treatments-xml/data/03/8E/87/038E87DAFFF5E412FE95FB23FF78F858.xml

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<document id="183ED8B85060B9E1321DB5F0756BBD38" ID-CLB-Dataset="304811" ID-DOI="10.5252/geodiversitas2024v46a15" ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f" ID-ISSN="1638-9395" ID-Zenodo-Dep="13985760" ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1729758589633" checkinUser="plazi" docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel &amp; Goret, Bernard" docDate="2024" docId="038E87DAFFF5E412FE95FB23FF78F858" docLanguage="en" docName="geodiversitas.2024.46.15.pdf" docOrigin="Geodiversitas 46 (15)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Timbellus occidentalis Merle, Pacaud, Ledon &amp; Goret, 2024, n. sp." docType="treatment" docUuid="53A32E50-A05D-4A09-B8B1-6CA776FDC29F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="513" masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3" masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe" masterLastPageNumber="551" masterPageNumber="495" pageNumber="512" updateTime="1732586620909" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="A56F75E939DC1764E5FFF0E39C45395B">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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<mods:namePart id="7076FA73D6342B9F57E95C30EF92F369">Merle, Didier</mods:namePart>
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<mods:affiliation id="3345C67AF01223CA7CADDAD67085DEF5">Muséum national dhistoire naturelle, Département Origines et Évolution (CR 2 P - MNHN, CNRS, UPMC, Sorbonne Université), 57 rue Cuvier, 75231 Paris cedex 05 (France) didier. merle @ mnhn. fr (corresponding author) jean-michel. pacaud @ mnhn. fr dnledon @ yahoo. fr</mods:affiliation>
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(
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A-C; 26A, B)
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(
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, pl. 13, fig. 29 [
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<emphasis id="B953EADEFFF5E415FF3CFA22FF60F9BF" box="[132,225,1524,1548]" italics="true" pageId="19" pageNumber="512">Timbellus</emphasis>
cf.
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Merle
<emphasis id="B953EADEFFF5E415FE79FA23FE6EF9BF" box="[449,495,1524,1548]" italics="true" pageId="19" pageNumber="512">et al.</emphasis>
2011: 446
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, pl. 101, figs 1, 2.
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. —
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Calcaire
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; middle
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(uppermost
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.F.A90533 (ex A24100 ex galerie de zoologie coll.,
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coll.), H:
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(
<figureCitation id="131C2A49FFF5E415FEC3F9ADFE62F920" box="[379,483,1659,1683]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de lOrme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="19" pageNumber="512">Figs 8A, B</figureCitation>
;
<figureCitation id="131C2A49FFF5E415FE56F9ADFD9AF920" box="[494,539,1659,1683]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="19" pageNumber="512">26A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9836CCFFF5E415FF3CF940FBC7FC12" blockId="19.[131,777,1580,2027]" lastBlockId="19.[813,1456,852,929]" pageId="19" pageNumber="512">
<materialsCitation id="3B4F3C91FFF5E415FF3CF940FD32F97B" ID-GBIF-Occurrence="4958671386" pageId="19" pageNumber="512">
<emphasis id="B953EADEFFF5E415FF3CF940FEA7F91D" bold="true" box="[132,294,1686,1710]" pageId="19" pageNumber="512">
<typeStatus id="549C886EFFF5E415FF3CF940FF61F91D" box="[132,224,1686,1710]" pageId="19" pageNumber="512" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F330765CFFF5E415FF50F940FEA7F91D" box="[232,294,1686,1710]" name="France" pageId="19" pageNumber="512">France</collectingCountry>
</emphasis>
• 1 spm; same as for the holotype;
