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<document id="20677F8C1ED4FC1A7454D5A458AB70B2" ID-DOI="10.21248/contrib.entomol.67.2.325-352" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742448" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267809870" checkinUser="felipe" docAuthor="Zorn, Carsten, Kobayashi, Hirokazu &amp; Wada, Kaoru" docDate="2017" docId="03826012FFD1FFAE2B730091CA22FDEB" docLanguage="en" docName="ContribEntomol.67.2.325-352.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Anomala recordata Zorn &amp; Kobayashi &amp; Wada 2017, spec. nov." docType="treatment" docUuid="53189F43-2F86-4D8B-B41D-9E5E4BD47013" docUuidSource="ZooBank" docVersion="3" lastPageNumber="334" masterDocId="FFBB186AFFD7FFA7280F027BCF12FFB3" masterDocTitle="Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records" masterLastPageNumber="352" masterPageNumber="325" pageNumber="331" updateTime="1699436963292" updateUser="plazi">
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<mods:title id="F76E7601627D3CD4F92494AE6016BC4F">Notes on the genus Anomala SAMOUELLE, 1819 (Coleoptera, Scarabaeidae, Rutelinae) in Vietnam and neighboring regions: eight new species and faunistic records</mods:title>
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<mods:namePart id="C1484947EBA127417DF4B778AFFFBB75">Zorn, Carsten</mods:namePart>
</mods:name>
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<mods:namePart id="D9A313DD46A070D3C73B123702A39763">Kobayashi, Hirokazu</mods:namePart>
</mods:name>
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<mods:namePart id="22440E180F38C92711697078F4E2161C">Wada, Kaoru</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8AD033E0ADB454840A2DE035BE02F2F">text</mods:typeOfResource>
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<mods:titleInfo id="FE8C5D728896DACDB75B06E0986F8F4C">
<mods:title id="2F4C06AE2A24021519B38CEB3CB3E9FF">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="CCB1411F20F4D09B662DD45BB7DCC9E6">
<mods:date id="EA647B61957CCB6EE16844595810A81B">2017</mods:date>
<mods:detail id="3B3C2B3069824B77A440902DAB4253F3" type="volume">
<mods:number id="CB7DFED898361A5E6477151A5B537D13">67</mods:number>
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<mods:identifier id="58A3E1945804F01637223430DDBD0D91" type="DOI">10.21248/contrib.entomol.67.2.325-352</mods:identifier>
<mods:identifier id="85FFAADB06D7CF80CC043689CE8A5DDD" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="D3C0C9AD506DAAAAC22555BEE3F16661" type="Zenodo-Dep">5742448</mods:identifier>
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<treatment id="03826012FFD1FFAE2B730091CA22FDEB" LSID="urn:lsid:zoobank.org:act:53189F43-2F86-4D8B-B41D-9E5E4BD47013" httpUri="http://treatment.plazi.org/id/03826012FFD1FFAE2B730091CA22FDEB" lastPageId="9" lastPageNumber="334" pageId="6" pageNumber="331">
<subSubSection id="C331828FFFD1FFA12B730091CA93FC95" pageId="6" pageNumber="331" type="nomenclature">
<paragraph id="8B94D104FFD1FFA12B730091CBBDFCB6" blockId="6.[892,1409,746,838]" box="[892,1199,746,773]" pageId="6" pageNumber="331">
<heading id="D0DC6668FFD1FFA12B730091CBBDFCB6" box="[892,1199,746,773]" fontSize="12" level="2" pageId="6" pageNumber="331" reason="2">
<taxonomicName id="4C2BAA87FFD1FFA12B730091CB52FCB6" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[892,1088,746,773]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="331" phylum="Arthropoda" rank="species" species="recordata" status="sp. nov.">
<emphasis id="B95F0D16FFD1FFA12B730091CB52FCB6" box="[892,1088,746,773]" italics="true" pageId="6" pageNumber="331">Anomala recordata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD1FFA12C490091CBBDFCB6" box="[1094,1199,746,773]" pageId="6" pageNumber="331" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B94D104FFD1FFA12B730169CA93FC95" blockId="6.[892,1409,746,838]" box="[892,1409,786,806]" pageId="6" pageNumber="331">
<uri id="FFBADD06FFD1FFA12B730169CA93FC95" box="[892,1409,786,806]" pageId="6" pageNumber="331">
urn:lsid:zoobank.org:act:
<uuid id="FF8DEBD1FFD1FFA12C210169CA93FC95" box="[1070,1409,786,806]" pageId="6" pageNumber="331">53189F43-2F86-4D8B-B41D-9E5E4BD47013</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFD1FFA12B730149CCDCFCF5" box="[892,974,818,838]" pageId="6" pageNumber="331" type="description">
