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<document id="F15796873A0820CF7C883557CAAA7DE7" ID-DOI="10.5281/zenodo.190342" ID-GBIF-Dataset="901a3155-73ef-489d-941b-050044f8b3d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="190342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461112263521" checkinUser="plazi" docAuthor="Praschag, Peter, Holloway, Rohan, Georges, Arthur, Päckert, Martin, Hundsdörfer, Anna K. &amp; Fritz, Uwe" docDate="2009" docId="03FF6A4BFFDE823FFF6DB70EFF2F7A7E" docLanguage="en" docName="zt02233p068.pdf" docOrigin="Zootaxa 2233" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Batagur affinis subsp. edwardmolli Praschag, Holloway, Georges, Päckert, Hundsdörfer &amp; Fritz, 2009, subsp. nov." docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFC61233FFD98237FFFAB13EFFD8784A" masterDocTitle="A new subspecies of Batagur affinis (Cantor, 1847), one of the worlds most critically endangered chelonians (Testudines: Geoemydidae)" masterLastPageNumber="68" masterPageNumber="57" pageNumber="64" updateTime="1698594356309" updateUser="plazi">
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<mods:title id="3811DAD20596D457E8C2035F29D83D18">A new subspecies of Batagur affinis (Cantor, 1847), one of the worlds most critically endangered chelonians (Testudines: Geoemydidae)</mods:title>
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<mods:namePart id="7BAE28DEBFF9DEEC67263B587FD84CFD">Praschag, Peter</mods:namePart>
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<mods:namePart id="B5D2CFD73B8FE6227DF99FE6AF4E3C26">Holloway, Rohan</mods:namePart>
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<mods:namePart id="15C017DBA0ABC2CB4B4286DC656BB028">Georges, Arthur</mods:namePart>
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<mods:namePart id="5F2DEED1FAEC7EFB78F8A081AE5B69D4">Päckert, Martin</mods:namePart>
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<mods:namePart id="47ED85C447F0B73D8B51C912F59D4788">Hundsdörfer, Anna K.</mods:namePart>
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<mods:namePart id="F298474A6B127CC31C08C3403B5ED885">Fritz, Uwe</mods:namePart>
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<mods:date id="5EA00ECEDA7B6DF2FDC6737F0EFEB2BF">2009</mods:date>
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<subSubSection id="C34C88D6FFDE8230FF6DB70EFDA47E00" box="[151,636,1584,1610]" pageId="7" pageNumber="64" type="nomenclature">
<paragraph id="8BE9DB5DFFDE8230FF6DB70EFDA47E00" blockId="7.[151,636,1584,1610]" box="[151,636,1584,1610]" pageId="7" pageNumber="64">
<heading id="D0A16C31FFDE8230FF6DB70EFDA47E00" bold="true" box="[151,636,1584,1610]" fontSize="11" level="1" pageId="7" pageNumber="64" reason="1">
<emphasis id="B922074FFFDE8230FF6DB70EFDA47E00" bold="true" box="[151,636,1584,1610]" pageId="7" pageNumber="64">
<taxonomicName id="4C56A0DEFFDE8230FF6DB70EFE347E00" box="[151,492,1584,1610]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="64" phylum="Chordata" rank="subSpecies" species="affinis" status="subsp. nov." subSpecies="edwardmolli">
<emphasis id="B922074FFFDE8230FF6DB70EFE347E00" bold="true" box="[151,492,1584,1610]" italics="true" pageId="7" pageNumber="64">Batagur affinis edwardmolli</emphasis>
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<taxonomicNameLabel id="A211BA34FFDE8230FE09B70EFDA47E00" box="[499,636,1584,1610]" pageId="7" pageNumber="64" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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<subSubSection id="C34C88D6FFDE8230FF6DB747FAF47F64" pageId="7" pageNumber="64" type="materials_examined">
<paragraph id="8BE9DB5DFFDE8230FF6DB747FAB87EF0" blockId="7.[151,1437,1657,2032]" pageId="7" pageNumber="64">
<emphasis id="B922074FFFDE8230FF6DB747FEC87ED9" bold="true" box="[151,272,1657,1683]" pageId="7" pageNumber="64">
<typeStatus id="54ED65FFFFDE8230FF6DB747FED37ED9" box="[151,267,1657,1683]" pageId="7" pageNumber="64" type="holotype">Holotype</typeStatus>
.
