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<document id="6FCCF6EBAE0465C51E3E03FDDEE0E986" ID-CLB-Dataset="29039" ID-DOI="10.11646/zootaxa.4459.1.1" ID-GBIF-Dataset="9acf71bc-6414-4878-8af0-a64d3ec1fea6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458302" ID-ZooBank="633A9498-FCC6-4439-BE33-A45B9E51B550" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539287574730" checkinUser="plazi" docAuthor="Grandinete, Yuri Campanholo, Noll, Fernando Barbosa &amp; Carpenter, James" docDate="2018" docId="03C5878FFFEAF31BFF309602D239FE2A" docLanguage="en" docName="zootaxa.4459.1.1.pdf" docOrigin="Zootaxa 4459 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes sculleni Bohart 1950" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFFCFFF7FFF3F300FFA79155D339FFDF" masterDocTitle="Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World" masterLastPageNumber="52" masterPageNumber="1" pageNumber="26" updateTime="1698577660185" updateUser="ExternalLinkService">
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<mods:title id="40BD7E39798E2C47FD66458FB0C3C7D2">Taxonomic Review of Eumenes Latreille, 1802 (Hymenoptera, Vespidae, Eumeninae) from the New World</mods:title>
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<mods:name id="FB2F14BFF7BCEDC8E895AFA24B6C2FA9" type="personal">
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<mods:namePart id="B8DC81AB20662E1F2D026ABEDA7F2E42">Grandinete, Yuri Campanholo</mods:namePart>
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<mods:name id="9DA4E7A3D83BF6659DECC0A2DDB38B8D" type="personal">
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<mods:namePart id="3A58FD0215F7710100E44E5F7E01B339">Noll, Fernando Barbosa</mods:namePart>
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<mods:namePart id="E0723AFC1679D2D1E31DC5F4A5C6E905">Carpenter, James</mods:namePart>
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<mods:typeOfResource id="E26213A2008B67A283843F8D400C280D">text</mods:typeOfResource>
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<mods:title id="046B7172A49D2490E2CA271500855FE0">Zootaxa</mods:title>
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<mods:part id="F1E3A96558A8F956B9F89122A071A193">
<mods:date id="D98E71B2C34A12E9728DC1323330C6CF">2018</mods:date>
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<mods:number id="13A8AD3A84C9BE1582DCCF77960B82BA">2018-08-14</mods:number>
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<mods:detail id="D8B7C0C8F41D2CB3D6E8D2ED1E6BFBD6" type="volume">
<mods:number id="9212FAA8395318A54D9E3388F3FA2C4F">4459</mods:number>
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<mods:number id="C2DF57065D4DD2BBB5E6D5EBF1114514">1</mods:number>
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<mods:classification id="1E3FD149FBA3071486E1645F3E1B21CC">journal article</mods:classification>
<mods:identifier id="76DE4ABE7CBA8F4560FC4D22B281A536" type="CLB-Dataset">29039</mods:identifier>
<mods:identifier id="691E32F5CCBA5856A5151C041BB4C801" type="DOI">10.11646/zootaxa.4459.1.1</mods:identifier>
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<mods:identifier id="F69DAF1ABEA6C799EEFD3AE3DF4EEAF3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C5878FFFEAF31BFF309602D239FE2A" ID-DOI="http://doi.org/10.5281/zenodo.5986384" ID-GBIF-Taxon="148692486" ID-Zenodo-Dep="5986384" LSID="urn:lsid:plazi:treatment:03C5878FFFEAF31BFF309602D239FE2A" httpUri="http://treatment.plazi.org/id/03C5878FFFEAF31BFF309602D239FE2A" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26">
<subSubSection id="C3766512FFEAF319FF309602D10CF86B" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BD33699FFEAF319FF309602D12FF8AD" blockId="25.[151,534,1879,1906]" box="[151,534,1879,1906]" pageId="25" pageNumber="26">
<heading id="D09B81F5FFEAF319FF309602D12FF8AD" bold="true" box="[151,534,1879,1906]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="4C6C4D1AFFEAF319FF309602D12FF8AD" ID-CoL="3CDNX" authority="Bohart, 1950" authorityName="Bohart" authorityYear="1950" box="[151,534,1879,1906]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFEAF319FF309602D12FF8AD" bold="true" box="[151,534,1879,1906]" pageId="25" pageNumber="26">
<emphasis id="B918EA8BFFEAF319FF309602D255F8AE" bold="true" box="[151,364,1879,1905]" italics="true" pageId="25" pageNumber="26">Eumenes sculleni</emphasis>