<collectionCode id="ED36AE09FFF5E415FDEDF940FD27F91E" box="[597,678,1686,1709]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A90534 (ex A24100 ex galerie de zoologie coll., Le Marchand coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FD0EF967FD88F950" ID-GBIF-Occurrence="4958671351" pageId="19" pageNumber="512">
• 4 spm; idem;
<collectionCode id="ED36AE09FFF5E415FF04F91DFE8DF951" box="[188,268,1739,1762]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.B63570 (de Morgan coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FDB1F91DFEB2F94E" ID-GBIF-Occurrence="4958671308" pageId="19" pageNumber="512">
• 1 spm,
<collectionCode id="ED36AE09FFF5E415FDEEF91DFD27F951" box="[598,678,1739,1762]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A90532 (de Morgan coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FE84F930FE09F881" ID-GBIF-Occurrence="4958671427" pageId="19" pageNumber="512">
• 1 spm;
<collectingRegion id="49E3F82EFFF5E415FE20F933FD85F94E" box="[408,516,1765,1789]" country="France" name="Haute-Normandie" pageId="19" pageNumber="512">Normandy</collectingRegion>
, Manche, Hautteville-Bocage; middle Eocene (upper Lutetian);
<collectionCode id="ED36AE09FFF5E415FD99F8D6FDF9F8A4" box="[545,632,1792,1815]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A90531 (ex B63609, de Morgan coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FE36F8CDFE90F8FE" ID-GBIF-Occurrence="4958671365" pageId="19" pageNumber="512">
• 1 spm; idem;
<collectionCode id="ED36AE09FFF5E415FD91F8CDFDFCF881" box="[553,637,1819,1842]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.B63609 (de Morgan coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FEA2F8E3FC88F8D4" ID-GBIF-Occurrence="4958671434" pageId="19" pageNumber="512">
• 35 spm; idem;
<collectionCode id="ED36AE09FFF5E415FE72F8E3FDA1F8FF" box="[458,544,1845,1868]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.B63610 (de Morgan coll.) Gourbesville, Manche, middle Eocene (upper Lutetian)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FF3EF8BCFD52F832" ID-GBIF-Occurrence="4958671340" box="[134,723,1897,1922]" pageId="19" pageNumber="512">
• 3 spm; idem;
<collectionCode id="ED36AE09FFF5E415FE9EF8BCFEFBF832" box="[294,378,1898,1921]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A24128 (Munier-Chalmas coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FD65F8BCFD61F805" ID-GBIF-Occurrence="4958671389" pageId="19" pageNumber="512">
• 10 spm;
<collectingRegion id="49E3F82EFFF5E415FF06F852FF73F82F" box="[190,242,1924,1948]" country="France" name="Pays de la Loire" pageId="19" pageNumber="512">Loire</collectingRegion>
Basin, Loire-Atlantique, Saffré (Bois-Gouët); middle Eocene (lower Bartonian);
<collectionCode id="ED36AE09FFF5E415FE2DF849FE68F805" box="[405,489,1951,1974]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A29658 (Pacaud coll.)
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FD52F849FB92FCDF" ID-GBIF-Occurrence="4958671304" pageId="19" pageNumber="512">
• 1 spm; idem;
<collectionCode id="ED36AE09FFF5E415FEB8F86FFED4F863" box="[256,341,1977,2000]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.A24031 (Vasseur coll.), figured in
<bibRefCitation id="EFB64B3DFFF5E415FD76F86FFF76F858" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." pageId="19" pageNumber="512" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFF5E415FD76F86FFC88F862" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[718,777,1977,2001]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFF5E415FF3CF802FF34F858" box="[132,181,2003,2027]" italics="true" pageId="19" pageNumber="512">et al.</emphasis>
(2011
</bibRefCitation>
: pl. 101, fig 1) as
<taxonomicName id="4C274D4FFFF5E415FE0AF805FCE8FCDF" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[434,873,852,2027]" class="Gastropoda" family="Muricidae" genus="Timbellus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="512" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF5E415FE0AF805FD8EF858" box="[434,527,2003,2027]" italics="true" pageId="19" pageNumber="512">Timbellus</emphasis>
cf.
