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<document id="04BC2F0084104795C13B947ABBD9C00E" ID-CLB-Dataset="36900" ID-DOI="10.11646/zootaxa.3765.1.2" ID-GBIF-Dataset="d7ac8ea4-79e3-4375-a4f6-a5bf6922b5e1" ID-ISSN="1175-5326" ID-Zenodo-Dep="285560" ID-ZooBank="BBD41CC5-D3E4-4FEF-B06D-6977693270AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460384167946" checkinUser="plazi" docAuthor="Mapouyat, Lissa, Hirschfeld, Mareike, Rödel, Mark-Oliver, Liedtke, H. Christoph, Loader, Simon P., Gonwouo, L. Nono, Dahmen, Matthias, Doherty-Bone, Thomas M. &amp; Barej, Michael F." docDate="2014" docId="038987F9FF89FFE5FAC25225FDCDFD09" docLanguage="en" docName="zt03765p053.pdf" docOrigin="Zootaxa 3765 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptodactylodon mertensi Perret 1959" docType="treatment" docVersion="8" lastPageNumber="39" masterDocId="FFB0FF81FF81FFEFFA555533FFE7FFE6" masterDocTitle="The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae)" masterLastPageNumber="53" masterPageNumber="29" pageNumber="37" updateTime="1698297238185" updateUser="ExternalLinkService">
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<mods:title id="8F193FF29EED3A8B5866CE336BB6DBCB">The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae)</mods:title>
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<mods:namePart id="12D4A5A9B23C251A02A143F606D4B61B">Hirschfeld, Mareike</mods:namePart>
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<mods:namePart id="2996032DBCD58C813439A8A2A76A6B18">Rödel, Mark-Oliver</mods:namePart>
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<mods:namePart id="73A18AF20687446FB1164E7784166596">Liedtke, H. Christoph</mods:namePart>
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<mods:namePart id="71FF84B156E406B0BC28D8A528853383">Loader, Simon P.</mods:namePart>
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<mods:namePart id="B87F1C5231786AE0ED9BA55F063DA82E">Gonwouo, L. Nono</mods:namePart>
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<mods:namePart id="E7CB3AFD6E72A2D37494E60AADE6BCCA">Dahmen, Matthias</mods:namePart>
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<mods:namePart id="BC815129704D0FF3BDECB8727594D9EE">Doherty-Bone, Thomas M.</mods:namePart>
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<treatment id="038987F9FF89FFE5FAC25225FDCDFD09" ID-DOI="http://doi.org/10.5281/zenodo.5626129" ID-GBIF-Taxon="127667042" ID-Zenodo-Dep="5626129" LSID="urn:lsid:plazi:treatment:038987F9FF89FFE5FAC25225FDCDFD09" httpUri="http://treatment.plazi.org/id/038987F9FF89FFE5FAC25225FDCDFD09" lastPageId="10" lastPageNumber="39" pageId="8" pageNumber="37">
<subSubSection id="C33A6564FF89FFE5FAC25225FDCDFD09" lastPageId="10" lastPageNumber="39" pageId="8" pageNumber="37" type="nomenclature">
<paragraph id="8B9F36EFFF89FFE7FAC25225FDBEF8C9" blockId="8.[151,601,1814,1839]" box="[151,601,1814,1839]" pageId="8" pageNumber="37">
<heading id="D0D78183FF89FFE7FAC25225FDBEF8C9" bold="true" box="[151,601,1814,1839]" fontSize="10" level="2" pageId="8" pageNumber="37" reason="2">
<taxonomicName id="4C204D6CFF89FFE7FAC25225FDBEF8C9" ID-CoL="3TN2Q" authority="Perret, 1959" authorityName="Perret" authorityYear="1959" box="[151,601,1814,1839]" class="Amphibia" family="Arthroleptidae" genus="Leptodactylodon" kingdom="Animalia" order="Anura" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="mertensi">
<emphasis id="B954EAFDFF89FFE7FAC25225FDBEF8C9" bold="true" box="[151,601,1814,1839]" pageId="8" pageNumber="37">
<emphasis id="B954EAFDFF89FFE7FAC25225FE26F8C9" bold="true" box="[151,449,1814,1839]" italics="true" pageId="8" pageNumber="37">Leptodactylodon mertensi</emphasis>
Perret, 1959
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</heading>
</paragraph>
