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<document id="C3F0C4AA08987E724CAB1ADA9350D8D1" ID-CLB-Dataset="29118" ID-DOI="10.11646/zootaxa.4455.1.1" ID-GBIF-Dataset="1586b06b-82e0-4be6-837e-0d11ca1a5214" ID-ISSN="1175-5326" ID-Zenodo-Dep="1456911" ID-ZooBank="70E92EDF-E11B-40A7-9573-0AC9F10E623E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539260167405" checkinUser="plazi" docAuthor="Neal, Lenka, Brasier, Madeleine J. &amp; Wiklund, Helena" docDate="2018" docId="03BDA104A3206954FF64FD91FF7625C6" docLanguage="en" docName="zootaxa.4455.1.1.pdf" docOrigin="Zootaxa 4455 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macellicephala mirabilis" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FF84D97CA326695DFFF3FFADFFAB244E" masterDocTitle="Six new species of Macellicephala (Annelida: Polynoidae) from the Southern Ocean and south Atlantic with re-description of type species" masterLastPageNumber="34" masterPageNumber="1" pageNumber="7" updateTime="1698575057705" updateUser="ExternalLinkService">
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<mods:title id="C329F1106EB0B9C82B2AA5F25E24831B">Six new species of Macellicephala (Annelida: Polynoidae) from the Southern Ocean and south Atlantic with re-description of type species</mods:title>
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<mods:namePart id="56DAD70DC958B0DFF35F94D0442B4995">Neal, Lenka</mods:namePart>
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<mods:namePart id="46A170E08AC9B61C22E893291BF75776">Brasier, Madeleine J.</mods:namePart>
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<mods:namePart id="AE9F8364F1915B367980004739D92AA4">Wiklund, Helena</mods:namePart>
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<treatment id="03BDA104A3206954FF64FD91FF7625C6" ID-DOI="http://doi.org/10.5281/zenodo.5994473" ID-GBIF-Taxon="148691072" ID-Zenodo-Dep="5994473" LSID="urn:lsid:plazi:treatment:03BDA104A3206954FF64FD91FF7625C6" httpUri="http://treatment.plazi.org/id/03BDA104A3206954FF64FD91FF7625C6" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
<subSubSection id="C30E4399A320695BFF64FD91FDB32636" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BAB1012A320695BFF64FD91FD0B2619" blockId="6.[151,672,572,632]" box="[151,672,572,599]" pageId="6" pageNumber="7">
<heading id="D0E3A77EA320695BFF64FD91FD0B2619" bold="true" box="[151,672,572,599]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="4C146B91A320695BFF64FD91FD0B2619" ID-CoL="3WXTT" authority="(McIntosh, 1885)" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[151,672,572,599]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A320695BFF64FD91FD0B2619" bold="true" box="[151,672,572,599]" pageId="6" pageNumber="7">
<emphasis id="B960CC00A320695BFF64FD91FE682618" bold="true" box="[151,451,572,598]" italics="true" pageId="6" pageNumber="7">Macellicephala mirabilis</emphasis>
(
<bibRefCitation id="EF856DE3A320695BFE21FD90FD3C2619" box="[466,663,573,599]" pageId="6" pageNumber="7" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh, 1885</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAB1012A320695BFF64FDF2FDB32636" blockId="6.[151,672,572,632]" box="[151,536,607,632]" pageId="6" pageNumber="7">
(
<figureCitation id="132F0C97A320695BFF6DFDF2FEBE2636" box="[158,277,607,632]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="6" pageNumber="7">Figure 2A</figureCitation>
,
<figureCitation id="132F0C97A320695BFED3FDF2FE202636" box="[288,395,607,632]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Figures 3</figureCitation>
,
<figureCitation id="132F0C97A320695BFE64FDCDFE032636" box="[407,424,608,632]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="6" pageNumber="7">4</figureCitation>
;
<tableCitation id="C69625A9A320695BFE47FDF2FDA12636" box="[436,522,607,632]" captionStart="TABLE 2" captionStartId="12.[147,221,540,559]" captionText="TABLE 2. Comparison of characters of all known Macellicephala species based on the examination of representatives of species described in this study and literature survey for other species." pageId="6" pageNumber="7">Table 2</tableCitation>
.)
</paragraph>
</subSubSection>
<subSubSection id="C30E4399A320695BFF64FD05FD072776" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BAB1012A320695BFF64FD05FC722692" blockId="6.[151,1324,680,824]" pageId="6" pageNumber="7">
<taxonomicName id="4C146B91A320695BFF64FD05FD5426F0" ID-CoL="6VTR7" authority="in Mclntosh (1885): 121" authorityName="in Mclntosh (1885)" authorityPageNumber="121" authorityYear="1885" box="[151,767,680,702]" class="Polychaeta" family="Polynoidae" genus="Polynoe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis" subGenus="Macellicephala">
<emphasis id="B960CC00A320695BFF64FD05FDAA26F0" box="[151,513,680,702]" italics="true" pageId="6" pageNumber="7">Polynoe (Macellicephala) mirabilis</emphasis>
in Mclntosh (1885): 121
</taxonomicName>
, p.16:
<figureCitation id="132F0C97A320695BFCB7FD05FCD426F0" box="[836,895,680,702]" captionStart="FIGURE 1" captionStartId="5.[151,250,1556,1578]" captionText="FIGURE 1. Molecular phylogenetic tree showing only Polynoidae part of the tree (outgroups not shown) of combined 18S, 16S and COI data. Node values are bootstrap support (BS) from the Maximum Likelihood analyses and posterior probability values (PP) from the Bayesian analyses, expressed as BS/PP. Purple colour indicates members of subfamily Macellicephalinae, while green colour shows Branchinotogluminae. The two Macellicephala clades recovered in the analyses are labelled with A and B." pageId="6" pageNumber="7">Fig. 1</figureCitation>
, p.