<emphasis id="B953EADEFFF5E415FD8EF805FD1CF858" box="[566,669,2003,2027]" italics="true" pageId="19" pageNumber="512">tripteroides</emphasis>
(
<bibRefCitation id="EFB64B3DFFF5E415FD12F805FCE3FCDF" author="LAMARCK J. - B." box="[682,866,852,2027]" pageId="19" pageNumber="512" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="131C2A49FFF5E415FCCFFC82FC44FCDF" box="[887,965,852,876]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de lOrme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="19" pageNumber="512">Figs 8C</figureCitation>
;
<figureCitation id="131C2A49FFF5E415FC69FC82FC7BFCDF" box="[977,1018,852,876]" captionStart="FIG" captionStartId="45.[132,143,1622,1639]" captionTargetBox="[132,1455,215,1562]" captionTargetId="figure-370@45.[700,1259,799,1371]" captionTargetPageId="45" captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13985825" httpUri="https://zenodo.org/record/13985825/files/figure.png" pageId="19" pageNumber="512">26B</figureCitation>
) •
</materialsCitation>
<materialsCitation id="3B4F3C91FFF5E415FBA2FC82FBC7FC12" ID-GBIF-Occurrence="4958671313" pageId="19" pageNumber="512">
1 spm; idem;
<collectionCode id="ED36AE09FFF5E415FB1DFC83FB78FCDF" box="[1189,1273,853,876]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="512">MNHN</collectionCode>
.F.J06383 (Vasseur coll.), figured in
<bibRefCitation id="EFB64B3DFFF5E415FC69FCB9FB00FC34" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[977,1153,879,903]" pageId="19" pageNumber="512" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFF5E415FC69FCB9FB8BFC34" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[977,1034,879,903]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFF5E415FBA9FCB9FBC1FC34" box="[1041,1088,879,903]" italics="true" pageId="19" pageNumber="512">et al.</emphasis>
(2011
</bibRefCitation>
: pl. 101, fig 2) as
<taxonomicName id="4C274D4FFFF5E415FA8DFCB9FBC3FC12" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Muricidae" genus="Timbellus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="512" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF5E415FA8DFCB9FA13FC34" box="[1333,1426,879,903]" italics="true" pageId="19" pageNumber="512">Timbellus</emphasis>
cf.
<emphasis id="B953EADEFFF5E415FC95FC5FFC15FC12" box="[813,916,905,929]" italics="true" pageId="19" pageNumber="512">tripteroides</emphasis>
(
<bibRefCitation id="EFB64B3DFFF5E415FC19FC5FFBBBFC12" author="LAMARCK J. - B." box="[929,1082,905,929]" pageId="19" pageNumber="512" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
)
</taxonomicName>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF5E415FC95FC6FFC48FBB5" pageId="19" pageNumber="512" type="etymology">
<paragraph id="8B9836CCFFF5E415FC95FC6FFC48FBB5" blockId="19.[813,1455,953,1030]" pageId="19" pageNumber="512">
<smallCapsWord id="8D7EA010FFF5E415FC95FC6FFC26FC7C" baselines="970,971" box="[813,935,953,976]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="19" pageNumber="512">ETYMOLOGY</smallCapsWord>
. — From the Latin
<taxonomicName id="4C274D4FFFF5E415FBC2FC6FFB64FC62" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1146,1253,953,977]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="19" pageNumber="512" phylum="Mollusca" rank="species" species="occidentalis">
<emphasis id="B953EADEFFF5E415FBC2FC6FFB64FC62" box="[1146,1253,953,977]" italics="true" pageId="19" pageNumber="512">occidentalis</emphasis>
</taxonomicName>
meaning west, considering the geographic range of this species in
<collectingRegion id="49E3F82EFFF5E415FB45FC05FAE9FC58" box="[1277,1384,979,1003]" country="France" name="Haute-Normandie" pageId="19" pageNumber="512">Normandy</collectingRegion>
and in the
<collectingRegion id="49E3F82EFFF5E415FCEBFC38FC07FBB5" box="[851,902,1006,1030]" country="France" name="Pays de la Loire" pageId="19" pageNumber="512">Loire</collectingRegion>
Basin.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF5E415FC95FBC8FC41FB29" pageId="19" pageNumber="512" type="materials_examined">
