treatments-xml/data/03/97/1F/03971F35FFF3FFB9995228F6FD03FCA5.xml

<document id="ECBA9908A847CB75E0DBB263C9974F1F" ID-CLB-Dataset="298344" ID-DOI="10.5852/ejt.2024.935.2557" ID-GBIF-Dataset="b9f405d9-04c7-4cfe-9f28-b6edc3416cb0" ID-ISSN="2118-9773" ID-Zenodo-Dep="11526096" ID-ZooBank="6C66A1E4-7EA4-45B7-B261-5D3ED749568E" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1717844083798" checkinUser="plazi" docAuthor="Lin, Chien-Hsiang, Steurbaut, Etienne &amp; Nolf, Dirk" docDate="2024" docId="03971F35FFF3FFB9995228F6FD03FCA5" docLanguage="en" docName="EJT.2024.935.203-240.pdf" docOrigin="European Journal of Taxonomy 935" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2557/11559" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Haemulon ypresiensis G.Cuvier 1829" docType="treatment" docVersion="3" lastPageNumber="228" masterDocId="FFAE674DFFE4FFA09B062942FFADFFE2" masterDocTitle="Early Eocene fish otoliths from the eastern and southern USA" masterLastPageNumber="240" masterPageNumber="203" pageNumber="226" updateTime="1718049141783" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8C111695D0DC7B4A48450A23D187F5F2">Early Eocene fish otoliths from the eastern and southern USA</mods:title>
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<mods:namePart id="50754384E0585601D3FF6E265A6F3705">Lin, Chien-Hsiang</mods:namePart>
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<mods:affiliation id="6EF33C7899DDDC70A11157212DCE0114">Biodiversity Research Center, Academia Sinica, 128 Academia Road, Seca 2, Nankang, Taipei 11529, Taiwan.</mods:affiliation>
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<mods:namePart id="D93D91952A70174F02B3A3FDBC1DF53A">Steurbaut, Etienne</mods:namePart>
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<mods:affiliation id="D07848B19FA9BF242C98AD8AA1824EB2">Royal Belgian Institute of Natural Sciences, 29 Vautier Street, 1000 Brussels, Belgium. &amp; Department of Earth and Environmental Sciences, KU Leuven, Celestijnenlaan 200 E, 3001 Leuven, Belgium.</mods:affiliation>
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<mods:affiliation id="D68518BE8A008F6CB331372391984802">Royal Belgian Institute of Natural Sciences, 29 Vautier Street, 1000 Brussels, Belgium.</mods:affiliation>
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<treatment id="03971F35FFF3FFB9995228F6FD03FCA5" ID-DOI="http://doi.org/10.5281/zenodo.11552989" ID-GBIF-Taxon="232726621" ID-Zenodo-Dep="11552989" LSID="urn:lsid:zoobank.org:act:946D0B8E-FDC0-4DA4-8A24-44CBBE0F9BA0" httpUri="http://treatment.plazi.org/id/03971F35FFF3FFB9995228F6FD03FCA5" lastPageId="25" lastPageNumber="228" pageId="23" pageNumber="226">
<subSubSection id="C324FDA8FFF3FFB7995228F6FC4DFE2D" box="[596,992,435,463]" pageId="23" pageNumber="226" type="nomenclature">
<paragraph id="8B81AE23FFF3FFB7995228F6FC4DFE2D" blockId="23.[360,1227,435,533]" box="[596,992,435,463]" pageId="23" pageNumber="226">
<emphasis id="B94A7231FFF3FFB7995228F6FCD3FE2C" bold="true" box="[596,894,435,462]" pageId="23" pageNumber="226">
“ 
<taxonomicName id="4C3ED5A0FFF3FFB7996528F1FCD3FE2C" authorityName="G.Cuvier" authorityYear="1829" box="[611,894,435,462]" family="Haemulidae" genus="Haemulon" kingdom="Animalia" order="Perciformes" pageId="23" pageNumber="226" phylum="Chordata" rank="species" species="ypresiensis">
<emphasis id="B94A7231FFF3FFB7996528F1FD49FE2C" bold="true" box="[611,740,435,462]" italics="true" pageId="23" pageNumber="226">Haemulon</emphasis>
” 
<emphasis id="B94A7231FFF3FFB799FC28F6FCD3FE2C" bold="true" box="[762,894,436,462]" italics="true" pageId="23" pageNumber="226">ypresiensis</emphasis>
</taxonomicName>
</emphasis>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="C324FDA8FFF3FFB79A6E2895FCCDFDF7" pageId="23" pageNumber="226" type="description">
<paragraph id="8B81AE23FFF3FFB79A6E2895FB66FE10" blockId="23.[360,1227,435,533]" box="[360,1227,471,498]" pageId="23" pageNumber="226">
<uri id="FFAFA221FFF3FFB79A6E2895FB66FE10" box="[360,1227,471,498]" pageId="23" pageNumber="226">
urn:lsid:zoobank.org:act: 
<uuid id="FF9894F6FFF3FFB799812895FB66FE10" box="[647,1227,471,498]" pageId="23" pageNumber="226">AA387688-D529-4AF7-B3DA-C6E3C2A82413</uuid>
</uri>
</paragraph>
<paragraph id="8B81AE23FFF3FFB799D228B8FCCDFDF7" blockId="23.[360,1227,435,533]" box="[724,864,506,533]" pageId="23" pageNumber="226">
<figureCitation id="1305B2A6FFF3FFB799D228B8FCCDFDF7" box="[724,864,506,533]" captionStart="Fig" captionStartId="24.[189,232,1569,1595]" captionTargetBox="[189,1398,264,1522]" captionTargetId="figure-17@24.[189,1398,264,1522]" captionTargetPageId="24" captionText="Fig. 10. Fish otoliths from the US lower Eocene formations. A. Lactarius amplus Stinton, 1978, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10757). B–C. Lactarius kokeni (Dante &amp; Frizzell, 1965), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10758, P 10759). D. Anisotremus rambo, 2022, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10763). E–F. Orthopristis burlesonis (Dante &amp; Frizzell, 1965), Pamunkey River, Hanovertown, Potapaco Member, Virginia (IRSNB P 10764–10765). G–H. “Haemulon” ypresiensis sp. nov., Pamunkey River, Hanovertown, Potapaco Member, Virginia. G. Holotype (IRSNB P 10766). H. Paratype (IRSNB P 10767). I–J. Ekokenia eporrecta (Koken, 1888), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10768, P 10769). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11526116" httpUri="https://zenodo.org/record/11526116/files/figure.png" pageId="23" pageNumber="226">Fig. 10G–H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324FDA8FFF3FFB79BBB2B7EFDF4FD29" pageId="23" pageNumber="226" type="diagnosis">
