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<document id="041EC229C3E26E8BAFDB189EDC2C0DFB" ID-CLB-Dataset="8352" ID-DOI="10.5281/zenodo.4526404" ID-GBIF-Dataset="3cea39ec-86b6-419b-8cdb-5baeedc4cc43" ID-ISSN="0253-116X" ID-Zenodo-Dep="4526404" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612224402507" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03C587F68414FFA4FF123ECDFC49FA1A" docLanguage="en" docName="LinzerbiolBeitr.49.2.1029-1039.pdf" docOrigin="Linzer biologische Beiträge 49 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) brevalatus Assing 2017, nov.sp." docType="treatment" docVersion="8" lastPageNumber="1035" masterDocId="FFFCFF8E8417FFA2FFB13A12FFEBFFAA" masterDocTitle="On the micropterous Quedius (Raphirus) species with a punctate scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="1039" masterPageNumber="1029" pageNumber="1032" updateTime="1698926418021" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5F99A6DB40BE11072805B8F54DDD3AD4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E123FAAB7AC33BB37ECCCD598375CCB">
<mods:title id="63325C29991E63C7290410411E77DB69">On the micropterous Quedius (Raphirus) species with a punctate scutellum of Turkey (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="0B647E99238E3FDE309E4EFB3853BEE7" type="personal">
<mods:role id="2E784BC5F85542EE5418775E6B432CE8">
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<mods:namePart id="93EFFF9025FD00A6BE4C0BC40826F245">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="E71F27A2E78E091F7DE4AE7FF400FF83">text</mods:typeOfResource>
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<mods:title id="3ABBD6B944854DE3260AA4454E45865D">Linzer biologische Beiträge</mods:title>
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<mods:part id="2C26F49922246334B252C2FC1A5D4B58">
<mods:date id="31CD20A06939A40857636E9ECB0D499B">2017</mods:date>
<mods:detail id="50D8FFE3F14872A203CC3EE1D5072B7F" type="pubDate">
<mods:number id="4C55E9F2D1914F551D2F238DB7867B84">2017-12-11</mods:number>
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<mods:detail id="4FB47F0E0955CD5FB18A121E264E625C" type="volume">
<mods:number id="F0E5FBDC4C63324D2946004256E2550A">49</mods:number>
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<mods:number id="3A61123E556C29849269A0F37479123F">2</mods:number>
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<mods:start id="0E10A8C12D3AF179CFAAD121DD921151">1029</mods:start>
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<mods:classification id="2CF37512016C7839AE53373F1CC2D57B">journal article</mods:classification>
<mods:identifier id="FD6591A90DDB458D48A3B196C86BE4DD" type="CLB-Dataset">8352</mods:identifier>
<mods:identifier id="753E1F3AE24213BEC73D7CC003D6F704" type="DOI">10.5281/zenodo.4526404</mods:identifier>
<mods:identifier id="0AFE9E2FB828C2EBDE7854EE3665C6C6" type="GBIF-Dataset">3cea39ec-86b6-419b-8cdb-5baeedc4cc43</mods:identifier>
<mods:identifier id="C917091F82895616389BD38AD15B493E" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="F7702AF468FD1E11B8CDF13806C53A39" type="Zenodo-Dep">4526404</mods:identifier>
</mods:mods>
<treatment id="03C587F68414FFA4FF123ECDFC49FA1A" ID-DOI="http://doi.org/10.5281/zenodo.4527168" ID-GBIF-Taxon="177313201" ID-Zenodo-Dep="4527168" LSID="urn:lsid:plazi:treatment:03C587F68414FFA4FF123ECDFC49FA1A" httpUri="http://treatment.plazi.org/id/03C587F68414FFA4FF123ECDFC49FA1A" lastPageId="6" lastPageNumber="1035" pageId="3" pageNumber="1032">
<subSubSection id="C376656B8414FFA1FF123ECDFCD2FB5C" box="[163,825,1247,1270]" pageId="3" pageNumber="1032" type="nomenclature">
<paragraph id="8BD336E08414FFA1FF123ECDFCD2FB5C" blockId="3.[163,825,1247,1270]" box="[163,825,1247,1270]" pageId="3" pageNumber="1032">
<heading id="D09B818C8414FFA1FF123ECDFCD2FB5C" box="[163,825,1247,1270]" fontSize="8" level="2" pageId="3" pageNumber="1032" reason="2">
<emphasis id="B918EAF28414FFA1FF123ECDFDD3FB5F" bold="true" box="[163,568,1247,1269]" pageId="3" pageNumber="1032">
<taxonomicName id="4C6C4D638414FFA1FF123ECDFE0EFB5F" ID-CoL="4R32R" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[163,485,1247,1269]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1032" phylum="Arthropoda" rank="species" species="brevalatus" status="sp. nov." subGenus="Raphirus">
<emphasis id="B918EAF28414FFA1FF123ECDFF11FB5F" bold="true" box="[163,250,1247,1269]" italics="true" pageId="3" pageNumber="1032">Quedius</emphasis>
(
<emphasis id="B918EAF28414FFA1FEB83ECDFE81FB5F" bold="true" box="[265,362,1247,1269]" italics="true" pageId="3" pageNumber="1032">Raphirus</emphasis>
)