<paragraph id="8B9836CCFFF5E415FC95FBC8FBF5FBFC" blockId="19.[811,1456,1053,1104]" pageId="19" pageNumber="512">
<smallCapsWord id="8D7EA010FFF5E415FC95FBC8FCDCFB80" baselines="1071,1071" box="[813,861,1054,1077]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="19" pageNumber="512">TYPE</smallCapsWord>
<smallCapsWord id="8D7EA010FFF5E415FCDAFBF5FC3DFB87" baselines="1072" box="[866,956,1059,1076]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="19" pageNumber="512">HORIZON</smallCapsWord>
. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see
<bibRefCitation id="EFB64B3DFFF5E415FC0EFBEEFBE6FBE3" author="BIGNOT G. &amp; HOMMERIL P. &amp; LARSONNEUR C." box="[950,1127,1080,1104]" pageId="19" pageNumber="512" pagination="405 - 416" refId="ref42941" refString="BIGNOT G., HOMMERIL P. &amp; LARSONNEUR C. 1968. - Le Lutetien au large du Cotentin, in Colloque sur l'Eocene (Paris, mai 1968). Memoires BRGM 58: 405 - 416." type="journal article" year="1968">
Bignot
<emphasis id="B953EADEFFF5E415FC47FBEEFBAFFBE3" box="[1023,1070,1080,1104]" italics="true" pageId="19" pageNumber="512">et al.</emphasis>
1968
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9836CCFFF5E415FC95FBBEFC41FB29" blockId="19.[813,1457,1127,1178]" pageId="19" pageNumber="512">
<smallCapsWord id="8D7EA010FFF5E415FC95FBBEFCDEFBCE" baselines="1145,1145" box="[813,863,1128,1151]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="19" pageNumber="512">TYPE</smallCapsWord>
<smallCapsWord id="8D7EA010FFF5E415FCD0FBBBFC47FBCD" baselines="1146" box="[872,966,1133,1150]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="19" pageNumber="512">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFF5E415FBBAFBBEFBC9FBCC" box="[1026,1096,1128,1151]" name="France" pageId="19" pageNumber="512">France</collectingCountry>
,
<collectingRegion id="49E3F82EFFF5E415FBEDFBB1FB44FBCC" box="[1109,1221,1127,1151]" country="France" name="Haute-Normandie" pageId="19" pageNumber="512">Normandy</collectingRegion>
, Manche, Fresville, see
<bibRefCitation id="EFB64B3DFFF5E415FC96FB54FC3DFB29" author="APGN &amp; ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE" box="[814,956,1154,1178]" pageId="19" pageNumber="512" refId="ref42597" refString="APGN (ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE) 2024. - Faluns cretaces et tertiaires de Fresville, BNO 0182. Available at: http: // www. apgn. fr / apgn / carte _ site / fic _ s 50 _ 75. pdf (last access on 03 / 2024)." type="book" year="2024">APGN (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF5E415FC95FB64FC3FFAAB" pageId="19" pageNumber="512" type="distribution">
<paragraph id="8B9836CCFFF5E415FC95FB64FC3FFAAB" blockId="19.[813,1456,1201,1305]" pageId="19" pageNumber="512">
<smallCapsWord id="8D7EA010FFF5E415FC95FB64FC3BFB7B" baselines="1219,1220" box="[813,954,1202,1225]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="19" pageNumber="512">DISTRIBUTION</smallCapsWord>
. —
<collectingRegion id="49E3F82EFFF5E415FC5EFB67FBD0FB7A" box="[998,1105,1201,1225]" country="France" name="Haute-Normandie" pageId="19" pageNumber="512">Normandy</collectingRegion>
(Calcaire de Fresville and Faluns de Hautteville) middle Eocene (uppermost Lutetian) and
<collectingRegion id="49E3F82EFFF5E415FAEAFB1AFA04FB57" box="[1362,1413,1228,1252]" country="France" name="Pays de la Loire" pageId="19" pageNumber="512">Loire</collectingRegion>
Basin (Sables du Bois-Gouët), lowermost Bartonian see
<bibRefCitation id="EFB64B3DFFF5E415FA82FB30FC39FAAA" author="MATHELIN J. - C. &amp; BIGNOT G." pageId="19" pageNumber="512" pagination="55 - 70" refId="ref46304" refString="MATHELIN J. - C. &amp; BIGNOT G. 1989. - Le falun de Foulangues et ses relations stratigraphique et paleogeographique avec le Biarritzien stratotypique, in 114 eme Congres national des Societes savantes. Geologie du Bassin Parisien. CTHS, Paris: 55 - 70." type="book chapter" year="1989">Mathelin &amp; Bignot (1989)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFF5E412FC95FA82FF78F858" lastPageId="20" lastPageNumber="513" pageId="19" pageNumber="512" type="description">