<paragraph id="8B81AE23FFF3FFB79BBB2B7EFE9BFDB4" blockId="23.[189,310,572,598]" box="[189,310,572,598]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2B7EFE9BFDB4" bold="true" box="[189,310,572,598]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2B7EFE9BFDB4" bold="true" box="[189,310,572,598]" pageId="23" pageNumber="226">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2B28FDF4FD29" blockId="23.[189,1398,618,715]" pageId="23" pageNumber="226">OL/OH = 1.3–1.5, OsL/CaL = 0.4–0.7. Oblong otoliths with weak postero-dorsal angle. Margins crenulated. Sulcus well-divided. Ostium wide, opens antero-dorsally. Cauda elongate, straight, but strongly bent ventrally at posterior.</paragraph>
</subSubSection>
<subSubSection id="C324FDA8FFF3FFB79BBB2BB0FE82FCBC" pageId="23" pageNumber="226" type="etymology">
<paragraph id="8B81AE23FFF3FFB79BBB2BB0FEE9FCEE" blockId="23.[189,324,754,780]" box="[189,324,754,780]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2BB0FEE9FCEE" bold="true" box="[189,324,754,780]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2BB0FEE9FCEE" bold="true" box="[189,324,754,780]" pageId="23" pageNumber="226">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2A63FE82FCBC" blockId="23.[189,1398,801,862]" pageId="23" pageNumber="226">
‘ 
<emphasis id="B94A7231FFF3FFB79BC12A60FEE4FCD9" box="[199,329,802,827]" italics="true" pageId="23" pageNumber="226">Ypresiensis</emphasis>
, 
<emphasis id="B94A7231FFF3FFB79A5C2A60FEC3FCD9" box="[346,366,802,827]" italics="true" pageId="23" pageNumber="226">is</emphasis>
, 
<emphasis id="B94A7231FFF3FFB79A792A60FE21FCD9" box="[383,396,802,827]" italics="true" pageId="23" pageNumber="226">e</emphasis>
’ = ‘from the Ypresian’. Refers to the geological stage from where the fossils were collected.
</paragraph>
</subSubSection>
<subSubSection id="C324FDA8FFF3FFB79BBB2AC7FB67FA3D" pageId="23" pageNumber="226" type="materials_examined">
<paragraph id="8B81AE23FFF3FFB79BBB2AC7FE41FC7D" blockId="23.[189,492,901,927]" box="[189,492,901,927]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2AC7FE41FC7D" bold="true" box="[189,492,901,927]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2AC7FE41FC7D" bold="true" box="[189,492,901,927]" pageId="23" pageNumber="226">Type material examined</emphasis>
</heading>
</paragraph>
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<materialsCitation id="3B56A47EFFF3FFB79BEC2AF6FDC0FBF6" collectionCode="IRSNB" country="United States of America" county="Left otolith" location="Hanovertown" municipality="Pamunkey River" pageId="23" pageNumber="226" specimenCode="P 10766" specimenCount="1" stateProvince="Virginia" typeStatus="holotype">
<emphasis id="B94A7231FFF3FFB79BEC2AF6FEF1FC2C" bold="true" box="[234,348,948,974]" pageId="23" pageNumber="226">
<typeStatus id="54851081FFF3FFB79BEC2AF6FEF1FC2C" box="[234,348,948,974]" pageId="23" pageNumber="226" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F329EEB3FFF3FFB79BBB2A9AFDF7FC13" box="[189,602,983,1010]" name="United States of America" pageId="23" pageNumber="226">UNITED STATES OF AMERICA</collectingCountry>
• 
<collectingCounty id="62E0D6AFFFF3FFB799712A95FCAFFC13" box="[631,770,983,1009]" pageId="23" pageNumber="226">Left otolith</collectingCounty>
; 
<collectingRegion id="49FA60C1FFF3FFB798172A95FCD9FC13" box="[785,884,983,1009]" country="United States of America" name="Virginia" pageId="23" pageNumber="226">Virginia</collectingRegion>
, 
<collectingMunicipality id="6BE53459FFF3FFB798842A95FBE6FC13" box="[898,1099,983,1009]" pageId="23" pageNumber="226">Pamunkey River</collectingMunicipality>
, 
<location id="8EE1F8F8FFF3FFB79F5C2A9AFB50FC10" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79F5C2A9AFB50FC10" box="[1114,1277,984,1010]" country="United States of America" county="Left otolith" municipality="Pamunkey River" name="Hanovertown" pageId="23" pageNumber="226" stateProvince="Virginia">Hanovertown</location>
, 
<location id="8EE1F8F8FFF3FFB79E0D2A9AFE89FBF6" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79E0D2A9AFE89FBF6" country="United States of America" county="Left otolith" municipality="Pamunkey River" name="Potapaco Member" pageId="23" pageNumber="226" stateProvince="Virginia">Potapaco Member</location>
; 
<figureCitation id="1305B2A6FFF3FFB79A362AB8FE31FBF6" box="[304,412,1018,1044]" captionStart="Fig" captionStartId="24.[189,232,1569,1595]" captionTargetBox="[189,1398,264,1522]" captionTargetId="figure-17@24.[189,1398,264,1522]" captionTargetPageId="24" captionText="Fig. 10. Fish otoliths from the US lower Eocene formations. A. Lactarius amplus Stinton, 1978, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10757). B–C. Lactarius kokeni (Dante &amp; Frizzell, 1965), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10758, P 10759). D. Anisotremus rambo, 2022, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10763). E–F. Orthopristis burlesonis (Dante &amp; Frizzell, 1965), Pamunkey River, Hanovertown, Potapaco Member, Virginia (IRSNB P 10764–10765). G–H. “Haemulon” ypresiensis sp. nov., Pamunkey River, Hanovertown, Potapaco Member, Virginia. G. Holotype (IRSNB P 10766). H. Paratype (IRSNB P 10767). I–J. Ekokenia eporrecta (Koken, 1888), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10768, P 10769). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11526116" httpUri="https://zenodo.org/record/11526116/files/figure.png" pageId="23" pageNumber="226">Fig. 10G</figureCitation>
