169 lines
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169 lines
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<document id="D67B5EAE47A6F1F2187DC9EE38C76547" ID-CLB-Dataset="93239" ID-DOI="10.11646/zootaxa.5162.4.1" ID-GBIF-Dataset="7d343017-483e-43be-9439-d06e748fc54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6810290" ID-ZooBank="DD2279FA-E8F1-4951-A5CA-91082E875580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657270124299" checkinUser="plazi" docAuthor="Fortey, Richard A., Wernette, Shelly J. & Hughes, Nigel C." docDate="2022" docId="03CB87846E711D2CFF3AC6F34AE8F044" docLanguage="en" docName="zootaxa.5162.4.1.pdf" docOrigin="Zootaxa 5162 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Reedocalymene Kobayashi 1951" docType="treatment" docVersion="6" lastPageNumber="337" masterDocId="FFF2FFFC6E551D08FFADC4414A1BF54C" masterDocTitle="Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China" masterLastPageNumber="356" masterPageNumber="301" pageNumber="337" updateTime="1699356965331" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="06887037C64E20D6DD584C39696D1F79">
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<mods:title id="35ABE57186F39295011BF9BDD9093004">Revision of F. R. C. Reed’s Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<mods:name id="05D61789437563C41C5B3972E90E9339" type="personal">
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<mods:roleTerm id="D1165DD1FDFD86C365EFDE8C42E785A2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E594B297A26C07695787E57CE5441C94">Fortey, Richard A.</mods:namePart>
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<mods:affiliation id="1F0BA36C9A20DC0C836130E9B6CFAC16">Department of Earth Sciences, Museum of Natural History, Cromwell Road, London, SW 7 5 BD, UK.</mods:affiliation>
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</mods:name>
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<mods:name id="E70250784772F3C6F5BCCDD9E2D1A7A3" type="personal">
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<mods:roleTerm id="0479E411ACC176B76BBCE19E122E3222">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4B12A602F6F129382363FD3497565DA8">Wernette, Shelly J.</mods:namePart>
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<mods:affiliation id="04F328F77265818576DBC392FF1621F4">Department of Earth and Planetary Sciences, University of California, Riverside, CA 92521, USA. & Department of Geography and Environmental Studies, Texas State University, San Marcos, TX 78666, USA</mods:affiliation>
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</mods:name>
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<mods:name id="CA3AE59D953096BD26C5126E9027DA9F" type="personal">
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<mods:roleTerm id="19A578276582181A6930FA5C40AEFBA2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="939E1ED25A6E8BDDCF48A494F05475C0">Hughes, Nigel C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A7D425089333FD9760BA817AD70D8C28">text</mods:typeOfResource>
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<mods:title id="A559BA369460DB50B11F62CEC2DADD62">Zootaxa</mods:title>
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<mods:part id="92515963B59A76F660B55B51C3A0370B">
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<mods:date id="724B25661C8144E3138D2ADA2513A87F">2022</mods:date>
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<mods:number id="0777F90ACCE3561B716DCD9133AAA837">2022-07-08</mods:number>
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<mods:detail id="B59FA1D8D6AE0B1FD5AE99A184940F96" type="volume">
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<mods:number id="D3E186EFF1C8EC45FF3B5A669C1458BE">5162</mods:number>
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<mods:detail id="992303AACF2AE3C1D33B887F0BDC233D" type="issue">
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<mods:number id="20CBD84B1FC9DA1B0CCF1686EAE98923">4</mods:number>
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<mods:identifier id="84CE7C91D20B3C3469B515F91F222E83" type="DOI">10.11646/zootaxa.5162.4.1</mods:identifier>
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<treatment id="03CB87846E711D2CFF3AC6F34AE8F044" ID-DOI="http://doi.org/10.5281/zenodo.6810352" ID-GBIF-Taxon="197214826" ID-Zenodo-Dep="6810352" LSID="urn:lsid:plazi:treatment:03CB87846E711D2CFF3AC6F34AE8F044" httpUri="http://treatment.plazi.org/id/03CB87846E711D2CFF3AC6F34AE8F044" lastPageNumber="337" pageId="36" pageNumber="337">
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<subSubSection id="C37865196E711D2CFF3AC6F34830F780" box="[151,555,690,717]" pageId="36" pageNumber="337" type="nomenclature">