<emphasis id="B918EAF28414FFA1FEC83ECDFE0EFB5F" bold="true" box="[377,485,1247,1269]" italics="true" pageId="3" pageNumber="1032">brevalatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57898414FFA1FE5D3EF2FDD3FB5F" box="[492,568,1248,1269]" pageId="3" pageNumber="1032" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13572A658414FFA1FDF43EF2FD77FB5C" box="[581,668,1248,1270]" captionStart="Figs 1-10" captionStartId="4.[163,203,1491,1513]" captionTargetBox="[163,1066,196,1462]" captionTargetId="figure-1@4.[163,1070,196,1481]" captionTargetPageId="4" captionText="Figs 1-10: Quedius brevalatus from Turkey (1-6) and Q. boopoides from Germany (7-10): (1, 7) forebody; (2-3) aedeagus in lateral and in ventral view (in transparent light); (4, 8) median lobe of aedeagus in lateral view (dry preparation); (5, 9) median lobe of aedeagus in ventral view; (6, 10) paramere. Scale bars: 1, 7: 1.0 mm; 2-6, 8-10: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526408" httpUri="https://zenodo.org/record/4526408/files/figure.png" pageId="3" pageNumber="1032">Figs 1-6</figureCitation>
,
<figureCitation id="13572A658414FFA1FD163EF2FD0FFB5F" box="[679,740,1248,1269]" captionStart="Figs 11-21" captionStartId="5.[163,203,1496,1518]" captionTargetBox="[167,1060,204,1475]" captionTargetId="figure-1@5.[163,1070,196,1481]" captionTargetPageId="5" captionText="Figs 11-21: Quedius brevalatus from Armenia (11-14) and Turkey (15-16), and Q. omissus from Turkey (17-21): (11-12, 19-20) median lobe of aedeagus in lateral and in ventral view; (13-15, 21) paramere; (16) female abdominal segments IX-X; (17-18) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="3" pageNumber="1032">11-13</figureCitation>
,
<figureCitation id="13572A658414FFA1FD5E3EF2FCC5FB5C" box="[751,814,1248,1270]" captionStart="Map 1" captionStartId="3.[163,208,1139,1160]" captionTargetBox="[163,1063,703,1130]" captionTargetId="figure-217@3.[163,1063,703,1130]" captionTargetPageId="3" captionText="Map 1: Distribution of micropterous Quedius (Raphirus) species with punctate scutellum in Turkey: Quedius reitteri (black circles), Q. omissus (white circle); Q. brevalatus (white triangle); Q. sp. (white star)." figureDoi="http://doi.org/10.5281/zenodo.4526406" httpUri="https://zenodo.org/record/4526406/files/figure.png" pageId="3" pageNumber="1032">Map 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376656B8414FFA1FF123F18FBF1FAD1" pageId="3" pageNumber="1032" type="materials_examined">
<paragraph id="8BD336E08414FFA1FF123F18FBF1FAD1" blockId="3.[163,1071,1289,1403]" pageId="3" pageNumber="1032">
<materialsCitation id="3B043CBD8414FFA1FF123F18FE56FAE7" ID-GBIF-Occurrence="3033390302" collectingDate="2017-07-12" collectionCode="T, V" country="Armenia" elevation="1670" latitude="39.52861" location="ARMENIA" longLatPrecision="19" longitude="46.309166" pageId="3" pageNumber="1032" specimenCount="2" typeStatus="holotype">
<collectionCode id="ED7DAE258414FFA1FF123F18FF5AFAB5" box="[163,177,1290,1311]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="1032">T</collectionCode>
y p e m a t e r i a l:
<typeStatus id="54D788428414FFA1FE293F18FE06FA8A" box="[408,493,1290,1312]" pageId="3" pageNumber="1032" type="holotype">Holotype</typeStatus>
3: &quot;
<collectingCountry id="F37B76708414FFA1FDAC3F18FD62FAB5" box="[541,649,1290,1311]" name="Armenia" pageId="3" pageNumber="1032">ARMENIA</collectingCountry>
[48] – NW Goris, W Verishen,
<geoCoordinate id="EE5850278414FFA1FC0E3F18FBC0FAB5" box="[959,1067,1290,1311]" degrees="39" direction="north" minutes="31" orientation="latitude" pageId="3" pageNumber="1032" precision="15" seconds="43" value="39.52861">39°31'43''N</geoCoordinate>
,
<geoCoordinate id="EE5850278414FFA1FF123F33FEE1FA9D" box="[163,266,1313,1335]" degrees="46" direction="east" minutes="18" orientation="longitude" pageId="3" pageNumber="1032" precision="15" seconds="33" value="46.309166">46°18'33''E</geoCoordinate>
<quantity id="4C949B058414FFA1FEA33F33FEB1FA9C" box="[274,346,1313,1335]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="1032" unit="m" value="1670.0">
<elevation id="0041D1D38414FFA1FEA33F33FEB1FA9C" box="[274,346,1313,1335]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="1032" unit="m" value="1670.0">1670 m</elevation>
</quantity>
, forest margin,
<date id="FFD210208414FFA1FE5C3F33FDB5FA9C" box="[493,606,1313,1334]" pageId="3" pageNumber="1032" value="2017-07-12">
<collectingDate id="EF96E9C88414FFA1FE5C3F33FDB5FA9C" box="[493,606,1313,1334]" pageId="3" pageNumber="1032" value="2017-07-12">12.VII.2017</collectingDate>
</date>
,
<collectionCode id="ED7DAE258414FFA1FDD83F33FD97FA9C" box="[617,636,1313,1334]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="1032" type="Museum">V</collectionCode>
. Assing /
<typeStatus id="54D788428414FFA1FD6B3F33FCD3FA9D" box="[730,824,1313,1335]" pageId="3" pageNumber="1032" type="holotype">Holotypus</typeStatus>
3
<taxonomicName id="4C6C4D638414FFA1FCEB3F33FBE6FA9C" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[858,1037,1313,1334]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1032" phylum="Arthropoda" rank="species" species="brevalatus" status="sp. nov.">
<emphasis id="B918EAF28414FFA1FCEB3F33FBE6FA9C" box="[858,1037,1313,1334]" italics="true" pageId="3" pageNumber="1032">Quedius brevalatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57898414FFA1FBA53F33FF5EFAE7" pageId="3" pageNumber="1032" rank="species">sp. n.</taxonomicNameLabel>
det.