<figureCitation id="132F0C97A320695BFC57FD05FC7F26F0" box="[932,980,680,702]" captionStart="FIGURE 12" captionStartId="21.[151,250,1213,1235]" captionTargetBox="[152,1435,193,1192]" captionTargetId="figure@21.[152,1435,193,1192]" captionTargetPageId="21" captionText="FIGURE 12. Macellicephala brenesorum sp. nov. (holotype, NHMUK.2018.830): AC) mid body parapodium; DE) detail of neurochaetae. Scale bars: A) 500 µm; B) 250 µm; C) 250 µm; D) 150 µm; E)100 µm." httpUri="https://zenodo.org/record/1456933/files/figure.png" pageId="6" pageNumber="7">12A</figureCitation>
:
<figureCitation id="132F0C97A320695BFC2EFD05FBE326F0" box="[989,1096,680,702]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="18.[151,250,1844,1866]" captionStartId-1="19.[151,250,1235,1257]" captionStartId-2="20.[151,250,1905,1927]" captionTargetBox-0="[151,1435,735,1822]" captionTargetBox-1="[152,1435,193,1213]" captionTargetBox-2="[170,1416,944,1879]" captionTargetId-0="figure@18.[151,1435,735,1822]" captionTargetId-1="figure@19.[152,1435,193,1213]" captionTargetId-2="figure@20.[170,1416,944,1879]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionTargetPageId-2="20" captionText-0="FIGURE 9. Macellicephala patersoni sp. nov. (holotype, NHMUK.2018.1009): A) specimen in dorsal view; B) specimen in ventral view; C) detail of prostomium and anterior end; D) detail of everted proboscis; E) detail of pygidium in dorsal view, showing two large loops; F) detail of pygidium in ventral view. Scale bars: AB) 5 mm; C) 1.5 mm; D) 2.5 mm; E) 1.5mm." captionText-1="FIGURE 10. Macellicephala patersoni sp. nov. (holotype, NHMUK.2018.1009): A) mid body parapodium; B) notochaetae; C) detail of notochaetae; DH) neurochaetae. Scale bars: A) 500 µm; B) 300 µm; CE) 100 µm; FG) 50 µm and H) 15 µm." captionText-2="FIGURE 11. Macellicephala brenesorum sp. nov. (holotype, NHMUK.2018.830): A) specimen in dorsal view; B) specimen in ventral view; C) detail of prostomium and anterior end; D) detail of proboscis; E) detail of pygidium in dorsal view; F) detail of pygidium in ventral view. Scale bars: AE) 2.5 mm; F) 1.250 mm." httpUri-0="https://zenodo.org/record/1456927/files/figure.png" httpUri-1="https://zenodo.org/record/1456929/files/figure.png" httpUri-2="https://zenodo.org/record/1456931/files/figure.png" pageId="6" pageNumber="7">Figs. 911</figureCitation>
[original description].
<taxonomicName id="4C146B91A320695BFF64FD6BFD232692" ID-CoL="3WXTT" authority="in Pettibone (1976): 10" authorityName="in Pettibone" authorityPageNumber="10" authorityYear="1976" box="[151,648,710,732]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A320695BFF64FD6BFE3C2692" box="[151,407,710,732]" italics="true" pageId="6" pageNumber="7">Macellicephala mirabilis</emphasis>
in
<bibRefCitation id="EF856DE3A320695BFE45FD6BFDCF2692" box="[438,612,710,732]" pageId="6" pageNumber="7" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
: 10
</taxonomicName>
, p. 11:
<figureCitation id="132F0C97A320695BFD21FD6BFC862692" box="[722,813,710,732]" captionStart="FIGURE 1" captionStartId="5.[151,250,1556,1578]" captionText="FIGURE 1. Molecular phylogenetic tree showing only Polynoidae part of the tree (outgroups not shown) of combined 18S, 16S and COI data. Node values are bootstrap support (BS) from the Maximum Likelihood analyses and posterior probability values (PP) from the Bayesian analyses, expressed as BS/PP. Purple colour indicates members of subfamily Macellicephalinae, while green colour shows Branchinotogluminae. The two Macellicephala clades recovered in the analyses are labelled with A and B." pageId="6" pageNumber="7">Fig. 1af</figureCitation>
[re-description].
</paragraph>
<paragraph id="8BAB1012A320695BFF64FD4BFC8F26B5" blockId="6.[151,1324,680,824]" box="[151,804,741,763]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB04373CA320695BFF64FD4BFC8F26B5" box="[151,804,741,763]" pageId="6" pageNumber="7">
Not
<taxonomicName id="4C146B91A320695BFF30FD48FC8B26B5" ID-CoL="6VTR7" authority="in Mclntosh (1905): 59" authorityName="in Mclntosh" authorityPageNumber="59" authorityYear="1905" box="[195,800,741,763]" class="Polychaeta" family="Polynoidae" genus="Polynoe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis" subGenus="Macellicephala">
<emphasis id="B960CC00A320695BFF30FD48FD8626B5" box="[195,557,741,763]" italics="true" pageId="6" pageNumber="7">Polynoe (Macellicephala) mirabilis</emphasis>
in
<treatmentCitation id="0AB53603A320695BFDBEFD48FC8B26B5" author="" box="[589,800,741,763]" page="59" pageId="6" pageNumber="7">
<bibRefCitation id="EF856DE3A320695BFDBEFD48FD5126B5" box="[589,762,741,763]" pageId="6" pageNumber="7" refId="ref17672" refString="Mclntosh, W. C. (1905) Marine Annelids (Polychaeta) of South Africa, Part 2. Marine Investigations in South Africa, 3, 57 - 92." type="journal article">Mclntosh (1905)</bibRefCitation>
: 59
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.