<paragraph id="8B9836CCFFF5E415FC95FA82FC3CFADF" blockId="19.[810,1457,1364,2028]" box="[813,957,1364,1390]" pageId="19" pageNumber="512">
<smallCapsWord id="8D7EA010FFF5E415FC95FA82FC3CFADF" baselines="1383,1383" box="[813,957,1364,1390]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="19" pageNumber="512">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFF5E415FC95FAA5FAD0F85F" blockId="19.[810,1457,1364,2028]" pageId="19" pageNumber="512">
Smooth, bulbous protoconch of 1.25 whorls. Teleoconch up to
<quantity id="4CDF9B29FFF5E415FCF3FA45FC2EFA1D" box="[843,943,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="19" pageNumber="512" unit="mm" value="26.6">26.6 mm</quantity>
in height, up to
<quantity id="4CDF9B29FFF5E415FBD2FA45FB4EFA1D" box="[1130,1231,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="19" pageNumber="512" unit="mm" value="15.3">15.3 mm</quantity>
in width, biconic in profile, composed of five whorls. Moderately high spire. Last whorl up to 85% total length. Apical angle 64° excluding spines, 91° including spines. Spiral sculpture with weak primary cords in adult specimens. First to fourth whorl, no cord. Fifth whorl: P1 to P3. Sixth whorl: P1 to P6 weak, s2 and no other cord below, ADP and MP occasional. On first whorl, eight protovarices. From second to last whorl, three varices and one intervarix intercalated Higher relief of intervarices on P2-P3. Varices lamellose, winged on last whorls. Appearance of short, straight P1 spine on third whorl. Sinus of P1 spine narrow and pointing adapically. On last whorl alate varices. Ovate aperture up to 9% of diameter and up to 80% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with denticles ID (occasional), D1 to D6, D1 and D2 more strongly developed. Outer lip not crenulate. Siphonal canal narrow, open, up to 49% of apertural length. Pseudo-umbilicus narrow.
</paragraph>
<paragraph id="8B9836CCFFF2E412FEE8FF0FFED8FF5B" blockId="20.[336,345,217,390]" box="[336,345,217,232]" pageId="20" pageNumber="513">4</paragraph>
<paragraph id="8B9836CCFFF2E412FEE8FEA1FED8FE35" blockId="20.[336,345,217,390]" box="[336,345,375,390]" pageId="20" pageNumber="513">3</paragraph>
<paragraph id="8B9836CCFFF2E412FE88FE27FEBEFDB4" blockId="20.[304,345,497,583]" box="[304,319,497,519]" pageId="20" pageNumber="513">cm</paragraph>
<paragraph id="8B9836CCFFF2E412FE88FDDAFEBEFDAB" blockId="20.[304,345,497,583]" box="[304,319,524,536]" pageId="20" pageNumber="513">in</paragraph>
<paragraph id="8B9836CCFFF2E412FE88FDCBFED8FD94" blockId="20.[304,345,497,583]" box="[304,345,536,583]" pageId="20" pageNumber="513">Width 2</paragraph>
<paragraph id="8B9836CCFFF2E412FEE8FD61FED8FD75" blockId="20.[336,345,695,710]" box="[336,345,695,710]" pageId="20" pageNumber="513">1</paragraph>
<paragraph id="8B9836CCFFF2E412FEE8FC8CFA87FCCC" blockId="20.[336,1286,858,929]" box="[336,1286,858,895]" pageId="20" pageNumber="513">0 0 1 2 3 4 5 6 7</paragraph>
<paragraph id="8B9836CCFFF2E412FD41FC44FCF8FC12" blockId="20.[336,1286,858,929]" box="[761,889,914,929]" pageId="20" pageNumber="513">Total length in cm</paragraph>
<caption id="DF586644FFF2E412FF3CFC1AFC26FC47" ID-DOI="http://doi.org/10.5281/zenodo.13985778" ID-Zenodo-Dep="13985778" httpUri="https://zenodo.org/record/13985778/files/figure.png" pageId="20" pageNumber="513" startId="20.[132,143,972,989]" targetBox="[353,1282,220,871]" targetPageId="20" targetType="figure">
<paragraph id="8B9836CCFFF2E412FF3CFC1AFC26FC47" blockId="20.[132,1455,972,1012]" pageId="20" pageNumber="513">
<smallCapsWord id="8D7EA010FFF2E412FF3CFC1AFF1CFC6F" baselines="985,985" box="[132,157,972,989]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="20" pageNumber="513">FIG</smallCapsWord>
. 9. — Bivariate plot of total length against width comparing