<collectionCode id="ED7DAE258414FFA1FF503F2AFF1FFAE7" box="[225,244,1336,1357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="1032" type="Museum">V</collectionCode>
.
<bibRefCitation id="EFFD4B118414FFA1FF4C3F25FE9AFAE6" author="ASSING V" box="[253,369,1335,1357]" pageId="3" pageNumber="1032" pagination="207 - 228" refId="ref5116" refString="ASSING V. (2017): On Quedius coloratus FAUVEL, 1875 and allied species, with an appendix on Quedius species collected in Greece with subterranean pitfall traps, and a new synonymy (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 49 (1): 207 - 228." type="journal article" year="2017">Assing 2017</bibRefCitation>
&quot; (cAss)
</materialsCitation>
.
<materialsCitation id="3B043CBD8414FFA1FE773F2AFC5AFAE7" ID-GBIF-Occurrence="3033390304" box="[454,945,1335,1362]" collectingDate="2017-07-12" collectionCode="T, V" country="Armenia" elevation="1670" latitude="39.52861" location="ARMENIA" longLatPrecision="19" longitude="46.309166" pageId="3" pageNumber="1032" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<typeStatus id="54D788428414FFA1FE773F2AFDC9FAE7" box="[454,546,1336,1357]" pageId="3" pageNumber="1032" type="paratype">Paratypes</typeStatus>
: 533,
<specimenCount id="9D6AFD698414FFA1FDEE3F25FD61FAF8" box="[607,650,1335,1362]" pageId="3" pageNumber="1032" type="female">6 ♀♀</specimenCount>
: same data as holotype (cAss)
</materialsCitation>
;
<materialsCitation id="3B043CBD8414FFA1FC0B3F25FDB3FACE" ID-GBIF-Occurrence="3033390311" collectingDate="2017-07-12" collectionCode="MNB" collectorName="Schulke" country="Armenia" elevation="1670" latitude="39.52861" location="ARMENIA" longLatPrecision="19" longitude="46.309166" pageId="3" pageNumber="1032" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
633,
<specimenCount id="9D6AFD698414FFA1FC5E3F25FBF1FAF8" box="[1007,1050,1335,1362]" pageId="3" pageNumber="1032" type="female">3 ♀♀</specimenCount>
: same data as holotype, but leg.
<collectorName id="269953368414FFA1FE723F5CFDE7FACE" box="[451,524,1358,1380]" pageId="3" pageNumber="1032">Schülke</collectorName>
(
<collectionCode id="ED7DAE258414FFA1FDA93F5CFDBBFACE" box="[536,592,1358,1380]" pageId="3" pageNumber="1032">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B043CBD8414FFA1FDD03F5CFBFDFAD1" ID-GBIF-Occurrence="3033390301" collectingDate="2006-07-25" collectionCode="V" country="Armenia" elevation="2510" latitude="40.625275" location="Rize" longLatPrecision="19" longitude="40.7575" municipality="Ovitdagi Gec." pageId="3" pageNumber="1032" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
233 [1 teneral],
<specimenCount id="9D6AFD698414FFA1FCB23F5CFCCDFAC2" box="[771,806,1358,1384]" pageId="3" pageNumber="1032" type="female">4♀♀</specimenCount>
: &quot;TR [5a]-
<location id="8EB3603B8414FFA1FC2E3F5CFC20FACE" LSID="urn:lsid:plazi:treatment:03C587F68414FFA4FF123ECDFC49FA1A:8EB3603B8414FFA1FC2E3F5CFC20FACE" box="[927,971,1358,1380]" country="Armenia" latitude="40.625275" longLatPrecision="19" longitude="40.7575" municipality="Ovitdagi Gec." name="Rize" pageId="3" pageNumber="1032">Rize</location>
, ca.
