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<document id="33F3E7A4C7A552525875D7EDE28833D0" ID-DOI="10.11646/zootaxa.5374.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146516" ID-ZooBank="E9F72C3C-A005-415D-8D36-2C0EA9036C26" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1700187360043" checkinUser="plazi" docAuthor="Vilela, Diogo Silva, Jacques, Gabriel De Castro &amp; Souza, Marcos Magalhães De" docDate="2023" docId="03812674FFA2FF9C7CC0D017FDE5F868" docLanguage="en" docName="zootaxa.5374.2.6.pdf" docOrigin="Zootaxa 5374 (2)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.2.6/52289" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Minagrion veredae Vilela &amp; Jacques &amp; Souza 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="259" masterDocId="FFB85E0CFFA3FF987C57D73CFFDDFF92" masterDocTitle="A new species of Minagrion Santos, 1965 (Odonata: Coenagrionidae) from the Cerrado of Northern Minas Gerais state, Brazil" masterLastPageNumber="262" masterPageNumber="255" pageNumber="256" updateTime="1700760234968" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods id="22FDCCE240B37EB6DAE0BDA6F1CEDFFD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="887298FA5C9993EBE2432052BCB267EB">
<mods:title id="FF3A04A882CDF15E500B305D41FA57AD">A new species of Minagrion Santos, 1965 (Odonata: Coenagrionidae) from the Cerrado of Northern Minas Gerais state, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="1118318FDE7A717224F8916A1CE704B0" type="personal">
<mods:role id="83EB9233C9F0F639277058BF7F61FF66">
<mods:roleTerm id="67CE3993815BE1170D128F8D7A32E9ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B25BCE19B3752087226589A30865AA0">Vilela, Diogo Silva</mods:namePart>
<mods:nameIdentifier id="5EC4EDF84C5221A3E2BD792C4F1DABAF" type="ORCID">0000-0001-6510-7018</mods:nameIdentifier>
<mods:affiliation id="E1AE1992AC7226FE1A9E6D3302C4DB1E">Instituto Federal de Educação, Ciência e Tecnologia do Sul de Minas – Campus Inconfidentes, Inconfidentes, Minas Gerais, Brazil</mods:affiliation>
<mods:nameIdentifier id="612A206D07CD81146A5E5F1BB520E8D5" type="email">diogo.vilela@ifsuldeminas.edu.br</mods:nameIdentifier>
</mods:name>
<mods:name id="9720B6C4C8FC4952E13453C909D37295" type="personal">
<mods:role id="B60051E4AE1C4B63746D9B97F17EA805">
<mods:roleTerm id="1EA4CFB9CB371C72DE9421672B41132E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="51F6B0945332326959A58119389A1C06">Jacques, Gabriel De Castro</mods:namePart>
<mods:nameIdentifier id="DD3CCF9A3FCE69E4A8F3E6F59AE39FF7" type="ORCID">0000-0002-9619-6065</mods:nameIdentifier>
<mods:affiliation id="2562E284B894BFF0208699E52CBD707A">Instituto Federal de Educação, Ciência e Tecnologia de Minas Gerais – Campus Bambuí, Bambuí, Minas Gerais, Brazil</mods:affiliation>
<mods:nameIdentifier id="82A0ADAB8799E2BE791EBA868D300E2D" type="email">gabriel.jacques@ifmg.edu.br</mods:nameIdentifier>
</mods:name>
<mods:name id="134AC5970282A2DF492A3164DD508003" type="personal">
<mods:role id="09B8EB718D403CC8EC54FEF9849DB872">
<mods:roleTerm id="FE838E1F4351E8907C73203B4B8E94C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD9CBDEC63941C47400402CEDEE124E2">Souza, Marcos Magalhães De</mods:namePart>
<mods:nameIdentifier id="99E0ECF4D1F773479B87142CB9A8944D" type="ORCID">0000-0003-0415-1714</mods:nameIdentifier>
<mods:nameIdentifier id="50392ACB618E3B62244F4F4E8736BF10" type="email">marcos.souza@ifsuldeminas.edu.br</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="E87A2609A0CF3ECEECA6CCB7495837E7">text</mods:typeOfResource>
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<mods:title id="8606E1578565F277D3BCC1DECA97F55F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B755B993C3780DBE39B10789FDD8639F">
<mods:date id="94AF1D02EE3EC1E7E70D4033ED1C9840">2023</mods:date>
<mods:detail id="36B16224F8A44BD16D9519F9D3837D1A" type="pubDate">
<mods:number id="5143CD9B7CB2CE30303337D6A3CB2591">2023-11-16</mods:number>
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<mods:detail id="889551FDAFABFC3FBA634BFB65D1B4B9" type="volume">
<mods:number id="B81A9513EEB86190FFD3EA1E3A568816">5374</mods:number>
</mods:detail>
<mods:detail id="264B17B4FE61F48330A7501C39F26A94" type="issue">
<mods:number id="7D2711AC56690F1A3FE11B795767A5A0">2</mods:number>
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<mods:start id="499FA34EF1F6E59C91F2F074BD86E041">255</mods:start>
<mods:end id="F0D908FC9F6004C7FB8B731C9535650B">262</mods:end>
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<mods:url id="A559A6C9174D5AF288B40BFF7B3E9309">https://mapress.com/zt/article/download/zootaxa.5374.2.6/52289</mods:url>
</mods:location>
<mods:classification id="39F49A988681FD41136108B1BF8A4DF2">journal article</mods:classification>
<mods:identifier id="A3DC1DE52829CFE809A3AE5339CB8640" type="DOI">10.11646/zootaxa.5374.2.6</mods:identifier>
<mods:identifier id="D9883672BDC5C5B48AA5A41D8763C9E3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="616240DA2DAFBE2BEC019D6AFF3F26B6" type="Zenodo-Dep">10146516</mods:identifier>
<mods:identifier id="FB54137F1A430C01DD2F385119B1E358" type="ZooBank">E9F72C3C-A005-415D-8D36-2C0EA9036C26</mods:identifier>
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<treatment id="03812674FFA2FF9C7CC0D017FDE5F868" LSID="urn:lsid:plazi:treatment:03812674FFA2FF9C7CC0D017FDE5F868" httpUri="http://treatment.plazi.org/id/03812674FFA2FF9C7CC0D017FDE5F868" lastPageId="4" lastPageNumber="259" pageId="1" pageNumber="256">
<subSubSection id="C332C4E9FFA2FF997CC0D017FD79F8D7" box="[151,676,1835,1862]" pageId="1" pageNumber="256" type="nomenclature">
