<documentid="183ED8B85060B9E1321DB5F0756BBD38"ID-CLB-Dataset="304811"ID-DOI="10.5252/geodiversitas2024v46a15"ID-GBIF-Dataset="fe16163e-5e9d-4c80-89c8-fce3fb2e064f"ID-ISSN="1638-9395"ID-Zenodo-Dep="13985760"ID-ZooBank="urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713"IM.bibliography_approvedBy="valdenar"IM.illustrations_approvedBy="valdenar"IM.materialsCitations_approvedBy="valdenar"IM.metadata_approvedBy="valdenar"IM.tables_requiresApprovalFor="GgImagineBatch"IM.taxonomicNames_approvedBy="valdenar"IM.treatmentCitations_approvedBy="valdenar"IM.treatments_approvedBy="valdenar"checkinTime="1729758589633"checkinUser="plazi"docAuthor="Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard"docDate="2024"docId="038E87DAFFF5E412FE95FB23FF78F858"docLanguage="en"docName="geodiversitas.2024.46.15.pdf"docOrigin="Geodiversitas 46 (15)"docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a15.pdf"docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article"docStyleId="F830B10FF475E64C1F1601E3B32DDC00"docStyleName="Geodiversitas.2018-.journal_article"docStyleVersion="4"docTitle="Timbellus occidentalis Merle, Pacaud, Ledon & Goret, 2024, n. sp."docType="treatment"docUuid="53A32E50-A05D-4A09-B8B1-6CA776FDC29F"docUuidSource="ZooBank"docVersion="9"lastPageNumber="513"masterDocId="FFB7FFA2FFE6E406FFB8FFD6FF81FFB3"masterDocTitle="New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe"masterLastPageNumber="551"masterPageNumber="495"pageNumber="512"updateTime="1732586620909"updateUser="ExternalLinkService"zenodo-license-document="CC0-1.0"zenodo-license-figures="CC0-1.0">
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<figureCitationid="131C2A49FFF5E415FED9FAC2FE20FA9D"box="[353,417,1300,1326]"captionStart="FIG"captionStartId="19.[132,143,711,728]"captionTargetBox="[132,1455,215,669]"captionTargetId="figure-804@19.[160,986,229,644]"captionTargetPageId="19"captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes."figureDoi="http://doi.org/10.5281/zenodo.13985776"httpUri="https://zenodo.org/record/13985776/files/figure.png"pageId="19"pageNumber="512">Figs 8</figureCitation>
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.F.A90533 (ex A24100 ex galerie de zoologie coll.,
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;
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.F.A90534 (ex A24100 ex galerie de zoologie coll., Le Marchand coll.)
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.F.B63610 (de Morgan coll.) – Gourbesville, Manche, middle Eocene (upper Lutetian)
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.F.A24031 (Vasseur coll.), figured in
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<bibRefCitationid="EFB64B3DFFF5E415FD12F805FCE3FCDF"author="LAMARCK J. - B."box="[682,866,852,2027]"pageId="19"pageNumber="512"refId="ref45696"refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p."type="book"year="1822">Lamarck, 1822</bibRefCitation>
)
</taxonomicName>
(
<figureCitationid="131C2A49FFF5E415FCCFFC82FC44FCDF"box="[887,965,852,876]"captionStart="FIG"captionStartId="19.[132,143,711,728]"captionTargetBox="[132,1455,215,669]"captionTargetId="figure-804@19.[160,986,229,644]"captionTargetPageId="19"captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes."figureDoi="http://doi.org/10.5281/zenodo.13985776"httpUri="https://zenodo.org/record/13985776/files/figure.png"pageId="19"pageNumber="512">Figs 8C</figureCitation>
;
<figureCitationid="131C2A49FFF5E415FC69FC82FC7BFCDF"box="[977,1018,852,876]"captionStart="FIG"captionStartId="45.[132,143,1622,1639]"captionTargetBox="[132,1455,215,1562]"captionTargetId="figure-370@45.[700,1259,799,1371]"captionTargetPageId="45"captionText="FIG. 26. — New species of Timbellus De Gregorio, 1885: A, B, Timbellus occidentalis n. sp.: A, holotype, MNHN.F.A90533 (Le Marchand coll.), Fresville, France, upper Lutetian;B, paratype,MNHN.F.A24031 (Vasseur coll.),Saffré (Bois-Gouët),France,Bartonian;C, D, T.calciacus n. sp.: C, holotype,MNHN.F.A90536 (Merle coll.), Chaussy (Les Garennes), France, Lutetian; D, paratype, MNHN.F.A90537 (Merle coll.), Chaussy (Les Garennes); E, F, T. longicanalis n. sp.: E, holotype, MNHN.F.A24101 (Le Marchand coll.), Fresville, France, upper Lutetian; F, paratype, MNHN.F.A24100, Fresville; G-I, T. magnei n. name: G, MNHN.F.A32508 (Merle coll.), juvenile specimen, Gan (Tuilerie), France; H, MNHN.F.A90545 (Merle coll.), juvenile specimen with protoconch, Gan (Tuilerie); I, MNHN.F.A72443 (Varone coll.), Gan (Tuilerie); J-L, T. curvispina n. sp.: J, holotype, MNHN.F.A90541 (MNHN coll.), Parnes, France, Lutetian; K, paratype, MNHN.F.A90543 (Pons coll.), Fontenay-en-Vexin, France, Lutetian; L, paratype, MNHN.F.A73848 (Pons coll.), Fontenay-en-Vexin; M, T. sixi n. sp., holotype, MNHN.F.A90544 (Six coll.), Trosly-Breuil (Fond Couturier), France, Ypresian; N-P, T. radulfiensis n. sp.: N, holotype, MNHN.F.A77754 (MNHN coll.), Rauville-la-Place, France, Priabonian; O, paratype, MNHN.F.A77755 (MNHN coll.), Rauville-la-Place; P, paratype, MNHN.F.A90559 (Vasseur coll.), Rauville-la-Place. Scale bars: A,B, N-P, 5 mm; C, D, 3 mm; E-G, I-L, 10 mm; H, 2 mm."figureDoi="http://doi.org/10.5281/zenodo.13985825"httpUri="https://zenodo.org/record/13985825/files/figure.png"pageId="19"pageNumber="512">26B</figureCitation>
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.F.J06383 (Vasseur coll.), figured in
<bibRefCitationid="EFB64B3DFFF5E415FC69FCB9FB00FC34"author="MERLE D. & GARRIGUES B. & POINTIER J. - P."box="[977,1153,879,903]"pageId="19"pageNumber="512"refId="ref47234"refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p."type="book"year="2011">
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<bibRefCitationid="EFB64B3DFFF5E415FC19FC5FFBBBFC12"author="LAMARCK J. - B."box="[929,1082,905,929]"pageId="19"pageNumber="512"refId="ref45696"refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p."type="book"year="1822">Lamarck, 1822</bibRefCitation>
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. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see
<bibRefCitationid="EFB64B3DFFF5E415FC0EFBEEFBE6FBE3"author="BIGNOT G. & HOMMERIL P. & LARSONNEUR C."box="[950,1127,1080,1104]"pageId="19"pageNumber="512"pagination="405 - 416"refId="ref42941"refString="BIGNOT G., HOMMERIL P. & LARSONNEUR C. 1968. - Le Lutetien au large du Cotentin, in Colloque sur l'Eocene (Paris, mai 1968). Memoires BRGM 58: 405 - 416."type="journal article"year="1968">
<bibRefCitationid="EFB64B3DFFF5E415FC96FB54FC3DFB29"author="APGN & ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE"box="[814,956,1154,1178]"pageId="19"pageNumber="512"refId="ref42597"refString="APGN (ASSOCIATION PATRIMOINE GEOLOGIQUE DE NORMANDIE) 2024. - Faluns cretaces et tertiaires de Fresville, BNO 0182. Available at: http: // www. apgn. fr / apgn / carte _ site / fic _ s 50 _ 75. pdf (last access on 03 / 2024)."type="book"year="2024">APGN (2024)</bibRefCitation>
(Calcaire de Fresville and Faluns de Hautteville) middle Eocene (uppermost Lutetian) and
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Basin (Sables du Bois-Gouët), lowermost Bartonian see
<bibRefCitationid="EFB64B3DFFF5E415FA82FB30FC39FAAA"author="MATHELIN J. - C. & BIGNOT G."pageId="19"pageNumber="512"pagination="55 - 70"refId="ref46304"refString="MATHELIN J. - C. & BIGNOT G. 1989. - Le falun de Foulangues et ses relations stratigraphique et paleogeographique avec le Biarritzien stratotypique, in 114 eme Congres national des Societes savantes. Geologie du Bassin Parisien. CTHS, Paris: 55 - 70."type="book chapter"year="1989">Mathelin & Bignot (1989)</bibRefCitation>
in width, biconic in profile, composed of five whorls. Moderately high spire. Last whorl up to 85% total length. Apical angle 64° excluding spines, 91° including spines. Spiral sculpture with weak primary cords in adult specimens. First to fourth whorl, no cord. Fifth whorl: P1 to P3. Sixth whorl: P1 to P6 weak, s2 and no other cord below, ADP and MP occasional. On first whorl, eight protovarices. From second to last whorl, three varices and one intervarix intercalated Higher relief of intervarices on P2-P3. Varices lamellose, winged on last whorls. Appearance of short, straight P1 spine on third whorl. Sinus of P1 spine narrow and pointing adapically. On last whorl alate varices. Ovate aperture up to 9% of diameter and up to 80% of length of last whorl (including siphonal canal). Columellar lip smooth, narrow, slightly erect anteriorly, forming curved inductura at base. Parietal lip slightly adherent. Outer lip with denticles ID (occasional), D1 to D6, D1 and D2 more strongly developed. Outer lip not crenulate. Siphonal canal narrow, open, up to 49% of apertural length. Pseudo-umbilicus narrow.
