treatments-xml/data/03/B7/5F/03B75F43F056FF83FF4876E0331E2616.xml

<document id="1B930495326A7C1198D7A5B2DA6D35C9" ID-DOI="10.11646/zootaxa.5567.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14703735" ID-ZooBank="AF6254F6-CE26-41FE-BB85-A3FFD16FAA39" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737371447294" checkinUser="plazi" docAuthor="VIRAKTAMATH, C. A., WEBB, M. D. &amp; YESHWANTH, H. M." docDate="2025" docId="03B75F43F056FF83FF4876E0331E2616" docLanguage="en" docName="zootaxa.5567.1.1.pdf" docOrigin="Zootaxa 5567 (1)" docSource="https://doi.org/10.11646/zootaxa.5567.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Monobazus longulus VIRAKTAMATH, WEBB &amp; YESHWANTH, 2025, sp. nov." docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="FF8E273BF049FF9CFFDF7756364D2343" masterDocTitle="Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species" masterLastPageNumber="105" masterPageNumber="1" pageNumber="32" updateTime="1737389185710" updateUser="ExternalLinkService">
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<mods:title id="337B5CCA86240D9ABB3EA998A6962461">Review of the Monobazus-group leafhopper genera (Hemiptera: Cicadellidae: Deltocephalinae) from the Indian subcontinent with description of five new genera and twenty-eight new species</mods:title>
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<mods:namePart id="0440B9922FECB5470D0F25B6EB019DC4">VIRAKTAMATH, C. A.</mods:namePart>
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<mods:namePart id="4F830D4D78AEE5EE9E68673736B79BAA">WEBB, M. D.</mods:namePart>
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<mods:date id="EB625E04B08F62DF4562DE2B0CFA934D">2025</mods:date>
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<subSubSection id="C304BDDEF056FF83FF4876E037B72292" box="[151,506,438,465]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BA1EE55F056FF83FF4876E037B72292" blockId="31.[151,506,438,501]" box="[151,506,438,465]" pageId="31" pageNumber="32">
<heading id="D0E95939F056FF83FF4876E037B72292" bold="true" box="[151,506,438,465]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
<emphasis id="B96A3247F056FF83FF4876E037B72292" bold="true" box="[151,506,438,465]" pageId="31" pageNumber="32">
<taxonomicName id="4C1E95D6F056FF83FF4876E037D82292" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[151,405,438,465]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longulus" status="sp. nov.">
<emphasis id="B96A3247F056FF83FF4876E037D82292" bold="true" box="[151,405,438,465]" italics="true" pageId="31" pageNumber="32">Monobazus longulus</emphasis>
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<taxonomicNameLabel id="A2598F3CF056FF83FE4376E137B72292" box="[412,506,439,465]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C304BDDEF056FF83FF48768D37AC273E" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA1EE55F056FF83FF48768D37D722B6" blockId="31.[151,506,438,501]" box="[151,410,475,501]" pageId="31" pageNumber="32">
<figureCitation id="1325F2D0F056FF83FF48768D369122B6" box="[151,220,475,501]" captionStart="FIGURES 4" captionStartId="45.[151,264,1920,1944]" captionTargetBox="[157,1430,186,1891]" captionTargetId="figure-17@45.[151,1435,181,1896]" captionTargetPageId="45" captionText="FIGURES 4A–P. Monobazus species, habitus, dorsal and lateral view. A–D, Monobazus confuscus (Pruthi), male (AB) and female (CD); E–F, M. deccanensis sp. nov. holotype male; G–J, M. dissimilis (Distant), male (GH) and female (IJ); K–L, M. himalayensis Distant, male; M–N, M. longulus sp. nov., holotype male; O–P, M. paulus sp. nov., paratype male." figureDoi="http://doi.org/10.5281/zenodo.14703751" httpUri="https://zenodo.org/record/14703751/files/figure.png" pageId="31" pageNumber="32">Figs 4</figureCitation>
MN, 9M, 38A–F
</paragraph>
<paragraph id="8BA1EE55F056FF83FF487575331E2122" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF4875753744217F" box="[151,265,547,572]" italics="true" pageId="31" pageNumber="32">Diagnosis</emphasis>
. Coloration similar to 
<taxonomicName id="4C1E95D6F056FF83FDD675753564217F" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[521,809,547,572]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="oppressa">
<emphasis id="B96A3247F056FF83FDD675753564217F" box="[521,809,547,572]" italics="true" pageId="31" pageNumber="32">Mavromoustaca oppressa</emphasis>
</taxonomicName>
, but slightly paler less bold. Aedeagus similar to that in 
<taxonomicName id="4C1E95D6F056FF83FF48751137042122" authorityName="Pruthi" authorityYear="1936" box="[151,329,583,609]" class="Insecta" family="Cicadellidae" genus="Thamnotettix" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="brunnescens">
<emphasis id="B96A3247F056FF83FF48751137042122" box="[151,329,583,609]" italics="true" pageId="31" pageNumber="32">M. brunnescens</emphasis>
</taxonomicName>
but ventral process on shaft arising more proximally, divergent from shaft and much longer.