<paragraph id="8B979762FFA2FF997CC0D017FD79F8D7" blockId="1.[151,676,1835,1898]" box="[151,676,1835,1862]" pageId="1" pageNumber="256">
<heading id="D0DF200EFFA2FF997CC0D017FD79F8D7" bold="true" box="[151,676,1835,1862]" fontSize="11" level="1" pageId="1" pageNumber="256" reason="1">
<emphasis id="B95C4B70FFA2FF997CC0D017FD79F8D7" bold="true" box="[151,676,1835,1862]" pageId="1" pageNumber="256">
<taxonomicName id="4C28ECE1FFA2FF997CC0D017FEA3F8D4" authority="Vilela &amp; Jacques &amp; Souza, 2023" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[151,382,1835,1862]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="veredae" status="sp. nov.">
<emphasis id="B95C4B70FFA2FF997CC0D017FEA3F8D4" bold="true" box="[151,382,1835,1862]" italics="true" pageId="1" pageNumber="256">Minagrion veredae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26FF60BFFA2FF997DD2D017FE3EF8D7" box="[389,483,1835,1861]" pageId="1" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
Vilela &amp; Souza
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C332C4E9FFA2FF997CC0D073FECFF8F8" box="[151,274,1871,1898]" pageId="1" pageNumber="256" type="description">
<paragraph id="8B979762FFA2FF997CC0D073FECFF8F8" blockId="1.[151,676,1835,1898]" box="[151,274,1871,1898]" pageId="1" pageNumber="256">
(
<figureCitation id="13138BE7FFA2FF997CC8D073FED4F8F8" box="[159,265,1871,1898]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="1.[151,250,1657,1681]" captionStartId-1="2.[151,250,1949,1973]" captionStartId-2="3.[151,250,1192,1216]" captionStartId-3="4.[151,250,1729,1753]" captionStartId-4="5.[151,250,1943,1967]" captionTargetBox-0="[151,1421,667,1620]" captionTargetBox-1="[157,1436,884,1906]" captionTargetBox-2="[175,1413,192,1157]" captionTargetBox-3="[161,1429,353,1698]" captionTargetBox-4="[162,1429,189,1907]" captionTargetId-0="figure-201@1.[151,1435,667,1631]" captionTargetId-1="figure-315@2.[151,1436,884,1925]" captionTargetId-2="figure-17@3.[164,1423,181,1168]" captionTargetId-3="figure-99@4.[151,1436,345,1706]" captionTargetId-4="figure-17@5.[151,1436,181,1919]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." captionText-1="FIGURE 2. Male S1 tubercle in lateral view: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." captionText-2="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." captionText-3="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." captionText-4="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi-0="http://doi.org/10.5281/zenodo.10146518" figureDoi-1="http://doi.org/10.5281/zenodo.10146520" figureDoi-2="http://doi.org/10.5281/zenodo.10146522" figureDoi-3="http://doi.org/10.5281/zenodo.10146524" figureDoi-4="http://doi.org/10.5281/zenodo.10146526" httpUri-0="https://zenodo.org/record/10146518/files/figure.png" httpUri-1="https://zenodo.org/record/10146520/files/figure.png" httpUri-2="https://zenodo.org/record/10146522/files/figure.png" httpUri-3="https://zenodo.org/record/10146524/files/figure.png" httpUri-4="https://zenodo.org/record/10146526/files/figure.png" pageId="1" pageNumber="256">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C332C4E9FFA2FF9A7CC0D0ABFCCCFF46" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256" type="materials_examined">
<paragraph id="8B979762FFA2FF997CC0D0ABFC8EF847" blockId="1.[151,1436,1943,2044]" pageId="1" pageNumber="256">
<materialsCitation id="3B409D3FFFA2FF997CC0D0ABFC8EF847" collectingDate="2023-04-15" collectionCode="IFSULDEMINAS" collectorName="P. S. Vital" country="Brazil" elevation="780" latitude="-15.1" location="Grande Sertao Veredas National Park" longLatPrecision="1286" longitude="-45.8" pageId="1" pageNumber="256" specimenCode="Z329" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais State" typeStatus="holotype">
<emphasis id="B95C4B70FFA2FF997CC0D0ABFED6F823" bold="true" box="[151,267,1943,1969]" pageId="1" pageNumber="256">
<typeStatus id="549329C0FFA2FF997CC0D0ABFEDBF823" box="[151,262,1943,1969]" pageId="1" pageNumber="256" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D2E5CEBFFA2FF997D42D0A5FEF7F826" box="[277,298,1945,1972]" pageId="1" pageNumber="256" type="male">♂</specimenCount>
,
<collectingCountry id="F33FD7F2FFA2FF997D6DD0ABFE43F823" box="[314,414,1943,1969]" name="Brazil" pageId="1" pageNumber="256">BRAZIL</collectingCountry>
(
<specimenCode id="DB8E3F19FFA2FF997DE7D0ABFE2DF823" box="[432,496,1943,1969]" pageId="1" pageNumber="256">Z329</specimenCode>
),
<collectingRegion id="49EC5980FFA2FF997E50D0ABFD39F823" box="[519,740,1943,1969]" country="Brazil" name="Minas Gerais" pageId="1" pageNumber="256">Minas Gerais State</collectingRegion>
,
<location id="8EF7C1B9FFA2FF997EA5D0ABFB7CF823" LSID="urn:lsid:plazi:treatment:03812674FFA2FF9C7CC0D017FDE5F868:8EF7C1B9FFA2FF997EA5D0ABFB7CF823" box="[754,1185,1943,1969]" country="Brazil" latitude="-15.1" longLatPrecision="1286" longitude="-45.8" name="Grande Sertao Veredas National Park" pageId="1" pageNumber="256" stateProvince="Minas Gerais State">Grande Sertão Veredas National Park</location>
,
<date id="FF96B1A2FFA2FF9978F8D0ABFAFAF820" box="[1199,1319,1943,1970]" pageId="1" pageNumber="256" value="2023-04-15">
<collectingDate id="EFD2484AFFA2FF9978F8D0ABFAFAF820" box="[1199,1319,1943,1970]" pageId="1" pageNumber="256" value="2023-04-15">15.iv.2023</collectingDate>
</date>
, (
<geoCoordinate id="EE1CF1A5FFA2FF997968D0ABFA4AF823" box="[1343,1431,1943,1970]" degrees="15" direction="south" minutes="6" orientation="latitude" pageId="1" pageNumber="256" precision="925" value="-15.1">15°6’ S</geoCoordinate>