<paragraphid="8B9836CCFFF2E412FD41FC44FCF8FC12"blockId="20.[336,1286,858,929]"box="[761,889,914,929]"pageId="20"pageNumber="513">Total length in cm</paragraph>
<bibRefCitationid="EFB64B3DFFF2E412FCFDFC1AFC40FC6E"author="LAMARCK J. - B."box="[837,961,972,989]"pageId="20"pageNumber="513"refId="ref45696"refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p."type="book"year="1822">Lamarck,1822</bibRefCitation>
)
</taxonomicName>
from the middle Lutetian of Paris Basin (65 complete specimens) and
<taxonomicNameid="4C274D4FFFF2E412FF5DFC35FEDCFC47"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[229,349,995,1012]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="n. sp.">
<bibRefCitationid="EFB64B3DFFF2E412FF3CFB25FEADFABD"author="LAMARCK J. - B."box="[132,300,1267,1294]"pageId="20"pageNumber="513"refId="ref45696"refString="LAMARCK J. - B. 1822. - Histoire naturelle des animaux sans vertebres presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent et Supplement a divers types de gasteropodes et de Trachelipodes comprenant l'indication des coquilles fossiles qui ne furent point citees dans leurs genres respectifs. Tome 7. Lamarck, Paris, 711 p."type="book"year="1822">Lamarck, 1822</bibRefCitation>
</taxonomicName>
by
<bibRefCitationid="EFB64B3DFFF2E412FEE2FB22FD1FFABD"author="COSSMANN M. & PISSARRO G."box="[346,670,1267,1294]"pageId="20"pageNumber="513"pagination="61 - 140"refId="ref43809"refString="COSSMANN M. & PISSARRO G. 1901. - Faune eocenique du Cotentin. 2 eme article. Bulletin de la Societe Geologique de Normandie 20: 11 - 90. [Also published as Cossmann M. & Pissarro G. 1901. - Faune eocenique du Cotentin (Mollusques), Tome 1, Fascicule 2, Imprimerie du Journal Le Havre, Le Havre, 61 - 140]."type="book chapter"year="1901">Cossmann & Pissarro (1901)</bibRefCitation>
and later with some doubts by
<bibRefCitationid="EFB64B3DFFF2E412FE3DFAC5FDDEFA9D"author="MERLE D. & GARRIGUES B. & POINTIER J. - P."box="[389,607,1299,1326]"pageId="20"pageNumber="513"refId="ref47234"refString="MERLE D., GARRIGUES B. & POINTIER J. - P. 2011. - Fossil & Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p."type="book"year="2011">
<taxonomicNameid="4C274D4FFFF2E412FE3DFAC5FE49FA9E"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[389,456,1299,1325]"class="Gastropoda"family="Muricidae"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="543"phylum="Mollusca"rank="subFamily"status="subfam. nov."subFamily="Merle">Merle</taxonomicName>
such as: a bulbous protoconch, an elongate shape, developed internal denticles, and few and weak cords. However,
<taxonomicNameid="4C274D4FFFF2E412FF49FA42FE00FA1E"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[241,385,1427,1453]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="n. sp.">
is still juvenile, does not bear internal denticles, and their axial wings are only developed on the last whorl (
<figureCitationid="131C2A49FFF2E412FE9BF9C4FE17F99E"box="[291,406,1554,1581]"captionStart="FIG"captionStartId="19.[132,143,711,728]"captionTargetBox="[132,1455,215,669]"captionTargetId="figure-804@19.[160,986,229,644]"captionTargetPageId="19"captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes."figureDoi="http://doi.org/10.5281/zenodo.13985776"httpUri="https://zenodo.org/record/13985776/files/figure.png"pageId="20"pageNumber="513">Fig. 8D, E</figureCitation>
<figureCitationid="131C2A49FFF2E412FF35F984FF49F9DF"box="[141,200,1618,1644]"captionStart="FIG"captionStartId="20.[132,143,972,989]"captionTargetBox="[353,1282,220,871]"captionTargetId="figure-560@20.[347,911,217,783]"captionTargetPageId="20"captionText="FIG. 9. — Bivariate plot of total length against width comparing Timbellus tripteroides (Lamarck,1822) from the middle Lutetian of Paris Basin (65 complete specimens) and T. occidentalis n. sp. from the upper Lutetian of Normandy (18 complete specimens)."figureDoi="http://doi.org/10.5281/zenodo.13985778"httpUri="https://zenodo.org/record/13985778/files/figure.png"pageId="20"pageNumber="513">Fig. 9</figureCitation>