<paragraph id="8B94D104FFD1FFA12B730149CCDCFCF5" blockId="6.[892,1409,746,838]" box="[892,974,818,838]" pageId="6" pageNumber="331">
(
<figureCitation id="1310CD81FFD1FFA12B8F0149CCBDFCF5" box="[896,943,818,838]" captionStart="Figs 1–9" captionStartId="7.[174,214,2023,2045]" captionTargetBox="[214,1362,260,1985]" captionTargetId="figure-122@7.[203,1392,255,2011]" captionTargetPageId="7" captionText="Figs 1–9: Anomala spp., habitus – 1. A. okushimai spec. nov., holotype. 2. A. immeliorata spec. nov., holotype. 3. A. confrater spec. nov., holotype.4. A. imperspicabilis spec.nov., holotype.5. A. ordinata spec.nov., holotype. 6. A. recordata spec.nov., holotype. 7. A. asaitoae spec. nov., holotype. 8. A. asaitoae spec. nov., paratype (female). 9. A. herbacea spec. nov., holotype. Scale = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5742452" httpUri="https://zenodo.org/record/5742452/files/figure.png" pageId="6" pageNumber="331">Figs 6</figureCitation>
,
<figureCitation id="1310CD81FFD1FFA12BB80149CCD8FCF5" box="[951,970,818,838]" captionStart="Figs 14–17" captionStartId="11.[174,214,2052,2074]" captionTargetBox="[179,1404,258,2008]" captionTargetId="figure-99@11.[174,1421,251,2026]" captionTargetPageId="11" captionText="Figs 14–17: Anomala spp., aedeagus dorsal (a), ventral (b) and lateral (c). – 14. A. ordinata spec. nov., holotype. 15. A. recordata" figureDoi="http://doi.org/10.5281/zenodo.5742456" httpUri="https://zenodo.org/record/5742456/files/figure.png" pageId="6" pageNumber="331">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFD1FFA12B59011BCB98FC6B" pageId="6" pageNumber="331" type="materials_examined">
<paragraph id="8B94D104FFD1FFA12B59011BCB98FC6B" blockId="6.[854,1459,863,984]" pageId="6" pageNumber="331">
<emphasis id="B95F0D16FFD1FFA12B59011BCCC1FCC4" bold="true" box="[854,979,863,888]" pageId="6" pageNumber="331">Type series</emphasis>
:
<materialsCitation id="3B43DB59FFD1FFA12BEB011BCBCDFC2B" collectingDate="2009-04-15" collectingDateMax="2009-04-30" collectingDateMin="2009-04-15" collectionCode="NSMT" country="Vietnam" location="Quang Nam" pageId="6" pageNumber="331" specimenCount="1" stateProvince="Quang Nam" typeStatus="holotype">
<emphasis id="B95F0D16FFD1FFA12BEB011BCB58FCCB" bold="true" box="[996,1098,864,888]" pageId="6" pageNumber="331">
<typeStatus id="54906FA6FFD1FFA12BEB011BCB58FCCB" box="[996,1098,864,888]" pageId="6" pageNumber="331" type="holotype">Holotype</typeStatus>
</emphasis>
: , C.
<collectingCountry id="F33C9194FFD1FFA12CAD011BCA0FFCCB" box="[1186,1309,864,888]" name="Vietnam" pageId="6" pageNumber="331">VIETNAM</collectingCountry>
:
<collectingRegion id="49EF1FE6FFD1FFA12D24011BCAA0FCC4" box="[1323,1458,864,888]" country="Vietnam" name="Quang Nam" pageId="6" pageNumber="331">Quang Nam</collectingRegion>
<collectingRegion id="49EF1FE6FFD1FFA12B5901FBCCA7FC2B" box="[854,949,896,920]" country="Vietnam" name="Da Nang" pageId="6" pageNumber="331">Da Nang</collectingRegion>
<date id="FF95F7C4FFD1FFA12BB401FBCB6BFC2B" box="[955,1145,896,920]" pageId="6" pageNumber="331" value="2009-04-15" valueMax="2009-04-30" valueMin="2009-04-15">
<collectingDate id="EFD10E2CFFD1FFA12BB401FBCB6BFC2B" box="[955,1145,896,920]" pageId="6" pageNumber="331" value="2009-04-15" valueMax="2009-04-30" valueMin="2009-04-15">15-30. APR. 2009</collectingDate>
</date>
. (
<collectionCode id="ED3A49C1FFD1FFA12C8501FBCBCBFC2B" box="[1162,1241,896,920]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="6" pageNumber="331">NSMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B43DB59FFD1FFA12CE601FBCB97FC6B" collectingDate="2010-04-15" collectionCode="MFPC" country="Vietnam" elevation="2200" location="Ngoe Thinh" pageId="6" pageNumber="331" specimenCount="1" stateProvince="Kon Tum" typeStatus="paratype">
<typeStatus id="54906FA6FFD1FFA12CE601FBCA5BFC2B" box="[1257,1353,896,920]" pageId="6" pageNumber="331" type="paratype">Paratype</typeStatus>
: 1 , near
<location id="8EF487DFFFD1FFA12B5901DBCCC8FC0B" LSID="urn:lsid:plazi:treatment:03826012FFD1FFAE2B730091CA22FDEB:8EF487DFFFD1FFA12B5901DBCCC8FC0B" box="[854,986,928,952]" country="Vietnam" name="Ngoe Thinh" pageId="6" pageNumber="331" stateProvince="Kon Tum">Ngoe Thinh</location>
, alt.