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</paragraph>
<paragraph id="8BAB1012A320695BFF64FCA8FCCF2754" blockId="6.[151,1324,680,824]" box="[151,868,772,794]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB04373CA320695BFF64FCA8FCCF2754" box="[151,868,772,794]" pageId="6" pageNumber="7">
Not
<taxonomicName id="4C146B91A320695BFF30FCA9FD1F2754" ID-CoL="3WXTT" authority="in Pettibone (1976): 10" authorityName="in Pettibone" authorityPageNumber="10" authorityYear="1976" box="[195,692,772,794]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A320695BFF30FCA9FE692754" box="[195,450,772,794]" italics="true" pageId="6" pageNumber="7">Macellicephala mirabilis</emphasis>
in
<treatmentCitation id="0AB53603A320695BFE10FCA9FD1F2754" author="" box="[483,692,772,794]" page="10" pageId="6" pageNumber="7">
<bibRefCitation id="EF856DE3A320695BFE10FCA9FD3B2754" box="[483,656,772,794]" pageId="6" pageNumber="7" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
: 10
</treatmentCitation>
</taxonomicName>
, p. 12:
<figureCitation id="132F0C97A320695BFD0CFCA9FCCA2754" box="[767,865,772,794]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Fig. 3ae</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB1012A320695BFF64FC8EFD072776" blockId="6.[151,1324,680,824]" box="[151,684,802,824]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB04373CA320695BFF64FC8EFD072776" box="[151,684,802,824]" pageId="6" pageNumber="7">
Not
<taxonomicName id="4C146B91A320695BFF30FC8FFD032776" ID-CoL="3WXTT" authority="in Monro (1936): 100" authorityName="in Monro" authorityPageNumber="100" authorityYear="1936" box="[195,680,802,824]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="7" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A320695BFF30FC8FFE692776" box="[195,450,802,824]" italics="true" pageId="6" pageNumber="7">Macellicephala mirabilis</emphasis>
in
<treatmentCitation id="0AB53603A320695BFE10FC8EFD032776" author="" box="[483,680,802,824]" page="100" pageId="6" pageNumber="7">
<bibRefCitation id="EF856DE3A320695BFE10FC8EFDDE2776" box="[483,629,802,824]" pageId="6" pageNumber="7" refId="ref17794" refString="Monro, C. C. A. (1936) Polychaete Worms, II. Discovery Reports, 12, 59 - 198." type="journal article">Monro (1936)</bibRefCitation>
: 100
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30E4399A3206955FF64FCC8FC0B2260" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BAB1012A320695BFF64FCC8FB2E27EC" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
<materialsCitation id="3B7C1A4FA320695BFF64FCC8FB2E27EC" ID-GBIF-Occurrence="1917073255" collectingDate="1874-07-10" collectionCode="BMNH" country="New Zealand" county="Challenger station" elevation="1280" latitude="-37.566666" location="Material" longLatPrecision="1180" longitude="179.36667" municipality="North Island" pageId="6" pageNumber="7" specimenCode="BMNH 1885" specimenCount="1" typeStatus="holotype">
<emphasis id="B960CC00A320695BFF64FCC8FE2B2730" bold="true" box="[151,384,869,894]" pageId="6" pageNumber="7">
<location id="8ECB46C9A320695BFF64FCC8FF542730" LSID="urn:lsid:plazi:treatment:03BDA104A3206954FF64FD91FF7625C6:8ECB46C9A320695BFF64FCC8FF542730" box="[151,255,869,894]" country="New Zealand" county="Challenger station" latitude="-37.566666" longLatPrecision="1180" longitude="179.36667" municipality="North Island" name="Material" pageId="6" pageNumber="7">Material</location>
examined.
</emphasis>
<typeStatus id="54AFAEB0A320695BFE74FCC8FE442730" box="[391,495,869,894]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
of
<bibRefCitation id="EF856DE3A320695BFDE6FCC8FD7F2730" box="[533,724,869,894]" pageId="6" pageNumber="7" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
,
<specimenCode id="DBB2B869A320695BFD13FCCBFCDC2730" box="[736,887,869,894]" collectionCode="BMNH" pageId="6" pageNumber="7">BMNH 1885</specimenCode>
: 12: 1: 100,
<collectingCountry id="F3035082A320695BFBF1FCCBFB322730" box="[1026,1177,869,894]" name="New Zealand" pageId="6" pageNumber="7">New Zealand</collectingCountry>
(off
<collectingMunicipality id="6BCF8A68A320695BFB22FCC8FACB2730" box="[1233,1376,869,894]" pageId="6" pageNumber="7">North Island</collectingMunicipality>
, blue mud),
<collectingCounty id="62CA689EA320695BFF13FC24FE0627EC" box="[224,429,905,930]" pageId="6" pageNumber="7">Challenger station</collectingCounty>
169, collected on
<collectingDate id="EFEECF3AA320695BFD88FC27FD5627EC" box="[635,765,906,930]" pageId="6" pageNumber="7" value="1874-07-10">
<date id="FFAA36D2A320695BFD88FC27FD5627EC" box="[635,765,906,930]" pageId="6" pageNumber="7" value="1874-07-10">10/07/1874</date>
</collectingDate>
,
<geoCoordinate id="EE2076D5A320695BFCFBFC27FCC927EC" box="[776,866,906,930]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="925" value="-37.566666">37°34'S</geoCoordinate>
,
<geoCoordinate id="EE2076D5A320695BFC9DFC27FC7327EC" box="[878,984,906,930]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="925" value="179.36667">179°22'E</geoCoordinate>
,
<elevation id="0039F721A320695BFC10FC27FB9327EF" box="[995,1080,906,930]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="6" pageNumber="7" unit="m" value="1280.0">
<quantity id="4CECBDF7A320695BFC10FC27FB9327EF" box="[995,1080,906,930]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="6" pageNumber="7" unit="m" value="1280.0">1280 m</quantity>
</elevation>
depth.