; 
<collectionCode id="ED2F36E6FFF3FFB79AAE2AB8FE53FBF6" box="[424,510,1018,1044]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
<specimenCode id="DB980658FFF3FFB799032AB9FDC4FBF6" box="[517,617,1018,1045]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">P 10766</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BEC2D7EFE20FA92" blockId="23.[188,1399,1084,1392]" pageId="23" pageNumber="226">
<materialsCitation id="3B56A47EFFF3FFB79BEC2D7EFDB3FB7E" collectionCode="IRSNB" country="United States of America" county="Left" location="UNITED STATES OF AMERICA" municipality="Pamunkey River" pageId="23" pageNumber="226" specimenCode="P 10767" specimenCount="2" stateProvince="Virginia" typeStatus="paratype">
<emphasis id="B94A7231FFF3FFB79BEC2D7EFECAFBB4" bold="true" box="[234,359,1084,1110]" pageId="23" pageNumber="226">
<typeStatus id="54851081FFF3FFB79BEC2D7EFECAFBB4" box="[234,359,1084,1110]" pageId="23" pageNumber="226" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<quantity id="4CC603C6FFF3FFB79A712D7EFE18FBB4" box="[375,437,1084,1110]" metricMagnitude="0" metricUnit="m" metricValue="1.8288" pageId="23" pageNumber="226" unit="in" value="72.0">72 in</quantity>
total) 
<collectingCountry id="F329EEB3FFF3FFB79BBB2D1DFDE2FB9B" box="[189,591,1119,1145]" name="United States of America" pageId="23" pageNumber="226">UNITED STATES OF AMERICA</collectingCountry>
• 
<specimenCount id="9D3865AAFFF3FFB799632D1DFD7FFB9B" box="[613,722,1119,1145]" pageId="23" pageNumber="226">2 otoliths</specimenCount>
of which one is figured: 
<figureCitation id="1305B2A6FFF3FFB798F52D1DFBF3FB9B" box="[1011,1118,1119,1145]" captionStart="Fig" captionStartId="24.[189,232,1569,1595]" captionTargetBox="[189,1398,264,1522]" captionTargetId="figure-17@24.[189,1398,264,1522]" captionTargetPageId="24" captionText="Fig. 10. Fish otoliths from the US lower Eocene formations. A. Lactarius amplus Stinton, 1978, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10757). B–C. Lactarius kokeni (Dante &amp; Frizzell, 1965), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10758, P 10759). D. Anisotremus rambo, 2022, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10763). E–F. Orthopristis burlesonis (Dante &amp; Frizzell, 1965), Pamunkey River, Hanovertown, Potapaco Member, Virginia (IRSNB P 10764–10765). G–H. “Haemulon” ypresiensis sp. nov., Pamunkey River, Hanovertown, Potapaco Member, Virginia. G. Holotype (IRSNB P 10766). H. Paratype (IRSNB P 10767). I–J. Ekokenia eporrecta (Koken, 1888), Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10768, P 10769). 1 = ventral view; 2 = inner view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11526116" httpUri="https://zenodo.org/record/11526116/files/figure.png" pageId="23" pageNumber="226">Fig. 10H</figureCitation>
; same collection data as for holotype; 
<collectionCode id="ED2F36E6FFF3FFB79A592DC0FE18FB7E" box="[351,437,1154,1180]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
<specimenCode id="DB980658FFF3FFB79ABB2DC1FDB3FB7E" box="[445,542,1154,1181]" collectionCode="IRSNB" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">P 10767</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB7993F2DC0FEBEFB5D" collectionCode="IRSNB" collectorName="Bashi Formation" country="United States of America" location="Gallagher Creek" municipality="Meridian" pageId="23" pageNumber="226" specimenCount="10" stateProvince="Mississippi" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB7993F2DC0FD17FB7E" box="[569,698,1154,1180]" pageId="23" pageNumber="226">10 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB799C12DC0FCFDFB7E" box="[711,848,1154,1180]" country="United States of America" name="Mississippi" pageId="23" pageNumber="226">Mississippi</collectingRegion>
, 
<collectingMunicipality id="6BE53459FFF3FFB7985A2DC0FC66FB7E" box="[860,971,1154,1180]" pageId="23" pageNumber="226">Meridian</collectingMunicipality>
, 
<location id="8EE1F8F8FFF3FFB798D12DC0FB31FB7E" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB798D12DC0FB31FB7E" box="[983,1180,1154,1180]" country="United States of America" municipality="Meridian" name="Gallagher Creek" pageId="23" pageNumber="226" stateProvince="Mississippi">Gallagher Creek</location>
, 
<geologicalTimeScale id="F91E2E88FFF3FFB79FAF2DC0FAC3FB7E" box="[1193,1390,1154,1180]" pageId="23" pageNumber="226">Bashi Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB79BBB2DE7FEBEFB5D" box="[189,275,1189,1215]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB79A2C2DE7FB71FB5D" box="[298,1244,1189,1216]" collectionCode="IRSNB" collectorName="Red Hot Truck Stop &amp; Bashi Formation" country="United States of America" location="Meridian" pageId="23" pageNumber="226" specimenCount="36" stateProvince="Mississippi" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB79A2C2DE7FE04FB5D" box="[298,425,1189,1215]" pageId="23" pageNumber="226">36 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB79AB52DE7FD91FB5D" box="[435,572,1189,1215]" country="United States of America" name="Mississippi" pageId="23" pageNumber="226">Mississippi</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB799412DE7FD1BFB5D" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB799412DE7FD1BFB5D" box="[583,694,1189,1215]" country="United States of America" name="Meridian" pageId="23" pageNumber="226" stateProvince="Mississippi">Meridian</location>
, Red Hot Truck Stop, 