<taxonomicName id="4C274D4FFFF2E412FD33FC1AFC44FC6E" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" box="[651,965,972,989]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FD33FC1AFCBCFC6E" box="[651,829,972,989]" italics="true" pageId="20" pageNumber="513">Timbellus tripteroides</emphasis>
(
<bibRefCitation id="EFB64B3DFFF2E412FCFDFC1AFC40FC6E" author="LAMARCK J. - B." box="[837,961,972,989]" pageId="20" pageNumber="513" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck,1822</bibRefCitation>
)
</taxonomicName>
from the middle Lutetian of Paris Basin (65 complete specimens) and
<taxonomicName id="4C274D4FFFF2E412FF5DFC35FEDCFC47" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[229,349,995,1012]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
<emphasis id="B953EADEFFF2E412FF5DFC35FEDCFC47" box="[229,349,995,1012]" italics="true" pageId="20" pageNumber="513">T. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FEDAFC35FE13FC47" box="[354,402,995,1012]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
from the upper Lutetian of Normandy (18 complete specimens).
</paragraph>
</caption>
<paragraph id="8B9836CCFFF2E412FF3CFB62FE9EFB7F" blockId="20.[130,777,1204,2027]" box="[132,287,1204,1230]" pageId="20" pageNumber="513">
<smallCapsWord id="8D7EA010FFF2E412FF3CFB62FE9EFB7F" baselines="1223,1223" box="[132,287,1204,1230]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="20" pageNumber="513">COMPARISONS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFF2E412FF3CFB02FF78F858" blockId="20.[130,777,1204,2027]" pageId="20" pageNumber="513">
This species was previously attributed to
<taxonomicName id="4C274D4FFFF2E412FDF1FB02FEADFABD" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FDF1FB02FC88FB5D" box="[585,777,1236,1262]" italics="true" pageId="20" pageNumber="513">Murex tripteroides</emphasis>
<bibRefCitation id="EFB64B3DFFF2E412FF3CFB25FEADFABD" author="LAMARCK J. - B." box="[132,300,1267,1294]" pageId="20" pageNumber="513" refId="ref45696" refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p." type="book" year="1822">Lamarck, 1822</bibRefCitation>
</taxonomicName>
by
<bibRefCitation id="EFB64B3DFFF2E412FEE2FB22FD1FFABD" author="COSSMANN M. &amp; PISSARRO G." box="[346,670,1267,1294]" pageId="20" pageNumber="513" pagination="61 - 140" refId="ref43809" refString="COSSMANN M. &amp; PISSARRO G. 1901. - Faune eocenique du Cotentin. 2 eme article. Bulletin de la Societe Geologique de Normandie 20: 11 - 90. [Also published as Cossmann M. &amp; Pissarro G. 1901. - Faune eocenique du Cotentin (Mollusques), Tome 1, Fascicule 2, Imprimerie du Journal Le Havre, Le Havre, 61 - 140]." type="book chapter" year="1901">Cossmann &amp; Pissarro (1901)</bibRefCitation>
and later with some doubts by
<bibRefCitation id="EFB64B3DFFF2E412FE3DFAC5FDDEFA9D" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[389,607,1299,1326]" pageId="20" pageNumber="513" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFF2E412FE3DFAC5FE49FA9E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[389,456,1299,1325]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFF2E412FE77FAC2FD89FA9E" box="[463,520,1299,1325]" italics="true" pageId="20" pageNumber="513">et al.</emphasis>
(2011)
</bibRefCitation>
. According to those authors, there are apparently strong similarities with
<taxonomicName id="4C274D4FFFF2E412FF3EFA82FE9BFADE" authorityName="Lamarck" authorityYear="1822" box="[134,282,1363,1389]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FF3EFA82FE9BFADE" box="[134,282,1363,1389]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
such as: a bulbous protoconch, an elongate shape, developed internal denticles, and few and weak cords. However,
<taxonomicName id="4C274D4FFFF2E412FF49FA42FE00FA1E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[241,385,1427,1453]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
<emphasis id="B953EADEFFF2E412FF49FA42FE00FA1E" box="[241,385,1427,1453]" italics="true" pageId="20" pageNumber="513">T. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FE30FA42FE43FA1D" box="[392,450,1428,1454]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
