251 lines
36 KiB
XML
251 lines
36 KiB
XML
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<document id="38F29843EA8FE0B7E3A1B63480B05B5B" ID-DOI="10.5281/zenodo.197230" ID-GBIF-Dataset="73882c0e-4d4c-4800-bfec-0fd95f877e0e" ID-ISSN="1175-5326" ID-Zenodo-Dep="197230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460262189519" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid & Pavićević, Dragan" docDate="2010" docId="039D4825F662FFC6FF12FD37FC00FE8D" docLanguage="en" docName="zt02565p041.pdf" docOrigin="Zootaxa 2565" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachytrachis tumidus Ingrisch & Pavićević, 2010, sp. n." docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFA4305DF643FFE3FF85FFB8FFB2FFE1" masterDocTitle="Seven new Tettigoniidae (Orthoptera) and a new Blattellidae (Blattodea) from the Durmitor area of Montenegro with notes on previously known taxa" masterLastPageNumber="41" masterPageNumber="1" pageNumber="34" updateTime="1698244838667" updateUser="plazi">
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<mods:mods id="37558351D154D2E535DCC0B7090E5B05" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="166D154AA9339511E596D60B3FB8B96E">
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<mods:title id="6661B5D5AD284A41433C5E91CC4622CC">Seven new Tettigoniidae (Orthoptera) and a new Blattellidae (Blattodea) from the Durmitor area of Montenegro with notes on previously known taxa</mods:title>
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</mods:titleInfo>
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<mods:name id="EBF31D8028BCBA7B6F9CC0705CF7063D" type="personal">
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<mods:role id="0EF68227462400635ACE35BA8FAF8546">
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<mods:roleTerm id="336F8AC3E8C983B87FB454243D5801C8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9C94C0067415D3385708BEC5DC4224F9">Ingrisch, Sigfrid</mods:namePart>
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</mods:name>
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<mods:name id="4A5BC74AB1ADE2E62243568DF53D108F" type="personal">
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<mods:role id="FB7115AB858F2DB7BBC46263967BC93F">
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<mods:roleTerm id="1EA6F159C1AF3C2E8EFB9424468AD023">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0835877E36071B5DB89383ED38E24EC6">Pavićević, Dragan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="307C953A0D40625693D337666CE22FD6">text</mods:typeOfResource>
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<mods:relatedItem id="9B01A296A1607E12A7A88EFEEF3AFCBA" type="host">
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<mods:titleInfo id="C5D8167937241E3AFB22FF4AAD0F5E99">
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<mods:title id="AACE7AF079F3C7765650D1032768AF8A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6CEA21622592C223D9BC3E3BDCA71EF3">
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<mods:date id="4621F6770605D68BA40227DD6D178A5A">2010</mods:date>
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<mods:detail id="2A59BDCC59428A0CF233CA9256E55225" type="volume">
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<mods:number id="5E121DC9A9AEDAC64508CE7FA5002013">2565</mods:number>
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</mods:detail>
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<mods:extent id="F050DC1C6B74C9BE9A1F499E13B0CB69" unit="page">
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<mods:start id="A0A56BC1023F5D86B5B0D08FFCF24519">1</mods:start>
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<mods:end id="3B66759257E729DFCE847E24D5D9BC61">41</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="DF7B54C6D810D2EAAB1BEE0F258AFAA9">journal article</mods:classification>
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<mods:identifier id="F25298DD77CC74D56445836BA5914B9E" type="DOI">10.5281/zenodo.197230</mods:identifier>
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<mods:identifier id="BFD37EDF6BCA53138E9EF80BAAA224D0" type="GBIF-Dataset">73882c0e-4d4c-4800-bfec-0fd95f877e0e</mods:identifier>
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<mods:identifier id="E57CFBEC529FB49C81B11DCA0C0B2650" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B0E46386E4225CBFD0B4EF447F69FC12" type="Zenodo-Dep">197230</mods:identifier>
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</mods:mods>