,
<geoCoordinate id="EE1CF1A5FFA2FF997CC0D087FEDEF844" box="[151,259,1979,2006]" degrees="45" direction="west" minutes="48" orientation="longitude" pageId="1" pageNumber="256" precision="925" value="-45.8">45°48’ W</geoCoordinate>
,
<elevation id="00057051FFA2FF997D59D087FE18F847" box="[270,453,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="9.0" metricValueMin="6.6" pageId="1" pageNumber="256" unit="m" value="780.0" valueMax="900.0" valueMin="660.0">
<quantity id="4CD03A87FFA2FF997D59D087FE41F847" box="[270,412,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="7.8" metricValueMax="9.0" metricValueMin="6.6" pageId="1" pageNumber="256" unit="m" value="780.0" valueMax="900.0" valueMin="660.0">660 to 900m</quantity>
asl
</elevation>
),
<collectorName id="26DDF2B4FFA2FF997D8ED087FD9DF847" box="[473,576,1979,2005]" pageId="1" pageNumber="256">P.S. Vital</collectorName>
leg.,
<collectionCode id="ED390FA7FFA2FF997E29D087FC92F847" box="[638,847,1979,2005]" pageId="1" pageNumber="256">IFSULDEMINAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B979762FFA2FF9A7C90D0E3FCCCFF46" blockId="1.[151,1436,1943,2044]" lastBlockId="2.[151,1437,150,861]" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256">
<materialsCitation id="3B409D3FFFA2FF997C90D0E3FA89F86B" box="[199,1364,2015,2044]" collectingDate="2023-04-15" collectionCode="IFSULDEMINAS" collectorName="E. D. F. Ferreira" country="Brazil" elevation="780" latitude="-15.1" location="Grande Sertao Veredas National Park" longLatPrecision="1286" longitude="-45.8" pageId="1" pageNumber="256" specimenCode="Z326" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<emphasis id="B95C4B70FFA2FF997C90D0E3FE9BF86B" bold="true" box="[199,326,2015,2041]" pageId="1" pageNumber="256">
<typeStatus id="549329C0FFA2FF997C90D0E3FE9CF86B" box="[199,321,2015,2041]" pageId="1" pageNumber="256" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Same data as holotype, except:
<specimenCount id="9D2E5CEBFFA2FF997EB2D0DDFCD5F86E" box="[741,776,2017,2044]" count="1" pageId="1" pageNumber="256" type="male">1♂</specimenCount>
(
<specimenCode id="DB8E3F19FFA2FF997F76D0E3FCBDF86B" box="[801,864,2015,2041]" pageId="1" pageNumber="256">Z326</specimenCode>
),
<collectorName id="26DDF2B4FFA2FF997F29D0DCFBF2F86B" box="[894,1071,2015,2042]" pageId="1" pageNumber="256">E.D.F. Ferreira</collectorName>
leg.,
<collectionCode id="ED390FA7FFA2FF997828D0E3FA8DF86B" box="[1151,1360,2015,2041]" pageId="1" pageNumber="256">IFSULDEMINAS</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B409D3FFFA2FF9A7933D0DDFC83FF22" collectingDate="2022-11-05" collectionCode="IFSULDEMINAS" collectorName="L. Millani" country="Brazil" elevation="780" lastPageId="2" lastPageNumber="257" latitude="-15.1" location="Grande Sertao Veredas National Park" longLatPrecision="1286" longitude="-45.8" pageId="1" pageNumber="256" specimenCode="Z112, Z140" specimenCount="2" specimenCount-male="2" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D2E5CEBFFA2FF997933D0DDFA41F86E" box="[1380,1436,2017,2044]" count="2" pageId="1" pageNumber="256" type="male">2♂♂</specimenCount>
(
<specimenCode id="DB8E3F19FFA1FF9A7CC8D7AAFF00FF22" box="[159,221,150,176]" pageId="2" pageNumber="257">Z112</specimenCode>
,
<specimenCode id="DB8E3F19FFA1FF9A7CBAD7AAFEF1FF22" box="[237,300,150,176]" pageId="2" pageNumber="257">Z140</specimenCode>
),
<date id="FF96B1A2FFA1FF9A7D14D7AAFE63FF23" box="[323,446,150,177]" pageId="2" pageNumber="257" value="2022-11-05">
<collectingDate id="EFD2484AFFA1FF9A7D14D7AAFE63FF23" box="[323,446,150,177]" pageId="2" pageNumber="257" value="2022-11-05">05.xi.2022</collectingDate>
</date>
,
<collectorName id="26DDF2B4FFA1FF9A7D9AD7ABFD9FFF22" box="[461,578,150,177]" pageId="2" pageNumber="257">L. Millani</collectorName>
leg.,
<collectionCode id="ED390FA7FFA1FF9A7EDED7AAFC87FF22" box="[649,858,150,176]" pageId="2" pageNumber="257">IFSULDEMINAS</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B409D3FFFA1FF9A7F3DD7A4FEB1FF46" collectingDate="2023-01-28" collectionCode="IFSULDEMINAS, FAAL" collectorName="T. P. Gouvea &amp; M. M. Souza" country="Brazil" elevation="780" latitude="-15.1" location="Grande Sertao Veredas National Park" longLatPrecision="1286" longitude="-45.8" pageId="2" pageNumber="257" specimenCode="Z195, Z198, Z313" specimenCount="3" specimenCount-male="3" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D2E5CEBFFA1FF9A7F3DD7A4FC7FFF21" box="[874,930,152,179]" count="2" pageId="2" pageNumber="257" type="male">2♂♂</specimenCount>
(
<specimenCode id="DB8E3F19FFA1FF9A7FE2D7AAFC2EFF23" box="[949,1011,150,177]" pageId="2" pageNumber="257">Z195</specimenCode>
,
<specimenCode id="DB8E3F19FFA1FF9A7854D7AAFB9CFF22" box="[1027,1089,150,176]" pageId="2" pageNumber="257">Z198</specimenCode>
),
<date id="FF96B1A2FFA1FF9A780DD7AAFB1BFF22" box="[1114,1222,150,176]" pageId="2" pageNumber="257" value="2023-01-28">
<collectingDate id="EFD2484AFFA1FF9A780DD7AAFB1BFF22" box="[1114,1222,150,176]" pageId="2" pageNumber="257" value="2023-01-28">28.i.2023</collectingDate>
</date>
,
<collectorName id="26DDF2B4FFA1FF9A7882D7ABFABCFF22" box="[1237,1377,150,177]" pageId="2" pageNumber="257">T.P. Gouvêa</collectorName>
leg.,
<collectionCode id="ED390FA7FFA1FF9A7CC0D786FEB5FF46" box="[151,360,186,212]" pageId="2" pageNumber="257">IFSULDEMINAS</collectionCode>
;
</materialsCitation>
