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<document id="1676CB4C3F8ABF8D53C2A47225FE53A9" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. & Hughes, Nigel C." docDate="2022" docId="03CB87846E7D1D21FF3AC2954BF3F31D" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Yanhaoia wynnei" docType="treatment" docVersion="6" lastPageNumber="342" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="341" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="45E7E96E27C24936649EDD012BB97093">Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:namePart id="4A4DA4D899220FE22DE20E39E9E1AAC2">Fortey, Richard A.</mods:namePart>
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<mods:affiliation id="9569D3D9EA798E02D563D772E952DBAF">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="FD44191BAD7D3084F2D46608B66FC693">Wernette, Shelly J.</mods:namePart>
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<mods:affiliation id="33137E518B333EE67818E88679EDD5DE">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. & Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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<mods:namePart id="20492B29600BD2718CD9A89384B3BF38">Hughes, Nigel C.</mods:namePart>
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<mods:title id="5857C831004E2C02C24050AF5DD10E13">Zootaxa</mods:title>
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<mods:date id="817548EC8FAEAA4896919A3533BC278C">2022</mods:date>
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<mods:number id="5AA51A200BF58CB75AB37634C8745468">2022-07-08</mods:number>
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<mods:number id="9553962B4268F65A39B20A732E94B8F9">5162</mods:number>
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<mods:number id="DB443E71D521A7A9F46CA9BCF0C92009">4</mods:number>
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<treatment id="03CB87846E7D1D21FF3AC2954BF3F31D" ID-DOI="http://doi.org/10.5281/zenodo.6810372" ID-GBIF-Taxon="197214809" ID-Zenodo-Dep="6810372" LSID="urn:lsid:plazi:treatment:03CB87846E7D1D21FF3AC2954BF3F31D" httpUri="http://treatment.plazi.org/id/03CB87846E7D1D21FF3AC2954BF3F31D" lastPageId="41" lastPageNumber="342" pageId="40" pageNumber="341">
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<subSubSection id="C37865196E7D1D20FF3AC2954810F3A2" box="[151,523,1748,1775]" pageId="40" pageNumber="341" type="nomenclature">
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<paragraph id="8BDD36926E7D1D20FF3AC2954810F3A2" blockId="40.[151,523,1748,1811]" box="[151,523,1748,1775]" pageId="40" pageNumber="341">
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<heading id="D09581FE6E7D1D20FF3AC2954810F3A2" bold="true" box="[151,523,1748,1775]" fontSize="11" level="1" pageId="40" pageNumber="341" reason="1">
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<taxonomicName id="4C624D116E7D1D20FF3AC2954810F3A2" authority="(Reed, 1915)" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[151,523,1748,1775]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="wynnei">
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<emphasis id="B916EA806E7D1D20FF3AC2954810F3A2" bold="true" box="[151,523,1748,1775]" pageId="40" pageNumber="341">
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<emphasis id="B916EA806E7D1D20FF3AC2954B7CF3A3" bold="true" box="[151,359,1748,1775]" italics="true" pageId="40" pageNumber="341">Yanhaoia wynnei</emphasis>
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(
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<bibRefCitation id="EFF34B636E7D1D20FEDAC2954818F3A2" author="Reed, F. R. C." box="[375,515,1748,1774]" pageId="40" pageNumber="341" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E7D1D20FF3AC2B84E06F238" pageId="40" pageNumber="341" type="description">
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<paragraph id="8BDD36926E7D1D20FF3AC2B84B0CF25F" blockId="40.[151,523,1748,1811]" box="[151,279,1785,1811]" pageId="40" pageNumber="341">
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<figureCitation id="13592A176E7D1D20FF3AC2B84B0CF25F" box="[151,279,1785,1811]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="40" pageNumber="341">Figs 13.4-9</figureCitation>
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</paragraph>
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<paragraph id="8BDD36926E7D1D20FF3AC37C4962F219" blockId="40.[151,1053,1853,1908]" box="[151,889,1853,1877]" pageId="40" pageNumber="341">
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1915
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<taxonomicName id="4C624D116E7D1D20FF5AC37C484BF219" authorityName="Reed" authorityYear="1915" box="[247,592,1853,1877]" class="Trilobita" family="Phacopidae" genus="Phacops" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="dagon" subGenus="Pterygometopus">
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<emphasis id="B916EA806E7D1D20FF5AC37C4B55F219" box="[247,334,1853,1877]" italics="true" pageId="40" pageNumber="341">Phacops</emphasis>
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(
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<emphasis id="B916EA806E7D1D20FEF0C37F4819F219" box="[349,514,1854,1877]" italics="true" pageId="40" pageNumber="341">Pterygometopus</emphasis>
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)
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<emphasis id="B916EA806E7D1D20FDBCC37C484BF219" box="[529,592,1853,1877]" italics="true" pageId="40" pageNumber="341">dagon</emphasis>
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</taxonomicName>
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; Reed, p. 53–54, pl. 9, fig. 3.