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<treatment id="039D4825F662FFC6FF12FD37FC00FE8D" ID-DOI="http://doi.org/10.5281/zenodo.6204501" ID-GBIF-Taxon="119393612" ID-Zenodo-Dep="6204501" LSID="urn:lsid:plazi:treatment:039D4825F662FFC6FF12FD37FC00FE8D" httpUri="http://treatment.plazi.org/id/039D4825F662FFC6FF12FD37FC00FE8D" lastPageId="37" lastPageNumber="38" pageId="33" pageNumber="34">
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<subSubSection id="C32EAAB8F662FFC2FF12FD37FE5FFD2E" pageId="33" pageNumber="34" type="nomenclature">
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<paragraph id="8B8BF933F662FFC2FF12FD37FE55FD48" blockId="33.[151,493,655,719]" box="[151,487,655,681]" pageId="33" pageNumber="34">
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<heading id="D0C34E5FF662FFC2FF12FD37FE55FD48" bold="true" box="[151,487,655,681]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
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<emphasis id="B9402521F662FFC2FF12FD37FE55FD48" bold="true" box="[151,487,655,681]" pageId="33" pageNumber="34">
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<taxonomicName id="4C3482B0F662FFC2FF12FD37FE2CFD48" ID-CoL="4BVSZ" box="[151,414,655,681]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
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<emphasis id="B9402521F662FFC2FF12FD37FE2CFD48" bold="true" box="[151,414,655,681]" italics="true" pageId="33" pageNumber="34">Pachytrachis tumidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273985AF662FFC2FE20FD37FE55FD48" box="[421,487,655,681]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8BF933F662FFC2FF12FD0DFE5FFD2E" blockId="33.[151,493,655,719]" box="[151,493,693,719]" pageId="33" pageNumber="34">
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<figureCitation id="130FE5B6F662FFC2FF12FD0DFE9FFD2E" box="[151,301,693,719]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="33" pageNumber="34">Figs. 17A–E</figureCitation>
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,
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<figureCitation id="130FE5B6F662FFC2FEBFFD0DFE21FD2E" box="[314,403,693,719]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">18A–K</figureCitation>
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,
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<figureCitation id="130FE5B6F662FFC2FE1BFD0DFE0BFD2E" box="[414,441,693,719]" captionStart="FIGURE 19" captionStartId="36.[151,255,1927,1951]" captionTargetBox="[334,1251,198,1908]" captionTargetId="figure@36.[324,1263,194,1914]" captionTargetPageId="36" captionText="FIGURE 19. Oscillograms for comparison of calling songs of Pachytrachis species: A – B, P. gracilis (Brunner, 1861); C – D, P. striolatu s (Fieber, 1853); E – F, P. t u m id u s sp. n. — A, C, E, series of syllables; B, D, F, one syllable in higher time resolution." httpUri="https://zenodo.org/record/197249/files/figure.png" pageId="33" pageNumber="34">19</figureCitation>
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E–F
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EAAB8F662FFC2FF12FCBDFF07FC3B" pageId="33" pageNumber="34" type="materials_examined">
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<paragraph id="8B8BF933F662FFC2FF12FCBDFA3FFCA5" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<typeStatus id="548F4791F662FFC2FF12FCBDFEB6FCFE" box="[151,260,773,799]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
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(male):
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<collectingCountry id="F323B9A3F662FFC2FEE3FCBDFE48FCFE" box="[358,506,773,799]" name="Serbia" pageId="33" pageNumber="34">Montenegro</collectingCountry>
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: Durmitor, Komarnica Canyon, Krivača (= village of Komarnica),
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<quantity id="4CCC54D6F662FFC2FA86FCBDFA2AFCFE" box="[1283,1432,773,799]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="33" pageNumber="34" unit="m" value="1025.0" valueMax="1100.0" valueMin="950.0">950–1100 m</quantity>
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,
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<date id="FF8ADFF3F662FFC2FF12FC92FE96FCA5" box="[151,292,810,836]" pageId="33" pageNumber="34" value="1990-08-09">9.VIII.1990</date>
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, leg. D. Pavićević, in Coll. Zoologisches Forschungsmuseum Alexander Koenig Bonn (
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<collectionCode id="ED2561F6F662FFC2FAACFC92FA33FCA5" LSID="urn:lsid:biocol.org:col:34613" box="[1321,1409,810,836]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="33" pageNumber="34">ZFMK</collectionCode>
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).