<locationDeviation id="9A612FC28414FFA1FC443F5CFF17FAD1" location="Rize" pageId="3" pageNumber="1032">
<quantity id="4C949B058414FFA1FC443F5CFBC5FACE" box="[1013,1070,1358,1380]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="1032" unit="km" value="50.0">50 km</quantity>
SSE Rize
</locationDeviation>
,
<collectingMunicipality id="6BB7AC9A8414FFA1FEB53F77FE6CFAD0" box="[260,391,1381,1403]" pageId="3" pageNumber="1032">Ovitdağı Geç.</collectingMunicipality>
,
<quantity id="4C949B058414FFA1FE3E3F76FE3DFAD0" box="[399,470,1380,1402]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="3" pageNumber="1032" unit="m" value="2510.0">
<elevation id="0041D1D38414FFA1FE3E3F76FE3DFAD0" box="[399,470,1380,1402]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="3" pageNumber="1032" unit="m" value="2510.0">2510 m</elevation>
</quantity>
, sifted,
<geoCoordinate id="EE5850278414FFA1FDAC3F77FD6BFAD0" box="[541,640,1381,1402]" degrees="40" direction="north" minutes="37" orientation="latitude" pageId="3" pageNumber="1032" precision="15" seconds="31" value="40.625275">40°37'31N</geoCoordinate>
,
<geoCoordinate id="EE5850278414FFA1FD3B3F77FD00FAD0" box="[650,747,1380,1402]" degrees="40" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="1032" precision="15" seconds="27" value="40.7575">40°45'27E</geoCoordinate>
,
<date id="FFD210208414FFA1FD453F76FC8EFAD0" box="[756,869,1380,1402]" pageId="3" pageNumber="1032" value="2006-07-25">
<collectingDate id="EF96E9C88414FFA1FD453F76FC8EFAD0" box="[756,869,1380,1402]" pageId="3" pageNumber="1032" value="2006-07-25">25.VII.2006</collectingDate>
</date>
,
<collectionCode id="ED7DAE258414FFA1FCDF3F77FC6AFAD0" box="[878,897,1381,1402]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="1032" type="Museum">V</collectionCode>
. Assing&quot; (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C376656B8414FFA4FF123F9BFC7EFDAC" lastPageId="6" lastPageNumber="1035" pageId="3" pageNumber="1032" type="description">
<paragraph id="8BD336E08414FFA1FF123F9BFF0CFA73" blockId="3.[163,1070,1417,1497]" pageId="3" pageNumber="1032">E t y m o l o g y: The specific epithet is an adjective composed of the Latin adjectives brevis (short) and alatus (winged). It alludes to the reduced length of the elytra and hind wings.</paragraph>
<caption id="DF1366688413FFA6FF123FC1FD7DF987" ID-DOI="http://doi.org/10.5281/zenodo.4526408" ID-Zenodo-Dep="4526408" httpUri="https://zenodo.org/record/4526408/files/figure.png" pageId="4" pageNumber="1033" startId="4.[163,203,1491,1513]" targetBox="[163,1066,196,1462]" targetPageId="4">
<paragraph id="8BD336E08413FFA6FF123FC1FD7DF987" blockId="4.[163,1071,1491,1581]" pageId="4" pageNumber="1033">
<emphasis id="B918EAF28413FFA6FF123FC1FF15FA42" bold="true" box="[163,254,1491,1513]" pageId="4" pageNumber="1033">Figs 1-10</emphasis>
:
<taxonomicName id="4C6C4D638413FFA6FEBA3FC1FE55FA42" authorityName="Assing" authorityYear="2017" box="[267,446,1491,1512]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1033" phylum="Arthropoda" rank="species" species="brevalatus">
<emphasis id="B918EAF28413FFA6FEBA3FC1FE55FA42" box="[267,446,1491,1512]" italics="true" pageId="4" pageNumber="1033">Quedius brevalatus</emphasis>
</taxonomicName>
from Turkey (
<emphasis id="B918EAF28413FFA6FDFA3FC1FD82FA42" bold="true" box="[587,617,1491,1512]" pageId="4" pageNumber="1033">1-6</emphasis>
) and
<emphasis id="B918EAF28413FFA6FD103FC6FCF7FA42" box="[673,796,1491,1513]" italics="true" pageId="4" pageNumber="1033">Q. boopoides</emphasis>
from Germany (
<emphasis id="B918EAF28413FFA6FC0A3FC1FC0FFA42" bold="true" box="[955,996,1491,1512]" pageId="4" pageNumber="1033">7-10</emphasis>
): (
<emphasis id="B918EAF28413FFA6FBB33FC1FBCDFA42" bold="true" box="[1026,1062,1491,1512]" pageId="4" pageNumber="1033">1, 7</emphasis>
) forebody; (
<emphasis id="B918EAF28413FFA6FEB83FF8FEC3FA55" bold="true" box="[265,296,1514,1535]" pageId="4" pageNumber="1033">2-3</emphasis>
) aedeagus in lateral and in ventral view (in transparent light); (
<emphasis id="B918EAF28413FFA6FCC33FF8FC7EFA55" bold="true" box="[882,917,1514,1535]" pageId="4" pageNumber="1033">4, 8</emphasis>
) median lobe of aedeagus in lateral view (dry preparation); (
<emphasis id="B918EAF28413FFA6FD883C13FDB6F9BF" bold="true" box="[569,605,1536,1558]" pageId="4" pageNumber="1033">5, 9</emphasis>
) median lobe of aedeagus in ventral view; (
<emphasis id="B918EAF28413FFA6FC463C12FBCCF9BF" bold="true" box="[1015,1063,1536,1557]" pageId="4" pageNumber="1033">6, 10</emphasis>
) paramere. Scale bars: 1, 7: 1.0 mm; 2-6, 8-10: 0.2 mm.