<quantity id="4CD37CE1FFD1FFA12C0301DBCB4FFC04" box="[1036,1117,928,952]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="331" unit="m" value="2200.0">
<elevation id="00063637FFD1FFA12C0301DBCB4FFC04" box="[1036,1117,928,952]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="331" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<collectingRegion id="49EF1FE6FFD1FFA12C6701DBCA1FFC04" box="[1128,1293,928,951]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="331">Kon Tum Prov.</collectingRegion>
, 1~
<date id="FF95F7C4FFD1FFA12D2201DBCABDFC0B" box="[1325,1455,928,952]" pageId="6" pageNumber="331" value="2010-04-15">
<collectingDate id="EFD10E2CFFD1FFA12D2201DBCABDFC0B" box="[1325,1455,928,952]" pageId="6" pageNumber="331" value="2010-04-15">15-IV-2010</collectingDate>
</date>
, native collector leg. (
<collectionCode id="ED3A49C1FFD1FFA12C3E01BBCB6CFC6B" box="[1073,1150,960,984]" pageId="6" pageNumber="331">MFPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFD1FFAE2B590184CE69FA64" lastPageId="9" lastPageNumber="334" pageId="6" pageNumber="331" type="description">
<paragraph id="8B94D104FFD1FFA12B590184CA5FFB84" blockId="6.[854,1459,1023,2104]" pageId="6" pageNumber="331">
<emphasis id="B95F0D16FFD1FFA12B590184CCCAFBA4" bold="true" box="[854,984,1023,1047]" pageId="6" pageNumber="331">Description</emphasis>
: Body shape elongate ovoid, strongly convex; body length: 18.0-19.0 mm, width: 9.0-
<quantity id="4CD37CE1FFD1FFA12CFB065BCA5BFB84" box="[1268,1353,1056,1080]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="331" unit="mm" value="9.5">9.5 mm</quantity>
.
</paragraph>
<paragraph id="8B94D104FFD1FFA12B590644CB39FA24" blockId="6.[854,1459,1023,2104]" pageId="6" pageNumber="331">
<emphasis id="B95F0D16FFD1FFA12B590644CC87FBE4" bold="true" box="[854,917,1087,1111]" pageId="6" pageNumber="331">Color</emphasis>
. Dorsal surface entirely vivid metallic green with weak iridescent shine except as follows: footstalk of antenna reddish brown, pronotum with orange band along lateral margin, pygidium with orange spot at apex, elytra with a transverse zig-zag band before the middle, consisting of three pairs of orange patches appearing on primary costae 2, 3 and 4, the latter encompassing parts of the neighboring interstices; ventral surface metallic green except as follows: sides of sternal segments orange, femora orange, abdominal ventrites reddish copper with orange lateral spots.
</paragraph>
<paragraph id="8B94D104FFD1FFA12B5907DBCCDAF944" blockId="6.[854,1459,1023,2104]" pageId="6" pageNumber="331">
<emphasis id="B95F0D16FFD1FFA12B5907DBCC82FA0B" bold="true" box="[854,912,1440,1464]" pageId="6" pageNumber="331">Head</emphasis>
with sparse erect pale yellow setae (
<quantity id="4CD37CE1FFD1FFA12D0107DBCA85FA04" box="[1294,1431,1440,1464]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="6" pageNumber="331" unit="mm" value="0.375" valueMax="0.5" valueMin="0.25">0.25-0.5 mm</quantity>
in length) along eyes; interocular distance equals 0.84 times the maximum transverse head width. Clypeus trapezoidal with rounded front angles and anterior margin; densely, somewhat rugosely punctate, punctures round (ca.
<quantity id="4CD37CE1FFD1FFA12B85043BCCFAF9E4" box="[906,1000,1600,1624]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="331" unit="mm" value="0.05">0.05 mm</quantity>
in diameter); anterior margin distinctly reflexed; fronto-clypeal suture weakly sinuate. Frons densely punctate, the punctures round in the middle, becoming larger anteriad, sparser posteriad. Vertex irregularly scattered with small, somewhat transverse punctures.
</paragraph>
<paragraph id="8B94D104FFD1FFA12B59057BCB21F78B" blockId="6.[854,1459,1023,2104]" pageId="6" pageNumber="331">
<emphasis id="B95F0D16FFD1FFA12B59057BCCD5F8AB" bold="true" box="[854,967,1792,1816]" pageId="6" pageNumber="331">Pronotum</emphasis>
1.8 times as wide as long; sides arcuate in the middle; convergent and straight in apical 2/5, subparallel basad, widest at base; lateral marginal line almost reaching the hind angles; basal marginal line weakly indicated medially of humerus; front angles produced and acute, hind angles obtuse, rounded off; disc densely covered with transverse moderately small punctures (ca.