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<paragraph id="8BDD36926E711D2CFF3AC6F34830F780" blockId="36.[151,555,690,717]" box="[151,555,690,717]" pageId="36" pageNumber="337">
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<heading id="D09581FE6E711D2CFF3AC6F34830F780" bold="true" box="[151,555,690,717]" fontSize="11" level="1" pageId="36" pageNumber="337" reason="1">
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<taxonomicName id="4C624D116E711D2CFF3AC6F34830F780" ID-CoL="77H3" authority="Kobayashi, 1951" authorityName="Kobayashi" authorityYear="1951" box="[151,555,690,717]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFF3AC6F34830F780" bold="true" box="[151,555,690,717]" pageId="36" pageNumber="337">
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<emphasis id="B916EA806E711D2CFF3AC6F34B4FF781" bold="true" box="[151,340,690,717]" italics="true" pageId="36" pageNumber="337">Reedocalymene</emphasis>
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<bibRefCitation id="EFF34B636E711D2CFEF6C6F34830F780" author="Kobayashi, T." box="[347,555,690,716]" pageId="36" pageNumber="337" pagination="1 - 87" refId="ref45054" refString="Kobayashi, T. (1951) On the Ordovician trilobites in central China. Journal of the Faculty of Science Imperial University of Tokyo, Section II, Geology, 8, 1 - 87." type="journal article" year="1951">Kobayashi, 1951</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E711D2CFF3AC6B64E72F643" box="[151,1129,759,783]" pageId="36" pageNumber="337" type="type_taxon">
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<paragraph id="8BDD36926E711D2CFF3AC6B64E72F643" blockId="36.[151,1129,759,783]" box="[151,1129,759,783]" pageId="36" pageNumber="337">
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<materialsCitation id="3B0A3CCF6E711D2CFF3AC6B64E7EF643" ID-GBIF-Occurrence="3861306321" box="[151,1125,759,783]" location="Pupaio Formation" pageId="36" pageNumber="337" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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<emphasis id="B916EA806E711D2CFF3AC6B64B38F643" bold="true" box="[151,291,759,783]" pageId="36" pageNumber="337">
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<typeStatus id="54D988306E711D2CFF3AC6B64AD7F643" box="[151,204,759,783]" pageId="36" pageNumber="337">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C624D116E711D2CFE87C6B6486AF643" ID-CoL="Q6CB" authority="Reed, 1917" authorityName="Reed" authorityYear="1917" box="[298,625,759,783]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
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<emphasis id="B916EA806E711D2CFE87C6B64BEEF643" box="[298,501,759,783]" italics="true" pageId="36" pageNumber="337">Calymene unicornis</emphasis>
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<bibRefCitation id="EFF34B636E711D2CFE56C6B6486AF643" author="Reed, F. R. C." box="[507,625,759,783]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed, 1917</bibRefCitation>
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</taxonomicName>
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,
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<location id="8EBD60496E711D2CFDD1C6B6492FF643" LSID="urn:lsid:plazi:treatment:03CB87846E711D2CFF3AC6F34AE8F044:8EBD60496E711D2CFDD1C6B6492FF643" box="[636,820,759,783]" name="Pupaio Formation" pageId="36" pageNumber="337" stateProvince="Yunnan">Pupaio Formation</location>
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,
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<collectingRegion id="49A6F8706E711D2CFC90C6B94996F643" box="[829,909,760,783]" country="China" name="Yunnan" pageId="36" pageNumber="337">Yunnan</collectingRegion>
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, original designation
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37865196E711D2CFF3AC77C4AE8F044" pageId="36" pageNumber="337" type="discussion">
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<paragraph id="8BDD36926E711D2CFF3AC77C4AE8F044" blockId="36.[151,1437,829,1288]" pageId="36" pageNumber="337">
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<emphasis id="B916EA806E711D2CFF3AC77C4B07F61B" bold="true" box="[151,284,829,855]" pageId="36" pageNumber="337">Discussion.</emphasis>
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<bibRefCitation id="EFF34B636E711D2CFE89C77C4BD9F61B" author="Reed, F. R. C." box="[292,450,829,855]" pageId="36" pageNumber="337" pagination="1 - 69" refId="ref46843" refString="Reed, F. R. C. (1917) Ordovician and Silurian fossils from Yun-nan. Palaeontologia Indica, 6 (3), 1 - 69." type="journal article" year="1917">Reed’s (1917)</bibRefCitation>