<paragraph id="8B9F36EFFF89FFE6FAC2526CFE8AFDFE" blockId="8.[151,1436,1886,2019]" lastBlockId="9.[151,1437,151,645]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
We examined 48 tadpoles, all from Mount Manengouba: ZMB 78499 (two tadpoles, Gosner stages 25 &amp; 40, Ebonemin,
<geoCoordinate id="EE145028FF89FFE7FB4D52B1FE54F87C" box="[280,435,1922,1947]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="1" value="5.0259504">5°1’33.42” N</geoCoordinate>
,
<geoCoordinate id="EE145028FF89FFE7FBEB52B1FDBBF87D" box="[446,604,1922,1947]" direction="east" orientation="longitude" pageId="8" pageNumber="37" precision="1" value="9.762983">9°45’46.74”E</geoCoordinate>
,
<quantity id="4CD89B0AFF89FFE7F83D52B1FD58F87C" box="[616,703,1922,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.3559999999999999" pageId="8" pageNumber="37" unit="m" value="1356.0">1356 m</quantity>
,
<date id="FF9E102FFF89FFE7F89E52B0FC41F87D" box="[715,934,1922,1947]" pageId="8" pageNumber="37" value="2010-12-13">13 December 2010</date>
); ZMB 78500 (eight tadpoles, Gosner stages 25 &amp; 42, Ebonemin,
<geoCoordinate id="EE145028FF89FFE7FBDD5295FDEBF859" box="[392,524,1958,1983]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="1" value="5.0129166">5°0’46.5”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF89FFE7F84C5295FD50F859" box="[537,695,1958,1983]" direction="east" orientation="longitude" pageId="8" pageNumber="37" precision="1" value="9.762983">9°45’46.74”E</geoCoordinate>
,
<quantity id="4CD89B0AFF89FFE7F8915294FCFBF858" box="[708,796,1959,1983]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="8" pageNumber="37" unit="m" value="1372.0">1372 m</quantity>
,
<date id="FF9E102FFF89FFE7F97F5295FBE0F859" box="[810,1031,1958,1983]" pageId="8" pageNumber="37" value="2010-12-16">16 December 2010</date>
); ZMB 78501 (one tadpole, Gosner stage 25, Ebonemin,
<geoCoordinate id="EE145028FF89FFE7FBD452F9FDE2F805" box="[385,517,1994,2019]" direction="north" orientation="latitude" pageId="8" pageNumber="37" precision="1" value="5.0129166">5°0’46.5”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF89FFE7F84552F9FD76F805" box="[528,657,1994,2019]" direction="east" orientation="longitude" pageId="8" pageNumber="37" precision="1" value="9.768277">9°46’5.8”E</geoCoordinate>
,
<quantity id="4CD89B0AFF89FFE7F8C952F8FD14F804" box="[668,755,1995,2019]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="8" pageNumber="37" unit="m" value="1372.0">1372 m</quantity>
,
<date id="FF9E102FFF89FFE7F8AB52F8FC49F805" box="[766,942,1994,2019]" pageId="8" pageNumber="37" value="2011-10-07">7 October 2011</date>
); ZMB 78503 (one tadpole Gosner stage 25, Ebonemin,
<geoCoordinate id="EE145028FF88FFE6FB4D55A4FE68FF56" box="[280,399,151,176]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="5.018389">5°1’6.2”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6FBCF55A4FDCEFF56" box="[410,553,151,176]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.771333">9°46’16.8”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6F86055ABFD6AFF49" box="[565,653,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.4469999999999998" pageId="9" pageNumber="38" unit="m" value="1447.0">1447 m</quantity>
,
<date id="FF9E102FFF88FFE6F8CC55ABFCADFF56" box="[665,842,151,176]" pageId="9" pageNumber="38" value="2011-10-09">9 October 2011</date>
); ZMB 78504 (two tadpoles, Gosner stages 25 &amp; 27, Caldera,
<geoCoordinate id="EE145028FF88FFE6FAA8558FFE65FF33" box="[253,386,188,213]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="5.0406113">5°2’26.2”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6FBDB558FFDEEFF32" box="[398,521,188,212]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="15" value="9.808333">9°48’30”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6F840558FFD89FF32" box="[533,622,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.889" pageId="9" pageNumber="38" unit="m" value="1889.0">1889 m</quantity>