</paragraph>
</caption>
<caption id="DF1366688412FFA7FF123FCAFEB3F998" ID-DOI="http://doi.org/10.5281/zenodo.4526410" ID-Zenodo-Dep="4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="5" pageNumber="1034" startId="5.[163,203,1496,1518]" targetBox="[167,1060,204,1475]" targetPageId="5">
<paragraph id="8BD336E08412FFA7FF123FCAFEB3F998" blockId="5.[163,1071,1496,1587]" pageId="5" pageNumber="1034">
<emphasis id="B918EAF28412FFA7FF123FCAFEE2FA44" bold="true" box="[163,265,1496,1518]" pageId="5" pageNumber="1034">Figs 11-21</emphasis>
:
<taxonomicName id="4C6C4D638412FFA7FEA63FCBFE21FA44" authorityName="Assing" authorityYear="2017" box="[279,458,1497,1518]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1034" phylum="Arthropoda" rank="species" species="brevalatus">
<emphasis id="B918EAF28412FFA7FEA63FCBFE21FA44" box="[279,458,1497,1518]" italics="true" pageId="5" pageNumber="1034">Quedius brevalatus</emphasis>
</taxonomicName>
from Armenia (
<emphasis id="B918EAF28412FFA7FDD53FCBFD71FA44" bold="true" box="[612,666,1497,1518]" pageId="5" pageNumber="1034">11-14</emphasis>
) and Turkey (
<emphasis id="B918EAF28412FFA7FC933FCBFCBCFA44" bold="true" box="[802,855,1497,1518]" pageId="5" pageNumber="1034">15-16</emphasis>
), and
<taxonomicName id="4C6C4D638412FFA7FC243FCBFC10FA44" authorityName="COIFFAIT" authorityYear="1977" box="[917,1019,1497,1518]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1034" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="B918EAF28412FFA7FC243FCBFC10FA44" box="[917,1019,1497,1518]" italics="true" pageId="5" pageNumber="1034">Q. omissus</emphasis>
</taxonomicName>
from Turkey (
<emphasis id="B918EAF28412FFA7FF423FFDFEC3F9AE" bold="true" box="[243,296,1519,1540]" pageId="5" pageNumber="1034">17-21</emphasis>
): (
<emphasis id="B918EAF28412FFA7FEF23FFDFE51F9AE" bold="true" box="[323,442,1519,1540]" pageId="5" pageNumber="1034">11-12, 19-20</emphasis>
) median lobe of aedeagus in lateral and in ventral view; (
<emphasis id="B918EAF28412FFA7FC7F3FFDFBCCF9AE" bold="true" box="[974,1063,1519,1540]" pageId="5" pageNumber="1034">13-15, 21</emphasis>
) paramere; (
<emphasis id="B918EAF28412FFA7FEBD3C14FEC8F9B1" bold="true" box="[268,291,1542,1563]" pageId="5" pageNumber="1034">16</emphasis>
) female abdominal segments IX-X; (
<emphasis id="B918EAF28412FFA7FDCF3C14FD58F9B1" bold="true" box="[638,691,1542,1563]" pageId="5" pageNumber="1034">17-18</emphasis>
) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BD336E08411FFA4FF123ADFFCD3FEF6" blockId="6.[163,1070,205,349]" pageId="6" pageNumber="1035">D e s c r i p t i o n: Measurements (in mm) and ratios (range, arithmetic mean) (n = 18): TL: 5.5-7.3, 6.7; HL: 0.70-0.77, 0.74; HW: 0.84-0.90, 0.86; PL: 0.97-1.09, 1.03; PW: 1.00-1.14, 1.06; EL: 0.49-0.56, 0.52; HTiL: 0.70-0.84, 0.77; HTaL: 0.60-0.70, 0.64; ML: 0.90-1.02, 0.95; HW/HL: 1.13-1.23, 1.17; PW/HW: 1.19-1.26, 1.23; PL/PW: 0.94- 0.99, 0.97; EL/PL: 0.48-0.53, 0.51; HTiL/HTaL: 1.14-1.26, 1.20.</paragraph>
<paragraph id="8BD336E08411FFA4FF123B7CFC17FE6B" blockId="6.[163,1070,366,449]" pageId="6" pageNumber="1035">Coloration: body black; abdominal segments IX and X anteriorly somewhat paler; legs yellowish-brown, with the metatibiae at least partly infuscate; antennae pale-brown to blackish-brown with the basal antennomeres more or less extensively yellowish-red.</paragraph>
<paragraph id="8BD336E08411FFA4FF123BC0FC7EFDAC" blockId="6.[163,1071,466,518]" pageId="6" pageNumber="1035">
Elytra (
<figureCitation id="13572A658411FFA4FF433BC0FEDBFE42" box="[242,304,466,488]" captionStart="Figs 1-10" captionStartId="4.[163,203,1491,1513]" captionTargetBox="[163,1066,196,1462]" captionTargetId="figure-1@4.[163,1070,196,1481]" captionTargetPageId="4" captionText="Figs 1-10: Quedius brevalatus from Turkey (1-6) and Q. boopoides from Germany (7-10): (1, 7) forebody; (2-3) aedeagus in lateral and in ventral view (in transparent light); (4, 8) median lobe of aedeagus in lateral view (dry preparation); (5, 9) median lobe of aedeagus in ventral view; (6, 10) paramere. Scale bars: 1, 7: 1.0 mm; 2-6, 8-10: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526408" httpUri="https://zenodo.org/record/4526408/files/figure.png" pageId="6" pageNumber="1035">Fig. 1</figureCitation>
) only half as long as pronotum (see ratio EL/PL). Hind wings reduced. Posterior margin of tergite VIII with narrow rudiment of a palisade fringe.