Bohart, 1950
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD33699FFEAF319FF3096CAD10CF86B" blockId="25.[151,857,1950,2034]" box="[151,565,1950,1972]" pageId="25" pageNumber="26">
<taxonomicName id="4C6C4D1AFFEAF319FF3096CAD2DBF86B" authority="Bohart, 1950" authorityName="Bohart" authorityYear="1950" box="[151,482,1950,1972]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFEAF319FF3096CAD2DBF86B" bold="true" box="[151,482,1950,1972]" pageId="25" pageNumber="26">
<emphasis id="B918EA8BFFEAF319FF3096CAD269F86B" bold="true" box="[151,336,1950,1972]" italics="true" pageId="25" pageNumber="26">Eumenes sculleni</emphasis>
Bohart, 1950
</emphasis>
</taxonomicName>
, 63: 78.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFEAF31AFF6096E8D16AFF34" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8BD33699FFEAF319FF6096E8D1ABF80C" blockId="25.[151,857,1950,2034]" box="[199,658,1981,2003]" pageId="25" pageNumber="26">
<emphasis id="B918EA8BFFEAF319FF6096E8D203F80C" bold="true" box="[199,314,1981,2003]" pageId="25" pageNumber="26">Type data:</emphasis>
Holotype male at CAS (n° 6143).
</paragraph>
<paragraph id="8BD33699FFEAF319FF609689D060F82D" blockId="25.[151,857,1950,2034]" box="[199,857,2012,2034]" pageId="25" pageNumber="26">
<materialsCitation id="3B043CC4FFEAF319FF609689D060F82D" ID-GBIF-Occurrence="1917355192" box="[199,857,2012,2034]" collectorName="Charleston Mts., C" elevation="2286" location="Charleston Mts." pageId="25" pageNumber="26" specimenCount="1" stateProvince="Nevada" typeStatus="holotype">
<emphasis id="B918EA8BFFEAF319FF609689D263F82D" bold="true" box="[199,346,2012,2034]" pageId="25" pageNumber="26">
<typeStatus id="54D7883BFFEAF319FF609689D3C5F82D" box="[199,252,2012,2034]" pageId="25" pageNumber="26">Type</typeStatus>
locality:
</emphasis>
“
<collectorName id="2699534FFFEAF319FECE9689D113F82D" box="[361,554,2012,2034]" pageId="25" pageNumber="26">
<location id="8EB36042FFEAF319FECE9689D136F82D" LSID="urn:lsid:plazi:treatment:03C5878FFFEAF31BFF309602D239FE2A:8EB36042FFEAF319FECE9689D136F82D" box="[361,527,2012,2034]" name="Charleston Mts." pageId="25" pageNumber="26" stateProvince="Nevada">Charleston Mts.</location>
, C
</collectorName>
1ark Co.,
<collectingRegion id="49A8F87BFFEAF319FD299689D1E7F82D" box="[654,734,2012,2034]" country="United States of America" name="Nevada" pageId="25" pageNumber="26">Nevada</collectingRegion>
,
<elevation id="0041D1AAFFEAF319FD4F9689D072F82D" box="[744,843,2012,2034]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="25" pageNumber="26" unit="ft" value="7500.0">
<quantity id="4C949B7CFFEAF319FD4F9689D072F82D" box="[744,843,2012,2034]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="25" pageNumber="26" unit="ft" value="7500.0">7500 feet</quantity>
</elevation>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BD33699FFE9F31AFF6091CDD283FF71" blockId="26.[199,595,152,266]" box="[199,442,152,174]" pageId="26" pageNumber="27">Bohart, 1951: 885 (cat).</paragraph>
<paragraph id="8BD33699FFE9F31AFF6091E3D2D7FF13" blockId="26.[199,595,152,266]" box="[199,494,182,204]" pageId="26" pageNumber="27">Krombein, 1979: 1508 (cat.).</paragraph>
<paragraph id="8BD33699FFE9F31AFF609180D16AFF34" blockId="26.[199,595,152,266]" box="[199,595,213,235]" pageId="26" pageNumber="27">
<bibRefCitation id="EFFD4B68FFE9F31AFF609180D2BEFF34" box="[199,391,213,235]" pageId="26" pageNumber="27" refId="ref43608" refString="MacLachlan, W. B. (1980) A key to and notes on the Eumenes of America North of Mexico (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 53 (3), 617 - 621." type="journal article">MacLachlan, 1980</bibRefCitation>
, 53: 618, 619 (key).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE9F31AFF6091A1D170FED5" box="[199,585,244,266]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BD33699FFE9F31AFF6091A1D170FED5" blockId="26.[199,595,152,266]" box="[199,585,244,266]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF6091A1D26AFED5" bold="true" box="[199,339,244,266]" pageId="26" pageNumber="27">Distribution:</emphasis>
<collectingCountry id="F37B7609FFE9F31AFEFE91A0D295FED6" box="[345,428,244,266]" name="United States of America" pageId="26" pageNumber="27">U. S. A.</collectingCountry>
(CA, NV, UT).
