3BA527BD262DDD882938">
<heading id="489D85ECCC7C9E9C5394C4ACB7972F6A" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="913CBBD17EB657CA2310F637D9D0672A">Specimens have been collected from leaf litter, underneath embedded rocks, and using buried pipe traps.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2DEK" type="notes">
<paragraph id="971E838AB67B54CFA764FBC5A6D94B66">
<heading id="0B89A916F2841604F006F45ADB7800CC" reason="title">Notes.</heading>
</paragraph>
<paragraph id="E2F447949EADCC1964939AE983FA47FD">
Barr’s concept of 
<taxonomicName id="610B1E1B501BB279E8575169824CA0BC" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="00A27840186AE6BA4F3BDD2E84EE707B" italics="true">S. jeanneli</emphasis>
</taxonomicName>
involved at least three species: the one whose male genitalia he illustrated and which we consider 
<taxonomicName id="504DD279B0864B227A7EA95FACFEF224" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="17C483EFF657DFCFC17BF305F19A98D3" italics="true">S. jeanneli</emphasis>
</taxonomicName>
, a larger species whose median lobe has a ventral medial tuft of long curved setae (
<taxonomicName id="77421BC642296D3C14FC418F91ED509B" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="213E5C0FEBBF04B45D29B95414E96518" italics="true">Serranillus</emphasis>
sp.
</taxonomicName>
“ North Carolina, Riley Knob ”), and a closely related species that occurs in northern 
<collectingRegion id="529EF9410FBE67DD81BE4F2E05A6BD02" country="United States of America" name="Georgia">Georgia</collectingRegion>
and the southern edge of the Great Smoky Mountains (
<taxonomicName id="4EC0A158F21B2389190C6DA0E60EC6E1" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="92298DE984C8F63B5122E55B5A56309F" italics="true">Serranillus</emphasis>
sp.
</taxonomicName>
“ North Carolina, Miller Cove ”). We have studied all of Barr’s anilline genitalia slide mounts, and the ones he identified as 
<taxonomicName id="7BF732F7A044D929112CD2C28DFDDABE" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="6E5C21DD2C827FBD50FE7AB368AC0A6A" italics="true">S. jeanneli</emphasis>
</taxonomicName>
were either those matching our concept or those belonging to 
<emphasis id="C80E44748CF05C523B51393CB330E091" italics="true">S.</emphasis>
sp. “ North Carolina, Miller Cove. ” The latter species does not occur at the type locality. Apparently, Barr never dissected 
<emphasis id="F844BE66B8690B41E7CE96DBBB55EF2F" italics="true">S.</emphasis>
sp. “ North Carolina, Riley Knob ”, the larger species that occurs at Coweeta, although his description implies that he considered it to be the same species whose genitalia he illustrated.
</paragraph>
<paragraph id="17DA37BBA591BEA50B7FEEBDB76085D9">
<bibRefCitation id="ECFA5382A95A906E03BC3DBCB3A40354" author="Barr Jr TC" firstAuthor="Barr" journalOrPublisher="Special Bulletin of the Japanese Society of Coleopterology" pagination="239-248" refId="B9" refString="Barr Jr TC (1995) Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species. Special Bulletin of the Japanese Society of Coleopterology 4: 239–248." title="Notes on some anillines (Coleoptera, Carabidae, Bembidiinae) from Southeastern United States, with descriptions of a new genus and two new species." volume="4" year="1995">Barr (1995)</bibRefCitation>
designated a 
<typeStatus id="6158523CFEB3E48F749B0C56F6CE3ACA" type="holotype">holotype</typeStatus>
and 
<specimenCount id="CD637DBE16472B5460B59A10D798BAA8" count="4" type="generic" typeStatus="paratype">four paratypes</specimenCount>
for 
<taxonomicName id="C8C1CD32D62EE490C3A9E40C1B830C66" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="52F428B4409DE5CBE87F59C3B8A2937B" italics="true">S. jeanneli</emphasis>
</taxonomicName>
and stated they were deposited at 
<named-content id="ED60D67E7DEBF2776B4E535A178EFDBD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="ACB0F2902DE1942EF714C2A02C5F8A9D" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
. However, no record exists of these specimens being deposited, and they could not be found in the type collection or the general collection (R. Androw, R. Davidson and A. Seago, pers. comm., 
<date id="E2CC211C49A79A47A239B99A5A90C3DE" value="2024-01">January 2024</date>
). The specimens were also not found in any of the unprocessed material accessioned to 
<named-content id="AEDC79A772AEED9827786EB476C78092" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="45EC94BCDC857059E481CC0646C0783A" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
after Barr’s death. One dried-out vial of undetermined specimens was found with label data matching Barr’s type series. In it were six anillines, including 
<specimenCount id="F470DA14B4C4F1257A6F32BBC5D71D59" count="2" type="female">two female</specimenCount>
and 
<specimenCount id="D0C09B6A2D19AF33F4A59BF2AAE19577" count="1" type="male">one male</specimenCount>
<taxonomicName id="37623EB7A3B2FB389AA7D33DBA772D50" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="genus">
<emphasis id="57C08CD0768824DB51E9DAA5DE3D4211" italics="true">Serranillus</emphasis>
</taxonomicName>
. However, the male was not the species that Barr illustrated as 
<taxonomicName id="A7B49DF46E49100738B81C4EC944F04F" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="C573A5967E14661CFD97093ED6A78788" italics="true">S. jeanneli</emphasis>
</taxonomicName>
, but the larger species we call 
<emphasis id="FD3E19757E4D0AD04A8D008EC5261B5B" italics="true">S.</emphasis>
sp. “ 
<collectingRegion id="773940FFC8E30C3576463468BABDA838" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
, Riley Knob. ” To stabilize the name and clarify the identity of 
<taxonomicName id="D7AA4F896B96684B4EF79BBF3D2617D4" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="2618A6F4BFEE97ACB45854E3CF7E797D" italics="true">S. jeanneli</emphasis>
</taxonomicName>
, we have chosen as 
<typeStatus id="5D9BB57DC89610C7F382491A5649F1C8" type="neotype">neotype</typeStatus>
a male of the correct species that was collected by Barr on the same day, along the same forest road approximately 
<quantity id="2FAEE23F2AE05254677F12B89BC456FC" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
airline distance from the given type locality.
</paragraph>
<paragraph id="8DD6BF809144CCCA0F8DAB87C5359EEB">
We noted that the previously published sequences from individuals identified as 
<taxonomicName id="10E6E3ABCFE0D393956D54AADAEEB22B" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="BA6C9B5E3F1A280078C1935686C39ED6" italics="true">S. jeanneli</emphasis>
</taxonomicName>
(DNA 1084 and DNA 2309) were in a clade with 
<emphasis id="9D8CC30B198B0130BF2DB003FDC73A4E" italics="true">S.</emphasis>
sp. “ 
<collectingRegion id="F9DCBE045C09D38D8E339290A7A2718D" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
, Miller Cove ” in our phylogeny, and the genitalia were confirmed to match that species rather than our concept of 
<taxonomicName id="92F1C0AB78AA2B0AF33EA4284BF2A7F0" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="8EAF645C507B1083E8A4C503F522DC83" italics="true">S. jeanneli</emphasis>
</taxonomicName>
(D. Maddison pers. comm., 
<date id="D9290B6E25A94C66F782E085CF366D0C" value="2024-01">January 2024</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>