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</paragraph>
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<paragraph id="8B8BF933F662FFC2FF43FCE8FAB2FC6E" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<typeStatus id="548F4791F662FFC2FF43FCE8FE8FFC8B" box="[198,317,848,874]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
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: 1 Ƥ, same locality as
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<typeStatus id="548F4791F662FFC2FDD6FCE8FD0FFC8B" box="[595,701,848,874]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
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,
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<date id="FF8ADFF3F662FFC2FD4EFCE8FCEAFC8B" box="[715,856,848,874]" pageId="33" pageNumber="34" value="1990-08-09">9.VIII.1990</date>
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, leg. S. Ingrisch (
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<collectionCode id="ED2561F6F662FFC2FBABFCE8FB34FC8B" LSID="urn:lsid:biocol.org:col:34613" box="[1070,1158,848,874]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="33" pageNumber="34">ZFMK</collectionCode>
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); 1 Ƥ, do.,
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<date id="FF8ADFF3F662FFC2FA97FCE8FA2EFC8B" box="[1298,1436,848,874]" pageId="33" pageNumber="34" value="1989-08-08">8.VIII.1989</date>
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(CI); 2 Ƥ,
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<collectingCountry id="F323B9A3F662FFC2FE97FCCDFE16FC6E" box="[274,420,885,911]" name="Serbia" pageId="33" pageNumber="34">Montenegro</collectingCountry>
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,
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<quantity id="4CCC54D6F662FFC2FE2AFCCDFE4BFC6E" box="[431,505,885,911]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="33" pageNumber="34" unit="km" value="40.0">40 km</quantity>
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NW Nikšič, Krstac,
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<quantity id="4CCC54D6F662FFC2FD6AFCCDFCF9FC6E" box="[751,843,885,911]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="m" value="1000.0">1000 m</quantity>
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,
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<date id="FF8ADFF3F662FFC2FCD2FCCDFC56FC6E" box="[855,996,885,911]" pageId="33" pageNumber="34" value="1968-08-05">5.VIII.1968</date>
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, leg. F. Willemse (CW).
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</paragraph>
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<paragraph id="8B8BF933F662FFC2FF43FC22FF07FC3B" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<emphasis id="B9402521F662FFC2FF43FC22FEDFFC55" bold="true" box="[198,365,922,948]" pageId="33" pageNumber="34">
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<typeStatus id="548F4791F662FFC2FF43FC22FEB6FC55" box="[198,260,922,948]" pageId="33" pageNumber="34">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F323B9A3F662FFC2FEF3FC22FDBAFC55" box="[374,520,922,948]" name="Serbia" pageId="33" pageNumber="34">Montenegro</collectingCountry>
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, Durmitor, Komarnica Canyon, Krivača (= village of Komarnica),
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<quantity id="4CCC54D6F662FFC2FAADFC22FF03FC3B" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="33" pageNumber="34" unit="m" value="1025.0" valueMax="1100.0" valueMin="950.0">950–1100 m</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EAAB8F662FFC2FF43FC5DFEB5FBAB" pageId="33" pageNumber="34" type="description">
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<paragraph id="8B8BF933F662FFC2FF43FC5DFEB5FBAB" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<emphasis id="B9402521F662FFC2FF43FC5DFE37FC1E" bold="true" box="[198,389,997,1023]" pageId="33" pageNumber="34">Measurements.</emphasis>
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Body length male 19.5, female 16.5–26.0; pronotum length male 4.5, female 5.1–5.3; tegmen length male 3.1, female 1.8–2.0; hind femur male 17.8, female 19.0–20.5; ovipositor female 14.0–
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<quantity id="4CCC54D6F662FFC2FF12FB88FEB1FBAB" box="[151,259,1072,1098]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="33" pageNumber="34" unit="mm" value="14.5">14.5 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EAAB8F662FFC2FF43FBEDFDF9FB3E" pageId="33" pageNumber="34" type="diagnosis">
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<paragraph id="8B8BF933F662FFC2FF43FBEDFDF9FB3E" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<emphasis id="B9402521F662FFC2FF43FBEDFEF4FB8E" bold="true" box="[198,326,1109,1135]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