) illustrates the differences in size between
<taxonomicNameid="4C274D4FFFF2E412FDC0F985FC84F9DF"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[632,773,1618,1644]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="n. sp.">
<figureCitationid="131C2A49FFF2E412FDD7F944FD2FF91F"box="[623,686,1682,1708]"captionStart="FIG"captionStartId="16.[132,143,1467,1484]"captionTargetBox="[132,1455,215,1424]"captionTargetId="figure-382@16.[698,1267,772,1332]"captionTargetPageId="16"captionText="FIG. 6. — Neotype, historical figures, spiral cords and denticles of the outer lip in T. tripteroides (Lamarck, 1822): A-D, morphotype with rounded varices and poorly distinguishable P1 spine; E-H, morphotype with easily distinguishable P1 spine: A, original figure of Lamarck (1805: pl. 3, fig. 4); B, C, neotype designated herein, MNHN.F.A90546 (Club géologique d'Île-de-France leg.), Thiverval-Grignon (Falunière), Yvelines, Paris Basin, France, middle Eocene (Lutetian); D, MNHN.F.A90552 (Pacaud coll.), Chaussy (Les Garennes),Paris Basin, France,middle Eocene (Lutetian); E, F, figured specimen of Deshayes (1835: pl. 82, fig. 1-2); G, H, figured specimen of Cossmann & Pissarro (1911: pl. 35, fig. 169-1; MNHN.F.J10350 (Cossmann coll.), Chaussy (Les Garennes). Scale bars: 20 mm. Credits: B-D, P. Loubry MNHN/CNRS; G, H, J. Mouchart."figureDoi="http://doi.org/10.5281/zenodo.13985772"httpUri="https://zenodo.org/record/13985772/files/figure.png"pageId="20"pageNumber="513">Figs 6</figureCitation>
;
<figureCitationid="131C2A49FFF2E412FD04F944FD7EF91F"box="[700,767,1682,1709]"captionStart="FIG"captionStartId="19.[132,143,711,728]"captionTargetBox="[132,1455,215,669]"captionTargetId="figure-804@19.[160,986,229,644]"captionTargetPageId="19"captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes."figureDoi="http://doi.org/10.5281/zenodo.13985776"httpUri="https://zenodo.org/record/13985776/files/figure.png"pageId="20"pageNumber="513">8D, E</figureCitation>
) are usually more marked than those of
<taxonomicNameid="4C274D4FFFF2E412FD89F965FD45F97F"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[561,708,1714,1740]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="n. sp.">
<figureCitationid="131C2A49FFF2E412FF35F904FF4CF95F"box="[141,205,1746,1772]"captionStart="FIG"captionStartId="19.[132,143,711,728]"captionTargetBox="[132,1455,215,669]"captionTargetId="figure-804@19.[160,986,229,644]"captionTargetPageId="19"captionText="FIG. 8. — Spiral cords and internal denticles of the outer lip in Timbellus occidentalisn. sp.(A-C) and T. tripteroides (Lamarck,1822) for comparison (D-E): A, B, holotype,MNHN.F.A90533 (Le Marchand coll.), Fresville,Manche,Normandy,France, middle Eocene (upper Lutetian);C, paratype,MNHN.F.A24031 (Vasseur coll.), Saffré (Bois-Gouët), Loire-Atlantique, Loire Basin, France, middle Eocene (Bartonian); D, E, MNHN.F.A90540 (Merle coll.), Beynes (Ferme de l’Orme), Yvelines, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-C, P. Loubry; D, E, L. Cazes."figureDoi="http://doi.org/10.5281/zenodo.13985776"httpUri="https://zenodo.org/record/13985776/files/figure.png"pageId="20"pageNumber="513">Fig. 8</figureCitation>
A-C) Finally,
<taxonomicNameid="4C274D4FFFF2E412FEDCF905FE79F95F"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[356,504,1746,1772]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="n. sp.">
<taxonomicNameid="4C274D4FFFF2E412FD95F8E7FC84F8F8"authorityName="Merle & Pacaud & Ledon & Goret"authorityYear="2024"box="[557,773,1841,1867]"class="Gastropoda"family="Muricidae"genus="Timbellus"kingdom="Animalia"order="Neogastropoda"pageId="20"pageNumber="513"phylum="Mollusca"rank="species"species="occidentalis"status="sp. nov.">