is smaller and does not exceed
<quantity id="4CDF9B29FFF2E412FF3BFA65FF50FA7E" box="[131,209,1459,1485]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="20" pageNumber="513" unit="mm" value="26.0">26 mm</quantity>
for five teleoconch whorls. At the same size and same number of whorls
<taxonomicName id="4C274D4FFFF2E412FEFDFA02FE50FA5E" authorityName="Lamarck" authorityYear="1822" box="[325,465,1491,1517]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FEFDFA02FE50FA5E" box="[325,465,1491,1517]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
is still juvenile, does not bear internal denticles, and their axial wings are only developed on the last whorl (
<figureCitation id="131C2A49FFF2E412FE9BF9C4FE17F99E" box="[291,406,1554,1581]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de lOrme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="20" pageNumber="513">Fig. 8D, E</figureCitation>
). Adults of
<taxonomicName id="4C274D4FFFF2E412FDB6F9C5FD1DF99F" authorityName="Lamarck" authorityYear="1822" box="[526,668,1554,1580]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FDB6F9C5FD1DF99F" box="[526,668,1554,1580]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
can reach more than
<quantity id="4CDF9B29FFF2E412FF41F9E4FEC6F9FE" box="[249,327,1586,1613]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="20" pageNumber="513" unit="mm" value="60.0">60 mm</quantity>
height at 7-8 whorls. A bivariate diagram (
<figureCitation id="131C2A49FFF2E412FF35F984FF49F9DF" box="[141,200,1618,1644]" captionStart="FIG" captionStartId="20.[132,143,972,989]" captionTargetBox="[353,1282,220,871]" captionTargetId="figure-560@20.[347,911,217,783]" captionTargetPageId="20" captionText="FIG. 9. — Bivariate plot of total length against width comparing Timbellus tripteroides (Lamarck,1822) from the middle Lutetian of Paris Basin (65 complete specimens) and T. occidentalis n. sp. from the upper Lutetian of Normandy (18 complete specimens)." figureDoi="http://doi.org/10.5281/zenodo.13985778" httpUri="https://zenodo.org/record/13985778/files/figure.png" pageId="20" pageNumber="513">Fig. 9</figureCitation>
) illustrates the differences in size between
<taxonomicName id="4C274D4FFFF2E412FDC0F985FC84F9DF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[632,773,1618,1644]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
<emphasis id="B953EADEFFF2E412FDC0F985FC84F9DF" box="[632,773,1618,1644]" italics="true" pageId="20" pageNumber="513">T. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FF3AF9A5FF3AF93E" box="[130,187,1651,1677]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
from
<collectingRegion id="49E3F82EFFF2E412FF42F9A4FEEEF93F" box="[250,367,1650,1676]" country="France" name="Haute-Normandie" pageId="20" pageNumber="513">Normandy</collectingRegion>
and
<taxonomicName id="4C274D4FFFF2E412FE26F9A5FDA8F93F" authorityName="Lamarck" authorityYear="1822" box="[414,553,1650,1676]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FE26F9A5FDA8F93F" box="[414,553,1650,1676]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
from the Paris Basin. In addition, the spiral cords of
<taxonomicName id="4C274D4FFFF2E412FE69F945FDDEF91F" authorityName="Lamarck" authorityYear="1822" box="[465,607,1682,1708]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FE69F945FDDEF91F" box="[465,607,1682,1708]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFF2E412FDD7F944FD2FF91F" box="[623,686,1682,1708]" captionStart="FIG" captionStartId="16.[132,143,1467,1484]" captionTargetBox="[132,1455,215,1424]" captionTargetId="figure-382@16.[698,1267,772,1332]" captionTargetPageId="16" captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann &amp; Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart." figureDoi="http://doi.org/10.5281/zenodo.13985772" httpUri="https://zenodo.org/record/13985772/files/figure.png" pageId="20" pageNumber="513">Figs 6</figureCitation>
;