</paragraph>
</subSubSection>
<subSubSection id="C376656B8411FFA4FF123805FCBAFD86" box="[163,849,535,557]" pageId="6" pageNumber="1035" type="synonymic_list">
<paragraph id="8BD336E08411FFA4FF123805FCBAFD86" blockId="6.[163,849,535,557]" box="[163,849,535,557]" pageId="6" pageNumber="1035">
Other external characters as in
<emphasis id="B918EAF28411FFA4FE6C3805FD4BFD86" box="[477,672,535,556]" italics="true" pageId="6" pageNumber="1035">
<taxonomicName id="4C6C4D638411FFA4FE6C3805FDDAFD86" authorityName="STEPHENS" authorityYear="1829" box="[477,561,535,556]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1036" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
boopoides
</emphasis>
MUNSTER, 1923.
</paragraph>
</subSubSection>
<subSubSection id="C376656B8411FFA4FF12382CFE18FC91" pageId="6" pageNumber="1035" type="description">
<paragraph id="8BD336E08411FFA4FF12382CFD9AFDD8" blockId="6.[163,1070,574,627]" pageId="6" pageNumber="1035">
3: protarsomeres I-IV strongly dilated; sternite VIII broadly triangularly excised posteriorly; aedeagus shaped as in
<figureCitation id="13572A658411FFA4FE7F384FFDCFFDD8" box="[462,548,604,627]" captionStart="Figs 1-10" captionStartId="4.[163,203,1491,1513]" captionTargetBox="[163,1066,196,1462]" captionTargetId="figure-1@4.[163,1070,196,1481]" captionTargetPageId="4" captionText="Figs 1-10: Quedius brevalatus from Turkey (1-6) and Q. boopoides from Germany (7-10): (1, 7) forebody; (2-3) aedeagus in lateral and in ventral view (in transparent light); (4, 8) median lobe of aedeagus in lateral view (dry preparation); (5, 9) median lobe of aedeagus in ventral view; (6, 10) paramere. Scale bars: 1, 7: 1.0 mm; 2-6, 8-10: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526408" httpUri="https://zenodo.org/record/4526408/files/figure.png" pageId="6" pageNumber="1035">Figs 2-6</figureCitation>
,
<figureCitation id="13572A658411FFA4FD9E384EFD87FDD8" box="[559,620,604,626]" captionStart="Figs 11-21" captionStartId="5.[163,203,1496,1518]" captionTargetBox="[167,1060,204,1475]" captionTargetId="figure-1@5.[163,1070,196,1481]" captionTargetPageId="5" captionText="Figs 11-21: Quedius brevalatus from Armenia (11-14) and Turkey (15-16), and Q. omissus from Turkey (17-21): (11-12, 19-20) median lobe of aedeagus in lateral and in ventral view; (13-15, 21) paramere; (16) female abdominal segments IX-X; (17-18) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="6" pageNumber="1035">11-15</figureCitation>
.