<quantity id="4CD37CE1FFD1FFA12B8C059BCCF6F84B" box="[899,996,2016,2040]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="331" unit="mm" value="0.05">0.05 mm</quantity>
), each puncture bearing a microscopic seta; with sparse erect yellow setae (
<quantity id="4CD37CE1FFD1FFA12CBD0A7BCA29F7A4" box="[1202,1339,2048,2072]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="5.0" metricValueMin="3.7" pageId="6" pageNumber="331" unit="mm" value="0.435" valueMax="0.5" valueMin="0.37">0.37-0.5 mm</quantity>
in length) along lateral margin.
</paragraph>
<caption id="DF54818CFFD0FFA028A1059CCA6CF784" ID-DOI="http://doi.org/10.5281/zenodo.5742452" ID-Zenodo-Dep="5742452" httpUri="https://zenodo.org/record/5742452/files/figure.png" pageId="7" pageNumber="332" startId="7.[174,214,2023,2045]" targetBox="[214,1362,260,1985]" targetPageId="7">
<paragraph id="8B94D104FFD0FFA028A1059CCA6CF784" blockId="7.[174,1422,2023,2104]" pageId="7" pageNumber="332">
<emphasis id="B95F0D16FFD0FFA028A1059CCE11F84F" bold="true" box="[174,259,2023,2045]" pageId="7" pageNumber="332">Figs 1–9</emphasis>
:
<taxonomicName id="4C2BAA87FFD0FFA0291D059CCE89F84E" box="[274,411,2023,2045]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95F0D16FFD0FFA0291D059CCE7BF84F" box="[274,361,2023,2044]" italics="true" pageId="7" pageNumber="332">Anomala</emphasis>
spp.
</taxonomicName>
, habitus –
<emphasis id="B95F0D16FFD0FFA02A02059CCD0BF84F" bold="true" box="[525,537,2023,2044]" pageId="7" pageNumber="332">1</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02A280593CDB5F84F" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[551,679,2023,2045]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="okushimai" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02A280593CDB5F84F" box="[551,679,2023,2045]" italics="true" pageId="7" pageNumber="332">A. okushimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02ABE0593CC06F84E" box="[689,788,2024,2045]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA02B8C059CCC9DF84F" bold="true" box="[899,911,2023,2044]" pageId="7" pageNumber="332">2</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02B910593CB39F84F" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[926,1067,2023,2045]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="immeliorata" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02B910593CB39F84F" box="[926,1067,2023,2045]" italics="true" pageId="7" pageNumber="332">A. immeliorata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02C3A0593CB8AF84E" box="[1077,1176,2024,2045]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA02D08059CCA01F84F" bold="true" box="[1287,1299,2023,2044]" pageId="7" pageNumber="332">3</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02D110593CA9FF84F" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[1310,1421,2023,2045]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="confrater" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02D110593CA9FF84F" box="[1310,1421,2023,2045]" italics="true" pageId="7" pageNumber="332">A. confrater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA028A10A7ECE1FF7A9" box="[174,269,2053,2074]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA0297D0A7FCE6CF7AA" bold="true" box="[370,382,2052,2073]" pageId="7" pageNumber="332">4</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA029860A7ECD3DF7A9" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[393,559,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="imperspicabilis" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA029860A7ECD3DF7A9" box="[393,559,2053,2074]" italics="true" pageId="7" pageNumber="332">A. imperspicabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02A3C0A7ECD80F7A9" box="[563,658,2053,2074]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA02AF80A7ECC11F7A9" bold="true" box="[759,771,2053,2074]" pageId="7" pageNumber="332">5</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02B030A7ECC6AF7A9" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[780,888,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="ordinata" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02B030A7ECC6AF7A9" box="[780,888,2053,2074]" italics="true" pageId="7" pageNumber="332">A. ordinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02B730A7ECCC8F7A9" box="[892,986,2053,2074]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA02C4F0A7FCB5EF7AA" bold="true" box="[1088,1100,2052,2073]" pageId="7" pageNumber="332">6</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02C590A7ECBDBF7A9" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[1110,1225,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="recordata" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02C590A7ECBDBF7A9" box="[1110,1225,2053,2074]" italics="true" pageId="7" pageNumber="332">A. recordata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02CC10A7ECA3FF7A9" box="[1230,1325,2053,2074]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA028A10A59CFA8F784" bold="true" box="[174,186,2082,2103]" pageId="7" pageNumber="332">7</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA028CA0A59CE3FF78B" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[197,301,2082,2104]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="asaitoae" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA028CA0A59CE3FF78B" box="[197,301,2082,2104]" italics="true" pageId="7" pageNumber="332">A. asaitoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA0293D0A59CE83F784" box="[306,401,2082,2103]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFD0FFA029F60A59CD17F784" bold="true" box="[505,517,2082,2103]" pageId="7" pageNumber="332">8</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02A000A59CD65F78B" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[527,631,2082,2104]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="asaitoae" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02A000A59CD65F78B" box="[527,631,2082,2104]" italics="true" pageId="7" pageNumber="332">A. asaitoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02A720A59CDCEF784" box="[637,732,2082,2103]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, paratype (female).