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</paragraph>
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<paragraph id="8BDD36926E7D1D20FF3AC31D4E06F238" blockId="40.[151,1053,1853,1908]" box="[151,1053,1884,1908]" pageId="40" pageNumber="341">
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1915
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<taxonomicName id="4C624D116E7D1D20FF5AC31D48D1F238" authorityName="Reed" authorityYear="1915" box="[247,714,1884,1908]" class="Trilobita" family="Phacopidae" genus="Phacops" kingdom="Animalia" order="Phacopida" pageId="40" pageNumber="341" phylum="Arthropoda" rank="variety" species="dagon" subGenus="Pterygometopus" variety="wynnei">
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<emphasis id="B916EA806E7D1D20FF5AC31D4B55F238" box="[247,334,1884,1908]" italics="true" pageId="40" pageNumber="341">Phacops</emphasis>
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(
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<emphasis id="B916EA806E7D1D20FEF0C31C4819F238" box="[349,514,1885,1908]" italics="true" pageId="40" pageNumber="341">Pterygometopus</emphasis>
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)
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<emphasis id="B916EA806E7D1D20FDBCC31D484BF238" box="[529,592,1884,1908]" italics="true" pageId="40" pageNumber="341">dagon</emphasis>
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var.
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<emphasis id="B916EA806E7D1D20FD2FC31C48D1F238" box="[642,714,1885,1908]" italics="true" pageId="40" pageNumber="341">wynnei</emphasis>
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</taxonomicName>
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; Reed, p. 54–55, pl. 9, figs 7–15.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E7D1D21FF3AC3E348CBF429" lastPageId="41" lastPageNumber="342" pageId="40" pageNumber="341" type="materials_examined">
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<paragraph id="8BDD36926E7D1D21FF3AC3E34842F40D" blockId="40.[151,1436,1954,2017]" lastBlockId="41.[151,1437,150,1617]" lastPageId="41" lastPageNumber="342" pageId="40" pageNumber="341">
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<emphasis id="B916EA806E7D1D20FF3AC3E34B1DF2F0" bold="true" box="[151,262,1954,1980]" pageId="40" pageNumber="341">Material.</emphasis>
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<typeStatus id="54D988306E7D1D20FEA0C3E24B66F2F1" box="[269,381,1955,1981]" pageId="40" pageNumber="341" type="lectotype">Lectotype</typeStatus>
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(selected herein): partial exoskeleton from Hwe Mawng,
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<figureCitation id="13592A176E7D1D20FBAEC3E34E7CF2F0" box="[1027,1127,1954,1980]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="40" pageNumber="341">Fig. 13.8</figureCitation>
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(
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<bibRefCitation id="EFF34B636E7D1D20FBD8C3E34EE2F2F1" author="Reed, F. R. C." box="[1141,1273,1954,1981]" pageId="40" pageNumber="341" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 9, fig. 10),
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<collectionCode id="ED73AE576E7D1D20FF3AC3874ADFF2AC" box="[151,196,1990,2016]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="40" pageNumber="341">GSI</collectionCode>
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11576. Other material: incomplete cephalic shield from Hwe Mawng,
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<figureCitation id="13592A176E7D1D20FC71C3874E5BF2AD" box="[988,1088,1990,2017]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="40" pageNumber="341">Fig. 13.5</figureCitation>
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(
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<bibRefCitation id="EFF34B636E7D1D20FBE2C3874EC8F2AD" author="Reed, F. R. C." box="[1103,1235,1990,2017]" pageId="40" pageNumber="341" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 9, fig. 7),