<materialsCitation id="3B409D3FFFA1FF9A7D24D780FCCCFF46" box="[371,785,186,215]" collectingDate="2023-01-28" collectionCode="IFSULDEMINAS, FAAL" collectorName="T. P. Gouvea &amp; M. M. Souza" country="Brazil" elevation="780" latitude="-15.1" location="Grande Sertao Veredas National Park" longLatPrecision="1286" longitude="-45.8" pageId="2" pageNumber="257" specimenCode="Z195, Z198, Z313" specimenCount="3" specimenCount-male="3" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D2E5CEBFFA1FF9A7D24D780FE4BFF45" box="[371,406,188,215]" count="1" pageId="2" pageNumber="257" type="male">1♂</specimenCount>
(
<specimenCode id="DB8E3F19FFA1FF9A7DF1D786FE3BFF46" box="[422,486,186,212]" pageId="2" pageNumber="257">Z313</specimenCode>
),
<collectorName id="26DDF2B4FFA1FF9A7DAED787FD59FF46" box="[505,644,186,213]" pageId="2" pageNumber="257">M.M. Souza</collectorName>
leg.,
<collectionCode id="ED390FA7FFA1FF9A7E95D786FCD0FF46" box="[706,781,186,212]" pageId="2" pageNumber="257">FAAL</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C332C4E9FFA1FF9A7C90D7E2FA46FEF6" pageId="2" pageNumber="257" type="etymology">
<paragraph id="8B979762FFA1FF9A7C90D7E2FA46FEF6" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7C90D7E2FE93FF6A" bold="true" box="[199,334,222,248]" pageId="2" pageNumber="257">Etymology.</emphasis>
Named
<taxonomicName id="4C28ECE1FFA1FF9A7DFBD7E2FDD9FF6A" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[428,516,222,248]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="veredae">
<emphasis id="B95C4B70FFA1FF9A7DFBD7E2FDD9FF6A" box="[428,516,222,248]" italics="true" pageId="2" pageNumber="257">veredae</emphasis>
</taxonomicName>
(name in genitive case) in honor to the palm swamp environments, called
<emphasis id="B95C4B70FFA1FF9A7911D7E2FA41FF6A" box="[1350,1436,222,248]" italics="true" pageId="2" pageNumber="257">veredas</emphasis>
in
<collectingCountry id="F33FD7F2FFA1FF9A7CE3D63EFF26FE8E" box="[180,251,258,284]" name="Brazil" pageId="2" pageNumber="257">Brazil</collectingCountry>
, which harbors vast biodiversity and is currently being threatened by human activities. This name is also a reference to the
<typeStatus id="549329C0FFA1FF9A7D1ED61BFEA4FED3" box="[329,377,295,321]" pageId="2" pageNumber="257">type</typeStatus>
locality in
<collectingRegion id="49EC5980FFA1FF9A7DA3D61AFD18FED3" box="[500,709,294,321]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="257">Minas Gerais state</collectingRegion>
, the Grande Sertão Veredas National Park, who is named after the
<emphasis id="B95C4B70FFA1FF9A7CC0D676FF30FEF6" box="[151,237,330,356]" italics="true" pageId="2" pageNumber="257">veredas</emphasis>
, but also inspired by the homonymous book, written by the “Mineiro” João Guimarães Rosa (1908–1967).
</paragraph>
</subSubSection>
<subSubSection id="C332C4E9FFA1FF9B7C90D652FC5EF923" lastPageId="3" lastPageNumber="258" pageId="2" pageNumber="257" type="description">
<paragraph id="8B979762FFA1FF9A7C90D652FC52FE42" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7C90D652FED9FE1A" box="[199,260,366,392]" italics="true" pageId="2" pageNumber="257">Head</emphasis>
(
<figureCitation id="13138BE7FFA1FF9A7D42D652FE8AFE1A" box="[277,343,366,392]" captionStart="FIGURE 1" captionStartId="1.[151,250,1657,1681]" captionTargetBox="[151,1421,667,1620]" captionTargetId="figure-201@1.[151,1435,667,1631]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146518" httpUri="https://zenodo.org/record/10146518/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
). Dorsally black; postocular spots light blue, elongated, merging with the occipital bar, of same color; frons angulated; antefrons, postfrons, labrum and mandibles greenish; clypeus and postclypeus black; ventral face of the head grey or bluish adjacent to the eye, remainder black.
</paragraph>
<paragraph id="8B979762FFA1FF9A7C90D6E6FA90FD16" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7C90D6E6FEC5FE66" box="[199,280,474,500]" italics="true" pageId="2" pageNumber="257">Thorax</emphasis>
(
<figureCitation id="13138BE7FFA1FF9A7D7ED6E6FEB1FE66" box="[297,364,474,500]" captionStart="FIGURE 1" captionStartId="1.[151,250,1657,1681]" captionTargetBox="[151,1421,667,1620]" captionTargetId="figure-201@1.[151,1435,667,1631]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146518" httpUri="https://zenodo.org/record/10146518/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
). Anterior lobe of prothorax blue anteriorly, remainder black; medial lobe black dorsally with circular blue lateral spots; posterior lobe entire, with a medial convexity, black with blue on the lateral margins. Pterothorax black dorsally with metallic copper reflections; lower half of mesepisternum with a blue stripe; mesepimeron black, with a posterior blue spot; mesinfraepisternum black with a small pale yellow/green area; anterior portion of metepisternum pale yellow, remainder blue; remainder of pterothorax pale yellow/green.
</paragraph>
<paragraph id="8B979762FFA1FF9A7C90D5B3FECFFD62" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7C90D5B3FF21FD3A" box="[199,252,655,680]" italics="true" pageId="2" pageNumber="257">Legs</emphasis>
(
<figureCitation id="13138BE7FFA1FF9A7D58D5B2FE89FD3A" box="[271,340,654,680]" captionStart="FIGURE 1" captionStartId="1.[151,250,1657,1681]" captionTargetBox="[151,1421,667,1620]" captionTargetId="figure-201@1.[151,1435,667,1631]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146518" httpUri="https://zenodo.org/record/10146518/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
). Pale yellow; femoral armature black, spines decreasing in size towards the apex; tibiae pale yellow, spines subequal in size; tarsal apex and claws with dark apex, the supplementary tooth on tarsal claw well developed.