<emphasis id="B95F0D16FFD0FFA02B970A59CCB6F784" bold="true" box="[920,932,2082,2103]" pageId="7" pageNumber="332">9</emphasis>
.
<taxonomicName id="4C2BAA87FFD0FFA02BA00A59CB0CF784" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[943,1054,2082,2103]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="332" phylum="Arthropoda" rank="species" species="herbacea" status="sp. nov.">
<emphasis id="B95F0D16FFD0FFA02BA00A59CB0CF784" box="[943,1054,2082,2103]" italics="true" pageId="7" pageNumber="332">A. herbacea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFD0FFA02C2C0A59CB91F784" box="[1059,1155,2082,2103]" pageId="7" pageNumber="332" rank="species">spec. nov.</taxonomicNameLabel>
, holotype. Scale = 10 mm.
</paragraph>
</caption>
<caption id="DF54818CFFDFFFAF28DB0A7FCAA1F7A9" ID-DOI="http://doi.org/10.5281/zenodo.5742454" ID-Zenodo-Dep="5742454" box="[212,1459,2052,2075]" httpUri="https://zenodo.org/record/5742454/files/figure.png" pageId="8" pageNumber="333" startId="8.[212,252,2052,2074]" targetBox="[223,1440,268,2015]" targetPageId="8">
<paragraph id="8B94D104FFDFFFAF28DB0A7FCAA1F7A9" blockId="8.[212,1459,2052,2104]" box="[212,1459,2052,2075]" pageId="8" pageNumber="333">
<emphasis id="B95F0D16FFDFFFAF28DB0A7FCE2EF7AA" bold="true" box="[212,316,2052,2074]" pageId="8" pageNumber="333">Figs 10–13</emphasis>
:
<taxonomicName id="4C2BAA87FFDFFFAF29490A7ECED9F7A9" box="[326,459,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="333" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95F0D16FFDFFFAF29490A7ECE8FF7A9" box="[326,413,2053,2074]" italics="true" pageId="8" pageNumber="333">Anomala</emphasis>
spp.
</taxonomicName>
, aedeagus dorsal (
<emphasis id="B95F0D16FFDFFFAF2A790A7ECD93F7A9" bold="true" box="[630,641,2053,2074]" pageId="8" pageNumber="333">a</emphasis>
), ventral (
<emphasis id="B95F0D16FFDFFFAF2AEC0A7ECDE2F7A9" bold="true" box="[739,752,2053,2074]" pageId="8" pageNumber="333">b</emphasis>
) and lateral (
<emphasis id="B95F0D16FFDFFFAF2B610A7ECC6AF7A9" bold="true" box="[878,888,2053,2074]" pageId="8" pageNumber="333">c</emphasis>
). –
<emphasis id="B95F0D16FFDFFFAF2B930A7FCCA6F7AA" bold="true" box="[924,948,2052,2073]" pageId="8" pageNumber="333">10</emphasis>
.
<taxonomicName id="4C2BAA87FFDFFFAF2BB10A7ECB28F7A9" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[958,1082,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="333" phylum="Arthropoda" rank="species" species="okushimai" status="sp. nov.">
<emphasis id="B95F0D16FFDFFFAF2BB10A7ECB28F7A9" box="[958,1082,2053,2074]" italics="true" pageId="8" pageNumber="333">A. okushimai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDFFFAF2C300A7ECB8FF7A9" box="[1087,1181,2053,2074]" pageId="8" pageNumber="333" rank="species">spec. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="B95F0D16FFDFFFAF2D0C0A7FCA09F7AA" bold="true" box="[1283,1307,2052,2073]" pageId="8" pageNumber="333">11</emphasis>
.
<taxonomicName id="4C2BAA87FFDFFFAF2D2A0A7ECAA1F7A9" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[1317,1459,2053,2074]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="333" phylum="Arthropoda" rank="species" species="immeliorata" status="sp. nov.">A. immeliorata</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B94D104FFDFFFAF28DB0A59CA07F784" blockId="8.[212,1459,2052,2104]" box="[212,1301,2082,2104]" pageId="8" pageNumber="333">
spec. nov.,
<typeStatus id="54906FA6FFDFFFAF29340A59CE80F78B" box="[315,402,2082,2104]" pageId="8" pageNumber="333" type="holotype">holotype</typeStatus>
.
<emphasis id="B95F0D16FFDFFFAF29950A59CEA0F784" bold="true" box="[410,434,2082,2103]" pageId="8" pageNumber="333">12</emphasis>
.
<taxonomicName id="4C2BAA87FFDFFFAF29B20A59CD3EF784" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[445,556,2082,2103]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="333" phylum="Arthropoda" rank="species" species="confrater" status="sp. nov.">
<emphasis id="B95F0D16FFDFFFAF29B20A59CD3EF784" box="[445,556,2082,2103]" italics="true" pageId="8" pageNumber="333">A. confrater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDFFFAF2A3D0A59CD83F784" box="[562,657,2082,2103]" pageId="8" pageNumber="333" rank="species">spec. nov.</taxonomicNameLabel>
,
<typeStatus id="54906FA6FFDFFFAF2A960A59CDE2F78B" box="[665,752,2082,2104]" pageId="8" pageNumber="333" type="holotype">holotype</typeStatus>
.