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE9F31AFF309063D082FE64" pageId="26" pageNumber="27" type="discussion">
<paragraph id="8BD33699FFE9F31AFF309063D741FE48" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF309063D2F4FE90" bold="true" box="[151,461,310,335]" pageId="26" pageNumber="27">Comments and diagnosis:</emphasis>
<taxonomicName id="4C6C4D1AFFE9F31AFE719062D000FE90" authority="Bohart, 1950" authorityName="Bohart" authorityYear="1950" box="[470,825,310,335]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFE9F31AFE719062D1A3FE91" box="[470,666,310,334]" italics="true" pageId="26" pageNumber="27">Eumenes sculleni</emphasis>
Bohart, 1950
</taxonomicName>
, can be identified by the shiny integument, especially on the metasoma; the long golden pubescence along the body; the weak and very sparse punctation on the metasoma; the globose T2, with its lateral angle strongly concave and rising up abruptly.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF6090F7D082FE64" blockId="26.[151,1437,310,2027]" box="[199,955,418,443]" pageId="26" pageNumber="27">
Redescription.
<emphasis id="B918EA8BFFE9F31AFED790F7D06BFE64" bold="true" box="[368,850,418,443]" pageId="26" pageNumber="27">
Holotype
<taxonomicName id="4C6C4D1AFFE9F31AFE4390F6D06BFE64" authority="Bohart, 1950" authorityName="Bohart" authorityYear="1950" box="[484,850,418,443]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFE9F31AFE4390F6D188FE64" bold="true" box="[484,689,418,443]" italics="true" pageId="26" pageNumber="27">Eumenes sculleni</emphasis>
Bohart, 1950
</taxonomicName>
</emphasis>
(
<figureCitation id="13572A1CFFE9F31AFCC690F7D088FE64" box="[865,945,418,443]" captionStart="FIGURES 13–26" captionStartId="42.[151,264,1649,1671]" captionTargetBox="[152,1436,193,1628]" captionTargetId="figure@42.[151,1436,193,1628]" captionTargetPageId="42" captionText="FIGURES 13–26. Eumenes species, lateral view. 13) E. cubensis Cresson, 1865 (female); 14) E. flavitinctus Bohart, 1950 (Holotype, female); 15) E. fervens Say, 1824 (Lectotype, female); 16) E. pictus (Smith, 1857) (female); 17) E. sculleni Bohart, 1950 (Holotype, male); 18) E. smithii de Saussure, 1852 (female); 19) E. belfragei Cresson, 1872 (Lectotype, female); 20) E. verticalis Say, 1824 (female); 21) Eumenes verticalis coloradensis Cresson, 1875 (lectotype, female); 22) E. verticalis neoboreus Bequaert, 1944 (Holotype, male); 23) E. verticalis tricinctus Isely, 1917 (Holotype, female); 24) E. filiformis de Saussure, 1855 (famele); 25) E. rufomaculata Fox, 1899 (Lectotype, female); 26) Eumenes versicolor de Saussure, 1852 (Lectotype, female). Scale bar = 1mm." httpUri="https://zenodo.org/record/1458306/files/figure.png" pageId="26" pageNumber="27">Fig. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE9F31BFF609093D093FF27" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BD33699FFE9F31AFF609093D6A4FDDC" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF609093D291FE00" bold="true" box="[199,424,454,479]" pageId="26" pageNumber="27">Male. Color, head:</emphasis>
Black with yellow marks as follows: mandibles; labrum; clypeus; inter-antennal region; broad longitudinal band on scape; narrow and short band on upper half of gena, adjacent to the compound eye.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF30935BD6BCFC98" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF30935BD227FDF8" bold="true" box="[151,286,526,551]" pageId="26" pageNumber="27">Mesosoma:</emphasis>
Black, with yellow marks as follows: broad band on anterior portion of pronotum and a small mark on posterior area, right next to the tegula; small marks on each side of anterior half of mesoscutum; tegula (except for a black mark on central area); parategula; wide mark on upper half of mesepisternum, adjacent to the mesepimeron, and a small mark on basal half; spots on each side of scutellum; almost entirely metanotum; broad band along the lateral portion of the posterior face of propodeum. Wide marks on coxae and femora; entirely tibiae and tarsi, although the distal tarsi are dark brownish.