</materialsCitation>
</paragraph>
<paragraph id="8BAB1012A320695BFF34FC00FC672040" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
<emphasis id="B960CC00A320695BFF34FC00FD932788" bold="true" box="[199,568,941,966]" pageId="6" pageNumber="7">Additional material examined.</emphasis>
Two specimens of McIntosh (1905),
<collectionCode id="ED0588D7A320695BFC18FC03FB94278B" box="[1003,1087,942,965]" pageId="6" pageNumber="7">BMNH</collectionCode>
1924: 7: 21: 12,
<collectingCountry id="F3035082A320695BFAF9FC00FA372788" box="[1290,1436,941,966]" name="South Africa" pageId="6" pageNumber="7">South Africa</collectingCountry>
(
<collectingRegion id="49D0DEF0A320695BFF6DFC7FFF7227A4" box="[158,217,978,1002]" country="South Africa" name="Western Cape" pageId="6" pageNumber="7">Cape</collectingRegion>
of Good Hope),
<quantity id="4CECBDF7A320695BFE6BFC7CFDBC27A7" box="[408,535,977,1002]" metricMagnitude="2" metricUnit="m" metricValue="7.2" metricValueMax="9.0" metricValueMin="5.4" pageId="6" pageNumber="7" unit="m" value="720.0" valueMax="900.0" valueMin="540.0">540900 m</quantity>
depth. Two specimens of
<bibRefCitation id="EF856DE3A320695BFCCFFC7FFC7727A4" box="[828,988,977,1002]" pageId="6" pageNumber="7" refId="ref17794" refString="Monro, C. C. A. (1936) Polychaete Worms, II. Discovery Reports, 12, 59 - 198." type="journal article">Monro (1936)</bibRefCitation>
,
<collectionCode id="ED0588D7A320695BFC14FC7FFB9027A7" box="[999,1083,978,1001]" pageId="6" pageNumber="7">BMNH</collectionCode>
1936: 2: 8: 488,
<collectingCountry id="F3035082A320695BFB09FC7CFA3727A4" box="[1274,1436,977,1002]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="7">South Georgia</collectingCountry>
(mouth of Stromness Harbour), Discovery station
<quantity id="4CECBDF7A320695BFD3BFC5BFC2B2043" box="[712,896,1013,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.6107749999999998" metricValueMax="1.78" metricValueMin="1.4415499999999999" pageId="6" pageNumber="7" unit="m" value="161.0775" valueMax="178.0" valueMin="144.155">
144, 155
<quantity id="4CECBDF7A320695BFCCAFC58FC2B2043" box="[825,896,1013,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="7" unit="m" value="178.0">178 m</quantity>
</quantity>
depth.
</paragraph>
<paragraph id="8BAB1012A320695BFF34FBB4FECA208C" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
<emphasis id="B960CC00A320695BFF34FBB4FED5207C" bold="true" box="[199,382,1049,1074]" pageId="6" pageNumber="7">Re-description.</emphasis>
<typeStatus id="54AFAEB0A320695BFE7BFBB4FE5B207C" box="[392,496,1049,1074]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
in relatively poor state, but many characters observable; specimen complete but now in two fragments (
<figureCitation id="132F0C97A320695BFE58FB90FD812018" box="[427,554,1085,1110]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Fig. 3A, B</figureCitation>
) as a result of previous examinations, chaetae mostly broken off, all elytra missing.
<typeStatus id="54AFAEB0A320695BFF0CFBCCFECC2034" box="[255,359,1121,1146]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
<quantity id="4CECBDF7A320695BFE82FBCCFE682037" box="[369,451,1121,1146]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="7" unit="mm" value="25.0">25 mm</quantity>
long,
<quantity id="4CECBDF7A320695BFDE3FBCCFDFF2037" box="[528,596,1121,1146]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="7" unit="mm" value="6.0">6 mm</quantity>
wide including parapodia and
<quantity id="4CECBDF7A320695BFC4EFBCFFBAA2037" box="[957,1025,1122,1146]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="mm" value="4.0">4 mm</quantity>
wide excluding parapodia; with 18 segments (segment 1 = tentacular segment); body dorsoventrally flattened, tapering anteriorly and posteriorly; yellow in alcohol.
</paragraph>
<paragraph id="8BAB1012A320695BFF34FB60FD8B21AC" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
Prostomium bilobed with deep anterior notch; prostomial lobes pronounced, greatly extended, with angular (truncated) anterior margins (
<figureCitation id="132F0C97A320695BFE1FFB5CFDE22144" box="[492,585,1265,1290]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
,
<figureCitation id="132F0C97A320695BFDAAFB5CFD1E2144" box="[601,693,1265,1290]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
). Frontal filaments now missing, but once present according to
<bibRefCitation id="EF856DE3A320695BFF64FAB8FEFE2160" box="[151,341,1301,1326]" pageId="6" pageNumber="7" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
as slender, elongated filaments, inserted internally on the margin of truncated prostomial lobes. Eyes absent. Facial tubercle consisting of two disconnected inflated pads, each medial to bases of palps. Ceratophore of median antenna very long (approaching anterior margin of prostomial lobes), cylindrical, inserted posteromedially on prostomium (near base of anterior notch) (
<figureCitation id="132F0C97A320695BFCABFA2CFC1921D4" box="[856,946,1409,1434]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
,
<figureCitation id="132F0C97A320695BFC4CFA2CFBBC21D4" box="[959,1047,1409,1434]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
); style of median antenna smooth, slender, tapering and long (extending to segment 5). Palps shortest and thickest of head appendages (extending to about segment 3), tapered, smooth.