<emphasis id="B95F0D16FFDFFFAF2AF70A59CC02F784" bold="true" box="[760,784,2082,2103]" pageId="8" pageNumber="333">13</emphasis>
.
<taxonomicName id="4C2BAA87FFDFFFAF2B140A59CCD3F784" authority="Zorn &amp; Kobayashi &amp; Wada, 2017" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[795,961,2082,2103]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="333" phylum="Arthropoda" rank="species" species="imperspicabilis" status="sp. nov.">
<emphasis id="B95F0D16FFDFFFAF2B140A59CCD3F784" box="[795,961,2082,2103]" italics="true" pageId="8" pageNumber="333">A. imperspicabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26CB06DFFDFFFAF2BC90A59CB37F784" box="[966,1061,2082,2103]" pageId="8" pageNumber="333" rank="species">spec. nov.</taxonomicNameLabel>
,
<typeStatus id="54906FA6FFDFFFAF2C210A59CB97F78B" box="[1070,1157,2082,2104]" pageId="8" pageNumber="333" type="holotype">holotype</typeStatus>
. Scale =
<quantity id="4CD37CE1FFDFFFAF2CD90A59CA03F784" box="[1238,1297,2082,2103]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="333" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A10284CEE4FE84" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A10284CE0EFEA4" bold="true" box="[174,284,255,279]" pageId="9" pageNumber="334">Scutellum</emphasis>
1.4 times as wide as long, apex acute; punctures equal those of pronotum.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A1033BCEDFFD24" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A1033BCFE3FEEB" bold="true" box="[174,241,320,344]" pageId="9" pageNumber="334">Elytra</emphasis>
regularly striate-sulcate; primary costae distinctly convex, interstices weakly convex; with two vague secondary costae in the subsutural interstice; striae not marked by a regular row of punctures; entire surface covered with large, very transverse, microscopic-setabearing punctures, leaving out only the ridges of primary costae and the humeral protuberance; punctures adjacent to each other, partly coalescing and forming small transverse striae; epipleura ending before the apical curvature; marginal membrane starting approximately at level of anterior third of metacoxa.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A100E4CE1DFC8B" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A100E4CE08FD04" bold="true" box="[174,282,671,695]" pageId="9" pageNumber="334">Pygidium</emphasis>
triangular and convex, somewhat tumid; disc with reticulate striolation which is arranged concentrically around the tumidity; with sparse yellow setae (
<quantity id="4CD37CE1FFDEFFAE28BA017BCE2DFCA4" box="[181,319,768,792]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="7.5" metricValueMin="1.0" pageId="9" pageNumber="334" unit="mm" value="0.425" valueMax="0.75" valueMin="0.1">0.1-0.75 mm</quantity>
in length) in apical and baso-lateral portions.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A10144CD58FCC4" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A10144CE2FFCE4" bold="true" box="[174,317,831,855]" pageId="9" pageNumber="334">Mesoventrite</emphasis>
very narrow and somewhat concave between the approximated mesocoxae.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A10104CDE3FC4B" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A10104CE28FC24" bold="true" box="[174,314,895,919]" pageId="9" pageNumber="334">Metaventrite</emphasis>
with very dense, transversely confluent punctures at the sides, sparsely punctate in the middle; with pale yellow setae (
<quantity id="4CD37CE1FFDEFFAE29BC01BBCD58FC64" box="[435,586,960,984]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="7.5" metricValueMin="1.2" pageId="9" pageNumber="334" unit="mm" value="0.435" valueMax="0.75" valueMin="0.12">0.12-0.75 mm</quantity>
in length) at the sides which become sparser and shorter in the middle.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A10184CE18FB0B" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A10184CE8AFBA4" bold="true" box="[174,408,1023,1047]" pageId="9" pageNumber="334">Abdominal ventrites</emphasis>
with very transverse punctures which are separated in the middle and confluate laterally; 2nd to 4th ventrite with a transverse row of semierect brownish setae (
<quantity id="4CD37CE1FFDEFFAE2952061BCEF5FBC4" box="[349,487,1120,1144]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="6.3" metricValueMin="2.0" pageId="9" pageNumber="334" unit="mm" value="0.415" valueMax="0.63" valueMin="0.2">0.2-0.63 mm</quantity>
in length, indistinct in the middle) and irregular short setae at the sides; sides not carinate.