<geologicalTimeScale id="F91E2E88FFF3FFB798B32DE7FBD5FB5D" box="[949,1144,1189,1215]" pageId="23" pageNumber="226">Bashi Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB79F802DE7FB71FB5D" box="[1158,1244,1189,1215]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB79FF52DE7FCA4FB01" collectionCode="IRSNB" collectorName="Bashi Formation" country="United States of America" location="Cave Branch" pageId="23" pageNumber="226" specimenCount="13" stateProvince="Alabama" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB79FF52DE7FADFFB5D" box="[1267,1394,1189,1215]" pageId="23" pageNumber="226">13 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB79BBA2D8AFE84FB00" box="[188,297,1224,1250]" country="United States of America" name="Alabama" pageId="23" pageNumber="226">Alabama</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB79A332D8BFE7CFB00" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79A332D8BFE7CFB00" box="[309,465,1224,1251]" country="United States of America" name="Cave Branch" pageId="23" pageNumber="226" stateProvince="Alabama">Cave Branch</location>
, 
<geologicalTimeScale id="F91E2E88FFF3FFB79ADB2D8AFD0FFB00" box="[477,674,1224,1250]" pageId="23" pageNumber="226">Bashi Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB799B52D8BFCA4FB01" box="[691,777,1225,1251]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB798222D8BFE0FFAE4" collectionCode="IRSNB" collectorName="Bashi Formation" country="United States of America" location="Elba Dam on Pea River" pageId="23" pageNumber="226" specimenCount="1" stateProvince="Alabama" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB798222D8BFC27FB00" box="[804,906,1224,1251]" pageId="23" pageNumber="226">1 otolith</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB798932D8AFBAFFB00" box="[917,1026,1224,1250]" country="United States of America" name="Alabama" pageId="23" pageNumber="226">Alabama</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB79F082D8AFA8AFB00" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79F082D8AFA8AFB00" box="[1038,1319,1224,1251]" country="United States of America" name="Elba Dam on Pea River" pageId="23" pageNumber="226" stateProvince="Alabama">Elba Dam on Pea River</location>
, 
<geologicalTimeScale id="F91E2E88FFF3FFB79E322D8AFE95FAE4" pageId="23" pageNumber="226">Bashi Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB79A4A2DAEFE0FFAE4" box="[332,418,1260,1286]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB79AC02DAEFBF6FAE4" box="[454,1115,1260,1286]" collectionCode="IRSNB" collectorName="Bashi Formation" country="United States of America" location="Ozark" pageId="23" pageNumber="226" specimenCount="4" stateProvince="Alabama" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB79AC02DAEFD90FAE4" box="[454,573,1260,1286]" pageId="23" pageNumber="226">4 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB7994B2DAEFD17FAE4" box="[589,698,1260,1286]" country="United States of America" name="Alabama" pageId="23" pageNumber="226">Alabama</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB799CC2DAEFCB8FAE4" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB799CC2DAEFCB8FAE4" box="[714,789,1260,1286]" country="United States of America" name="Ozark" pageId="23" pageNumber="226" stateProvince="Alabama">Ozark</location>
, 
<geologicalTimeScale id="F91E2E88FFF3FFB798202DAEFC5DFAE4" box="[806,1008,1260,1286]" pageId="23" pageNumber="226">Bashi Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB79F032DAEFBF6FAE4" box="[1029,1115,1260,1286]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB79F792DAEFCE7FACB" collectionCode="IRSNB" collectorName="Hatchetigbee Formation" country="United States of America" location="Hatchetigbee Bluff" pageId="23" pageNumber="226" specimenCount="2" stateProvince="Alabama" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB79F792DAEFB5BFAE4" box="[1151,1270,1260,1286]" pageId="23" pageNumber="226">2 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB79E032DAEFADFFAE4" box="[1285,1394,1260,1286]" country="United States of America" name="Alabama" pageId="23" pageNumber="226">Alabama</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB79BBB2C4DFE0BFACB" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79BBB2C4DFE0BFACB" box="[189,422,1295,1321]" country="United States of America" name="Hatchetigbee Bluff" pageId="23" pageNumber="226" stateProvince="Alabama">Hatchetigbee Bluff</location>
, 
<geologicalTimeScale id="F91E2E88FFF3FFB79ABC2C4DFD70FACB" box="[442,733,1295,1321]" pageId="23" pageNumber="226">Hatchetigbee Formation</geologicalTimeScale>
; 
<collectionCode id="ED2F36E6FFF3FFB799F22C4DFCE7FACB" box="[756,842,1295,1321]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB798772C4DFE54FAAE" collectionCode="IRSNB" collectorName="Potapaco Member" country="United States of America" location="Rappahannock River" pageId="23" pageNumber="226" specimenCount="3" stateProvince="Virginia" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB798772C4DFC46FACB" box="[881,1003,1295,1321]" pageId="23" pageNumber="226">3 otoliths</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB798FB2C4DFBCDFACB" box="[1021,1120,1295,1321]" country="United States of America" name="Virginia" pageId="23" pageNumber="226">Virginia</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB79F742C4DFADFFACB" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB79F742C4DFADFFACB" box="[1138,1394,1295,1321]" country="United States of America" name="Rappahannock River" pageId="23" pageNumber="226" stateProvince="Virginia">Rappahannock River</location>
, Potapaco Member; 