</paragraph>
<paragraph id="8BAB1012A320695BFF34FA40FC99223C" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
Tentaculophores prominent, of equal size, cylindrical, inserted laterally to prostomium, achaetous; styles mostly missing, only style of a ventral cirrus present (on left side), smooth, very slender and long (longer than palps but shorter than style of median antenna) (
<figureCitation id="132F0C97A320695BFD35F998FC8A2200" box="[710,801,1589,1614]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
,
<figureCitation id="132F0C97A320695BFCDCF998FC222200" box="[815,905,1589,1614]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
). Pharynx not observed. Second segment with elytrophores, biramous parapodia, chaetae and ventral cirri.
</paragraph>
<paragraph id="8BAB1012A320695BFF34F9D0FE2822F4" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">Nine pairs of prominent, bulbous elytrophores, on segments 2, 4, 5, 7, 9, 11, 13, 15 and 17; all elytra missing. Dorsal ridges absent.</paragraph>
<paragraph id="8BAB1012A320695BFF34F968FD172394" blockId="6.[151,1437,869,2010]" pageId="6" pageNumber="7">
Parapodia biramous. Notopodia reduced to elongate acicular lobe, tip of notoacicula not penetrating epidermis (
<figureCitation id="132F0C97A320695BFF6CF944FF51234C" box="[159,250,1769,1794]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
). neuropodial large (
<figureCitation id="132F0C97A320695BFE12F944FD90234C" box="[481,571,1769,1794]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
), subtriangular and tapering to elongate acicular lobe, tip of neuroacicula not penetrating epidermis. Cirrigerous segments with prominent, bulbous dorsal cirrophores, inserted basally on notopodia; all dorsal styles now missing, but once present according to
<bibRefCitation id="EF856DE3A320695BFC1CF89CFB1B2304" box="[1007,1200,1841,1866]" pageId="6" pageNumber="7" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
as slender, smooth, cirriform appendages, far exceeding length of adjacent neuropodial; small, but distinct conical dorsal tubercles present. Ventral cirri smooth, tapering, present from segment 2; inserted basally on segment 2, where very long, about 1.5 times length of associated neuropodial lobe; inserted medially on subsequent segments, where short (not reaching the distal margin of neuropodial l lobe).
</paragraph>
<caption id="DF6B409AA321695AFF64F88DFDE423A0" httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1824,1846]" targetBox="[252,1336,206,1786]" targetPageId="7">
<paragraph id="8BAB1012A321695AFF64F88DFDE423A0" blockId="7.[151,1437,1824,2030]" pageId="7" pageNumber="8">
<emphasis id="B960CC00A321695AFF64F88DFEB82378" bold="true" box="[151,275,1824,1846]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A)
<taxonomicName id="4C146B91A321695AFA73F88CFF58231B" authorityName="in Monro" authorityYear="1936" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A321695AFA73F88CFF58231B" italics="true" pageId="7" pageNumber="8">M. mirabilis</emphasis>
</taxonomicName>
(holotype, BMNH: 1885: 12: 1: 100); B)
<taxonomicName id="4C146B91A321695AFD50F8EDFC9F231B" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[675,820,1855,1877]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="macintoshi" status="sp. nov.">
<emphasis id="B960CC00A321695AFD50F8EDFC9F231B" box="[675,820,1855,1877]" italics="true" pageId="7" pageNumber="8">M. macintoshi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFCCFF892FC26231B" box="[828,909,1855,1877]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFCCFF892FC26231B" bold="true" box="[828,909,1855,1877]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, BMNH: 1924: 7: 21: 12); C)
<taxonomicName id="4C146B91A321695AFAC2F8EDFA37231B" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[1329,1436,1856,1877]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="monroi" status="sp. nov.">
<emphasis id="B960CC00A321695AFAC2F8EDFA37231B" box="[1329,1436,1856,1877]" italics="true" pageId="7" pageNumber="8">M. monroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFF64F8F0FF43233D" box="[151,232,1885,1907]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFF64F8F0FF43233D" bold="true" box="[151,232,1885,1907]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D)
<taxonomicName id="4C146B91A321695AFA73F8F2FF7423DC" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="monroi" status="sp. nov.">
<emphasis id="B960CC00A321695AFA73F8F2FF7423DC" italics="true" pageId="7" pageNumber="8">M. monroi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFF1AF8D1FE9723DC" box="[233,316,1916,1938]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFF1AF8D1FE9723DC" bold="true" box="[233,316,1916,1938]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E)
<taxonomicName id="4C146B91A321695AFEABF831FDAA23FF" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[344,513,1947,1969]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="brenesorum" status="sp. nov.">
<emphasis id="B960CC00A321695AFEABF831FDAA23FF" box="[344,513,1947,1969]" italics="true" pageId="7" pageNumber="8">M. brenesorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFDE5F836FDDF23FF" box="[534,628,1947,1969]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFDE5F836FDDF23FF" bold="true" box="[534,628,1947,1969]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, NHMUK.2018.830); F)
<taxonomicName id="4C146B91A321695AFBE8F831FB0023FF" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[1051,1195,1948,1969]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="patersoni" status="sp. nov.">
<emphasis id="B960CC00A321695AFBE8F831FB0023FF" box="[1051,1195,1948,1969]" italics="true" pageId="7" pageNumber="8">M. patersoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFB33F836FAB423FF" box="[1216,1311,1947,1969]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFB33F836FAB423FF" bold="true" box="[1216,1311,1947,1969]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, NHMUK.2018.1009); G)
<taxonomicName id="4C146B91A321695AFE4BF817FD862381" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[440,557,1977,1999]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="gloveri" status="sp. nov.">
<emphasis id="B960CC00A321695AFE4BF817FD862381" box="[440,557,1977,1999]" italics="true" pageId="7" pageNumber="8">M. gloveri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFDB3F814FD362381" box="[576,669,1977,1999]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFDB3F814FD362381" bold="true" box="[576,669,1977,1999]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, NHMUK.2018.211); H)
<taxonomicName id="4C146B91A321695AFBCFF817FB182381" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[1084,1203,1977,1999]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="8" phylum="Annelida" rank="species" species="linseae" status="sp. nov.">
<emphasis id="B960CC00A321695AFBCFF817FB182381" box="[1084,1203,1977,1999]" italics="true" pageId="7" pageNumber="8">M. linseae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA321695AFB36F814FA8A2381" box="[1221,1313,1977,1999]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B960CC00A321695AFB36F814FA8A2381" bold="true" box="[1221,1313,1977,1999]" pageId="7" pageNumber="8">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, NHMUK.2018.9354). All scale bars 1 mm.