</paragraph>
<paragraph id="8BD336E08411FFA4FF12389BFF28FD12" blockId="6.[163,1071,643,696]" pageId="6" pageNumber="1035">
<specimenCount id="9D6AFD698411FFA4FF12389BFF5AFD34" box="[163,177,649,670]" pageId="6" pageNumber="1035" type="female">♀</specimenCount>
: protarsomeres I-IV weakly dilated; posterior abdominal segments shaped as in
<figureCitation id="13572A658411FFA4FBB73891FF28FD12" captionStart="Figs 11-21" captionStartId="5.[163,203,1496,1518]" captionTargetBox="[167,1060,204,1475]" captionTargetId="figure-1@5.[163,1070,196,1481]" captionTargetPageId="5" captionText="Figs 11-21: Quedius brevalatus from Armenia (11-14) and Turkey (15-16), and Q. omissus from Turkey (17-21): (11-12, 19-20) median lobe of aedeagus in lateral and in ventral view; (13-15, 21) paramere; (16) female abdominal segments IX-X; (17-18) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="6" pageNumber="1035">Fig. 16.</figureCitation>
</paragraph>
<paragraph id="8BD336E08411FFA4FF1238DBFE18FC91" blockId="6.[163,1070,713,827]" pageId="6" pageNumber="1035">
I n t r a s p e c i f i c v a r i a t i o n: The material from
<collectingCountry id="F37B76708411FFA4FC9738DBFC94FD75" box="[806,895,713,735]" name="Armenia" pageId="6" pageNumber="1035">Armenia</collectingCountry>
is distinguished from the Turkish specimens by a slightly smaller and more slender paramere (
<figureCitation id="13572A658411FFA4FC6638F5FF54FCB6" captionStart="Figs 11-21" captionStartId="5.[163,203,1496,1518]" captionTargetBox="[167,1060,204,1475]" captionTargetId="figure-1@5.[163,1070,196,1481]" captionTargetPageId="5" captionText="Figs 11-21: Quedius brevalatus from Armenia (11-14) and Turkey (15-16), and Q. omissus from Turkey (17-21): (11-12, 19-20) median lobe of aedeagus in lateral and in ventral view; (13-15, 21) paramere; (16) female abdominal segments IX-X; (17-18) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="6" pageNumber="1035">Figs 13- 15</figureCitation>
). The otherwise similar external and male sexual characters, however, suggest that both populations are conspecific.
</paragraph>
</subSubSection>
<subSubSection id="C376656B8411FFA4FF123959FE58FB45" pageId="6" pageNumber="1035" type="discussion">
<paragraph id="8BD336E08411FFA4FF123959FE58FB45" blockId="6.[163,1070,843,1264]" pageId="6" pageNumber="1035">
C o m p a r a t i v e n o t e s: Among the West Palaearctic
<taxonomicName id="4C6C4D638411FFA4FCE53959FC43FCCA" authorityName="STEPHENS" authorityYear="1829" box="[852,936,843,864]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1036" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAF28411FFA4FCE53959FC43FCCA" box="[852,936,843,864]" italics="true" pageId="6" pageNumber="1035">Quedius</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C6C4D638411FFA4FBA4395EFEE5FCD5" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1035" phylum="Arthropoda" rank="species" species="brevalatus">
<emphasis id="B918EAF28411FFA4FBA4395EFEE5FCD5" italics="true" pageId="6" pageNumber="1035">Q. brevalatus</emphasis>
</taxonomicName>
is most similar to
<emphasis id="B918EAF28411FFA4FE663978FD8AFCD5" box="[471,609,874,895]" italics="true" pageId="6" pageNumber="1035">Q. boopoides</emphasis>
both in external and sexual characters. It is distinguished from that species particularly by distinctly shorter elytra (
<emphasis id="B918EAF28411FFA4FC2D399BFBCCFC34" box="[924,1063,905,926]" italics="true" pageId="6" pageNumber="1035">Q. boopoides</emphasis>
: elytra 0.65-0.70 times as long as pronotum), reduced hind wings (shorter than elytra), darker abdominal segments IX-X, a less pronounced palisade fringe at the posterior margin of tergite VII, and by the shape of the aedeagus. In
<emphasis id="B918EAF28411FFA4FC9039F7FC40FC50" box="[801,939,997,1018]" italics="true" pageId="6" pageNumber="1035">Q. boopoides</emphasis>
, the median lobe is slightly larger, the apex of the median lobe is larger with the subapical tooth more distant from the apex and apically extended into a distinct carina, and the paramere is apically less acute. For comparison, illustrations of the forebody and of the aedeagus of
<emphasis id="B918EAF28411FFA4FF123E72FEC0FBDE" box="[163,299,1119,1141]" italics="true" pageId="6" pageNumber="1035">Q. boopoides</emphasis>
are provided in
<figureCitation id="13572A658411FFA4FE623E4DFDDCFBDE" box="[467,567,1119,1141]" captionStart="Figs 1-10" captionStartId="4.[163,203,1491,1513]" captionTargetBox="[163,1066,196,1462]" captionTargetId="figure-1@4.[163,1070,196,1481]" captionTargetPageId="4" captionText="Figs 1-10: Quedius brevalatus from Turkey (1-6) and Q. boopoides from Germany (7-10): (1, 7) forebody; (2-3) aedeagus in lateral and in ventral view (in transparent light); (4, 8) median lobe of aedeagus in lateral view (dry preparation); (5, 9) median lobe of aedeagus in ventral view; (6, 10) paramere. Scale bars: 1, 7: 1.0 mm; 2-6, 8-10: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526408" httpUri="https://zenodo.org/record/4526408/files/figure.png" pageId="6" pageNumber="1035">Figs 7-10</figureCitation>