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Females of the new species are similar to those of
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<emphasis id="B9402521F662FFC2FC1AFBEDFC49FB8F" box="[927,1019,1109,1134]" italics="true" pageId="33" pageNumber="34">P. f r a t e r</emphasis>
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(Bunner, 1882). They differ by the ovipositor which is little but distinctly curved ventrad and by the shape of the subgenital plate. Males differ from all known species of
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<taxonomicName id="4C3482B0F662FFC2FE64FB18FD31FB58" box="[481,643,1184,1209]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis id="B9402521F662FFC2FE64FB18FD31FB58" box="[481,643,1184,1209]" italics="true" pageId="33" pageNumber="34">Pachytrachis</emphasis>
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</taxonomicName>
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by the internal swelling of the cerci and by the compressed and widened apical parts of the titillators.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32EAAB8F662FFC1FF43FB52FE82FE56" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
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<paragraph id="8B8BF933F662FFC2FF43FB52FD3BFAAE" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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<emphasis id="B9402521F662FFC2FF43FB52FEECFAE5" bold="true" box="[198,350,1258,1284]" pageId="33" pageNumber="34">Description.</emphasis>
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Pronotum with disc convex, without median carina; completely rounded into paranota. Micropterous; tegmina of male reduced to stridulatory apparatus; tegmina of female reduced to scales, very little overlapping in middle (
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<figureCitation id="130FE5B6F662FFC2FE63FA8DFDCFFAAE" box="[486,637,1333,1359]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="33" pageNumber="34">Figs 17A–D</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8BF933F662FFC2FF43FAE2FC2BF9B5" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
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Male. Stridulatory file on underside of left tegmen curved, concave, with ca 75 moderately dense teeth (
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<figureCitation id="130FE5B6F662FFC2FF1AFA38FEBFFA7B" box="[159,269,1408,1434]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Fig. 18A</figureCitation>
|
|||
|
|
). Mirror on right tegmen wider than long; hind margin rounded (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FB81FA38FBDDFA7B" box="[1028,1135,1408,1434]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="33" pageNumber="34">Fig. 17B</figureCitation>
|
|||
|
|
). Tenth abdominal tergite faintly furrowed in midline; rather short, hardly longer than ninth tergite; apical margin truncate in middle, faintly concave at both sides (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FD85FA72FDDEFA05" box="[512,620,1482,1508]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Fig. 18F</figureCitation>
|
|||
|
|
). Epiproct small; broadly rounded; with a circular pit at base. Cerci faintly curved at base, in dorsal view otherwise straight, in lateral view slightly sinuate; without tooth but with a distinct internal swelling behind basal third; before apex very faintly swollen (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FBEEF9ADFAB4F9CE" box="[1131,1286,1557,1583]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Figs. 18E–F</figureCitation>
|
|||
|
|
). Subgenital plate with narrow median carina; apex roundly excised; styli thin.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BF933F662FFC2FF43F9D8FDABF925" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
|
|||
|
|
Titillators separate; curved in middle; apical parts compressed, band-shaped, brownish hyaline; apex rounded; dorsal margin little convex and finely crenulated; basal parts dark brown; very base ovoid and brownish hyaline (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FEF4F912FDBEF925" box="[369,524,1706,1732]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Figs. 18B–D</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BF933F662FFC2FF43F968FCD5F89E" blockId="33.[151,1437,771,2031]" pageId="33" pageNumber="34">
|
|||
|
|
Female. Tenth abdominal tergite with central area little prolonged behind but angularly excised in middle. Epiproct triangular with a circular pit on top. Cerci thin, apex pointing. Subgenital plate with central area flattened, lateral areas with little concave surface; lateral margins converging; apex roundly excised; lobes on both sides of excision rounded or in a freeze-dried specimen subacute (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FC65F8F8FBC6F8BB" box="[992,1140,1856,1882]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Figs. 18G–J</figureCitation>