<figureCitation id="131C2A49FFF2E412FD04F944FD7EF91F" box="[700,767,1682,1709]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de lOrme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="20" pageNumber="513">8D, E</figureCitation>
) are usually more marked than those of
<taxonomicName id="4C274D4FFFF2E412FD89F965FD45F97F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[561,708,1714,1740]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
<emphasis id="B953EADEFFF2E412FD89F965FD45F97F" box="[561,708,1714,1740]" italics="true" pageId="20" pageNumber="513">T. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FD74F965FC89F97E" box="[716,776,1715,1741]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="131C2A49FFF2E412FF35F904FF4CF95F" box="[141,205,1746,1772]" captionStart="FIG" captionStartId="19.[132,143,711,728]" captionTargetBox="[132,1455,215,669]" captionTargetId="figure-804@19.[160,986,229,644]" captionTargetPageId="19" captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de lOrme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes." figureDoi="http://doi.org/10.5281/zenodo.13985776" httpUri="https://zenodo.org/record/13985776/files/figure.png" pageId="20" pageNumber="513">Fig. 8</figureCitation>
A-C) Finally,
<taxonomicName id="4C274D4FFFF2E412FEDCF905FE79F95F" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[356,504,1746,1772]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="n. sp.">
<emphasis id="B953EADEFFF2E412FEDCF905FE79F95F" box="[356,504,1746,1772]" italics="true" pageId="20" pageNumber="513">T. occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FDB9F904FDBCF95F" box="[513,573,1746,1772]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
could be regarded as a progenetic species of
<taxonomicName id="4C274D4FFFF2E412FE1BF924FD89F8BF" box="[419,520,1778,1804]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFF2E412FE1BF924FD89F8BF" box="[419,520,1778,1804]" italics="true" pageId="20" pageNumber="513">Timbellus</emphasis>
</taxonomicName>
, because it associates a small size with adult characters comparable to
<taxonomicName id="4C274D4FFFF2E412FDC9F8C4FD7EF89F" authorityName="Lamarck" authorityYear="1822" box="[625,767,1810,1836]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FDC9F8C4FD7EF89F" box="[625,767,1810,1836]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
. Moreover, the two species never co-occur.
<taxonomicName id="4C274D4FFFF2E412FD95F8E7FC84F8F8" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[557,773,1841,1867]" class="Gastropoda" family="Muricidae" genus="Timbellus" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="occidentalis" status="sp. nov.">
<emphasis id="B953EADEFFF2E412FD95F8E7FC84F8F8" box="[557,773,1841,1867]" italics="true" pageId="20" pageNumber="513">Timbellus occidentalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFF2E412FF3AF884FF3FF8DF" box="[130,190,1874,1900]" pageId="20" pageNumber="513" rank="species">n. sp.</taxonomicNameLabel>
ranges between the uppermost Lutetian (
<collectingRegion id="49E3F82EFFF2E412FD3EF887FC81F8D8" box="[646,768,1873,1899]" country="France" name="Haute-Normandie" pageId="20" pageNumber="513">Normandy</collectingRegion>
) and the lowermost the Bartonian (Loire-Atlantique), whereas
<taxonomicName id="4C274D4FFFF2E412FF3AF844FE93F818" authorityName="Lamarck" authorityYear="1822" box="[130,274,1937,1963]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="20" pageNumber="513" phylum="Mollusca" rank="species" species="tripteroides">
<emphasis id="B953EADEFFF2E412FF3AF844FE93F818" box="[130,274,1937,1963]" italics="true" pageId="20" pageNumber="513">T. tripteroides</emphasis>
</taxonomicName>
ranges from the lower to the middle Lutetian in the Paris Basin and in the
<collectingRegion id="49E3F82EFFF2E412FE74F867FDC7F878" box="[460,582,1969,1995]" country="United Kingdom" name="Hampshire" pageId="20" pageNumber="513">Hampshire</collectingRegion>
Basin during the Bartonian.
</paragraph>
</subSubSection>
</treatment>
</document>