<emphasis id="B918EA8BFFE9F31AFD2F9397D037FD04" bold="true" box="[648,782,706,731]" pageId="26" pageNumber="27">Metasoma:</emphasis>
Black with yellow marks as follows: wide mark on each side of middle area of T1; narrow and transverse band on apex of T1; broad band on each side of middle area of T2, contiguous laterally with a broad band on apex. Broad band on apex of T3–T6 and S2–S6. T7 and S7 entirely black.
<emphasis id="B918EA8BFFE9F31AFF46927BD20FFC98" bold="true" box="[225,310,814,839]" pageId="26" pageNumber="27">Wings:</emphasis>
Hyaline, with costal, medial and submedial cells yellowish. Pterostigma and veins dark brownish.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF609207D6A4FBC0" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF609207D260FCB4" bold="true" box="[199,345,850,875]" pageId="26" pageNumber="27">Pubescence:</emphasis>
In lateral or oblique view, the pubescence that is under yellow marks along the integument might give the impression of being whitish, but all the pubescence along the body shows the same color pattern.
<emphasis id="B918EA8BFFE9F31AFAF49223D6A5FC50" bold="true" box="[1363,1436,886,911]" pageId="26" pageNumber="27">Head:</emphasis>
Very long yellowish pubescence mainly on frons and occipital region. Pubescence on scape long but shorter than on frons. Pubescence on base and lateral margins of clypeus as long as on frons, while on the apex it becomes shorter.
<emphasis id="B918EA8BFFE9F31AFF5092B7D247FC24" bold="true" box="[247,382,994,1019]" pageId="26" pageNumber="27">Mesosoma:</emphasis>
Long (as long as on frons) and yellowish pubescence homogeneously distributed. Long (shorter than the rest of mesosoma) on anterior face of coxae. Erect bristles all over the internal face of fore femur.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF30957FD219FB54" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF30957FD224FB9C" bold="true" box="[151,285,1066,1091]" pageId="26" pageNumber="27">Metasoma:</emphasis>
T1 with yellowish and long (as long as mesosoma). Base of T2 with yellowish and long (shorter than on T1) pubescence that becomes slightly shorter towards apex. T3–T7 and S2–S7 with short and yellowish pubescence.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF6095C3D257FABC" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF6095C3D11FFB70" bold="true" box="[199,550,1174,1199]" pageId="26" pageNumber="27">Surface of integument, head:</emphasis>
Clypeus with coarse punctation homogeneously distributed; frons and vertex with coarse and moderately dense punctation, slightly sparser behind the ocelli.
<emphasis id="B918EA8BFFE9F31AFBD995EFD63CFB0C" bold="true" box="[1150,1285,1210,1235]" pageId="26" pageNumber="27">Mesosoma:</emphasis>
Coarse and moderately dense (as on frons) covering all mesosoma.