</paragraph>
<paragraph id="8B979762FFA1FF9A7C90D5C7FE95FCCF" blockId="2.[151,1437,150,861]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7C90D5C7FED6FC86" box="[199,267,763,788]" italics="true" pageId="2" pageNumber="257">Wings</emphasis>
(
<figureCitation id="13138BE7FFA1FF9A7D4DD5C6FE87FC86" box="[282,346,762,788]" captionStart="FIGURE 1" captionStartId="1.[151,250,1657,1681]" captionTargetBox="[151,1421,667,1620]" captionTargetId="figure-201@1.[151,1435,667,1631]" captionTargetPageId="1" captionText="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146518" httpUri="https://zenodo.org/record/10146518/files/figure.png" pageId="2" pageNumber="257">Fig. 1</figureCitation>
). Hyaline, pterostigma dark brown, with pale contours, 1 cell long; 9 Px in HW, 10 Px in FW; CuA extending for 7 cells distal to vein descending from subnodus in FW,
<quantity id="4CD03A87FFA1FF9A7FF7D422FC16FCAA" box="[928,971,798,824]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="257" unit="in" value="8.0">8 in</quantity>
HW; MP reaching wing posterior margin in FW and HW.
</paragraph>
<caption id="DF57C7EAFFA1FF9A7CC0D0A1FAF3F827" ID-DOI="http://doi.org/10.5281/zenodo.10146520" ID-Zenodo-Dep="10146520" box="[151,1326,1949,1974]" httpUri="https://zenodo.org/record/10146520/files/figure.png" pageId="2" pageNumber="257" startId="2.[151,250,1949,1973]" targetBox="[157,1436,884,1906]" targetPageId="2" targetType="figure">
<paragraph id="8B979762FFA1FF9A7CC0D0A1FAF3F827" blockId="2.[151,1326,1949,1974]" box="[151,1326,1949,1974]" pageId="2" pageNumber="257">
<emphasis id="B95C4B70FFA1FF9A7CC0D0A1FECEF827" bold="true" box="[151,275,1949,1973]" pageId="2" pageNumber="257">FIGURE 2.</emphasis>
Male S1 tubercle in lateral view: (a)
<taxonomicName id="4C28ECE1FFA1FF9A7EDAD0A2FC92F827" authority="Vilela &amp; Jacques &amp; Souza, 2023" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[653,847,1949,1973]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="veredae" status="sp. nov.">
<emphasis id="B95C4B70FFA1FF9A7EDAD0A2FC92F827" box="[653,847,1949,1973]" italics="true" pageId="2" pageNumber="257">Minagrion veredae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26FF60BFFA1FF9A7F01D0A2FC78F827" box="[854,933,1950,1973]" pageId="2" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
HOLOTYPE; (b)
<taxonomicName id="4C28ECE1FFA1FF9A7836D0A2FAF5F827" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1121,1320,1949,1973]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="B95C4B70FFA1FF9A7836D0A2FAF5F827" box="[1121,1320,1949,1973]" italics="true" pageId="2" pageNumber="257">Minagrion waltheri</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF57C7EAFFA0FF9B7CC0D394FACDFB52" ID-DOI="http://doi.org/10.5281/zenodo.10146522" ID-Zenodo-Dep="10146522" box="[151,1296,1192,1217]" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="3" pageNumber="258" startId="3.[151,250,1192,1216]" targetBox="[175,1413,192,1157]" targetPageId="3" targetType="figure">
<paragraph id="8B979762FFA0FF9B7CC0D394FACDFB52" blockId="3.[151,1296,1192,1217]" box="[151,1296,1192,1217]" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7CC0D394FECEFB52" bold="true" box="[151,275,1192,1216]" pageId="3" pageNumber="258">FIGURE 3.</emphasis>
Lateral view of genital ligula: (a)
<taxonomicName id="4C28ECE1FFA0FF9B7E38D395FCECFB52" authority="Vilela &amp; Jacques &amp; Souza, 2023" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[623,817,1192,1216]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="veredae" status="sp. nov.">
<emphasis id="B95C4B70FFA0FF9B7E38D395FCECFB52" box="[623,817,1192,1216]" italics="true" pageId="3" pageNumber="258">Minagrion veredae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26FF60BFFA0FF9B7F60D395FC5BFB52" box="[823,902,1193,1216]" pageId="3" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
HOLOTYPE; (b)
<taxonomicName id="4C28ECE1FFA0FF9B7814D395FAD4FB52" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1091,1289,1192,1216]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="B95C4B70FFA0FF9B7814D395FAD4FB52" box="[1091,1289,1192,1216]" italics="true" pageId="3" pageNumber="258">Minagrion waltheri</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B979762FFA0FF9B7C90D237FB25FA03" blockId="3.[151,1437,1291,2037]" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7C90D237FEECFAB7" box="[199,305,1291,1317]" italics="true" pageId="3" pageNumber="258">Abdomen</emphasis>
(
<figureCitation id="13138BE7FFA0FF9B7D13D237FE6EFAB7" box="[324,435,1291,1317]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1657,1681]" captionStartId-1="2.[151,250,1949,1973]" captionTargetBox-0="[151,1421,667,1620]" captionTargetBox-1="[157,1436,884,1906]" captionTargetId-0="figure-201@1.[151,1435,667,1631]" captionTargetId-1="figure-315@2.[151,1436,884,1925]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Lateral view of male habitus: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." captionText-1="FIGURE 2. Male S1 tubercle in lateral view: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi-0="http://doi.org/10.5281/zenodo.10146518" figureDoi-1="http://doi.org/10.5281/zenodo.10146520" httpUri-0="https://zenodo.org/record/10146518/files/figure.png" httpUri-1="https://zenodo.org/record/10146520/files/figure.png" pageId="3" pageNumber="258">Figs. 1–2</figureCitation>
). S1–2 yellow/orange dorsally, duller colored towards the lateral and ventral portions; tubercle with digit-like shape with a darker and rounded apex bearing a patch of several apical setae; S3–6 yellow/ orange with a black apical ring; S7 yellow basally, darkening posteriorly; S8–9 black, with a large “T” shaped blue spot; S10 blue, the toothed crest forming a dorsal angled corner with the lateral margin (
<figureCitation id="13138BE7FFA0FF9B7824D24BFB31FA03" box="[1139,1260,1399,1425]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="3" pageNumber="258">Figs. 4a–c</figureCitation>
).