</paragraph>
<paragraph id="8BA1EE55F056FF83FF18753D37DD21EA" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF18753D370621C7" box="[199,331,619,644]" italics="true" pageId="31" pageNumber="32">Description</emphasis>
. Coloration similar to 
<taxonomicName id="4C1E95D6F056FF83FD8A753D34AF21C7" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[597,738,619,644]" class="Insecta" genus="Mavromoustaca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="oppressa">
<emphasis id="B96A3247F056FF83FD8A753D34AF21C7" box="[597,738,619,644]" italics="true" pageId="31" pageNumber="32">M. oppressa</emphasis>
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, but slightly paler and dark spots less bold. Face ventrad of antennal bases brown.
</paragraph>
<paragraph id="8BA1EE55F056FF83FF1875E537F121B2" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">Head as wide as pronotum. Crown 0.6× as long as interocular width. Pronotum 2.2× as wide as long and shorter than exposed mesonotum.</paragraph>
<paragraph id="8BA1EE55F056FF83FF1875AD320A208A" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF1875AD37232056" box="[199,366,763,789]" italics="true" pageId="31" pageNumber="32">Male genitalia</emphasis>
. Pygofer narrowed posteriorly, more than 2.5× as long as wide in lateral view; posterodorsal angle conically rounded; ventral process very short, arising at midlength on ventral margin, finger-like. Subgenital plate with macrosetae in distal half and also with hair-like setae, almost 7x as long as wide at base. Style with apophysis stout, almost straight, with rounded apex. Connective with stem as long as arms. Aedeagus with shaft of uniform width in lateral view, curved anterodorsally, with basal process arising from atrium, divergent from shaft at basal 0.33, forked almost at midlength and much longer than shaft; gonopore apical.
</paragraph>
<paragraph id="8BA1EE55F056FF83FF187485332520AE" blockId="31.[151,1437,547,1365]" box="[199,1384,979,1005]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF187485372420AF" box="[199,361,979,1004]" italics="true" pageId="31" pageNumber="32">Measurements</emphasis>
. Male 
<quantity id="4CE643B0F056FF83FE677485344320AE" box="[440,526,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="31" pageNumber="32" unit="mm" value="4.3">4.3 mm</quantity>
long, 
<quantity id="4CE643B0F056FF83FD8B748534E720AE" box="[596,682,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="31" pageNumber="32" unit="mm" value="1.1">1.1 mm</quantity>
across eyes and also across posterolateral angles of pronotum.
</paragraph>
<paragraph id="8BA1EE55F056FF83FF1874A137C02776" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF1874A137EF2752" box="[199,418,1015,1041]" italics="true" pageId="31" pageNumber="32">Material examined</emphasis>
. 
<materialsCitation id="3B76E408F056FF83FE6874A137C02776" collectingDate="1980-10-02" collectionCode="UASB" collectorName="H.S. Pruthi &amp; Km N. Kalka &amp; C. A. Viraktamath" country="India" location="Himachal Pradesh" pageId="31" pageNumber="32" specimenCount="1" specimenCount-male="1" stateProvince="Himachal Pradesh" typeStatus="holotype">
<typeStatus id="54A550F7F056FF83FE6874A134062752" box="[439,587,1015,1041]" pageId="31" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D1825DCF056FF83FD8774A134222752" box="[600,623,1015,1041]" count="1" pageId="31" pageNumber="32" type="male">♂</specimenCount>
, 
<collectingCountry id="F309AEC5F056FF83FD5E74A134982752" box="[641,725,1015,1041]" name="India" pageId="31" pageNumber="32">INDIA</collectingCountry>
: 
<collectorName id="26EB8B83F056FF83FD3974A1356E2752" box="[742,803,1015,1041]" name="H.S. Pruthi" pageId="31" pageNumber="32">H. P.</collectorName>
[
<collectingRegion id="49DA20B7F056FF83FCE974A132592752" box="[822,1044,1015,1041]" country="India" name="Himachal Pradesh" pageId="31" pageNumber="32">Himachal Pradesh</collectingRegion>
]: 5 
<collectorName id="26EB8B83F056FF83FBE874A032922752" box="[1079,1247,1014,1041]" pageId="31" pageNumber="32">Km N. Kalka</collectorName>
, 
<date id="FFA0C895F056FF83FB2E74A1331B2752" box="[1265,1366,1015,1041]" pageId="31" pageNumber="32" value="1980-10-02">
<collectingDate id="EFE4317DF056FF83FB2E74A1331B2752" box="[1265,1366,1015,1041]" pageId="31" pageNumber="32" value="1980-10-02">2.x.1980</collectingDate>
</date>
, 
<collectorName id="26EB8B83F056FF83FAB874A1376C2776" pageId="31" pageNumber="32">C.A. Viraktamath</collectorName>
(
<collectionCode id="ED0F7690F056FF83FEEE734D37CF2776" box="[305,386,1051,1077]" country="India" name="University of Agricultural Sciences" pageId="31" pageNumber="32">UASB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA1EE55F056FF83FF18736937AC273E" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF1873693772271A" box="[199,319,1087,1113]" italics="true" pageId="31" pageNumber="32">Etymology</emphasis>
. The specific epithet is the Latin adjective longulus, a-, -um (= rather long) referring to the long basal process of the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="C304BDDEF056FF83FF1873D1331E2616" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8BA1EE55F056FF83FF1873D1331E2616" blockId="31.[151,1437,547,1365]" pageId="31" pageNumber="32">
<emphasis id="B96A3247F056FF83FF1873D1377927E2" bold="true" box="[199,308,1159,1185]" pageId="31" pageNumber="32">Remarks</emphasis>
. 