<emphasis id="B918EA8BFFE9F31AFCA2958BD0B2FB28" bold="true" box="[773,907,1246,1271]" pageId="26" pageNumber="27">Metasoma:</emphasis>
T1 with reduced punctation, at least with some punctures more evident on apical half; T2 with reduced punctation on basal half, becoming more evident and denser towards apex; S2 with evident punctation scattered mainly on lateral margins; T3–T7 and S3–S7 without evident punctation.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF60943BD6A5F9C8" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF60943BD2B0FA58" bold="true" box="[199,393,1390,1415]" pageId="26" pageNumber="27">Structure, head:</emphasis>
Clypeus convex, longer than wide, with the apex concave forming two projections with well developed carinae on apex of each one. Inter-antennal region longitudinally cariniform, narrow than the antennal socket. Lateral ocelli closer to the compound eyes than to each other. Occipital carina not angled on middle of gena and very close to the compound eyes next the mandibles. F11 long, with the apex pointed, surpassing the apical edge of F8 (
<figureCitation id="13572A1CFFE9F31AFEB994ABD24BF9C8" box="[286,370,1534,1559]" captionStart="FIGURES 77–92" captionStartId="48.[151,264,1946,1968]" captionTargetBox="[151,1436,193,1925]" captionTargetId="figure@48.[151,1436,193,1925]" captionTargetPageId="48" captionText="FIGURES 77–92. First tergum, dorsal view. 77) Eumenes americanus; 78) E. bequaerti; 79) E. bollii; 80) E. consobrinus; 81) E. coyotae; 82) E. crucifera; 83) E. cubensis; 84) E. flavitinctus; 85) E. fraternus; 86) E. pictus; 87) E. sculleni; 88) E. smithii; 89) E. verticalis; 90) E. filiformis; 91) E. rufomaculactus; 92) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458318/files/figure.png" pageId="26" pageNumber="27">Fig. 87</figureCitation>
). Dorsal surface without longitudinal carina. Microscopic erect bristles on ventral surface of it.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF309777D70DF930" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF309777D227F9E4" bold="true" box="[151,286,1570,1595]" pageId="26" pageNumber="27">Mesosoma:</emphasis>
Pronotal carina on dorsal face of pronotum weakly developed, while on lateral face it is more developed. Pronotal fovea present. Mesepimeron slightly elevated regarding the mesepisternum level. Posterior projection of tegula weakly developed, rounded; parategula lamelliform; posterior face of propodeum strongly concave.
<emphasis id="B918EA8BFFE9F31AFEA497DBD2B0F978" bold="true" box="[259,393,1678,1703]" pageId="26" pageNumber="27">Metasoma:</emphasis>
T1 about 2.5 times longer than wide, strongly swollen in dorsal view (
<figureCitation id="13572A1CFFE9F31AFB1897DBD62CF978" box="[1215,1301,1678,1703]" captionStart="FIGURES 27–42" captionStartId="43.[151,264,1902,1924]" captionTargetBox="[151,1435,193,1881]" captionTargetId="figure@43.[151,1436,193,1881]" captionTargetPageId="43" captionText="FIGURES 27–42. First tergum, dorsal view. 27) Eumenes americanus; 28) E. bequaerti; 29) E. bollii; 30) E. consobrinus; 31) E. coyotae; 32) E. crucifera; 33) E. cubensis; 34) E. flavitinctus; 35) E. fraternus; 36) E. pictus; 37) E. sculleni; 38) E. smithii; 39) E. verticalis; 40) E. filiformis; 41) E. rufomaculactus; 42) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458308/files/figure.png" pageId="26" pageNumber="27">Fig. 37</figureCitation>
); T2 longer than wide, laterally globose (