|
|||
|
|
). Ovipositor with ventral margin little but distinctly concave; apex tapering (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FD68F8DDFCE9F89E" box="[749,859,1893,1919]" captionStart="FIGURE 18. A – K" captionStartId="35.[151,255,1868,1892]" captionTargetBox="[226,1343,194,1827]" captionTargetId="figure@35.[226,1344,194,1837]" captionTargetPageId="35" captionText="FIGURE 18. A – K, Pachytrachis tumidus sp. n., (A – F, male holotype; G – H, female paratype freeze-dried; I – K, female paratype air-dried); L – N, Pachytrachis frater (Brunner, 1882) female from Moraċa valley (CW). – A, stridulatory file; B – D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor." httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="33" pageNumber="34">Fig. 18K</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B8BF933F662FFC1FF43F832FE82FE56" blockId="33.[151,1437,771,2031]" lastBlockId="34.[151,1437,152,439]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
|||
|
|
<emphasis id="B9402521F662FFC2FF43F832FEE7F845" bold="true" box="[198,341,1930,1956]" pageId="33" pageNumber="34">Coloration.</emphasis>
|
|||
|
|
Dark brown with marmoration; sternites orange, yellowish or whitish brown. Frons brown marmorated (
|
|||
|
|
<figureCitation id="130FE5B6F662FFC2FEBCF808FE14F82B" box="[313,422,1968,1994]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="33" pageNumber="34">Fig. 17E</figureCitation>
|
|||
|
|
). Vertex and pronotal disc with fine white median line, on vertex accompanied by a cuneate black spot on both sides. Two other fine white lines from bases of antennae along dorsal margin of eyes to occiput; on dorsal side bordered or not by a narrow black band, on ventral side bordered by a wide black post-ocular band. Pronotum with disc brown; paranota black with broad white ventral margin including a spot suffused with brown (
|
|||
|
|
<figureCitation id="130FE5B6F661FFC1FE63FF5AFD30FF1D" box="[486,642,226,252]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="34" pageNumber="35">Figs. 17C–D</figureCitation>
|
|||
|
|
). Tegmina of male with disc dark brown, at apex with a white spot; lateral area blackish brown with a white spot at base (
|
|||
|
|
<figureCitation id="130FE5B6F661FFC1FC83FEB0FCC1FEC3" box="[774,883,264,290]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="34" pageNumber="35">Fig. 17A</figureCitation>
|
|||
|
|
). Tegmina of female with hidden part brownish white; exposed part blackish brown with internal and posterior margin on internal side white (
|
|||
|
|
<figureCitation id="130FE5B6F661FFC1FA9BFE95FA3DFEA6" box="[1310,1423,301,327]" captionStart="FIGURE 17. A – E" captionStartId="34.[151,259,1948,1972]" captionTargetBox="[151,1436,477,1935]" captionTargetId="figure@34.[151,1436,477,1937]" captionTargetPageId="34" captionText="FIGURE 17. A – E, Pachytrachis tumidus sp. n. from Krivaċa (A – B male holotype; C – E, female paratype); F – G Pachytrachis frater (Brunner, 1882) female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E – F, female face." httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="34" pageNumber="35">Fig. 17C</figureCitation>
|
|||
|
|
). Abdominal tergites dark brown; first three abdominal tergites with black lateral bands bordered by white bands on dorsal and ventral sides. Legs brown with blackish brown spots; hind femur with little distinct fishbone pattern.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF4BA9BBF661FFC1FF12F824FB7FF812" httpUri="https://zenodo.org/record/197247/files/figure.png" pageId="34" pageNumber="35" targetBox="[151,1436,477,1935]" targetPageId="34">
|
|||
|
|
<paragraph id="8B8BF933F661FFC1FF12F824FB7FF812" blockId="34.[151,1436,1948,2035]" pageId="34" pageNumber="35">
|
|||
|
|
<emphasis id="B9402521F661FFC1FF12F824FE9DF855" bold="true" box="[151,303,1948,1972]" pageId="34" pageNumber="35">FIGURE 17.</emphasis>
|
|||
|
|
A–E,
|
|||
|
|
<taxonomicName id="4C3482B0F661FFC1FEFCF824FDDDF852" box="[377,623,1948,1971]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9402521F661FFC1FEFCF824FDDDF852" box="[377,623,1948,1971]" italics="true" pageId="34" pageNumber="35">Pachytrachis tumidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9402521F661FFC1FDF2F824FD0BF855" bold="true" box="[631,697,1948,1972]" pageId="34" pageNumber="35">
|
|||
|
|
<taxonomicNameLabel id="A273985AF661FFC1FDF2F824FD0BF855" box="[631,697,1948,1972]" pageId="34" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
from Krivaċa (A–B male holotype; C–E, female paratype); F–G
|
|||
|
|
<taxonomicName id="4C3482B0F661FFC1FF12F804FD9AF832" authority="Brunner, 1882" authorityName="Brunner" authorityYear="1882" box="[151,552,1980,2003]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="frater">
|
|||
|
|