</paragraph>
<paragraph id="8B979762FFA0FF9B7C90D2A7FB71FA4B" blockId="3.[151,1437,1291,2037]" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7C90D2A7FEBFFA27" box="[199,354,1435,1461]" italics="true" pageId="3" pageNumber="258">Genital ligula</emphasis>
(
<figureCitation id="13138BE7FFA0FF9B7D26D2A7FE69FA27" box="[369,436,1435,1461]" captionStart="FIGURE 3" captionStartId="3.[151,250,1192,1216]" captionTargetBox="[175,1413,192,1157]" captionTargetId="figure-17@3.[164,1423,181,1168]" captionTargetPageId="3" captionText="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146522" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="3" pageNumber="258">Fig. 3</figureCitation>
). Segment one with a pair of digit-like lateral sclerotized processes, similar to lobes, in the flexure; inner fold vestigial. Segment two with a lanceolate apex, and two small lateral lobes.
</paragraph>
<paragraph id="8B979762FFA0FF9B7C90D2DFFB4AF9D7" blockId="3.[151,1437,1291,2037]" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7C90D2DFFE51FA6F" box="[199,396,1507,1533]" italics="true" pageId="3" pageNumber="258">Anal appendages</emphasis>
(
<figureCitation id="13138BE7FFA0FF9B7DC9D2DFFDD3FA6F" box="[414,526,1506,1533]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="4.[151,250,1729,1753]" captionStartId-1="5.[151,250,1943,1967]" captionTargetBox-0="[161,1429,353,1698]" captionTargetBox-1="[162,1429,189,1907]" captionTargetId-0="figure-99@4.[151,1436,345,1706]" captionTargetId-1="figure-17@5.[151,1436,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." captionText-1="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi-0="http://doi.org/10.5281/zenodo.10146524" figureDoi-1="http://doi.org/10.5281/zenodo.10146526" httpUri-0="https://zenodo.org/record/10146524/files/figure.png" httpUri-1="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">Figs. 4–5</figureCitation>
). Cercus forcipate, slightly longer than S10; dorsal plate bearing a row of four strong teeth, plus one smaller apical tooth; reaching the midlength of the basal plate; basal plate flat, broadening on the apical 1/3, ending on a rounded tip. Paraproct narrow, apex acute and directed upwards.
</paragraph>
<paragraph id="8B979762FFA0FF9B7C90D173FCD7F9FB" blockId="3.[151,1437,1291,2037]" box="[199,778,1615,1641]" pageId="3" pageNumber="258">Measurements. Abdomen 23.8; FW 14.6; HW 13.7.</paragraph>
<paragraph id="8B979762FFA0FF9B7C90D14EFC5EF923" blockId="3.[151,1437,1291,2037]" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7C90D14EFE06F91F" bold="true" box="[199,475,1650,1677]" pageId="3" pageNumber="258">
Variation in
<typeStatus id="549329C0FFA0FF9B7D09D14FFE0BF91F" box="[350,470,1651,1677]" pageId="3" pageNumber="258" type="paratype">paratypes</typeStatus>
.
</emphasis>
Overall coloration and morphology agree with the
<typeStatus id="549329C0FFA0FF9B784AD14FFB5EF91F" box="[1053,1155,1651,1677]" pageId="3" pageNumber="258" type="holotype">holotype</typeStatus>
. Measurements varied as follows: Total 30–31; abdomen 24–25; FW 14–14.5; HW 13–13.5.
</paragraph>
</subSubSection>
<subSubSection id="C332C4E9FFA0FF9C7C90D186FE9CFED3" lastPageId="4" lastPageNumber="259" pageId="3" pageNumber="258" type="diagnosis">
<paragraph id="8B979762FFA0FF9C7C90D186FE9CFED3" blockId="3.[151,1437,1291,2037]" lastBlockId="4.[151,1437,151,321]" lastPageId="4" lastPageNumber="259" pageId="3" pageNumber="258">
<emphasis id="B95C4B70FFA0FF9B7C90D186FE9FF946" bold="true" box="[199,322,1722,1748]" pageId="3" pageNumber="258">Diagnosis.</emphasis>
<taxonomicName id="4C28ECE1FFA0FF9B7D1DD187FDFFF947" authority="Vilela &amp; Jacques &amp; Souza, 2023" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[330,546,1723,1749]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="veredae" status="sp. nov.">
<emphasis id="B95C4B70FFA0FF9B7D1DD187FDFFF947" box="[330,546,1723,1749]" italics="true" pageId="3" pageNumber="258">Minagrion veredae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26FF60BFFA0FF9B7E7CD187FD5EF947" box="[555,643,1723,1749]" pageId="3" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
is closer to
<taxonomicName id="4C28ECE1FFA0FF9B7F47D187FBE8F947" authority="(Selys, 1876)" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[784,1077,1723,1749]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="B95C4B70FFA0FF9B7F47D187FC49F947" box="[784,916,1723,1749]" italics="true" pageId="3" pageNumber="258">M. waltheri</emphasis>
(Selys, 1876)
</taxonomicName>
and
<taxonomicName id="4C28ECE1FFA0FF9B7839D187FF04F96B" authority="(Selys, 1876)" baseAuthorityName="Selys" baseAuthorityYear="1876" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="mecistogastrum">
<emphasis id="B95C4B70FFA0FF9B7839D187FA9BF946" box="[1134,1350,1723,1748]" italics="true" pageId="3" pageNumber="258">M. mecistogastrum</emphasis>
(Selys, 1876)
</taxonomicName>
in terms of body morphology and coloration. It can be easily differentiated from
<taxonomicName id="4C28ECE1FFA0FF9B7833D1E3FAE7F96A" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1124,1338,1759,1784]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="mecistogastrum">
<emphasis id="B95C4B70FFA0FF9B7833D1E3FAE7F96A" box="[1124,1338,1759,1784]" italics="true" pageId="3" pageNumber="258">M. mecistogastrum</emphasis>
</taxonomicName>
by body size (the largest species of the genus, with an average body size of
<quantity id="4CD03A87FFA0FF9B7FC6D03EFC01F88F" box="[913,988,1794,1821]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="258" unit="mm" value="57.0">57mm</quantity>
), genital ligula and morphology of anal appendages. The following characters allow for the differentiation between
<taxonomicName id="4C28ECE1FFA0FF9B7F89D01BFB87F8D3" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[990,1114,1831,1857]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="veredae">