,
<date id="FF9E102FFF88FFE6F82E558FFCAAFF32" box="[635,845,188,213]" pageId="9" pageNumber="38" value="2011-09-02">2 September 2011</date>
); ZMB 78505 (two tadpoles, Gosner stage 25 &amp; 28, Caldera,
<geoCoordinate id="EE145028FF88FFE6FB5755ECFE61FF1E" box="[258,390,223,248]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="5.0406113">5°2’26.2”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6FBCD55D3FDF4FF1E" box="[408,531,224,248]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="15" value="9.808333">9°48’30”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6F87155D3FD66FF11" box="[548,641,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.889" pageId="9" pageNumber="38" unit="m" value="1889.0">1889 m</quantity>
,
<date id="FF9E102FFF88FFE6F8C655D3FCB9FF1E" box="[659,862,223,248]" pageId="9" pageNumber="38" value="2011-10-17">17 October 2011</date>
); ZMB 78507 (three tadpoles, Gosner stage 25, Manengouba village,
<geoCoordinate id="EE145028FF88FFE6FBC45437FDC3FEFB" box="[401,548,260,285]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="4.9473057">4°56’50.3”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6F8665437FD25FEFB" box="[563,706,260,285]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.879389">9°52’45.8”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6F8845437FCFAFEFA" box="[721,797,260,285]" metricMagnitude="2" metricUnit="m" metricValue="9.959999999999999" pageId="9" pageNumber="38" unit="m" value="996.0">996 m</quantity>
,
<date id="FF9E102FFF88FFE6F9795437FC2BFEFA" box="[812,972,260,284]" pageId="9" pageNumber="38" value="2011-06-28">28 June 2011</date>
); ZMB 78508–09 (six tadpoles and one tadpole, respectively, Gosner stage 25, M’Bouroukou,
<geoCoordinate id="EE145028FF88FFE6F9455414FC73FEA6" box="[784,916,295,320]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="5.0633893">5°3’48.2”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6F9F65414FBD4FEA6" box="[931,1075,295,320]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.866692">9°52’0.09”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6FE175414FB7BFED9" box="[1090,1180,295,320]" metricMagnitude="3" metricUnit="m" metricValue="1.518" pageId="9" pageNumber="38" unit="m" value="1518.0">1518 m</quantity>
,
<date id="FF9E102FFF88FFE6FEFE541BFA68FEA6" box="[1195,1423,295,320]" pageId="9" pageNumber="38" value="2011-09-22">22 September 2011</date>
); ZMB 78511 (one tadpole, Gosner stage 25, Nkack,
<geoCoordinate id="EE145028FF88FFE6F8B8547FFC96FE83" box="[749,881,332,357]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="5.0381665">5°2’17.4”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6F92A547FFBE8FE83" box="[895,1039,332,357]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.77425">9°46’27.3”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6FE48547FFB91FE82" box="[1053,1142,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.466" pageId="9" pageNumber="38" unit="m" value="1466.0">1466 m</quantity>
,
<date id="FF9E102FFF88FFE6FED1547FFAAEFE82" box="[1156,1353,332,357]" pageId="9" pageNumber="38" value="2011-10-12">12 October 2011</date>
); ZMB 78513 (five tadpoles, Gosner stage 25, Pastoral Nkongsamba,
<geoCoordinate id="EE145028FF88FFE6F932545CFBE2FE6E" box="[871,1029,367,392]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="4.987017">4°59’13.26”N</geoCoordinate>
<geoCoordinate id="EE145028FF88FFE6FE5A5443FB4EFE6E" box="[1039,1193,367,392]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.879416">9° 52’45.9”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6FEE2545CFAF7FE61" box="[1207,1296,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.459" pageId="9" pageNumber="38" unit="m" value="1459.0">1459 m</quantity>
,