. From the similar micropterous and sympatric
<taxonomicName id="4C6C4D638411FFA4FBA43E72FF18FB39" authorityName="COIFFAIT" authorityYear="1977" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1035" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="B918EAF28411FFA4FBA43E72FF18FB39" italics="true" pageId="6" pageNumber="1035">Q. omissus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D638411FFA4FEB33E6CFE65FB39" authorityName="Assing" authorityYear="2017" box="[258,398,1150,1171]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1035" phylum="Arthropoda" rank="species" species="brevalatus">
<emphasis id="B918EAF28411FFA4FEB33E6CFE65FB39" box="[258,398,1150,1171]" italics="true" pageId="6" pageNumber="1035">Q. brevalatus</emphasis>
</taxonomicName>
differs by shorter hind wings (
<taxonomicName id="4C6C4D638411FFA4FD673E6CFCA3FB39" authorityName="COIFFAIT" authorityYear="1977" box="[726,840,1150,1171]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1035" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="B918EAF28411FFA4FD673E6CFCA3FB39" box="[726,840,1150,1171]" italics="true" pageId="6" pageNumber="1035">Q. omissus</emphasis>
</taxonomicName>
: hind wing rudiments distinctly projecting beyond posterior margin of the elytra when unfolded) and above all by the shape of the aedeagus. For illustrations of the male primary sexual characters of
<taxonomicName id="4C6C4D638411FFA4FF123EC8FEF9FB45" authorityName="COIFFAIT" authorityYear="1977" box="[163,274,1242,1263]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1035" phylum="Arthropoda" rank="species" species="omissus">
<emphasis id="B918EAF28411FFA4FF123EC8FEF9FB45" box="[163,274,1242,1263]" italics="true" pageId="6" pageNumber="1035">Q. omissus</emphasis>
</taxonomicName>
see
<figureCitation id="13572A658411FFA4FE8E3EC8FE58FB45" box="[319,435,1242,1264]" captionStart="Figs 11-21" captionStartId="5.[163,203,1496,1518]" captionTargetBox="[167,1060,204,1475]" captionTargetId="figure-1@5.[163,1070,196,1481]" captionTargetPageId="5" captionText="Figs 11-21: Quedius brevalatus from Armenia (11-14) and Turkey (15-16), and Q. omissus from Turkey (17-21): (11-12, 19-20) median lobe of aedeagus in lateral and in ventral view; (13-15, 21) paramere; (16) female abdominal segments IX-X; (17-18) aedeagus in lateral and in ventral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4526410" httpUri="https://zenodo.org/record/4526410/files/figure.png" pageId="6" pageNumber="1035">Figs 17-21.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376656B8411FFA4FF123F13FC49FA1A" pageId="6" pageNumber="1035" type="distribution">
<paragraph id="8BD336E08411FFA4FF123F13FC49FA1A" blockId="6.[163,1070,1281,1456]" pageId="6" pageNumber="1035">
D i s t r i b u t i o n a n d n a t u r a l h i s t o r y: The species is currently known from the type locality near Goris, South
<collectingCountry id="F37B76708411FFA4FD273F0DFD1AFA9F" box="[662,753,1311,1333]" name="Armenia" pageId="6" pageNumber="1035">Armenia</collectingCountry>
, and from one locality in
<collectingRegion id="49A8F8028411FFA4FBB13F0DFF15FAFE" country="Turkey" name="Rize" pageId="6" pageNumber="1035">Rize province</collectingRegion>
, northeastern
<collectingCountry id="F37B76708411FFA4FE273F2CFE1AFAFE" box="[406,497,1342,1364]" name="Turkey" pageId="6" pageNumber="1035">Anatolia</collectingCountry>
. The material from the type locality was sifted from roots and litter beneath
<emphasis id="B918EAF28411FFA4FE243F4FFE00FAD8" box="[405,491,1373,1394]" italics="true" pageId="6" pageNumber="1035">Quercus</emphasis>
and
<emphasis id="B918EAF28411FFA4FDAF3F4FFDBAFAD8" box="[542,593,1373,1394]" italics="true" pageId="6" pageNumber="1035">Rosa</emphasis>
at the margin of an oak forest at an altitude of
<quantity id="4C949B058411FFA4FF123F69FF18FA3B" box="[163,243,1403,1425]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="6" pageNumber="1035" unit="m" value="1670.0">1670 m</quantity>
, the specimens from
<collectingCountry id="F37B76708411FFA4FE7F3F69FDFCFA3B" box="[462,535,1403,1425]" name="Turkey" pageId="6" pageNumber="1035">Turkey</collectingCountry>
were sifted from grass roots and moss around large rocks at an altitude of
<quantity id="4C949B058411FFA4FE343F88FE3FFA05" box="[389,468,1434,1456]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="6" pageNumber="1035" unit="m" value="2510.0">2510 m</quantity>
. One of the
<typeStatus id="54D788428411FFA4FDE13F88FD5AFA05" box="[592,689,1434,1455]" pageId="6" pageNumber="1035" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="F37B76708411FFA4FD5E3F88FCD3FA1A" box="[751,824,1434,1456]" name="Turkey" pageId="6" pageNumber="1035">Turkey</collectingCountry>
is teneral.
</paragraph>
</subSubSection>
</treatment>
</document>
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