<emphasis id="B9402521F661FFC1FF12F804FEDCF832" box="[151,366,1980,2003]" italics="true" pageId="34" pageNumber="35">Pachytrachis frater</emphasis>
|
|||
|
|
(Brunner, 1882)
|
|||
|
|
</taxonomicName>
|
|||
|
|
female from Moraċa valley, Montenegro (CW). – A, male left tegmen; B, right tegmen; C, female head pronotum and wings in dorsal view; D, G, do. lateral view; E–F, female face.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF4BA9BBF660FFC0FF12F8F4FDFCF805" httpUri="https://zenodo.org/record/197248/files/figure.png" pageId="35" pageNumber="36" targetBox="[226,1343,194,1827]" targetPageId="35">
|
|||
|
|
<paragraph id="8B8BF933F660FFC0FF12F8F4FDFCF805" blockId="35.[151,1436,1868,2020]" pageId="35" pageNumber="36">
|
|||
|
|
<emphasis id="B9402521F660FFC0FF12F8F4FE95F885" bold="true" box="[151,295,1868,1892]" pageId="35" pageNumber="36">FIGURE 18.</emphasis>
|
|||
|
|
A–K,
|
|||
|
|
<taxonomicName id="4C3482B0F660FFC0FEF5F8F5FDEAF885" box="[368,600,1869,1892]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
|
|||
|
|
<emphasis id="B9402521F660FFC0FEF5F8F5FDEAF885" box="[368,600,1869,1892]" italics="true" pageId="35" pageNumber="36">Pachytrachis tumidus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B9402521F660FFC0FDE4F8F4FD2FF885" bold="true" box="[609,669,1868,1892]" pageId="35" pageNumber="36">
|
|||
|
|
<taxonomicNameLabel id="A273985AF660FFC0FDE4F8F4FD2FF885" box="[609,669,1868,1892]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, (A–F, male holotype; G–H, female paratype freeze-dried; I–K, female paratype air-dried); L–N,
|
|||
|
|
<taxonomicName id="4C3482B0F660FFC0FE2BF8D5FC87F865" authority="Brunner, 1882" authorityName="Brunner" authorityYear="1882" box="[430,821,1901,1924]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="frater">
|
|||
|
|
<emphasis id="B9402521F660FFC0FE2BF8D5FD32F865" box="[430,640,1901,1924]" italics="true" pageId="35" pageNumber="36">Pachytrachis frater</emphasis>
|
|||
|
|
(Brunner, 1882)
|
|||
|
|
</taxonomicName>
|
|||
|
|
female from Moraċa valley (CW). – A, stridulatory file; B–D, titillators in different magnification (B, oblique lateral view; C, dorsal view; D, do. with baso-lateral sclerites); E, male abdominal apex with cerci in lateral view; F, do. in dorsal view; G, I, M, female subgenital plate in ventral view; H, J, L, do. in lateral view; K, N, ovipositor.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF4BA9BBF667FFC7FF12F83FFE8DF807" httpUri="https://zenodo.org/record/197249/files/figure.png" pageId="36" pageNumber="37" targetBox="[334,1251,198,1908]" targetPageId="36">
|
|||
|
|
<paragraph id="8B8BF933F667FFC7FF12F83FFE8DF807" blockId="36.[151,1436,1927,2022]" pageId="36" pageNumber="37">
|
|||
|
|
<emphasis id="B9402521F667FFC7FF12F83FFE9AF87E" bold="true" box="[151,296,1927,1951]" pageId="36" pageNumber="37">FIGURE 19.</emphasis>
|
|||
|
|
Oscillograms for comparison of calling songs of
|
|||
|
|
<taxonomicName id="4C3482B0F667FFC7FCC4F830FC7DF87E" box="[833,975,1928,1951]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9402521F667FFC7FCC4F830FC7DF87E" box="[833,975,1928,1951]" italics="true" pageId="36" pageNumber="37">Pachytrachis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species: A–B,
|
|||
|
|
<taxonomicName id="4C3482B0F667FFC7FBF1F830FA25F87E" authority="Brunner, 1861" authorityName="Brunner" authorityYear="1861" box="[1140,1431,1928,1951]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gracilis">
|
|||
|
|
<emphasis id="B9402521F667FFC7FBF1F830FB52F87E" box="[1140,1248,1928,1951]" italics="true" pageId="36" pageNumber="37">P. gracilis</emphasis>
|
|||
|
|
(Brunner, 1861)
|
|||
|
|
</taxonomicName>
|
|||
|
|
; C–D,
|
|||
|
|
<taxonomicName id="4C3482B0F667FFC7FF5DF812FEFDF820" box="[216,335,1962,1985]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="striolatu">
|
|||
|
|
<emphasis id="B9402521F667FFC7FF5DF812FEFDF820" box="[216,335,1962,1985]" italics="true" pageId="36" pageNumber="37">P. striolatu</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
s (Fieber, 1853); E–F,
|
|||
|
|
<emphasis id="B9402521F667FFC7FDC6F812FD02F820" box="[579,688,1962,1985]" italics="true" pageId="36" pageNumber="37">P. t u m id u s</emphasis>
|
|||
|
|
<emphasis id="B9402521F667FFC7FD3EF812FD4AF823" bold="true" box="[699,760,1962,1986]" pageId="36" pageNumber="37">sp. n.</emphasis>
|
|||
|
|
— A, C, E, series of syllables; B, D, F, one syllable in higher time resolution.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C32EAAB8F666FFC6FF43FF2FFC00FE8D" pageId="37" pageNumber="38" type="etymology">
|
|||
|
|
<paragraph id="8B8BF933F666FFC6FF43FF2FFBCEFF36" blockId="37.[151,1436,151,364]" pageId="37" pageNumber="38">
|
|||
|
|
<emphasis id="B9402521F666FFC6FF43FF2FFEE1FF50" bold="true" box="[198,339,151,177]" pageId="37" pageNumber="38">Etymology.</emphasis>
|
|||
|
|
Named for the internal swelling on the male cercus; from Latin tumid = swollen.