<collectionCode id="ED2F36E6FFF3FFB79AA52C70FE54FAAE" box="[419,505,1330,1356]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="3B56A47EFFF3FFB799142C70FE2BFA8D" collectionCode="IRSNB" country="United States of America" location="Loyola Retreat House" pageId="23" pageNumber="226" specimenCount="1" stateProvince="Maryland" typeStatus="paratype">
<specimenCount id="9D3865AAFFF3FFB799142C70FDD5FAAE" box="[530,632,1330,1356]" pageId="23" pageNumber="226">1 otolith</specimenCount>
; 
<collectingRegion id="49FA60C1FFF3FFB799822C70FD54FAAE" box="[644,761,1330,1356]" country="United States of America" name="Maryland" pageId="23" pageNumber="226">Maryland</collectingRegion>
, 
<location id="8EE1F8F8FFF3FFB798032C70FBA7FAAF" LSID="urn:lsid:plazi:treatment:03971F35FFF3FFB9995228F6FD03FCA5:8EE1F8F8FFF3FFB798032C70FBA7FAAF" box="[773,1034,1330,1357]" country="United States of America" name="Loyola Retreat House" pageId="23" pageNumber="226" stateProvince="Maryland">Loyola Retreat House</location>
, 
<locationDeviation id="9A33B701FFF3FFB79F132C71FB45FAAE" box="[1045,1256,1330,1357]" pageId="23" pageNumber="226">N of Popes Creek</locationDeviation>
, Woodstock Member; 
<collectionCode id="ED2F36E6FFF3FFB79A362C17FE2BFA8D" box="[304,390,1365,1391]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="23" pageNumber="226">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2CD5FE56FA52" blockId="23.[189,507,1430,1457]" box="[189,507,1430,1457]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2CD5FE56FA52" bold="true" box="[189,507,1430,1457]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2CD5FE56FA52" bold="true" box="[189,507,1430,1457]" pageId="23" pageNumber="226">Type locality and horizon</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2C87FB67FA3D" blockId="23.[189,1226,1477,1504]" box="[189,1226,1477,1504]" pageId="23" pageNumber="226">
<collectingCountry id="F329EEB3FFF3FFB79BBB2C87FE45FA3D" box="[189,488,1477,1503]" name="United States of America" pageId="23" pageNumber="226">United States of America</collectingCountry>
, Pamunkey River, Hanovertown (
<collectingRegion id="49FA60C1FFF3FFB798722C87FC77FA3D" box="[884,986,1477,1503]" country="United States of America" name="Virginia" pageId="23" pageNumber="226">Virginia</collectingRegion>
), Potapaco Member.
</paragraph>
</subSubSection>
<subSubSection id="C324FDA8FFF3FFB79BBB2F44FB44F870" pageId="23" pageNumber="226" type="description">
<paragraph id="8B81AE23FFF3FFB79BBB2F44FDBDF9C3" blockId="23.[189,528,1542,1569]" box="[189,528,1542,1569]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2F44FDBDF9C3" bold="true" box="[189,528,1542,1569]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2F44FDBDF9C3" bold="true" box="[189,528,1542,1569]" pageId="23" pageNumber="226">
Dimensions of the 
<typeStatus id="54851081FFF3FFB79AA22F45FDBDF9C3" box="[420,528,1543,1569]" pageId="23" pageNumber="226" type="holotype">holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2F77FC29F9B2" blockId="23.[189,900,1589,1616]" box="[189,900,1589,1616]" pageId="23" pageNumber="226">
Length = 
<quantity id="4CC603C6FFF3FFB79A292F77FE36F9B2" box="[303,411,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="3.33" pageId="23" pageNumber="226" unit="mm" value="3.33">3.33 mm</quantity>
; height = 
<quantity id="4CC603C6FFF3FFB799162F77FDD6F9B2" box="[528,635,1589,1616]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" pageId="23" pageNumber="226" unit="mm" value="2.19">2.19 mm</quantity>
; thickness = 
<quantity id="4CC603C6FFF3FFB798122F77FCD2F9B2" box="[788,895,1589,1616]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="23" pageNumber="226" unit="mm" value="0.76">0.76 mm</quantity>
.
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2F34FEE3F972" blockId="23.[189,334,1654,1680]" box="[189,334,1654,1680]" pageId="23" pageNumber="226">
<heading id="D0C9194FFFF3FFB79BBB2F34FEE3F972" bold="true" box="[189,334,1654,1680]" fontSize="11" level="3" pageId="23" pageNumber="226" reason="3">
<emphasis id="B94A7231FFF3FFB79BBB2F34FEE3F972" bold="true" box="[189,334,1654,1680]" pageId="23" pageNumber="226">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFF3FFB79BBB2FE7FB44F870" blockId="23.[188,1399,1701,1939]" pageId="23" pageNumber="226">This species is distinguished by oblong otoliths with crenulated margins and a moderately thick build. The inner face is convex and the outer face is concave. The sulcus is clearly devided with crest-like cristae forming a deep incised sulcus. The ostium opens widely and antero-dorsally, and an expansion is present at the ostial lobe. The junction of the ostial and caudal crista inferior is located more posteriorly with respect to the same junction in the crista superior. The cauda is straight for about two-thirds of its length and markedly bent ventrally at the posterior end. The dorsal area is narrower than the ventral one. There is a shallow dorsal depression located just above the crest-like caudal crista superior.</paragraph>
</subSubSection>
<caption id="DF41FEABFFFCFFB89BBB2F63FB12F8B7" ID-DOI="http://doi.org/10.5281/zenodo.11526116" ID-Zenodo-Dep="11526116" httpUri="https://zenodo.org/record/11526116/files/figure.png" pageId="24" pageNumber="227" startId="24.[189,232,1569,1595]" targetBox="[189,1398,264,1522]" targetPageId="24">
<paragraph id="8B81AE23FFFCFFB89BBB2F63FB12F8B7" blockId="24.[187,1399,1569,1877]" pageId="24" pageNumber="227">
<emphasis id="B94A7231FFFCFFB89BBB2F63FEB4F9D9" bold="true" box="[189,281,1569,1595]" pageId="24" pageNumber="227">Fig. 10.</emphasis>
Fish otoliths from the US lower Eocene formations. 