</paragraph>
</caption>
<caption id="DF6B409AA32E6955FF64FB34FA9720A2" httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1177,1199]" targetBox="[152,1435,193,1156]" targetPageId="8">
<paragraph id="8BAB1012A32E6955FF64FB34FA9720A2" blockId="8.[151,1436,1177,1260]" pageId="8" pageNumber="9">
<emphasis id="B960CC00A32E6955FF64FB34FEA720E1" bold="true" box="[151,268,1177,1199]" pageId="8" pageNumber="9">FIGURE 3</emphasis>
<taxonomicName id="4C146B91A32E6955FEE1FB34FDB920E1" authorityName="in Monro" authorityYear="1936" box="[274,530,1177,1199]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A32E6955FEE1FB34FDB920E1" box="[274,530,1177,1199]" italics="true" pageId="8" pageNumber="9">Macellicephala mirabilis</emphasis>
</taxonomicName>
(holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BAB1012A32E6955FF34FAB4FBE821D0" blockId="8.[151,1437,1305,2014]" pageId="8" pageNumber="9">
Notochaetae stouter than neurochaetae (
<figureCitation id="132F0C97A32E6955FD60FAB4FD45217F" box="[659,750,1305,1330]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="8" pageNumber="9">Fig. 4B</figureCitation>
), short to very long, few (about 7 present per ramus) often broken off entirely, smooth, with blunt tips. Neurochaetae numerous, upper and lower neurochaetae shorter, others very long; two forms present: either flattened, with only faint denticulation of each side (
<figureCitation id="132F0C97A32E6955FB7AFACCFB4B2137" box="[1161,1248,1377,1402]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="8" pageNumber="9">Fig. 4C</figureCitation>
) or with distinct spines, spines moderately separated (
<figureCitation id="132F0C97A32E6955FDC5FA28FD0521D0" box="[566,686,1413,1438]" captionStart="FIGURE 4" captionStartId="9.[151,250,1948,1970]" captionTargetBox="[152,1435,445,1924]" captionTargetId="figure@9.[152,1435,445,1924]" captionTargetPageId="9" captionText="FIGURE 4. Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100): A) Parapodium from 3 rd segment; B) detail of notochaetae; C) wide, flattened neurochaeta; D) narrow lower neurochaeta; E) detail of serration of lower neurochaeta. Scale bars: A) 500 µm; B) 250 µm, CE) 25 µm." httpUri="https://zenodo.org/record/1456917/files/figure.png" pageId="8" pageNumber="9">Fig. 4D, E</figureCitation>
); both forms tapering into sharp tip.
</paragraph>
<paragraph id="8BAB1012A32E6955FF34FA07FC0B2260" blockId="8.[151,1437,1305,2014]" pageId="8" pageNumber="9">
Two distinct white lines running through ventral side of neuropodia and ventrum of each segment (except for tentacular segment) (
<figureCitation id="132F0C97A32E6955FE76FA60FE4B21A8" box="[389,480,1485,1510]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="8" pageNumber="9">Fig. 3D</figureCitation>
). Nephridial papillae of segments 10, 11 and 12 prominent (
<figureCitation id="132F0C97A32E6955FB7FFA60FB4D21A8" box="[1164,1254,1485,1510]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="8" pageNumber="9">Fig. 3D</figureCitation>
), inconspicuous in other segments, present from segment 5. Reduced parapodia of segment 18 lateral to pygidium, consisting of notopodia only. Pygidium rounded. Anal cirri not observed (
<figureCitation id="132F0C97A32E6955FCCFF9B8FC382260" box="[828,915,1557,1582]" captionStart="FIGURE 3" captionStartId="8.[151,250,1177,1199]" captionTargetBox="[152,1435,193,1156]" captionTargetId="figure@8.[152,1435,193,1156]" captionTargetPageId="8" captionText="FIGURE 3 Macellicephala mirabilis (holotype, BMNH 1885: 12: 1: 100; specimen in two fragments): A) anterior fragment in dorsal view; B) posterior fragment of specimen in dorsal view; C) detail of prostomium in dorsal view; D) ventral view with nephridial papillae on segments 1012 marked by arrows; E) detail of pygidium in ventral view. All scale bars 1 mm." httpUri="https://zenodo.org/record/1456915/files/figure.png" pageId="8" pageNumber="9">Fig. 3E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E4399A32E6954FF34F994FCA1250E" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BAB1012A32E6955FF34F994FB242367" blockId="8.[151,1437,1305,2014]" pageId="8" pageNumber="9">
<emphasis id="B960CC00A32E6955FF34F994FE90221C" bold="true" box="[199,315,1593,1618]" pageId="8" pageNumber="9">Remarks.</emphasis>
During the examination of
<taxonomicName id="4C146B91A32E6955FD60F994FCEA221F" authorityName="McIntosh" authorityYear="1885" box="[659,833,1593,1617]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">