</paragraph>
<paragraph id="8B94D104FFDEFFAE28A106BBCE69FA64" blockId="9.[174,779,255,1496]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A106BBCFF2FB6B" bold="true" box="[174,224,1216,1240]" pageId="9" pageNumber="334">Legs</emphasis>
slender and long; protibia bidentate; terminal tooth prolonged and acute at apex in male; protarsal segment 5 thickened, ventrally with a deep concavity in the apical half; inner protarsal claw and outer mesotarsal claw apically incised; the upper branch of the inner protarsal claw small and slender, the lower branch broad, somewhat enlarged ventrally but without tooth; the upper branch of outer mesotarsal claw slightly more slender than the lower one.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDEFFAE28A10784CA0BFE04" pageId="9" pageNumber="334" type="diagnosis">
<paragraph id="8B94D104FFDEFFAE28A10784CA0BFE04" blockId="9.[174,779,1535,2104]" lastBlockId="9.[816,1422,256,440]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE28A10784CE09F9A4" bold="true" box="[174,283,1535,1559]" pageId="9" pageNumber="334">Diagnosis</emphasis>
: This species is a member of the group related to
<taxonomicName id="4C2BAA87FFDEFFAE2910045BCDD6F98B" authority="BURMEISTER, 1855" authorityName="BURMEISTER" authorityYear="1855" box="[287,708,1568,1592]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDEFFAE2910045BCEFCF98B" box="[287,494,1568,1592]" italics="true" pageId="9" pageNumber="334">Anomala spiloptera</emphasis>
<bibRefCitation id="EFBAACF5FFDEFFAE29F7045BCDD6F98B" author="BURMEISTER, H." box="[504,708,1568,1592]" pageId="9" pageNumber="334" refId="ref28524" refString="BURMEISTER, H. 1855: Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Coleoptera Lamellicornia Phyllophaga chaenochela. - Enslin, Berlin: x + 569 pp. - http: // www. biodiversitylibrary. org / item / 34016 # page / 9 / mode / 1 up [accessed 2017 / 06 / 26]." type="book" year="1855">BURMEISTER, 1855</bibRefCitation>
</taxonomicName>
, all of which share a distinctive structure of the aedeagus: the parameres are usually very elongate and are connected by a wide wrinkled membrane dorso-basally. The ventral plate is usually equipped with a long process resembling a narrow blade. Many species related to
<taxonomicName id="4C2BAA87FFDEFFAE2A5E04BBCDC6F96B" authorityName="BURMEISTER" authorityYear="1855" box="[593,724,1728,1752]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDEFFAE2A5E04BBCDC6F96B" box="[593,724,1728,1752]" italics="true" pageId="9" pageNumber="334">A. spiloptera</emphasis>
</taxonomicName>
have different color variants. Within this group,
<taxonomicName id="4C2BAA87FFDEFFAE2A85049BCC18F94B" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[650,778,1760,1784]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="recordata">
<emphasis id="B95F0D16FFDEFFAE2A85049BCC18F94B" box="[650,778,1760,1784]" italics="true" pageId="9" pageNumber="334">A. recordata</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="4C2BAA87FFDEFFAE29A3057BCC18F8AB" authority="OHAUS, 1914" authorityName="OHAUS" authorityYear="1914" box="[428,778,1792,1816]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="B95F0D16FFDEFFAE29A3057BCD67F8AB" box="[428,629,1792,1816]" italics="true" pageId="9" pageNumber="334">Anomala iridicollis</emphasis>
<bibRefCitation id="EFBAACF5FFDEFFAE2A72057BCC18F8AB" author="OHAUS, F." box="[637,778,1792,1816]" pageId="9" pageNumber="334" pagination="193 - 217" refId="ref29591" refString="OHAUS, F. 1914: XV. Beitrag zur Kenntnis der Ruteliden. - Entomologische Zeitung 75: 193 - 217 - https: // www. biodiversitylibrary. org / page / 8825521 # page / 205 / mode / 1 up [accessed 2017 / 10 / 12]." type="journal article" year="1914">OHAUS, 1914</bibRefCitation>
</taxonomicName>
as well as the dark greenish forms of
<taxonomicName id="4C2BAA87FFDEFFAE2A64055BCE4DF8EB" authority="NONFRIED, 1892" authorityName="NONFRIED" authorityYear="1892" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="viridicostata">
<emphasis id="B95F0D16FFDEFFAE2A64055BCC18F88B" box="[619,778,1824,1848]" italics="true" pageId="9" pageNumber="334">A. viridicostata</emphasis>
<bibRefCitation id="EFBAACF5FFDEFFAE28A1053BCE4DF8EB" author="NONFRIED, A. F." box="[174,351,1856,1880]" pageId="9" pageNumber="334" pagination="81 - 95" refId="ref29529" refString="NONFRIED, A. F. 1892: Verzeichnis der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden und Cerambyciden, nebst Beschreibung neuer Arten. - Entomologische Nachrichten (Berlin) 18: 81 - 95." type="journal article" year="1892">NONFRIED, 1892</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2BAA87FFDEFFAE2998053BCDFDF8EB" authority="BURMEISTER, 1855" authorityName="BURMEISTER" authorityYear="1855" box="[407,751,1856,1880]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDEFFAE2998053BCD09F8EB" box="[407,539,1856,1880]" italics="true" pageId="9" pageNumber="334">A. spiloptera</emphasis>