<emphasis id="B94A7231FFFCFFB8987E2F63FC20F9D9" bold="true" box="[888,909,1569,1595]" pageId="24" pageNumber="227">A</emphasis>
. 
<taxonomicName id="4C3ED5A0FFFCFFB8989A2F60FAA6F9D9" authority="Stinton, 1978" authorityName="Stinton" authorityYear="1978" box="[924,1291,1569,1595]" family="Lactariidae" genus="Lactarius" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="species" species="amplus">
<emphasis id="B94A7231FFFCFFB8989A2F60FBCAF9D9" box="[924,1127,1569,1595]" italics="true" pageId="24" pageNumber="227">Lactarius amplus</emphasis>
<bibRefCitation id="EFAFD3D2FFFCFFB89F6A2F63FAA6F9D9" author="Stinton F. C." box="[1132,1291,1569,1595]" pageId="24" pageNumber="227" pagination="127 - 189" refId="ref14706" refString="Stinton F. C. 1978. Fish otoliths from the English Eocene (part 3). Palaeontographical Society Monographs 555: 127 - 189. https: // doi. org / 10.1080 / 25761900.2022.12131741" type="journal article" year="1978">Stinton, 1978</bibRefCitation>
</taxonomicName>
, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10757). 
<emphasis id="B94A7231FFFCFFB898EA2F06FB8EF9BC" bold="true" box="[1004,1059,1604,1630]" pageId="24" pageNumber="227">B–C</emphasis>
. 
<taxonomicName id="4C3ED5A0FFFCFFB89F332F07FEC3F960" authority="(Dante &amp; Frizzell, 1965)" baseAuthorityName="Dante &amp; Frizzell" baseAuthorityYear="1965" family="Lactariidae" genus="Lactarius" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="species" species="kokeni">
<emphasis id="B94A7231FFFCFFB89F332F07FB51F9BC" box="[1077,1276,1604,1630]" italics="true" pageId="24" pageNumber="227">Lactarius kokeni</emphasis>
(Dante &amp; Frizzell, 1965)
</taxonomicName>
, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10758, P 10759). 
<emphasis id="B94A7231FFFCFFB89BB92FC9FF7FF947" bold="true" box="[191,210,1675,1701]" pageId="24" pageNumber="227">D</emphasis>
. 
<taxonomicName id="4C3ED5A0FFFCFFB89BD92FC9FE6EF946" authorityName="Lin &amp; Nolf" authorityYear="2022" box="[223,451,1674,1701]" family="Haemulidae" genus="Anisotremus" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="species" species="rambo">
<emphasis id="B94A7231FFFCFFB89BD92FC9FE6EF946" box="[223,451,1674,1701]" italics="true" pageId="24" pageNumber="227">Anisotremus rambo</emphasis>
</taxonomicName>
, 2022, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10763). 
<emphasis id="B94A7231FFFCFFB89BBD2FECFF42F92A" bold="true" box="[187,239,1710,1736]" pageId="24" pageNumber="227">E–F</emphasis>
. 
<taxonomicName id="4C3ED5A0FFFCFFB89A002FECFCC8F92B" authority="(Dante &amp; Frizzell, 1965)" baseAuthorityName="Dante &amp; Frizzell" baseAuthorityYear="1965" box="[262,869,1710,1737]" family="Haemulidae" genus="Orthopristis" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="species" species="burlesonis">
<emphasis id="B94A7231FFFCFFB89A002FECFDB3F92A" box="[262,542,1710,1736]" italics="true" pageId="24" pageNumber="227">Orthopristis burlesonis</emphasis>
(Dante &amp; Frizzell, 1965)
</taxonomicName>
, Pamunkey River, Hanovertown, Potapaco Member, Virginia (IRSNB P 10764–10765). 
<emphasis id="B94A7231FFFCFFB899D62F93FCA6F909" bold="true" box="[720,779,1745,1771]" pageId="24" pageNumber="227">G–H</emphasis>
. “ 
<taxonomicName id="4C3ED5A0FFFCFFB8982E2F93FC0FF909" authorityName="G.Cuvier" authorityYear="1829" box="[808,930,1745,1771]" family="Haemulidae" genus="Haemulon" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="B94A7231FFFCFFB8982E2F93FC0FF909" box="[808,930,1745,1771]" italics="true" pageId="24" pageNumber="227">Haemulon</emphasis>
</taxonomicName>
” 
<emphasis id="B94A7231FFFCFFB898BE2F90FB94F909" box="[952,1081,1746,1771]" italics="true" pageId="24" pageNumber="227">ypresiensis</emphasis>
sp. nov., Pamunkey River, Hanovertown, Potapaco Member, Virginia. 
<emphasis id="B94A7231FFFCFFB899C92FB6FD4BF8EC" bold="true" box="[719,742,1780,1806]" pageId="24" pageNumber="227">G</emphasis>
. Holotype (IRSNB P 10766). 