<emphasis id="B960CC00A32E6955FD60F994FCEA221F" box="[659,833,1593,1617]" italics="true" pageId="8" pageNumber="9">Macellicephala</emphasis>
</taxonomicName>
specimens collected as part of this study, several inconsistencies emerged with regards to the revision of the genus carried out by
<bibRefCitation id="EF856DE3A32E6955FBA3F9F0FABB2238" box="[1104,1296,1629,1654]" pageId="8" pageNumber="9" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
. Given that
<taxonomicName id="4C146B91A32E6955FF64F92CFDD622D4" authority="(McIntosh, 1885)" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[151,637,1665,1690]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A32E6955FF64F92CFE1922D7" box="[151,434,1665,1689]" italics="true" pageId="8" pageNumber="9">Macellicephala mirabilis</emphasis>
(
<bibRefCitation id="EF856DE3A32E6955FE32F92CFDDE22D4" box="[449,629,1665,1690]" pageId="8" pageNumber="9" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh, 1885</bibRefCitation>
)
</taxonomicName>
is the
<typeStatus id="54AFAEB0A32E6955FD3AF92FFD5222D4" box="[713,761,1666,1690]" pageId="8" pageNumber="9">type</typeStatus>
species of
<taxonomicName id="4C146B91A32E6955FC8BF92CFB8D22D7" authorityName="McIntosh" authorityYear="1885" box="[888,1062,1665,1689]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">
<emphasis id="B960CC00A32E6955FC8BF92CFB8D22D7" box="[888,1062,1665,1689]" italics="true" pageId="8" pageNumber="9">Macellicephala</emphasis>
</taxonomicName>
and this genus is in turn the
<typeStatus id="54AFAEB0A32E6955FA98F92FFA3722D4" box="[1387,1436,1666,1690]" pageId="8" pageNumber="9">type</typeStatus>
genus of
<taxonomicName id="4C146B91A32E6955FEF3F908FE7A22F0" authorityName="Hartmann-Schröder" authorityYear="1971" box="[256,465,1701,1726]" class="Polychaeta" family="Polynoidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="subFamily" subFamily="Macellicephalinae">Macellicephalinae</taxonomicName>
, it is important to clarify the definition of
<taxonomicName id="4C146B91A32E6955FC47F908FB7B22F3" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[948,1232,1701,1725]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A32E6955FC47F908FB7B22F3" box="[948,1232,1701,1725]" italics="true" pageId="8" pageNumber="9">Macellicephala mirabilis</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF856DE3A32E6955FB2CF908FA3722F0" box="[1247,1436,1701,1726]" pageId="8" pageNumber="9" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
based her re-description of
<taxonomicName id="4C146B91A32E6955FE38F967FDFD22AF" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[459,598,1737,1761]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A32E6955FE38F967FDFD22AF" box="[459,598,1737,1761]" italics="true" pageId="8" pageNumber="9">M. mirabilis</emphasis>
</taxonomicName>
on the following specimens found in the collections of the NHM London:
<typeStatus id="54AFAEB0A32E6955FF64F940FF522348" box="[151,249,1773,1798]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
of
<bibRefCitation id="EF856DE3A32E6955FEEEF940FE702348" box="[285,475,1773,1798]" pageId="8" pageNumber="9" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
, BMNH 1885: 12: 1: 100; two specimens of McIntosh (1905), BMNH 1924: 7: 21: 12 (see
<taxonomicName id="4C146B91A32E6955FF38F8BFFEC12367" authority="Neal &amp; Brasier &amp; Wiklund, 2018" authorityName="Neal &amp; Brasier &amp; Wiklund" authorityYear="2018" box="[203,362,1809,1833]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="macintoshi" status="sp. nov.">
<emphasis id="B960CC00A32E6955FF38F8BFFEC12367" box="[203,362,1809,1833]" italics="true" pageId="8" pageNumber="9">M. macintoshi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253717BA32E6955FE81F8BCFE672364" box="[370,460,1809,1834]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B960CC00A32E6955FE81F8BCFE672364" bold="true" box="[370,460,1809,1834]" pageId="8" pageNumber="9">sp. nov.</emphasis>
</taxonomicNameLabel>
) and two specimens of
<bibRefCitation id="EF856DE3A32E6955FD25F8BFFCDD2364" box="[726,886,1809,1834]" pageId="8" pageNumber="9" refId="ref17794" refString="Monro, C. C. A. (1936) Polychaete Worms, II. Discovery Reports, 12, 59 - 198." type="journal article">Monro (1936)</bibRefCitation>
, BMNH 1936: 2: 8: 488.