<emphasis id="B95C4B70FFA0FF9B7F89D01BFB87F8D3" box="[990,1114,1831,1857]" italics="true" pageId="3" pageNumber="258">M. veredae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C28ECE1FFA0FF9B78D9D01BFAD2F8D3" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[1166,1295,1831,1857]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="B95C4B70FFA0FF9B78D9D01BFAD2F8D3" box="[1166,1295,1831,1857]" italics="true" pageId="3" pageNumber="258">M. waltheri</emphasis>
</taxonomicName>
(the latter in parenthesis): cercus (
<figureCitation id="13138BE7FFA0FF9B7DD2D077FE3FF8F7" box="[389,482,1867,1893]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="3" pageNumber="258">Figs. 4b</figureCitation>
,
<figureCitation id="13138BE7FFA0FF9B7DB9D076FDD4F8F7" box="[494,521,1866,1893]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">5c</figureCitation>
) markedly forcipate, with the ventral margin deeply concave (less forcipate, with ventral margin mostly linear;
<figureCitation id="13138BE7FFA0FF9B7DB4D053FDE0F81B" box="[483,573,1903,1929]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="3" pageNumber="258">Figs. 4e</figureCitation>
,
<figureCitation id="13138BE7FFA0FF9B7E1ED052FDB5F81B" box="[585,616,1902,1929]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">5b</figureCitation>
); cercus dorsal plate bearing a row of four large strong teeth (
<figureCitation id="13138BE7FFA0FF9B794FD053FA50F81B" box="[1304,1421,1902,1929]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">Figs. 5a, c</figureCitation>
), plus one smaller apical tooth (cercus dorsal plate bearing a row of short teeth, lacking apical tooth;
<figureCitation id="13138BE7FFA0FF9B78BAD0AFFA9DF83F" box="[1261,1344,1938,1965]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">Fig. 5b</figureCitation>
); cercus basal plate flat (
<figureCitation id="13138BE7FFA0FF9B7D1AD08BFE42F843" box="[333,415,1974,2001]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">Fig. 5c</figureCitation>
), broadening on the apical 1/3, ending on a rounded tip (cercus basal plate flat, narrowing at the apical 1/3, ending on a blunt tip;
<figureCitation id="13138BE7FFA0FF9B7E07D0E7FD79F867" box="[592,676,2010,2037]" captionStart="FIGURE 5" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[162,1429,189,1907]" captionTargetId="figure-17@5.[151,1436,181,1919]" captionTargetPageId="5" captionText="FIGURE 5. Closeup of male cercus showing diagnostic characters: (a, c) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146526" httpUri="https://zenodo.org/record/10146526/files/figure.png" pageId="3" pageNumber="258">Fig. 5b</figureCitation>
); paraprocts (
<figureCitation id="13138BE7FFA0FF9B7F6AD0E7FC50F867" box="[829,909,2011,2037]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="3" pageNumber="258">Fig. 4a</figureCitation>
) narrow and acute (paraprocts thicker and also acute;
<figureCitation id="13138BE7FFA7FF9C7CB5D7ABFEE8FF23" box="[226,309,151,177]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="4" pageNumber="259">Fig. 4d</figureCitation>
); S10 toothed crest (
<figureCitation id="13138BE7FFA7FF9C7E4AD7ABFD4EFF23" box="[541,659,151,177]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="4" pageNumber="259">Figs. 4a–c</figureCitation>
) forming a dorsal angled corner with the lateral margin (S10 toothed crest not forming an angled corner, instead a rounded border;
<figureCitation id="13138BE7FFA7FF9C7F15D787FC68FF47" box="[834,949,187,213]" captionStart="FIGURE 4" captionStartId="4.[151,250,1729,1753]" captionTargetBox="[161,1429,353,1698]" captionTargetId="figure-99@4.[151,1436,345,1706]" captionTargetPageId="4" captionText="FIGURE 4. Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c) Minagrion veredae sp. nov. HOLOTYPE; (d–f) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="4" pageNumber="259">Figs. 4e–f</figureCitation>
); genital ligula (
<figureCitation id="13138BE7FFA7FF9C783ED787FB6AFF47" box="[1129,1207,187,213]" captionStart="FIGURE 3" captionStartId="3.[151,250,1192,1216]" captionTargetBox="[175,1413,192,1157]" captionTargetId="figure-17@3.[164,1423,181,1168]" captionTargetPageId="3" captionText="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146522" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="4" pageNumber="259">Fig. 3a</figureCitation>
) with a pair of digit-like lateral sclerotized processes (sclerotized processes small, not digit-like;
<figureCitation id="13138BE7FFA7FF9C7FB4D7E3FBE9FF6B" box="[995,1076,223,249]" captionStart="FIGURE 3" captionStartId="3.[151,250,1192,1216]" captionTargetBox="[175,1413,192,1157]" captionTargetId="figure-17@3.[164,1423,181,1168]" captionTargetPageId="3" captionText="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146522" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="4" pageNumber="259">Fig. 3b</figureCitation>
); segment two of ligula (
<figureCitation id="13138BE7FFA7FF9C7912D7E3FA49FF6B" box="[1349,1428,223,249]" captionStart="FIGURE 3" captionStartId="3.[151,250,1192,1216]" captionTargetBox="[175,1413,192,1157]" captionTargetId="figure-17@3.[164,1423,181,1168]" captionTargetPageId="3" captionText="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146522" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="4" pageNumber="259">Fig. 3a</figureCitation>
) with a lanceolate apex, and two small lateral lobes (segment two of ligula with a blunt apex, and two longer lateral lobes;