</emphasis>
Natural History Museum Vienna,
<collectionCode id="ED474398FFDE8230FD5EB744FD377EDE" box="[676,751,1658,1684]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="64">NMW</collectionCode>
38903, juvenile in alcohol (hatched and died in captivity), Sre Ambel River system, Koh Kong Province,
<collectingCountry id="F3419BCDFFDE8230FD44B79EFCE27EF0" box="[702,826,1696,1722]" name="Cambodia" pageId="7" pageNumber="64">Cambodia</collectingCountry>
; don. Head Start Centre Sre Ambel,
<date id="FFE8FD9DFFDE8230FB1CB79EFA847EF0" box="[1254,1372,1696,1722]" pageId="7" pageNumber="64" value="2009-07">July 2009</date>
.
</paragraph>
<paragraph id="8BE9DB5DFFDE8230FF3FB7F9FAF47F64" blockId="7.[151,1437,1657,2032]" pageId="7" pageNumber="64">
<emphasis id="B922074FFFDE8230FF3FB7F9FE927EAB" bold="true" box="[197,330,1735,1761]" pageId="7" pageNumber="64">
<typeStatus id="54ED65FFFFDE8230FF3FB7F9FE9D7EAB" box="[197,325,1735,1761]" pageId="7" pageNumber="64" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Museum of Zoology Dresden,
<collectionCode id="ED474398FFDE8230FD44B7F9FD277EAB" box="[702,767,1735,1761]" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="7" pageNumber="64">MTD</collectionCode>
47538, juvenile in alcohol, same data as
<typeStatus id="54ED65FFFFDE8230FB13B7F9FA8C7EAB" box="[1257,1364,1735,1761]" pageId="7" pageNumber="64" type="holotype">holotype</typeStatus>
. Field Museum of Natural History, Chicago,
<collectionCode id="ED474398FFDE8230FDA2B7D0FD757F42" LSID="urn:lsid:biocol.org:col:34795" box="[600,685,1774,1800]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="7" pageNumber="64">FMNH</collectionCode>
224093 (ex EOM 2398; specimen figured on Plate
<collectionCode id="ED474398FFDE8230FAF1B7D0FAE57F42" box="[1291,1341,1774,1800]" pageId="7" pageNumber="64">IIIA</collectionCode>
in Moll 1980), broken shell of adult female, Terengganu River,
<collectingCountry id="F3419BCDFFDE8230FCDBB62AFC487F64" box="[801,912,1812,1838]" name="Malaysia" pageId="7" pageNumber="64">Malaysia</collectingCountry>
; leg. Edward
<collectingCountry id="F3419BCDFFDE8230FBCAB62AFB907F64" box="[1072,1096,1812,1838]" name="Iceland" pageId="7" pageNumber="64">O</collectingCountry>
. Moll,
<date id="FFE8FD9DFFDE8230FB66B62AFAF07F64" box="[1180,1320,1812,1838]" pageId="7" pageNumber="64" value="1976-07-07">7 July 1976</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C88D6FFDE8230FF3FB604FE9F7FE8" pageId="7" pageNumber="64" type="etymology">
<paragraph id="8BE9DB5DFFDE8230FF3FB604FE9F7FE8" blockId="7.[151,1437,1657,2032]" pageId="7" pageNumber="64">
<emphasis id="B922074FFFDE8230FF3FB604FE827F1E" bold="true" box="[197,346,1850,1876]" pageId="7" pageNumber="64">Etymology.</emphasis>
The new subspecies is named in recognition of Professor Edward
<collectingCountry id="F3419BCDFFDE8230FB52B605FB1A7F1F" box="[1192,1218,1851,1877]" name="Iceland" pageId="7" pageNumber="64">O</collectingCountry>
. Moll, one of the foremost experts on river turtles, who substantially contributed to the knowledge of
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<emphasis id="B922074FFFDE8230FB75B65CFA9C7F31" box="[1167,1348,1890,1915]" italics="true" pageId="7" pageNumber="64">Batagur affinis</emphasis>
</taxonomicName>
and its natural history.