<emphasis id="B94A7231FFFCFFB89F592FB6FBDBF8EC" bold="true" box="[1119,1142,1780,1806]" pageId="24" pageNumber="227">H</emphasis>
. Paratype (IRSNB P 10767). 
<emphasis id="B94A7231FFFCFFB89A252E55FEE1F8D3" bold="true" box="[291,332,1815,1841]" pageId="24" pageNumber="227">I–J</emphasis>
. 
<taxonomicName id="4C3ED5A0FFFCFFB89A582E55FCA9F8D3" authority="(Koken, 1888)" baseAuthorityName="Koken" baseAuthorityYear="1888" box="[350,772,1815,1841]" class="Actinopterygii" family="Sciaenidae" genus="Ekokenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="24" pageNumber="227" phylum="Chordata" rank="species" species="eporrecta">
<emphasis id="B94A7231FFFCFFB89A582E55FDEAF8D3" box="[350,583,1815,1841]" italics="true" pageId="24" pageNumber="227">Ekokenia eporrecta</emphasis>
(
<bibRefCitation id="EFAFD3D2FFFCFFB8995C2E55FD56F8D3" author="Koken E." box="[602,763,1815,1841]" pageId="24" pageNumber="227" pagination="274 - 305" refId="ref13473" refString="Koken E. 1888. Neue Untersuchungen an tertiaren Fisch-Otolithen. Zeitschrift der Deutschen Geologischen Gesellschaft 40: 274 - 305." type="journal article" year="1888">Koken, 1888</bibRefCitation>
)
</taxonomicName>
, Taylor’s Branch of Two Mile Creek, Reklaw Fm., Texas (IRSNB P 10768, P 10769). 1 = ventral view; 2 = inner view. Scale bars = 1 mm.
</paragraph>
</caption>
<subSubSection id="C324FDA8FFFDFFB99BBB284BFD03FCA5" pageId="25" pageNumber="228" type="discussion">
<paragraph id="8B81AE23FFFDFFB99BBB284BFE82FEC1" blockId="25.[189,303,265,291]" box="[189,303,265,291]" pageId="25" pageNumber="228">
<heading id="D0C9194FFFFDFFB99BBB284BFE82FEC1" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="25" pageNumber="228" reason="3">
<emphasis id="B94A7231FFFDFFB99BBB284BFE82FEC1" bold="true" box="[189,303,265,291]" pageId="25" pageNumber="228">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFFDFFB99BBB2875FCBCFD8F" blockId="25.[188,1399,311,622]" pageId="25" pageNumber="228">
The otoliths of “ 
<emphasis id="B94A7231FFFDFFB99A8F287AFE0BFEB3" box="[393,422,312,337]" italics="true" pageId="25" pageNumber="228">H.</emphasis>
” 
<emphasis id="B94A7231FFFDFFB99ABB287AFD93FEB3" box="[445,574,312,337]" italics="true" pageId="25" pageNumber="228">ypresiensis</emphasis>
sp. nov. are most similar to those of “ 
<taxonomicName id="4C3ED5A0FFFDFFB99F1A2875FB3BFEB3" authorityName="G.Cuvier" authorityYear="1829" box="[1052,1174,311,337]" family="Haemulidae" genus="Haemulon" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="B94A7231FFFDFFB99F1A2875FB3BFEB3" box="[1052,1174,311,337]" italics="true" pageId="25" pageNumber="228">Haemulon</emphasis>
</taxonomicName>
” 
<emphasis id="B94A7231FFFDFFB99FA82875FAB3FEB3" box="[1198,1310,311,337]" italics="true" pageId="25" pageNumber="228">pulchrum</emphasis>
(Frost, 1934) and “ 
<taxonomicName id="4C3ED5A0FFFDFFB99A4E2819FE6FFE97" authorityName="G.Cuvier" authorityYear="1829" box="[328,450,347,373]" family="Haemulidae" genus="Haemulon" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="B94A7231FFFDFFB99A4E2819FE6FFE97" box="[328,450,347,373]" italics="true" pageId="25" pageNumber="228">Haemulon</emphasis>
</taxonomicName>
” 
<emphasis id="B94A7231FFFDFFB99AD12819FDEBFE97" box="[471,582,347,373]" italics="true" pageId="25" pageNumber="228">obliquum</emphasis>
(
<bibRefCitation id="EFAFD3D2FFFDFFB999532819FD5EFE97" author="Muller A." box="[597,755,347,373]" pageId="25" pageNumber="228" pagination="1 - 360" refId="ref13839" refString="Muller A. 1999. Ichthyofaunen aus dem atlantischen Tertiar der USA. Leipziger Geowissenschaften 9 / 10: 1 - 360." type="journal article" year="1999">Müller, 1999</bibRefCitation>
), based on their sulcus and outline shape. “ 
<taxonomicName id="4C3ED5A0FFFDFFB99FE92819FAC4FE97" authorityName="G.Cuvier" authorityYear="1829" box="[1263,1385,347,373]" family="Haemulidae" genus="Haemulon" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="228" phylum="Chordata" rank="genus">
<emphasis id="B94A7231FFFDFFB99FE92819FAC4FE97" box="[1263,1385,347,373]" italics="true" pageId="25" pageNumber="228">Haemulon</emphasis>
</taxonomicName>
” 
<emphasis id="B94A7231FFFDFFB99BBB283CFE80FE7A" box="[189,301,382,408]" italics="true" pageId="25" pageNumber="228">pulchrum</emphasis>
is a widely distributed Eocene European species, and the otoliths of the new species resemble those from the Lutetian of Osteroden, 
<collectingCountry id="F329EEB3FFFDFFB9999E28E0FCA8FE5E" box="[664,773,418,444]" name="Germany" pageId="25" pageNumber="228">Germany</collectingCountry>
(see 