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<collectionCode id="ED73AE576E7D1D20FAC2C3874F87F2AC" box="[1391,1436,1990,2016]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="40" pageNumber="341">GSI</collectionCode>
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11573; 2 incomplete cephalic shields from Mong Ha,
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<figureCitation id="13592A176E7C1D21FD51C4D64979F5FD" box="[764,866,151,177]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.6</figureCitation>
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and unfigured (
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<bibRefCitation id="EFF34B636E7C1D21FBB5C4D64E86F5FD" author="Reed, F. R. C." box="[1048,1181,150,177]" pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 9, fig 11,8 respectively),
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<collectionCode id="ED73AE576E7C1D21FF42C4FA4B07F599" box="[239,284,187,213]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="41" pageNumber="342">GSI</collectionCode>
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11577, 11574 respectively; librigenae from Mong Ha, Nawng Yun, and Hwe Mawng (
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<bibRefCitation id="EFF34B636E7C1D21FB42C4FA4F6FF599" author="Reed, F. R. C." box="[1263,1396,186,213]" pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 9, figs 9,13,14 respectively),
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<collectionCode id="ED73AE576E7C1D21FE4DC49E4816F5B5" box="[480,525,223,249]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="41" pageNumber="342">GSI</collectionCode>
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11575, 11579, 11580 respectively;_pygidia from Hpawkyi and Nawng Yun,
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<figureCitation id="13592A176E7C1D21FADDC49E4AD0F451" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.7</figureCitation>
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,
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<figureCitation id="13592A176E7C1D21FF62C5424AC6F451" box="[207,221,259,285]" captionStart="FIGURE 9" captionStartId="24.[151,250,1794,1818]" captionTargetBox="[217,1370,669,1771]" captionTargetId="figure-280@24.[217,1370,669,1771]" captionTargetPageId="24" captionText="FIGURE 9. Mioptychopyge thebawi (Reed) from the Hwe Mawng Beds (uppermost Katian) and Li-Lu Formation (Katian), northern Shan State, Myanmar. 1a, b. Cranidium (original of Reed 1915, pl. 6, fig. 5), GSI 11525, Nawng Yun, original and retrodeformation. 2. Hypostome (original of Reed 1915, pl. 6, fig. 9), GSI 11529, Lilu. 3a, b. Pygidium (original of Reed 1915, pl. 7, fig. 1), GSI 11533, Hwe Mawng, original and reetrodeformation. 4a, b. Cranidium (original of Reed 1915, pl. 6, fig. 7), GSI 11527, Hkawnhkok, original and retrodeformation. 5. Pygidium (original of Reed 1915, pl. 6, fig. 8), GSI 11528, Mong Ha. 6a, b. Pygidium (original of Reed 1915, pl. 6, fig. 12), GSI 11532, Hwe Mawng, original and retrodeformation. 7. Pygidium (original of Reed 1915, pl. 6, fig. 11), GSI 11531, Hwe Mawng. Scale bar = 5 mm for 1, 3–7; = 2 mm for 2." figureDoi="http://doi.org/10.5281/zenodo.6810314" httpUri="https://zenodo.org/record/6810314/files/figure.png" pageId="41" pageNumber="342">9</figureCitation>
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(
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<bibRefCitation id="EFF34B636E7C1D21FF46C5424B74F451" author="Reed, F. R. C." box="[235,367,258,285]" pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed, 1915</bibRefCitation>
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, pl. 9, figs 12,15)
|
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<collectionCode id="ED73AE576E7C1D21FD9AC542487FF451" box="[567,612,259,285]" country="India" httpUri="http://grbio.org/cool/jprb-qxkv" name="Geological Survey of India" pageId="41" pageNumber="342">GSI</collectionCode>
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11578, 11581. All specimens from Hwe Mawng Beds (Katian) at Hpakhi, Hwe Mawng, Mong Ha, or Nawng Yun.