<emphasis id="B96A3247F056FF83FE9E73D1343127E2" box="[321,636,1159,1185]" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="4C1E95D6F056FF83FE9E73D1378527E2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[321,456,1159,1185]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longulus">M. longulus</taxonomicName>
, 
<taxonomicName id="4C1E95D6F056FF83FE0D73D1343127E2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[466,636,1159,1185]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="biligiriensis">M. biligiriensis</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C1E95D6F056FF83FD6973D1351F27E2" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[694,850,1159,1185]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="sudindicus">
<emphasis id="B96A3247F056FF83FD6973D1351F27E2" box="[694,850,1159,1185]" italics="true" pageId="31" pageNumber="32">M. sudindicus</emphasis>
</taxonomicName>
have the ventral process of the aedeagal shaft arising very close to the base. 
<taxonomicName id="4C1E95D6F056FF83FE4773FD34532786" authorityName="VIRAKTAMATH &amp; WEBB &amp; YESHWANTH" authorityYear="2025" box="[408,542,1195,1221]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B96A3247F056FF83FE4773FD34532786" box="[408,542,1195,1221]" italics="true" pageId="31" pageNumber="32">M. longulus</emphasis>
</taxonomicName>
differs from the other two species in having the ventral process of the aedeagal shaft with long stem before it forks (
<figureCitation id="1325F2D0F056FF83FDF5739934C227AA" box="[554,655,1231,1257]" captionStart="FIGURES 38" captionStartId="79.[151,264,1894,1918]" captionTargetBox="[156,1430,186,1865]" captionTargetId="figure-17@79.[151,1435,181,1870]" captionTargetPageId="79" captionText="FIGURES 38A–F. Monobazus longulus sp. nov., male genitalia:A, genital capsule, lateral view; B, connective, dorsal view; C, style, dorsal view; D, aedeagal shaft, ventral view; E, aedeagal shaft, posterior view; F, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703853" httpUri="https://zenodo.org/record/14703853/files/figure.png" pageId="31" pageNumber="32">Fig. 38D</figureCitation>
) as in the case of 
<taxonomicName id="4C1E95D6F056FF83FC8C7399324727AA" box="[851,1034,1231,1257]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="himalayensis">
<emphasis id="B96A3247F056FF83FC8C7399324727AA" box="[851,1034,1231,1257]" italics="true" pageId="31" pageNumber="32">M. himalayensis</emphasis>
</taxonomicName>
and the male pygofer has very short digitate ventral process arising almost at midlength of ventral margin (
<figureCitation id="1325F2D0F056FF83FC7673A5325F264E" box="[937,1042,1267,1293]" captionStart="FIGURES 38" captionStartId="79.[151,264,1894,1918]" captionTargetBox="[156,1430,186,1865]" captionTargetId="figure-17@79.[151,1435,181,1870]" captionTargetPageId="79" captionText="FIGURES 38A–F. Monobazus longulus sp. nov., male genitalia:A, genital capsule, lateral view; B, connective, dorsal view; C, style, dorsal view; D, aedeagal shaft, ventral view; E, aedeagal shaft, posterior view; F, aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.14703853" httpUri="https://zenodo.org/record/14703853/files/figure.png" pageId="31" pageNumber="32">Fig. 38A</figureCitation>
). It differs from 
<taxonomicName id="4C1E95D6F056FF83FB1773A533CD264E" box="[1224,1408,1267,1293]" class="Insecta" family="Cicadellidae" genus="Monobazus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="himalayensis">
<emphasis id="B96A3247F056FF83FB1773A533CD264E" box="[1224,1408,1267,1293]" italics="true" pageId="31" pageNumber="32">M. himalayensis</emphasis>
</taxonomicName>
in having the ventral process of aedeagus arising from midventral margin of pygofer, the posterodorsal lobe of pygofer not produced into a hook-like process and the subgenital plate with macrosetae in addition to hair-like setae.
</paragraph>
</subSubSection>
</treatment>
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