<figureCitation id="13572A1CFFE9F31AFE7997E7D10BF914" box="[478,562,1714,1739]" captionStart="FIGURES 43–58" captionStartId="44.[151,264,1946,1968]" captionTargetBox="[202,1388,195,1934]" captionTargetId="figure@44.[201,1388,193,1934]" captionTargetPageId="44" captionText="FIGURES 43–58. Second tergum, dorsal view. 43) Eumenes americanus; 44) E. bequaerti; 45) E. bollii; 46) E. consobrinus; 47) E. coyotae; 48) E. crucifera; 49) E. cubensis; 50) E. flavitinctus; 51) E. fraternus; 52) E. pictus; 53) E. sculleni; 54) E. smithii; 55) E. verticalis; 56) E. filiformis; 57) E. rufomaculactus; 58) E. versicolor. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458310/files/figure.png" pageId="26" pageNumber="27">Fig. 53</figureCitation>
). Basal angle of T2 rising up abruptly, concave (
<figureCitation id="13572A1CFFE9F31AFBF197E7D792F914" box="[1110,1195,1714,1739]" captionStart="FIGURES 59–61" captionStartId="45.[151,264,1004,1026]" captionTargetBox="[151,1436,193,983]" captionTargetId="figure@45.[151,1436,193,983]" captionTargetPageId="45" captionText="FIGURES 59–61. Second tergum, lateral view. 59) Eumenes consobrinus (female); 60) Eumenes sculleni (female). 61) First and second terga of E. filiformis (female). Pre-apical projection on the pre-apical ventral surface of T1." httpUri="https://zenodo.org/record/1458312/files/figure.png" pageId="26" pageNumber="27">Fig. 60</figureCitation>
). Apex of T2 with an apical lamella that becomes reduced towards lateral margins. S7 flattened apically.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF6097AFD2D4F8A0" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF6097AFD245F8CC" bold="true" box="[199,380,1786,1811]" pageId="26" pageNumber="27">Male genitalia:</emphasis>
Aedeagus as in figure 103a, b. Paramere (
<figureCitation id="13572A1CFFE9F31AFCCE97AFD0EEF8CC" box="[873,983,1786,1811]" captionStart="FIGURES 102–109" captionStartId="50.[151,264,1926,1948]" captionTargetBox="[151,1436,193,1903]" captionTargetId="figure@50.[151,1436,193,1904]" captionTargetPageId="50" captionText="FIGURES 102–109. Male genitalia: a, b) aedeagus (lateral and ventral view, respectively); c) paramere, internal view. 102) E. pictus; 103) E. sculleni; 104) E. smithii; 105) E. verticalis; 106) E. filiformis; 107) E. rufomaculatus; 108) E. versicolor; 109) General scheme of the apical projection of the aedeagus: a) weakly concave/truncate; b) strongly concave. Scale bar = 1mm." httpUri="https://zenodo.org/record/1458322/files/figure.png" pageId="26" pageNumber="27">Fig. 103c</figureCitation>
) with long bristles on middle region of gonostyle; digitus broad and short, nor surpassing the distal lobe (fig. 120) with scattered short bristles; cuspis with erect bristles, longer than on digitus; volsella with scattered short bristles and long ones on ventral margin; distal lobe rounded, without bristles.
</paragraph>
<paragraph id="8BD33699FFE9F31AFF6096DFD262F818" blockId="26.[151,1437,310,2027]" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF6096DFD21EF87C" bold="true" box="[199,295,1930,1955]" pageId="26" pageNumber="27">Female:</emphasis>
Besides the usual color, pubescence, and punctation as in males. Clypeus broader than males, but still longer than wide.
</paragraph>
<paragraph id="8BD33699FFE9F31BFF609687D093FF27" blockId="26.[151,1437,310,2027]" lastBlockId="27.[151,1437,151,501]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
<emphasis id="B918EA8BFFE9F31AFF609687D278F834" bold="true" box="[199,321,2002,2027]" pageId="26" pageNumber="27">Variation:</emphasis>
Clypeus may have black marks on center area; yellow marks may be more developed; yellow marks on scutellum contiguous; yellow bands on middle area of T
<date id="FFD21059FFE8F31BFC2591CDD0FEFF6F" box="[898,967,152,176]" pageId="27" pageNumber="28">2 may</date>
be contiguous dorsally. Punctation on T1 and on basal half of T
<date id="FFD21059FFE8F31BFE2A91E9D2EBFF0A" box="[397,466,188,213]" pageId="27" pageNumber="28">2 may</date>
be more evident; punctation on frons and posterior face of propodeum may be denser. Concavity on posterior face of propodeum usually weaker on females.