|
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</paragraph>
|
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<paragraph id="8BDD36926E7C1D21FF6AC50B48CBF429" blockId="41.[151,1437,150,1617]" box="[199,720,330,357]" pageId="41" pageNumber="342">
|
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<emphasis id="B916EA806E7C1D21FF6AC50B4B41F428" bold="true" box="[199,346,330,356]" pageId="41" pageNumber="342">Occurrence.</emphasis>
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Hwe Mawng Beds,
|
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<typeStatus id="54D988306E7C1D21FD92C50A4874F429" box="[575,623,331,357]" pageId="41" pageNumber="342">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E7C1D21FF6AC52F4F18F115" pageId="41" pageNumber="342" type="description">
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<paragraph id="8BDD36926E7C1D21FF6AC52F4E31F6E9" blockId="41.[151,1437,150,1617]" pageId="41" pageNumber="342">
|
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<emphasis id="B916EA806E7C1D21FF6AC52F4B42F4C4" bold="true" box="[199,345,366,392]" pageId="41" pageNumber="342">Description.</emphasis>
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<bibRefCitation id="EFF34B636E7C1D21FECDC52E4BE5F4C5" author="Reed, F. R. C." box="[352,510,366,393]" pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed’s (1915)</bibRefCitation>
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general description can be supplemented by comments on particular features. The
|
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<typeStatus id="54D988306E7C1D21FF3AC5D24AE5F4E1" box="[151,254,403,429]" pageId="41" pageNumber="342" type="lectotype">lectotype</typeStatus>
|
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is distorted, but the cast shows the best surface detail. Other cephalic shields are internal moulds for the most part and as a result both the axial and glabellar furrows are deep and wide compared with their expression on the dorsal surface. The cephalic shield was originally somewhat less than twice as wide as long or narrower (
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<figureCitation id="13592A176E7C1D21FAC2C59A4AD7F755" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.4</figureCitation>
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). Three of the specimens displaying the glabella (Figs 13.4,6,8) show that the short S1 basal glabellar furrow is forked close to its base, and that the posterior fork outlines a basal lateral glabellar lobe –this is clearly seen on the
|
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<typeStatus id="54D988306E7C1D21FF3AC6064AE5F72D" box="[151,254,583,609]" pageId="41" pageNumber="342" type="lectotype">lectotype</typeStatus>
|
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and in
|
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|
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<figureCitation id="13592A176E7C1D21FEE2C6064BA2F72D" box="[335,441,583,609]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Figs 13.4</figureCitation>
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,
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<figureCitation id="13592A176E7C1D21FE10C6064BD0F72D" box="[445,459,583,609]" captionStart="FIGURE 6" captionStartId="19.[151,250,1801,1825]" captionTargetBox="[199,1388,181,1777]" captionTargetId="figure-18@19.[199,1388,181,1777]" captionTargetPageId="19" captionText="FIGURE 6. Birmanites birmanicus (Reed) from the Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 1. Hypostome (original of Reed 1915, pl. 5, fig. 17), GSI 11519, Hwe Mawng. 2a, b. Cephalon (original of Reed 1915, pl. 6, fig. 2), GSI 11522, Hwe Mawng, original and retrodeformation. 3. Partial specimen (original of Reed 1915, pl. 6, fig. 3), GSI 11523, Hpakhi. 4a, b. Pygidium (original of Reed 1915, pl. 6, fig. 4), GSI 11524, Hwe Mawng, original and retrodeformation. 5. Cranidium (original of Reed 1915, pl. 7, fig. 2), GSI 11534, Hpakhi. 6. Partial specimen (original of Reed 1915, pl. 5, figs 15,16), GSI 11518, Hpakhi. Scale bars = 2 mm for 1; = 5 mm for 2–6." figureDoi="http://doi.org/10.5281/zenodo.6810306" httpUri="https://zenodo.org/record/6810306/files/figure.png" pageId="41" pageNumber="342">6</figureCitation>