<bibRefCitation id="EFAFD3D2FFFDFFB9984A28E0FB9FFE5E" author="Schwarzhans W." box="[844,1074,418,444]" pageId="25" pageNumber="228" pagination="299 - 369" refId="ref14285" refString="Schwarzhans W. 2007. The otoliths from the middle Eocene of Osteroden near Bramsche, north-western Germany. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlungen 244: 299 - 369. https: // doi. org / 10.1127 / 0077 - 7749 / 2007 / 0244 - 0299" type="journal article" year="2007">Schwarzhans 2007</bibRefCitation>
: fig. 32; 
<bibRefCitation id="EFAFD3D2FFFDFFB99FA328E0FADEFE5E" author="Lin C. - H. &amp; Nolf D. &amp; Steurbaut E. &amp; Girone A." box="[1189,1395,418,444]" pageId="25" pageNumber="228" pagination="879 - 907" refId="ref13634" refString="Lin C. - H., Nolf D., Steurbaut E. &amp; Girone A. 2017 b. Fish otoliths from the Lutetian of the Aquitaine Basin (SW France), a breakthrough in the knowledge of the European Eocene ichthyofauna. Journal of Systematic Paleontology 15 (11): 879 - 907. https: // doi. org / 10.1080 / 14772019.2016.1246112" type="journal article" year="2017">
Lin 
<emphasis id="B94A7231FFFDFFB99FDC28E1FABFFE5E" box="[1242,1298,418,444]" italics="true" pageId="25" pageNumber="228">et al</emphasis>
. 2017b
</bibRefCitation>
: fig. 12h–l). However, the dorsal rim of the new species is flat, whereas in “ 
<emphasis id="B94A7231FFFDFFB99F3E2884FBF8FE3D" box="[1080,1109,454,479]" italics="true" pageId="25" pageNumber="228">H.</emphasis>
” 
<emphasis id="B94A7231FFFDFFB99F6C2887FB77FE3D" box="[1130,1242,453,479]" italics="true" pageId="25" pageNumber="228">pulchrum</emphasis>
, it is notably elevated and forms the highest part of the otoliths. On the other hand, “ 
<emphasis id="B94A7231FFFDFFB99F0B28A8FB87FDE1" box="[1037,1066,490,515]" italics="true" pageId="25" pageNumber="228">H.</emphasis>
” 
<emphasis id="B94A7231FFFDFFB99F3928ABFB03FDE1" box="[1087,1198,489,515]" italics="true" pageId="25" pageNumber="228">obliquum</emphasis>
, a middle to late Eocene American species (see 
<bibRefCitation id="EFAFD3D2FFFDFFB9992C2B4EFD5BFDC5" author="Lin C. - H. &amp; Nolf D." box="[554,758,524,551]" pageId="25" pageNumber="228" pagination="1 - 122" refId="ref13528" refString="Lin C. - H. &amp; Nolf D. 2022. Middle and late Eocene fish otoliths from the eastern and southern USA. European Journal of Taxonomy 814 (1): 1 - 122. https: // doi. org / 10.5852 / ejt. 2022.814.1745" type="journal article" year="2022">Lin &amp; Nolf 2022</bibRefCitation>
), has otoliths that are more similar to the new species, with the largest difference being that the cauda tip extends further backwards and is more markedly bent, reaching almost to the posterior rim of the otoliths.
</paragraph>
<paragraph id="8B81AE23FFFDFFB99BBB2BD7FD64FD4D" blockId="25.[189,713,661,687]" box="[189,713,661,687]" pageId="25" pageNumber="228">
<heading id="D0C9194FFFFDFFB99BBB2BD7FD64FD4D" bold="true" box="[189,713,661,687]" fontSize="11" level="3" pageId="25" pageNumber="228" reason="3">
<emphasis id="B94A7231FFFDFFB99BBB2BD7FD64FD4D" bold="true" box="[189,713,661,687]" pageId="25" pageNumber="228">Stratigraphic and geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B81AE23FFFDFFB99BBB2B86FDEDFCE0" blockId="25.[189,1399,708,770]" pageId="25" pageNumber="228">
Ypresian: Bashi Formation, 
<collectingRegion id="49FA60C1FFFDFFB999132B86FD36FD3C" box="[533,667,708,734]" country="United States of America" name="Mississippi" pageId="25" pageNumber="228">Mississippi</collectingRegion>
and 
<collectingRegion id="49FA60C1FFFDFFB999DA2B86FCE7FD3C" box="[732,842,708,734]" country="United States of America" name="Alabama" pageId="25" pageNumber="228">Alabama</collectingRegion>
; Hatchetigbee Formation, 
<collectingRegion id="49FA60C1FFFDFFB99F8B2B86FB56FD3C" box="[1165,1275,708,734]" country="United States of America" name="Alabama" pageId="25" pageNumber="228">Alabama</collectingRegion>
; Potapaco Member, 
<collectingRegion id="49FA60C1FFFDFFB99A2B2BAAFE21FCE0" box="[301,396,744,770]" country="United States of America" name="Virginia" pageId="25" pageNumber="228">Virginia</collectingRegion>
and 
<collectingRegion id="49FA60C1FFFDFFB99AC02BAAFD91FCE0" box="[454,572,744,770]" country="United States of America" name="Maryland" pageId="25" pageNumber="228">Maryland</collectingRegion>
.
</paragraph>
<paragraph id="8B81AE23FFFDFFB99BBB2A68FD03FCA5" blockId="25.[189,686,810,839]" box="[189,686,810,839]" pageId="25" pageNumber="228">
<emphasis id="B94A7231FFFDFFB99BBB2A68FD03FCA5" bold="true" box="[189,686,810,839]" italics="true" pageId="25" pageNumber="228">Remarks on taxa requiring comments</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>