<bibRefCitation id="EFBAACF5FFDEFFAE2A2C053BCDFDF8EB" author="BURMEISTER, H." box="[547,751,1856,1880]" pageId="9" pageNumber="334" refId="ref28524" refString="BURMEISTER, H. 1855: Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Coleoptera Lamellicornia Phyllophaga chaenochela. - Enslin, Berlin: x + 569 pp. - http: // www. biodiversitylibrary. org / item / 34016 # page / 9 / mode / 1 up [accessed 2017 / 06 / 26]." type="book" year="1855">BURMEISTER, 1855</bibRefCitation>
</taxonomicName>
. It is, however, clearly separated from those species by the unique shape of the aedeagus (fig. 15) which is nearly symmetrical (strongly asymmetrical in
<taxonomicName id="4C2BAA87FFDEFFAE2A7705DBCDE9F80B" authorityName="BURMEISTER" authorityYear="1855" box="[632,763,1952,1976]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDEFFAE2A7705DBCDE9F80B" box="[632,763,1952,1976]" italics="true" pageId="9" pageNumber="334">A. spiloptera</emphasis>
</taxonomicName>
), and the ventral plate has a long, strongly downwards curved process (short in
<taxonomicName id="4C2BAA87FFDEFFAE29CD059BCD52F84B" authorityName="OHAUS" authorityYear="1914" box="[450,576,2016,2040]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="B95F0D16FFDEFFAE29CD059BCD52F84B" box="[450,576,2016,2040]" italics="true" pageId="9" pageNumber="334">A. iridicollis</emphasis>
</taxonomicName>
, nearly straight in
<taxonomicName id="4C2BAA87FFDEFFAE28A10A7BCE5FF7AB" authorityName="NONFRIED" authorityYear="1892" box="[174,333,2048,2072]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="viridicostata">
<emphasis id="B95F0D16FFDEFFAE28A10A7BCE5FF7AB" box="[174,333,2048,2072]" italics="true" pageId="9" pageNumber="334">A. viridicostata</emphasis>
</taxonomicName>
). Moreover, males of
<taxonomicName id="4C2BAA87FFDEFFAE2A3B0A7BCDC1F7AB" authorityName="NONFRIED" authorityYear="1892" box="[564,723,2048,2072]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="viridicostata">
<emphasis id="B95F0D16FFDEFFAE2A3B0A7BCDC1F7AB" box="[564,723,2048,2072]" italics="true" pageId="9" pageNumber="334">A. viridicostata</emphasis>
</taxonomicName>
have protarsomere 5 distinctly more thickened, and in all specimens of
<taxonomicName id="4C2BAA87FFDEFFAE2BCF037BCB2CFEAB" authorityName="OHAUS" authorityYear="1914" box="[960,1086,256,280]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="B95F0D16FFDEFFAE2BCF037BCB2CFEAB" box="[960,1086,256,280]" italics="true" pageId="9" pageNumber="334">A. iridicollis</emphasis>
</taxonomicName>
examined, the elytra were deep bluish black with orange spots, with rather weak metallic shine.
<taxonomicName id="4C2BAA87FFDEFFAE2B78033BCB53FEEB" authorityName="Zorn &amp; Kobayashi &amp; Wada" authorityYear="2017" box="[887,1089,320,344]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="recordata">
<emphasis id="B95F0D16FFDEFFAE2B78033BCB53FEEB" box="[887,1089,320,344]" italics="true" pageId="9" pageNumber="334">Anomala recordata</emphasis>
</taxonomicName>
also has a rather broad subsutural interstice with two vague secondary costae. The subsutural interstice is distinctly narrower in
<taxonomicName id="4C2BAA87FFDEFFAE2D0B03FBCA95FE2B" authorityName="BURMEISTER" authorityYear="1855" box="[1284,1415,384,408]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="spiloptera">
<emphasis id="B95F0D16FFDEFFAE2D0B03FBCA95FE2B" box="[1284,1415,384,408]" italics="true" pageId="9" pageNumber="334">A. spiloptera</emphasis>
</taxonomicName>
, while it is broad but rather flat in
<taxonomicName id="4C2BAA87FFDEFFAE2C9A03DBCA01FE0B" authorityName="OHAUS" authorityYear="1914" box="[1173,1299,416,440]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="334" phylum="Arthropoda" rank="species" species="iridicollis">
<emphasis id="B95F0D16FFDEFFAE2C9A03DBCA01FE0B" box="[1173,1299,416,440]" italics="true" pageId="9" pageNumber="334">A. iridicollis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C331828FFFDEFFAE2B3F039BCA22FDEB" pageId="9" pageNumber="334" type="etymology">
<paragraph id="8B94D104FFDEFFAE2B3F039BCA22FDEB" blockId="9.[816,1422,480,600]" pageId="9" pageNumber="334">
<emphasis id="B95F0D16FFDEFFAE2B3F039BCCB5FE4B" bold="true" box="[816,935,480,504]" pageId="9" pageNumber="334">Etymology</emphasis>
: The specific epithet is formed by the Latin adjective “recordatus, -a, -um” and translates as “remembered” or “recollected”. This alludes to the number of similar species with which it might be confused.
</paragraph>
</subSubSection>
</treatment>
</document>
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