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(right). S2 is gently forwardly–directed, and the sigmoid form of gently backwardly-directed S3 is best shown in
|
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<figureCitation id="13592A176E7C1D21FE6CC62A4833F7C9" box="[449,552,619,645]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.6</figureCitation>
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. Certain species of
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<taxonomicName id="4C624D116E7C1D21FCACC62A4991F7C9" authorityName="Cooper" authorityYear="1931" box="[769,906,619,645]" class="Trilobita" family="Pterygometopidae" genus="Calyptaulax" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="genus">
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|||
|
|
<emphasis id="B916EA806E7C1D21FCACC62A4991F7C9" box="[769,906,619,645]" italics="true" pageId="41" pageNumber="342">Calyptaulax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
show a similar structure. The anterior cranidial border is shown on the
|
|||
|
|
<typeStatus id="54D988306E7C1D21FE0DC6CE481CF7E5" box="[416,519,655,681]" pageId="41" pageNumber="342" type="lectotype">lectotype</typeStatus>
|
|||
|
|
and on the cranidium
|
|||
|
|
<figureCitation id="13592A176E7C1D21FCA9C6CE4977F7E5" box="[772,876,655,681]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.4</figureCitation>
|
|||
|
|
, where it is narrow and rim-like, especially medially, and defined by a shallow furrow. Eyes are very large, half (exsag.) cranidial length (sag.), the deeply-defined palpebral rims flipped upwards from the level of the intraocular cheeks. The number of dorso-ventral lens files is not precisely determinable but certainly exceeds twenty, on high standing eyes. The free cheek illustrated by
|
|||
|
|
<bibRefCitation id="EFF34B636E7C1D21FACEC6BA4AC0F675" author="Reed, F. R. C." pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915</bibRefCitation>
|
|||
|
|
, pl. 9, fig. 9) suggests about 15 lenses per dorso-ventral file. The
|
|||
|
|
<typeStatus id="54D988306E7C1D21FC1FC75E4E02F675" box="[946,1049,799,825]" pageId="41" pageNumber="342" type="lectotype">lectotype</typeStatus>
|
|||
|
|
clearly shows deeply and coarsely pitted sculpture on the intraocular cheeks. The same sculpture extends on to the lateral parts of the fixed cheeks, but more feebly. There is no evidence of a genal spine. The incomplete six thoracic segments on the
|
|||
|
|
<typeStatus id="54D988306E7C1D21FB66C7264F29F6CD" box="[1227,1330,871,897]" pageId="41" pageNumber="342" type="lectotype">lectotype</typeStatus>
|
|||
|
|
show the same kind of sculpture anterior to the deep and narrow epifacetal pleural furrows.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDD36926E7C1D21FF6AC7EE4F18F115" blockId="41.[151,1437,150,1617]" pageId="41" pageNumber="342">
|
|||
|
|
Well-preserved subtrapezoidal pygidium is best shown by cast from external mould (
|
|||
|
|
<figureCitation id="13592A176E7C1D21FB25C7EE4EEAF685" box="[1160,1265,943,969]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.9</figureCitation>
|
|||
|
|
, right), 75% as long as wide. Eight (?faint ninth) axial rings, but internal mould (
|
|||
|
|
<figureCitation id="13592A176E7C1D21FCC1C79249C9F6A1" box="[876,978,979,1005]" captionStart="FIGURE 13" captionStartId="43.[151,250,1547,1571]" captionTargetBox="[151,1436,181,1520]" captionTargetId="figure-18@43.[151,1436,181,1524]" captionTargetPageId="43" captionText="FIGURE 13. (1) Hadromeros? submitis (Reed), Cranidium (original of Reed 1915, pl. 8, fig. 13), GSI 11560, Li-Lu Formation (Katian); Lilu, northern Shan State, Myanmar. (2) Placoparina? dravidicus (Reed), Pygidium (original of Reed 1915, pl. 8, fig. 12), GSI 11559, Lower Naungkangyi Beds; Mâ Shio, northern Shan State, Myanmar. (3) Sphaerocoryphe sp. (Reed), Cranidium (original of Reed 1906, pl. 5, fig. 26), GSI 8340, Naungkangyi Beds, Insang, northern part of Shan State, Myanmar. (4–9) Yanhaoia wynnei (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 4. Cranidium (original of Reed 1915, pl. 9, fig. 3), GSI 11569, Mân-ngai. 5. Partial cranidium (original of Reed 1915, pl. 9 fig. 7), GSI 11573, Hwe Mawng. 6. Partial cranidium (original of Reed 1915, pl. 9, fig. 11), GSI 11577, Mong Ha. 7. Pygidium (original of Reed 1915, pl. 9, fig. 12), GSI 11578, Hpakhi. 8. Cranidium and thorax (cast from lectotype, original of Reed 1915, pl. 9, fig. 10), GSI 11576, Hwe Mawng. 