</paragraph>
</subSubSection>
<subSubSection id="C3766512FFE8F31BFF609051D239FE2A" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BD33699FFE8F31BFF609051D239FE2A" blockId="27.[151,1437,151,501]" pageId="27" pageNumber="28">
<emphasis id="B918EA8BFFE8F31BFF609051D240FEC2" bold="true" box="[199,377,260,285]" pageId="27" pageNumber="28">Type material:</emphasis>
The holotype male (CAS) of
<taxonomicName id="4C6C4D1AFFE8F31BFD709050D775FEC2" authority="Bohart, 1950" authorityName="Bohart" authorityYear="1950" box="[727,1100,260,285]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE8F31BFD709050D097FEC3" box="[727,942,260,284]" italics="true" pageId="27" pageNumber="28">Eumenes verticalis</emphasis>
Bohart, 1950
</taxonomicName>
is in excellent condition and bears the labels: ‘Charlerston Mts. \ Kyle Canyon \ Clark Co., Nev.’ ‘About
<quantity id="4C949B7CFFE8F31BFC4C9072D706FE9F" box="[1003,1087,295,320]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="27" pageNumber="28" unit="ft" value="7500.0">7500 ft</quantity>
. \
<date id="FFD21059FFE8F31BFBFF9072D7D6FE9F" box="[1112,1263,295,320]" pageId="27" pageNumber="28" value="1942-07-25">July 25, 1942</date>
\ H. A. Sculen, Col.’ ‘[Red Label]
<taxonomicName id="4C6C4D1AFFE8F31BFECB9018D2E8FEBB" box="[364,465,333,356]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFE8F31BFECB9018D2E8FEBB" box="[364,465,333,356]" italics="true" pageId="27" pageNumber="28">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE8F31BFE409019D176FEBB" authorityName="Bohart" authorityYear="1950" box="[487,591,332,356]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE8F31BFE409019D176FEBB" box="[487,591,332,356]" italics="true" pageId="27" pageNumber="28">verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE8F31BFDC39019D182FEBB" authorityName="Bohart" authorityYear="1950" box="[612,699,332,356]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFE8F31BFDC39019D182FEBB" box="[612,699,332,356]" italics="true" pageId="27" pageNumber="28">sculleni</emphasis>
</taxonomicName>
\ R. Bohart’ ‘California Academy \ of Sciences \ Type No. 6143’. Four paratypes, one female and three males (UCD) are in good condition and bears the labels: [female] ‘Charlerston Mts. \ Kyle Canyon \ Clark Co., Nev. \ About
<quantity id="4C949B7CFFE8F31BFC9690C1D0BBFE72" box="[817,898,404,429]" metricMagnitude="3" metricUnit="m" metricValue="2.37744" pageId="27" pageNumber="28" unit="ft" value="7800.0">7800 ft</quantity>
\
<date id="FFD21059FFE8F31BFC3F90C1D709FE73" box="[920,1072,404,429]" pageId="27" pageNumber="28" value="1942-07-25">July 25, 1942</date>
\ H. A. Scullen, Col.’ ‘
<taxonomicName id="4C6C4D1AFFE8F31BFA9090C0D6A5FE73" box="[1335,1436,405,428]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B918EA8BFFE8F31BFA9090C0D6A5FE73" box="[1335,1436,405,428]" italics="true" pageId="27" pageNumber="28">Eumenes</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE8F31BFF0290EDD234FE0F" authorityName="Bohart" authorityYear="1950" box="[165,269,440,464]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="B918EA8BFFE8F31BFF0290EDD234FE0F" box="[165,269,440,464]" italics="true" pageId="27" pageNumber="28">verticalis</emphasis>
</taxonomicName>
\
<taxonomicName id="4C6C4D1AFFE8F31BFE8490EDD243FE0F" authorityName="Bohart" authorityYear="1950" box="[291,378,440,464]" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sculleni">
<emphasis id="B918EA8BFFE8F31BFE8490EDD243FE0F" box="[291,378,440,464]" italics="true" pageId="27" pageNumber="28">sculleni</emphasis>
</taxonomicName>
\ R. Bohart’; ‘Charleston Mt. Park \ Nev.
<quantity id="4C949B7CFFE8F31BFCF790EDD09AFE0F" box="[848,931,439,464]" metricMagnitude="3" metricUnit="m" metricValue="2.7432" pageId="27" pageNumber="28" unit="ft" value="9000.0">9000 ft</quantity>
.
<date id="FFD21059FFE8F31BFC0990EDD727FE0F" box="[942,1054,440,464]" pageId="27" pageNumber="28" value="1940-06-21">VI-21-40</date>
’; two males bear the same label as holotype.
</paragraph>
</subSubSection>
</treatment>
</document>
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