<date id="FF9E102FFF88FFE6FF4A5443FF35FE4A" pageId="9" pageNumber="38" value="2011-01-18">18 January 2011</date>
); ZMB 78514–16 (two tadpoles, eight tadpoles and six tadpoles, respectively; Gosner stages 25 &amp; 26, Pastoral Nkongsamba,
<geoCoordinate id="EE145028FF88FFE6FBCF5484FDCAFE36" box="[410,557,439,464]" direction="north" orientation="latitude" pageId="9" pageNumber="38" precision="1" value="4.97975">4°58’47.1”N</geoCoordinate>
,
<geoCoordinate id="EE145028FF88FFE6F86D5484FD20FE36" box="[568,711,439,464]" direction="east" orientation="longitude" pageId="9" pageNumber="38" precision="1" value="9.894972">9°53’41.9”E</geoCoordinate>
,
<quantity id="4CD89B0AFF88FFE6F8865484FCCDFE29" box="[723,810,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="9" pageNumber="38" unit="m" value="1225.0">1225 m</quantity>
,
<date id="FF9E102FFF88FFE6F963548BFBF4FE36" box="[822,1043,439,464]" pageId="9" pageNumber="38" value="2011-09-29">29 September 2011</date>
). The tadpoles occurred in small to mid-sized rivers between
<quantity id="4CD89B0AFF88FFE6FB9554EFFDEDFE12" box="[448,522,476,501]" metricMagnitude="2" metricUnit="m" metricValue="9.959999999999999" pageId="9" pageNumber="38" unit="m" value="996.0">996 m</quantity>
and
<quantity id="4CD89B0AFF88FFE6F81054EFFD7AFE12" box="[581,669,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.889" pageId="9" pageNumber="38" unit="m" value="1889.0">1889 m</quantity>
altitude. These riverbanks were covered with forest and farmbush vegetation (
<figureCitation id="131B2A6AFF88FFE6FB4E54CCFE87FDFE" box="[283,352,511,536]" captionStart="FIGURE 2" captionStartId="2.[151,250,1236,1258]" captionTargetBox="[151,1436,265,1214]" captionTargetId="figure@2.[151,1436,265,1215]" captionTargetPageId="2" captionText="FIGURE 2. Habitats of Leptodactylodon tadpoles; a: small forest fragment behind Manengouba-Village, habitat of L. mertensi and L. ornatus ornatus, Mt. Manengouba, N 4 ° 57 ' 12.6 &quot; E 9 ° 51 ' 33.36 &quot;, 1093 m; b: mountainous forest, habitat of L. erythrogaster, Mt. Manengouba, N 4 ° 57 ' 12.6 &quot; E 9 ° 51 ' 33.36 &quot;, 2100 m; c: strongly degraded farmbush area, habitat of L. bicolor, Mt. Manengouba, near Pola N 5 ° 3 ' 27.78 &quot; E 9 ° 50 ' 8.88 &quot;, 1742 m; d: small forest fragment between farms, habitat of L. mertensi, Mt. Manengouba, near Nkack, N 5 ° 0 ' 46.5 &quot; E 9 ° 46 ' 5.82 &quot;, 1372 m. The tadpoles were caught hiding under stones in river parts with little current." httpUri="https://zenodo.org/record/285562/files/figure.png" pageId="9" pageNumber="38">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B9F36EFFF88FFE6FA925717FD60FD63" blockId="9.[151,1437,151,645]" pageId="9" pageNumber="38">The description is based on ten genotyped tadpoles (ZMB 78500–01, 7850–304, 78507–09, 78511, 78513, 78515) at Gosner stage 25. Body length/total length ratio and description of tail tips was based on non-genotyped specimens of the same developmental stage.</paragraph>
<caption id="DF5F6667FF88FFE6FAC25309FCE8F989" httpUri="https://zenodo.org/record/285568/files/figure.png" pageId="9" pageNumber="38" targetBox="[211,1372,702,1569]" targetPageId="9">
<paragraph id="8B9F36EFFF88FFE6FAC25309FCE8F989" blockId="9.[151,1436,1594,1647]" pageId="9" pageNumber="38">
<emphasis id="B954EAFDFF88FFE6FAC25309FEF4F9A9" bold="true" box="[151,275,1594,1616]" pageId="9" pageNumber="38">FIGURE 8.</emphasis>
Dorsal (a) and lateral view (b) of
<taxonomicName id="4C204D6CFF88FFE6F8395309FC93F9B6" box="[620,884,1594,1616]" class="Amphibia" family="Arthroleptidae" genus="Leptodactylodon" kingdom="Animalia" order="Anura" pageId="9" pageNumber="38" phylum="Chordata" rank="species" species="mertensi">
<emphasis id="B954EAFDFF88FFE6F8395309FC93F9B6" box="[620,884,1594,1616]" italics="true" pageId="9" pageNumber="38">Leptodactylodon mertensi</emphasis>