9. Pygidium (original of Reed 1915, pl. 9, fig. 5), GSI 11571, Nawng Yu. (10–11) Dalmanitina? dagon (Reed) from the Li-Lu Formation (Katian) and Hwe Mawng Beds (uppermost Katian), northern Shan State, Myanmar. 10. Cranidium (refigured from Reed 1915, pl. 9, fig. 2), GSI 11568-11572, Mân-ngai. 11. Cephalon (original of Reed 1915, pl. 9, fig. 1), GSI 11568, Mân-ngai. (12) Illaenus sp. (Reed), Partial cranidium (refigured from Reed 1917, pl. 8, fig. 3), GSI 11906, Shihtien Formation (Darriwilian), Pupiao, Baoshan Prefecture, western Yunnan, China. Scale bar = 2 mm for 1, 3, 4, 6–9; = 5 mm for 2, 5, 10–12." figureDoi="http://doi.org/10.5281/zenodo.6810326" httpUri="https://zenodo.org/record/6810326/files/figure.png" pageId="41" pageNumber="342">Fig. 13.7</figureCitation>
|
|||
|
|
) would have displayed at least ten. Interpleural furrows much weaker than pleural furrows, defining six or seven ribs fading out on border, but again internal mould certainly shows more ribs extending to the posterior axial rings. Border is gently concave, most noticeably laterally. Posterior termination of pygidium behind axis shows a tendency to come to a median point.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37865196E7C1D21FF6AC0234BF3F31D" pageId="41" pageNumber="342" type="discussion">
|
|||
|
|
<paragraph id="8BDD36926E7C1D21FF6AC0234BF3F31D" blockId="41.[151,1437,150,1617]" pageId="41" pageNumber="342">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FF6AC0234B57F130" bold="true" box="[199,332,1122,1148]" pageId="41" pageNumber="342">Discussion.</emphasis>
|
|||
|
|
<bibRefCitation id="EFF34B636E7C1D21FEF9C0224BFBF131" author="Reed, F. R. C." box="[340,480,1122,1149]" pageId="41" pageNumber="342" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915)</bibRefCitation>
|
|||
|
|
distinguished
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FD28C02248CFF130" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[645,724,1123,1148]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FD28C02248CFF130" box="[645,724,1123,1148]" italics="true" pageId="41" pageNumber="342">wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as a subspecies of his taxon
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FBB0C0224F87F131" authorityName="Reed" authorityYear="1915" box="[1053,1436,1123,1149]" class="Trilobita" family="Phacopidae" genus="Phacops" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="dagon" subGenus="Pterygometopus">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FBB0C0224F87F131" box="[1053,1436,1123,1149]" italics="true" pageId="41" pageNumber="342">Phacops (Pterygometopus) dagon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from the Upper Naungkangyi Beds. Quite apart from their stratigraphical separation, the cranidium of
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FA88C0C64F70F1ED" authorityName="Reed" authorityYear="1915" box="[1317,1387,1159,1185]" class="Trilobita" family="Phacopidae" genus="Phacops" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="dagon">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FA88C0C64F70F1ED" box="[1317,1387,1159,1185]" italics="true" pageId="41" pageNumber="342">dagon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from that of
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FEF2C0EA4BB5F188" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[351,430,1195,1220]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FEF2C0EA4BB5F188" box="[351,430,1195,1220]" italics="true" pageId="41" pageNumber="342">wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in having a distinct and wider cephalic border (sag.), relatively wider basal part of the glabella, and the first glabellar furrow S1 has less distinct distal bifurcation. It is regarded as belonging to a different genus. Hence
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FE95C0B2488FF040" box="[312,660,1267,1293]" class="Trilobita" family="Pterygometopidae" genus="Pterygometopus" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="subSpecies" species="dagon" subSpecies="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FE95C0B2488FF040" box="[312,660,1267,1293]" italics="true" pageId="41" pageNumber="342">Pterygometopus dagon wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is here elevated to species rank, and not regarded as closely related to