|
|||
|
|
<emphasis id="B9402521F666FFC6FF43FF05FED5FF36" bold="true" box="[198,359,189,215]" pageId="37" pageNumber="38">Distribution.</emphasis>
|
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Mountains of western
|
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<collectingCountry id="F323B9A3F666FFC6FDF0FF05FCB5FF36" box="[629,775,189,215]" name="Serbia" pageId="37" pageNumber="38">Montenegro</collectingCountry>
|
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, possibly extending into
|
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|
|
<collectingCountry id="F323B9A3F666FFC6FBAEFF05FBCAFF36" box="[1067,1144,189,215]" name="Bosnia and Herzegovina" pageId="37" pageNumber="38">Bosna</collectingCountry>
|
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|
|
.
|
|||
|
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</paragraph>
|
|||
|
|
<paragraph id="8B8BF933F666FFC6FF43FF5AFC00FE8D" blockId="37.[151,1436,151,364]" pageId="37" pageNumber="38">
|
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|
<emphasis id="B9402521F666FFC6FF43FF5AFEE9FF1D" bold="true" box="[198,347,226,252]" pageId="37" pageNumber="38">Stridulation</emphasis>
|
|||
|
|
(
|
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|
|
<figureCitation id="130FE5B6F666FFC6FEE9FF5AFE71FF1D" box="[364,451,226,252]" captionStart="FIGURE 19" captionStartId="36.[151,255,1927,1951]" captionTargetBox="[334,1251,198,1908]" captionTargetId="figure@36.[324,1263,194,1914]" captionTargetPageId="36" captionText="FIGURE 19. Oscillograms for comparison of calling songs of Pachytrachis species: A – B, P. gracilis (Brunner, 1861); C – D, P. striolatu s (Fieber, 1853); E – F, P. t u m id u s sp. n. — A, C, E, series of syllables; B, D, F, one syllable in higher time resolution." httpUri="https://zenodo.org/record/197249/files/figure.png" pageId="37" pageNumber="38">Fig. 19</figureCitation>
|
|||
|
|
): The single male collected produced echemes with 3–4 syllables. In contrast, males of
|
|||
|
|
<taxonomicName id="4C3482B0F666FFC6FF3DFEB0FE10FEC0" box="[184,418,264,289]" class="Insecta" family="Tettigoniidae" genus="Pachytrachis" kingdom="Animalia" order="Orthoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="frater">
|
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|
|
<emphasis id="B9402521F666FFC6FF3DFEB0FE10FEC0" box="[184,418,264,289]" italics="true" pageId="37" pageNumber="38">Pachytrachis frater</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
collected near the Adriatic coast produced monosyllabic songs
|
|||
|
|
<bibRefCitation id="EFA584C2F666FFC6FB21FEB0FAF8FEC3" author="Heller" box="[1188,1354,264,290]" pageId="37" pageNumber="38" refString="Heller, K. G. (1988). Bioakustik der europaischen Laubheuschrecken. Verlag Josef Margraf, Weikersheim; 1 - 358." type="book chapter" year="1988">Heller (1988)</bibRefCitation>
|
|||
|
|
. In the new species, the echemes are repeated at intervals of 1–4 s. Stridulation is mainly in the ultrasonic range with a maximum at about 30 kHz (determined by an ultrasonic detector).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|