<figureCitation id="13138BE7FFA7FF9C7CB6D61BFEE9FED3" box="[225,308,295,321]" captionStart="FIGURE 3" captionStartId="3.[151,250,1192,1216]" captionTargetBox="[175,1413,192,1157]" captionTargetId="figure-17@3.[164,1423,181,1168]" captionTargetPageId="3" captionText="FIGURE 3. Lateral view of genital ligula: (a) Minagrion veredae sp. nov. HOLOTYPE; (b) Minagrion waltheri." figureDoi="http://doi.org/10.5281/zenodo.10146522" httpUri="https://zenodo.org/record/10146522/files/figure.png" pageId="4" pageNumber="259">Fig. 3b</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF57C7EAFFA7FF9C7CC0D1FDFDEDF96F" ID-DOI="http://doi.org/10.5281/zenodo.10146524" ID-Zenodo-Dep="10146524" httpUri="https://zenodo.org/record/10146524/files/figure.png" pageId="4" pageNumber="259" startId="4.[151,250,1729,1753]" targetBox="[161,1429,353,1698]" targetPageId="4" targetType="figure">
<paragraph id="8B979762FFA7FF9C7CC0D1FDFDEDF96F" blockId="4.[151,1437,1729,1790]" pageId="4" pageNumber="259">
<emphasis id="B95C4B70FFA7FF9C7CC0D1FDFECBF94B" bold="true" box="[151,278,1729,1753]" pageId="4" pageNumber="259">FIGURE 4.</emphasis>
Male cercus in lateral (left), dorsal (middle) and mediodorsal (right) views: (a–c)
<taxonomicName id="4C28ECE1FFA7FF9C782AD1FEFA9FF948" authority="Vilela &amp; Jacques &amp; Souza, 2023" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[1149,1346,1730,1754]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="veredae" status="sp. nov.">
<emphasis id="B95C4B70FFA7FF9C782AD1FEFA9FF948" box="[1149,1346,1730,1754]" italics="true" pageId="4" pageNumber="259">Minagrion veredae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26FF60BFFA7FF9C791CD1FEFA40F94B" box="[1355,1437,1730,1753]" pageId="4" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
HOLOTYPE; (d–f)
<taxonomicName id="4C28ECE1FFA7FF9C7D34D1DAFDF7F96C" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[355,554,1766,1790]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="waltheri">
<emphasis id="B95C4B70FFA7FF9C7D34D1DAFDF7F96C" box="[355,554,1766,1790]" italics="true" pageId="4" pageNumber="259">Minagrion waltheri</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C332C4E9FFA7FF9C7C90D06CFDE5F868" pageId="4" pageNumber="259" type="biology_ecology">
<paragraph id="8B979762FFA7FF9C7C90D06CFDE5F868" blockId="4.[151,1436,1872,2042]" pageId="4" pageNumber="259">
<emphasis id="B95C4B70FFA7FF9C7C90D06CFE2DF8F8" bold="true" box="[199,496,1872,1898]" pageId="4" pageNumber="259">Habitat and distribution.</emphasis>
The habitat of
<taxonomicName id="4C28ECE1FFA7FF9C7EC2D06DFCCCF8F8" authorityName="Vilela &amp; Jacques &amp; Souza" authorityYear="2023" box="[661,785,1872,1898]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="veredae">
<emphasis id="B95C4B70FFA7FF9C7EC2D06DFCCCF8F8" box="[661,785,1872,1898]" italics="true" pageId="4" pageNumber="259">M. veredae</emphasis>
</taxonomicName>
is similar to what is reported for other species of the genus, the palm swamp environments (
<bibRefCitation id="EFB9EA93FFA7FF9C7DA0D048FD59F81D" author="Santos, N. D." box="[503,644,1908,1935]" pageId="4" pageNumber="259" pagination="8 - 12" refId="ref3755" refString="Santos, N. D. (1965) Contribuicao ao conhecimento dos &quot; Odonata &quot; da regiao de Pocos de Caldas, MG. &quot; Minagrion &quot; gen. n., para &quot; Telagrion mecistogastrum &quot; Selys Longchamps 1865, com a descricao de uma nova especie. Atas da Sociedade de Biologia do Rio de Janeiro, 9 (1), 8 - 12." type="journal article" year="1965">Santos 1965</bibRefCitation>
;
<bibRefCitation id="EFB9EA93FFA7FF9C7ED8D048FC4EF81C" author="Machado, A. B. M. &amp; Bede, L. C." box="[655,915,1908,1934]" pageId="4" pageNumber="259" pagination="269 - 296" refId="ref3572" refString="Machado, A. B. M. &amp; Bede, L. C. (2016) Two new genera and nine new species of damselflies from a localized area in Minas Gerais, Brazil (Odonata: Zygoptera). International Journal of Odonatology, 18 (4), 269 - 296. https: // doi. org / 10.1080 / 13887890.2015.1072113" type="journal article" year="2016">Machado &amp; Bedê 2016</bibRefCitation>
;
<bibRefCitation id="EFB9EA93FFA7FF9C7FCBD048FB80F81C" author="Vilela, D. S. &amp; Anjos-Santos, D. &amp; Koroiva, R. &amp; Cordero-Rivera, A. &amp; Guillermo-Ferreira, R." box="[924,1117,1908,1934]" pageId="4" pageNumber="259" pagination="176 - 198" refId="ref3907" refString="Vilela, D. S., Anjos-Santos, D., Koroiva, R., Cordero-Rivera, A. &amp; Guillermo-Ferreira, R. (2020) Revision of the genus Minagrion Santos, 1965 (Odonata: Coenagrionidae). Zootaxa, 4786 (2), 176 - 198. https: // doi. org / 10.11646 / zootaxa. 4786.2.2" type="journal article" year="2020">
Vilela
<emphasis id="B95C4B70FFA7FF9C7FB3D049FBC6F81C" box="[996,1051,1908,1934]" italics="true" pageId="4" pageNumber="259">et al.</emphasis>
2020
</bibRefCitation>
;
<figureCitation id="13138BE7FFA7FF9C7831D048FB75F81C" box="[1126,1192,1908,1934]" captionStart="FIGURE 6" captionStartId="6.[151,250,684,708]" captionTargetBox="[151,1436,181,660]" captionTargetId="figure-18@6.[151,1436,181,660]" captionTargetPageId="6" captionText="FIGURE 6. Type locality of Minagrion veredae at the GSVNP." figureDoi="http://doi.org/10.5281/zenodo.10146529" httpUri="https://zenodo.org/record/10146529/files/figure.png" pageId="4" pageNumber="259">Fig. 6</figureCitation>
). These environments are composed of permanent wet soils, which provide suitable reproductive sites for these species throughout the year (Vilela
<emphasis id="B95C4B70FFA7FF9C7D77D081FE8CF844" box="[288,337,1980,2006]" italics="true" pageId="4" pageNumber="259">et al</emphasis>
. 2016). Nonetheless, the larval stages of
<taxonomicName id="4C28ECE1FFA7FF9C7F4CD081FC4EF844" authorityName="Santos" authorityYear="1965" box="[795,915,1981,2006]" class="Insecta" family="Coenagrionidae" genus="Minagrion" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="259" phylum="Arthropoda" rank="genus">
<emphasis id="B95C4B70FFA7FF9C7F4CD081FC4EF844" box="[795,915,1981,2006]" italics="true" pageId="4" pageNumber="259">Minagrion</emphasis>
</taxonomicName>
are still unknown, pending on further searches on veredas to fill this taxonomic gap.
</paragraph>
</subSubSection>
</treatment>
</document>
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