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FF1BC1564B72F07D" box="[182,361,1303,1329]" class="Trilobita" family="Pterygometopidae" genus="Pterygometopus" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="subSpecies" species="dagon" subSpecies="dagon">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FF1BC1564B72F07D" box="[182,361,1303,1329]" italics="true" pageId="41" pageNumber="342">P. dagon dagon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. On the other hand,
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FDF5C15648BCF07C" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[600,679,1303,1328]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FDF5C15648BCF07C" box="[600,679,1303,1328]" italics="true" pageId="41" pageNumber="342">wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is also different from chasmopine pterygometopids with typically inflated and enlarged lateral glabellar lobes. It more closely resembles genera such as
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FBCCC17A4ED7F019" authorityName="Delo" authorityYear="1935" box="[1121,1228,1339,1365]" class="Trilobita" family="Pterygometopidae" genus="Achatella" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FBCCC17A4ED7F019" box="[1121,1228,1339,1365]" italics="true" pageId="41" pageNumber="342">Achatella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which, according to
|
|||
|
|
<bibRefCitation id="EFF34B636E7C1D21FF18C11E4BBEF035" author="Swisher, R. E. & Westrop, S. R. & Amati, L." box="[181,421,1375,1401]" pageId="41" pageNumber="342" pagination="59 - 78" refId="ref47624" refString="Swisher, R. E., Westrop, S. R. & Amati, L. (2016) Systematics and biogeographical significance of the Upper Ordovician pterygometopine trilobite Achatella Delo, 1935. Journal of Paleontology, 90, 59 - 78. https: // doi. org / 10.1017 / jpa. 2015.71" type="journal article" year="2016">
|
|||
|
|
Swisher
|
|||
|
|
<emphasis id="B916EA806E7C1D21FEBAC11E4B4BF035" box="[279,336,1375,1401]" italics="true" pageId="41" pageNumber="342">et al.</emphasis>
|
|||
|
|
(2016)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, is a Baltic/Laurentian clade, and one with smaller eyes and generally distinct tuberculate sculpture, unlike
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FEF5C1C24BBCF0D0" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[344,423,1411,1436]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FEF5C1C24BBCF0D0" box="[344,423,1411,1436]" italics="true" pageId="41" pageNumber="342">wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The presence of three pairs of well-developed lateral glabellar furrows is a plesiomorphic character. Small lateral basal glabellar lobes which become eliminated in more advanced species is pterygometopid, and exceptionally large eyes are typical of several taxa. Although the
|
|||
|
|
<typeStatus id="54D988306E7C1D21FC19C18A49FFF0A9" box="[948,996,1483,1509]" pageId="41" pageNumber="342">type</typeStatus>
|
|||
|
|
species of
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FBCAC18A4ECBF0A9" authorityName="Zhou, Yuan & Zhou" authorityYear="1998" box="[1127,1232,1483,1509]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FBCAC18A4ECBF0A9" box="[1127,1232,1483,1509]" italics="true" pageId="41" pageNumber="342">Yanhaoia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is not completely known and is older,
|
|||
|
|
<taxonomicName id="4C624D116E7C1D21FE29C1AE4BE8F344" baseAuthorityName="Reed" baseAuthorityYear="1915" box="[388,499,1519,1545]" class="Trilobita" family="Pterygometopidae" genus="Yanhaoia" kingdom="Animalia" order="Phacopida" pageId="41" pageNumber="342" phylum="Arthropoda" rank="species" species="wynnei">
|
|||
|
|
<emphasis id="B916EA806E7C1D21FE29C1AE4B8FF344" box="[388,404,1519,1544]" italics="true" pageId="41" pageNumber="342">Y</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B916EA806E7C1D21FE09C1AE4BE8F344" box="[420,499,1519,1544]" italics="true" pageId="41" pageNumber="342">wynnei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
does share with it a narrow glabella posteriorly, and particularly large eyes, and, given the other similarities between Burmese and SW Chinese taxa, is assigned provisionally to this genus pending a revision of the whole group.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|