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species is the nominated type species of
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<taxonomicName id="4C624D116E711D2CFC3CC77C4E5AF61B" authorityName="Kobayashi" authorityYear="1951" box="[913,1089,829,855]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFC3CC77C4E5AF61B" box="[913,1089,829,855]" italics="true" pageId="36" pageNumber="337">Reedocalymene</emphasis>
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</taxonomicName>
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, although
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<bibRefCitation id="EFF34B636E711D2CFB15C77F4F32F614" author="Lu, Y. - H." box="[1208,1321,829,856]" pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu (1975)</bibRefCitation>
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described much better preserved material of the same genus. In a cladistic analysis of the Subfamily
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<taxonomicName id="4C624D116E711D2CFB3DC7204B09F6D3" authority="Turvey (2002)" authorityName="Turvey" authorityYear="2002" class="Trilobita" family="Calymenidae" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="subFamily" subFamily="Reedocalymeninae">
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Reedocalymeninae
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<bibRefCitation id="EFF34B636E711D2CFAC1C7234B09F6D3" author="Turvey, S. T." pageId="36" pageNumber="337" pagination="53 - 68" refId="ref47933" refString="Turvey, S. T. (2002) Phylogeny of the Reedocalymeninae (Trilobita): implications for Early Ordovician biogeography of Gondwana. In: Crame, A. J. & Owen, A. W. (Eds.), Palaeobiogeography and biodiversity change: the Ordovician and Mesozoic - Cenozoic radiations. Geological Society of London Special Publications, 194, 53 - 68." type="journal article" year="2002">Turvey (2002)</bibRefCitation>
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</taxonomicName>
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showed that
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<taxonomicName id="4C624D116E711D2CFE07C7C44841F6D3" authorityName="Kobayashi" authorityYear="1951" box="[426,602,901,927]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFE07C7C44841F6D3" box="[426,602,901,927]" italics="true" pageId="36" pageNumber="337">Reedocalymene</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C624D116E711D2CFD3FC7C449EBF6EC" authority="Kobayashi, 1951" authorityName="Kobayashi" authorityYear="1951" box="[658,1008,901,928]" class="Trilobita" family="Bathycheilidae" genus="Calymenesun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFD3FC7C44932F6D3" box="[658,809,901,927]" italics="true" pageId="36" pageNumber="337">Calymenesun</emphasis>
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<bibRefCitation id="EFF34B636E711D2CFC9CC7C449EBF6EC" author="Kobayashi, T." box="[817,1008,901,928]" pageId="36" pageNumber="337" pagination="1 - 87" refId="ref45054" refString="Kobayashi, T. (1951) On the Ordovician trilobites in central China. Journal of the Faculty of Science Imperial University of Tokyo, Section II, Geology, 8, 1 - 87." type="journal article" year="1951">Kobayashi, 1951</bibRefCitation>
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</taxonomicName>
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are closely related.
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<bibRefCitation id="EFF34B636E711D2CFB79C7C74F24F6EC" author="Lu, Y. - H." box="[1236,1343,901,928]" pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu (1975</bibRefCitation>
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, p. 448) had noted this previously, and provided some distinguishing characters, supplemented further by
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<bibRefCitation id="EFF34B636E711D2CFB62C7EB4F82F68F" author="Peng, S. - C. & Lin, T. - R. & Li, Y." box="[1231,1433,937,964]" pageId="36" pageNumber="337" pagination="63 - 75" refId="ref46395" refString="Peng, S. - C., Lin, T. - R. & Li, Y. (2000) Notes on the genus Reedocalymene Kobayashi, 1951 (Trilobita, Ordovician). Acta Palaeontologica Sinica, 39, 63 - 75." type="journal article" year="2000">
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Peng
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<emphasis id="B916EA806E711D2CFAA0C7EB4F25F68F" box="[1293,1342,937,963]" italics="true" pageId="36" pageNumber="337">et al</emphasis>
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. (2000)
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</bibRefCitation>
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. However, some of these are contradictory.