</taxonomicName>
with sketches of the mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8B9F36EFFF88FFE5FA9253A8FADEFE13" blockId="9.[151,1436,1691,2004]" lastBlockId="10.[151,1436,151,752]" lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38">
<emphasis id="B954EAFDFF88FFE6FA9253A8FE86F952" bold="true" box="[199,353,1691,1716]" pageId="9" pageNumber="38">Morphology.</emphasis>
Long but robust tadpole with long tail and very broad muscular tail axis; body elliptical in dorsal and robust and pointed in lateral view (
<figureCitation id="131B2A6AFF88FFE6F802538CFD7CF93E" box="[599,667,1727,1752]" captionStart="FIGURE 8" captionStartId="9.[151,250,1594,1616]" captionTargetBox="[211,1372,702,1569]" captionTargetId="figure@9.[207,1380,697,1573]" captionTargetPageId="9" captionText="FIGURE 8. Dorsal (a) and lateral view (b) of Leptodactylodon mertensi with sketches of the mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm." httpUri="https://zenodo.org/record/285568/files/figure.png" pageId="9" pageNumber="38">Fig. 8</figureCitation>
a, b); body length 28.3 ± 2.4% (N= 31) of total length; body height 42.2 ± 3.9% (N= 10) of body length; body width 54.0 ± 4.2% (N= 10) of body length, maximum body width on level of spiracle insertion; snout truncate to nearly rounded in dorsal view; nostrils oval and situated laterally; nostrils equidistant from eye and snout tip; eye diameter 12.7 ± 0.9% (N= 10) of body length; inter-orbital distance slightly smaller than inter-nostril; tail fins narrow; dorsal fin mostly originating posterior to tail base (exceptionally fin insertions at or slightly anterior to tail base were observed); narrow dorsal fins, reaching deepest point posterior to mid-tail, slightly deeper than ventral fin; ventral fin originates at level with tail base, runs almost parallel to tail axis; tail tip rounded; tail axis very broad and muscular in dorsal and lateral view; body height 91.0 ± 4.1% (N= 10) of total tail height; maximum height of tail axis 69.1 ± 5.9% (N= 9) of maximum tail height; vent tube dextral; lateral sacs present, extending from spiracle to end of body, covering lower two thirds of flanks; short sinister spiracle, translucent, opening lateral, not visible in dorsal view, originating at mid-body; mouth opening frontal; labial tooth row formula 0/0; both jaw sheaths completely keratinized; upper jaw narrow, almost rectangular, slightly bent, distinctly and uniformly serrated; lower jaw U-shaped, distinctly serrated with a lateral pair of caniniform projections (fangs), median part with six to seven needle-like cusps (
<figureCitation id="131B2A6AFF8BFFE5FE75547FFB84FE83" box="[1056,1123,332,357]" captionStart="FIGURE 8" captionStartId="9.[151,250,1594,1616]" captionTargetBox="[211,1372,702,1569]" captionTargetId="figure@9.[207,1380,697,1573]" captionTargetPageId="9" captionText="FIGURE 8. Dorsal (a) and lateral view (b) of Leptodactylodon mertensi with sketches of the mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm." httpUri="https://zenodo.org/record/285568/files/figure.png" pageId="10" pageNumber="39">Fig. 8</figureCitation>
d;
<bibRefCitation id="EFB14B1EFF8BFFE5FED4547FFA96FE82" author="Channing" box="[1153,1393,332,357]" pageId="10" pageNumber="39" refString="Channing, A., Rodel, M. - O. &amp; Channing, J. (2012) Tadpoles of Africa - The biology and identification of all known tadpoles in sub-Saharan Africa. Edition Chimaira, Frankfurt am Main, 404 pp." type="book" year="2012">
Channing
<emphasis id="B954EAFDFF8BFFE5FEA0547EFAC9FE82" box="[1269,1326,332,356]" italics="true" pageId="10" pageNumber="39">et al.</emphasis>
2012
</bibRefCitation>
: 8); two distinct serrations abaxial to fangs; posterior lip huge semicircular to oval bean shaped, covered with 21 papillae (