</paragraph>
<paragraph id="8BAB1012A32E6954FF34F898FE8424B6" blockId="8.[151,1437,1305,2014]" lastBlockId="9.[151,1436,151,392]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The material above was re-examined as part of this study and the definition of
<taxonomicName id="4C146B91A32E6955FBC1F898FAE02303" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[1074,1355,1845,1869]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="mirabilis">
<emphasis id="B960CC00A32E6955FBC1F898FAE02303" box="[1074,1355,1845,1869]" italics="true" pageId="8" pageNumber="9">Macellicephala mirabilis</emphasis>
</taxonomicName>
is here restricted to characters derived from the
<typeStatus id="54AFAEB0A32E6955FD85F8F4FD73233C" box="[630,728,1881,1906]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
BMNH 1885: 12: 1: 100, which was described in detail by
<bibRefCitation id="EF856DE3A32E6955FF64F8D0FEF023D8" box="[151,347,1917,1942]" pageId="8" pageNumber="9" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
. This form has very extended, anteriorly angular (truncated) prostomial lobes (
<figureCitation id="132F0C97A32E6955FB1DF8D0FAE223D8" box="[1262,1353,1917,1942]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
) rather than shorter, rounded or tapering form of prostomial lobes as seen in other species of
<taxonomicName id="4C146B91A32E6955FBBFF80CFB5123F7" authorityName="McIntosh" authorityYear="1885" box="[1100,1274,1953,1977]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="9" phylum="Annelida" rank="genus">
<emphasis id="B960CC00A32E6955FBBFF80CFB5123F7" box="[1100,1274,1953,1977]" italics="true" pageId="8" pageNumber="9">Macellicephala</emphasis>
</taxonomicName>
(see
<figureCitation id="132F0C97A32E6955FAC7F80CFADE23F7" box="[1332,1397,1953,1978]" captionStart="FIGURE 2" captionStartId="7.[151,250,1824,1846]" captionTargetBox="[252,1336,206,1786]" captionTargetId="figure@7.[228,1359,193,1803]" captionTargetPageId="7" captionText="FIGURE 2. Drawings of prostomia and anterior ends in dorsal view of species described and redescribed in this study: A) M. mirabilis (holotype, BMNH: 1885: 12: 1: 100); B) M. macintoshi sp. nov. (holotype, BMNH: 1924: 7: 21: 12); C) M. monroi sp. nov. (paratype, BMNH: 1930: 1930: 10. 8. 440), historic specimen collected from South Georgia by Monro (1930); D) M. monroi sp. nov. (holotype, NHMUK.2018.53), specimen collected from South Georgia during BIOPEARL I expedition to Scotia Sea; E) M. brenesorum sp. nov. (holotype, NHMUK.2018.830); F) M. patersoni sp. nov. (holotype, NHMUK.2018.1009); G) M. gloveri sp. nov. (holotype, NHMUK.2018.211); H) M. linseae sp. nov. (holotype, NHMUK.2018.9354). All scale bars 1 mm." httpUri="https://zenodo.org/record/1456913/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
for comparison). The description and drawing provided by
<bibRefCitation id="EF856DE3A32E6955FCFBF868FC6E2390" box="[776,965,1989,2014]" pageId="8" pageNumber="9" refId="ref17573" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by HMS Challenger during the years 1873 - 76. Report of the Scientific Results of the Voyage of HMS Challenger 1873 - 76, 12, 1 - 554." type="journal article">McIntosh (1885)</bibRefCitation>
clearly depicted slender elongated frontal filaments and dorsal cirri, which have since fallen off and were therefore not observed by
<bibRefCitation id="EF856DE3A32F6954FB8FFF3AFA9C24FE" box="[1148,1335,151,176]" pageId="9" pageNumber="10" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
or in our examination. Papillation medial to bases of elytrophores and dorsal cirrophores reported by
<bibRefCitation id="EF856DE3A32F6954FB5FFF11FAC3249B" box="[1196,1384,188,213]" pageId="9" pageNumber="10" refId="ref18546" refString="Pettibone, M. H. (1976) Revision of the Genus Macellicephala McIntosh and the Subfamily Macellicephalinae Hartmann- Schroder (Polychaeta, Polynoidae). Smithsonian Contributions to Zoology, 229, 1 - 71." type="journal article">Pettibone (1976)</bibRefCitation>
was not observed.
</paragraph>
<paragraph id="8BAB1012A32F6954FF34FEA8FCA1250E" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="10">
For a full comparison of all valid
<taxonomicName id="4C146B91A32F6954FDBFFEA9FD512552" authorityName="McIntosh" authorityYear="1885" box="[588,762,260,284]" class="Polychaeta" family="Polynoidae" genus="Macellicephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="10" phylum="Annelida" rank="genus">
<emphasis id="B960CC00A32F6954FDBFFEA9FD512552" box="[588,762,260,284]" italics="true" pageId="9" pageNumber="10">Macellicephala</emphasis>
</taxonomicName>
species based on examination of material presented in this study and published literature for the others see
<tableCitation id="C69625A9A32F6954FD42FE8AFCAC250E" box="[689,775,295,320]" captionStart="TABLE 2" captionStartId="12.[147,221,540,559]" captionText="TABLE 2. Comparison of characters of all known Macellicephala species based on the examination of representatives of species described in this study and literature survey for other species." pageId="9" pageNumber="10">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E4399A32F6954FF34FEE1FF7625C6" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BAB1012A32F6954FF34FEE1FF7625C6" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="10">
<emphasis id="B960CC00A32F6954FF34FEE1FECB252B" bold="true" box="[199,352,332,357]" pageId="9" pageNumber="10">Distribution.</emphasis>
Known only from its
<typeStatus id="54AFAEB0A32F6954FDACFEE0FD24252B" box="[607,655,333,357]" pageId="9" pageNumber="10">type</typeStatus>
locality:
<collectingCountry id="F3035082A32F6954FD0FFEE0FC3D252B" box="[764,918,332,357]" name="New Zealand" pageId="9" pageNumber="10">New Zealand</collectingCountry>
, off North Island,
<geoCoordinate id="EE2076D5A32F6954FB9BFEE1FB69252A" box="[1128,1218,332,356]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="-37.566666">37°34'S</geoCoordinate>
,
<geoCoordinate id="EE2076D5A32F6954FB3CFEE1FA92252A" box="[1231,1337,332,356]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="179.36667">179°22'E</geoCoordinate>
,
<quantity id="4CECBDF7A32F6954FAB5FEE1FA37252A" box="[1350,1436,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="9" pageNumber="10" unit="m" value="1280.0">1280 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>