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<bibRefCitation id="EFF34B636E711D2CFD28C78F48EAF6A4" author="Lu, Y. - H." box="[645,753,973,1000]" pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu (1975</bibRefCitation>
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, p. 447) stated of
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<taxonomicName id="4C624D116E711D2CFC6AC78C4E45F6AB" authorityName="Kobayashi" authorityYear="1951" box="[967,1118,973,999]" class="Trilobita" family="Bathycheilidae" genus="Calymenesun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFC6AC78C4E45F6AB" box="[967,1118,973,999]" italics="true" pageId="36" pageNumber="337">Calymenesun</emphasis>
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</taxonomicName>
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that “the pygidium is quite different from all others of the Calymenids [sic] in having a very narrow border.” On p. 448 he states (of similarity between the two genera) that they share “a broad border on the pygidium”. He stated also that an important feature is the shape of the glabella which is claimed as “semioval” in
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<taxonomicName id="4C624D116E711D2CFCCCC0784E0AF11F" authorityName="Kobayashi" authorityYear="1951" box="[865,1041,1081,1107]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B916EA806E711D2CFCCCC0784E0AF11F" box="[865,1041,1081,1107]" italics="true" pageId="36" pageNumber="337">Reedocalymene</emphasis>
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</taxonomicName>
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, as it is on
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<taxonomicName id="4C624D116E711D2CFB34C07B4F2CF11F" authority="Yi" authorityName="Yi" box="[1177,1335,1081,1107]" class="Trilobita" family="Calymenidae" genus="Reedocalymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="expansa">
|
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<emphasis id="B916EA806E711D2CFB34C07B4F0DF11F" box="[1177,1302,1082,1107]" italics="true" pageId="36" pageNumber="337">R. expansa</emphasis>
|
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Yi
|
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</taxonomicName>
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(see
|
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<bibRefCitation id="EFF34B636E711D2CFADBC07B4AC9F134" author="Lu, Y. - H." pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu, 1975</bibRefCitation>
|
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|
, pl. 46, fig. 4). However, the
|
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|
<typeStatus id="54D988306E711D2CFD88C01C4897F13B" box="[549,652,1117,1143]" pageId="36" pageNumber="337" type="lectotype">lectotype</typeStatus>
|
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|
of
|
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|
<taxonomicName id="4C624D116E711D2CFD19C01F4924F13B" authorityName="Reed" authorityYear="1917" box="[692,831,1118,1143]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="unicornis">
|
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|
|
<emphasis id="B916EA806E711D2CFD19C01F4924F13B" box="[692,831,1118,1143]" italics="true" pageId="36" pageNumber="337">R. unicornis</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
shows a wider part of the glabella at the level of the basal lobes, which is actually more like that of the type species of
|
|||
|
|
<emphasis id="B916EA806E711D2CFCD5C0C04E0DF1D7" box="[888,1046,1153,1179]" italics="true" pageId="36" pageNumber="337">
|
|||
|
|
<taxonomicName id="4C624D116E711D2CFCD5C0C04E09F1D7" authorityName="Kobayashi" authorityYear="1951" box="[888,1042,1153,1179]" class="Trilobita" family="Bathycheilidae" genus="Calymenesun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">Calymenesun</taxonomicName>
|
|||
|
|
,
|
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|
|
</emphasis>
|
|||
|
|
<taxonomicName id="4C624D116E711D2CFBB0C0C34EB8F1D7" authority="Sun" authorityName="Sun" box="[1053,1187,1153,1180]" class="Trilobita" family="Bathycheilidae" genus="Calymenesun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="tingi">
|
|||
|
|
<emphasis id="B916EA806E711D2CFBB0C0C34E6BF1D7" box="[1053,1136,1154,1180]" italics="true" pageId="36" pageNumber="337">C. tingi</emphasis>
|
|||
|
|
Sun
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFF34B636E711D2CFB1FC0C04F01F1D0" author="Lu, Y. - H." box="[1202,1306,1153,1180]" pageId="36" pageNumber="337" pagination="1 - 453" refId="ref45640" refString="Lu, Y. - H. (1975) Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica, New Series B, 11, 1 - 453." type="journal article" year="1975">Lu, 1975</bibRefCitation>
|
|||
|
|
, pl. 46, figs 9,12). As discussed by
|
|||
|
|
<bibRefCitation id="EFF34B636E711D2CFE3BC0E74846F1F3" author="Peng, S. - C. & Lin, T. - R. & Li, Y." box="[406,605,1189,1216]" pageId="36" pageNumber="337" pagination="63 - 75" refId="ref46395" refString="Peng, S. - C., Lin, T. - R. & Li, Y. (2000) Notes on the genus Reedocalymene Kobayashi, 1951 (Trilobita, Ordovician). Acta Palaeontologica Sinica, 39, 63 - 75." type="journal article" year="2000">
|
|||
|
|
Peng
|
|||
|
|
<emphasis id="B916EA806E711D2CFE79C0E74817F1F3" box="[468,524,1189,1215]" italics="true" pageId="36" pageNumber="337">et al.</emphasis>
|
|||
|
|
(2000)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
the distinction between the two genera, both erected by
|
|||
|
|
<bibRefCitation id="EFF34B636E711D2CFB60C0E44F83F18C" author="Kobayashi, T." box="[1229,1432,1189,1216]" pageId="36" pageNumber="337" pagination="1 - 87" refId="ref45054" refString="Kobayashi, T. (1951) On the Ordovician trilobites in central China. Journal of the Faculty of Science Imperial University of Tokyo, Section II, Geology, 8, 1 - 87." type="journal article" year="1951">Kobayashi (1951)</bibRefCitation>
|
|||
|
|
, is not clearcut, and they may eventually be synonymized. We follow these authors in recognizing the two genera
|
|||
|
|
<emphasis id="B916EA806E711D2CFF3AC0AF4AF7F04B" box="[151,236,1262,1287]" italics="true" pageId="36" pageNumber="337">pro tem</emphasis>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|