<bibRefCitation id="EFB14B1EFF8BFFE5FAAA54A7FE15FE4A" author="Channing" box="[255,498,404,429]" pageId="10" pageNumber="39" refString="Channing, A., Rodel, M. - O. &amp; Channing, J. (2012) Tadpoles of Africa - The biology and identification of all known tadpoles in sub-Saharan Africa. Edition Chimaira, Frankfurt am Main, 404 pp." type="book" year="2012">
Channing
<emphasis id="B954EAFDFF8BFFE5FB2054A6FE48FE4A" box="[373,431,404,428]" italics="true" pageId="10" pageNumber="39">et al.</emphasis>
2012
</bibRefCitation>
: 20); arrangement of papillae symmetric to vertical body axis; papillae arranged in two semicircular rows (
<figureCitation id="131B2A6AFF8BFFE5FBF75484FE02FE36" box="[418,485,439,464]" captionStart="FIGURE 8" captionStartId="9.[151,250,1594,1616]" captionTargetBox="[211,1372,702,1569]" captionTargetId="figure@9.[207,1380,697,1573]" captionTargetPageId="9" captionText="FIGURE 8. Dorsal (a) and lateral view (b) of Leptodactylodon mertensi with sketches of the mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm." httpUri="https://zenodo.org/record/285568/files/figure.png" pageId="10" pageNumber="39">Fig. 8</figureCitation>
c); straight to semicircular skin fold on lower lip, just posterior to lower jaw sheath; oral disc width 34.0 ± 2.6% (N= 10) of body length; mouth width 26.2 ± 3.2% (N= 10) of oral disc width.
</paragraph>
<paragraph id="8B9F36EFFF8BFFE5FA9254CCFB48FD62" blockId="10.[151,1436,151,752]" pageId="10" pageNumber="39">
The most advanced individuals (ZMB 78499, Gosner stage 40; ZMB 78500, three tadpoles of Gosner stages 40–42) measured
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(body length.
<quantity id="4CD89B0AFF8BFFE5F8945717FC8CFDDA" box="[705,875,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.2600000000000002" metricValueMax="1.3900000000000001" metricValueMin="1.1300000000000001" pageId="10" pageNumber="39" unit="mm" value="12.6" valueMax="13.9" valueMin="11.3">11.3–13.9 mm</quantity>
; tail length:
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). A few tadpoles of Gosner stages 25–26 even reached
<quantity id="4CD89B0AFF8BFFE5F8745774FD23FDB9" box="[545,708,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="4.779999999999999" metricValueMax="4.95" metricValueMin="4.61" pageId="10" pageNumber="39" unit="mm" value="47.8" valueMax="49.5" valueMin="46.1">46.1–49.5 mm</quantity>
total length.
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Channing
<emphasis id="B954EAFDFF8BFFE5F999577AFBE3FD86" box="[972,1028,584,608]" italics="true" pageId="10" pageNumber="39">et al.</emphasis>
(2012)
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reported
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total length.
<bibRefCitation id="EFB14B1EFF8BFFE5FAC2575FFEC9FD62" author="Amiet" box="[151,302,620,645]" pageId="10" pageNumber="39" refString="Amiet, J. - L. (1980) Revision du genre Leptodactylodon Andersson (Amphibia, Anura, Astylosterninae). Annales de la Faculte des Sciences de Yaounde, 27, 69 - 224." type="journal article" year="1980">Amiet (1980)</bibRefCitation>
measured metamorphs, still possessing
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tails, with
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SVL.
</paragraph>
<paragraph id="8B9F36EFFF8BFFE5FA9257BCFDCDFD09" blockId="10.[151,1436,151,752]" pageId="10" pageNumber="39">
<emphasis id="B954EAFDFF8BFFE5FA9257BCFDEDFD4E" bold="true" box="[199,522,655,680]" pageId="10" pageNumber="39">Coloration in preservation.</emphasis>
Pale brown body and tail; dorsal parts of body and entire tail axis heavily mottled with little dark spots; ventral parts lighter; fins with very few dark markings, translucent cream white; some individuals with cream-white snout.
</paragraph>
</subSubSection>
</treatment>
</document>
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