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<emphasis id="B922074FFFDE8230FF3FB691FE947F83" bold="true" box="[197,332,1967,1993]" pageId="7" pageNumber="64">Diagnosis.</emphasis>
Adults differ from nominotypical subspecies of
<taxonomicName id="4C56A0DEFFDE8230FC4AB691FBB47F82" box="[944,1132,1967,1992]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="7" pageNumber="64" phylum="Chordata" rank="species" species="affinis">
<emphasis id="B922074FFFDE8230FC4AB691FBB47F82" box="[944,1132,1967,1992]" italics="true" pageId="7" pageNumber="64">Batagur affinis</emphasis>
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by their distinctly more elongated head with upturned snout; males with chocolate brown to almost black head (east coast of peninsular
<collectingCountry id="F3419BCDFFD1823FFEE3B1A6FE5E78F8" box="[281,390,152,178]" name="Malaysia" pageId="8" pageNumber="65">Malaysia</collectingCountry>
) or sometimes rusty brown to reddish head (Sre Ambel River system,
<collectingCountry id="F3419BCDFFD1823FFB32B1A6FA9D78F8" box="[1224,1349,152,178]" name="Cambodia" pageId="8" pageNumber="65">Cambodia</collectingCountry>
), edges of mouth orange; iris golden or bright yellow. Females and juveniles with conspicuous whitish grey to silvery blotches in temporal and parietal region; hatchlings with distally yellow marginal scutes. For corresponding characters of
<taxonomicName id="4C56A0DEFFD1823FFECEB032FE64796F" box="[308,444,268,293]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="8" pageNumber="65" phylum="Chordata" rank="subSpecies" species="affinis" subSpecies="affinis">
<emphasis id="B922074FFFD1823FFECEB032FE64796F" box="[308,444,268,293]" italics="true" pageId="8" pageNumber="65">B. a. affinis</emphasis>
</taxonomicName>
, see
<tableCitation id="C6D4EEE6FFD1823FFE0DB032FD88796C" box="[503,592,268,294]" captionStart="TABLE 2" captionStartId="7.[151,239,267,291]" captionText="TABLE 2. External morphology and current distribution ranges of estuarine Batagur based on personal observations and" pageId="8" pageNumber="65">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C88D6FFD1823FFF3FB00CFF2F7A7E" pageId="8" pageNumber="65" type="description">
<paragraph id="8BE9DB5DFFD1823FFF3FB00CFB4579AD" blockId="8.[151,1437,152,564]" pageId="8" pageNumber="65">
<emphasis id="B922074FFFD1823FFF3FB00CFDD87906" bold="true" box="[197,512,306,332]" pageId="8" pageNumber="65">
Description of
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.
</emphasis>
Specimen slightly macerated; some epidermal scutes detached from shell. Carapace roundish when viewed from above, with weakly serrated central and posterior marginal scutes; medial keel distinct, with posteriorly directed, slightly pointed spines. Plastron anteriorly truncated, posteriorly with anal notch. Straight line carapace length approximately
<quantity id="4CAE76B8FFD1823FFBECB098FBB7798A" box="[1046,1135,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="65" unit="mm" value="86.0">86 mm</quantity>
, carapace width
<quantity id="4CAE76B8FFD1823FFAC4B098FA40798A" box="[1342,1432,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="8.4" pageId="8" pageNumber="65" unit="mm" value="84.0">84 mm</quantity>
; medial plastron length
<quantity id="4CAE76B8FFD1823FFE5EB0F3FE2179AD" box="[420,505,461,487]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="65" unit="mm" value="74.0">74 mm</quantity>
, maximum plastron length (to tips of anal scutes)
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.
</paragraph>
<paragraph id="8BE9DB5DFFD1823FFF3FB0CDFF2F7A7E" blockId="8.[151,1437,152,564]" pageId="8" pageNumber="65">
<emphasis id="B922074FFFD1823FFF3FB0CDFEF87A47" bold="true" box="[197,288,499,525]" pageId="8" pageNumber="65">Range:</emphasis>
East coast of peninsular
<collectingCountry id="F3419BCDFFD1823FFDA9B0CAFD197A44" box="[595,705,500,526]" name="Malaysia" pageId="8" pageNumber="65">Malaysia</collectingCountry>
and adjacent
<collectingCountry id="F3419BCDFFD1823FFC91B0CAFC027A44" box="[875,986,500,526]" name="Thailand" pageId="8" pageNumber="65">Thailand</collectingCountry>
; Sre Ambel River system,
<collectingCountry id="F3419BCDFFD1823FFADAB0CAFA447A44" box="[1312,1436,500,526]" name="Cambodia" pageId="8" pageNumber="65">Cambodia</collectingCountry>
(
<figureCitation id="136DC7D8FFD1823FFF65B324FF3F7A7E" box="[159,231,538,564]" captionStart="FIGURE 5" captionStartId="8.[151,255,1910,1934]" captionTargetBox="[166,1425,622,1881]" captionTargetId="figure@8.[166,1426,622,1882]" captionTargetPageId="8" captionText="FIGURE 5. Historical distribution of Batagur affinis affinis, B. a. edwardmolli, and B. baska (modified from Praschag et al. 2008). Note that the species are extirpated in most of their former ranges." httpUri="https://zenodo.org/record/190347/files/figure.png" pageId="8" pageNumber="65">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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