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<document id="0AC44F811151D8893D73F030791ADA0F" ID-CLB-Dataset="53857" ID-DOI="10.11646/zootaxa.5089.1.1" ID-GBIF-Dataset="67fbf2b0-b944-485d-90b5-976914d338ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="5836060" ID-ZooBank="E12F2B20-A84A-48E2-9C77-B281F1BFC62E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641892626941" checkinUser="plazi" docAuthor="Ortega-Salas, Héctor, González-Soriano, Enrique &amp; Jocque, Merlijn" docDate="2022" docId="039887EB3020FFEBFF73FB2EF143FC6B" docLanguage="en" docName="zootaxa.5089.1.1.pdf" docOrigin="Zootaxa 5089 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paraphlebia akan Ortega-Salas &amp; Gonzalez-Soriano 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="8" masterDocId="FFA1FF933026FFECFFE4FFC9F553FF86" masterDocTitle="Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="7" updateTime="1699298278583" updateUser="ExternalLinkService">
<mods:mods id="E345EA5009E5C7B3362418941FA058C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="42C35B253E7DFCA40D8A9AEF71183A4A">
<mods:title id="D3C7D75341B5A631B9754712D138D460">Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species</mods:title>
</mods:titleInfo>
<mods:name id="97A12E2639557F82D182C08E0700423D" type="personal">
<mods:role id="547E3ECF36C7B5E8146341A7029B8A47">
<mods:roleTerm id="36EFD00BCFE05F51A2AC9AB739D973F6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="139964232792EE81260AD826F1ECE3D2">Ortega-Salas, Héctor</mods:namePart>
<mods:nameIdentifier id="CC169D253653CD6A55FB4680315FC282" type="ORCID">0000-0002-5373-4839</mods:nameIdentifier>
<mods:nameIdentifier id="F83FA6628A54296A8D74415DC17F4B41" type="email">hector.ortegasalas@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name id="1B76E19AE7D3083C6F14570F9A9B1D40" type="personal">
<mods:role id="5B98C7652A1BEFD8E7BDD06152DA92BF">
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</mods:role>
<mods:namePart id="EA8B58ADD71214C01D1BD8F3253C72A2">González-Soriano, Enrique</mods:namePart>
<mods:nameIdentifier id="63FC3AB5F05B79E46AE6523C96D16C31" type="ORCID">0000-0002-4798-7274</mods:nameIdentifier>
<mods:nameIdentifier id="A1146B21030EFFC63DC5AFF520A553E1" type="email">esoriano@ib.unam.mx</mods:nameIdentifier>
</mods:name>
<mods:name id="EED40E950C21EAA7E9818F4B784BB263" type="personal">
<mods:role id="9042DE96E369CE1EAFC8F4F07664DBB4">
<mods:roleTerm id="9F881E9DF65DF6FCA187445303D62D52">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FDBEE31F115AE5EAEB691B80029ED71">Jocque, Merlijn</mods:namePart>
<mods:nameIdentifier id="FE1FEAF09861756880CF55997EF83744" type="ORCID">0000-0002-7196-7476</mods:nameIdentifier>
<mods:nameIdentifier id="BC9FD69EDFFA1C4E5E3D1D19C69B179C" type="email">merlijn.jocque@binco.eu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0C4A9E90D830828FFF7BD06FCE995C76">text</mods:typeOfResource>
<mods:relatedItem id="2437B3156699E19EF0A68D9E9870A128" type="host">
<mods:titleInfo id="3E8516AFDD9DBE48D4CAE2DDA1C1D059">
<mods:title id="BFBFEE3B6EEC009DA22D696EDB0AABEC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3456A45C25323D2DB9475EEB0254EA82">
<mods:date id="3DF6E3D08994EDDB8A4BE033FD9D214F">2022</mods:date>
<mods:detail id="40ABC57759DB3398575ED678D4D8004F" type="pubDate">
<mods:number id="DE1D9035BC41BDC483DC7C2537CE5E26">2022-01-11</mods:number>
</mods:detail>
<mods:detail id="136C52022E559F14CB569CA0103B303B" type="volume">
<mods:number id="1C666AC304652B2CAD2457157CE4D2C7">5089</mods:number>
</mods:detail>
<mods:detail id="151DDAC6AF5F734C53AD39C5B6A0F3F3" type="issue">
<mods:number id="B11BC7FB27279BD140C700A2CC6F0C4C">1</mods:number>
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<mods:start id="D35A6083A79D87D1C893826D497CDC11">1</mods:start>
<mods:end id="AF967910EEC100A6036EA6D428E2A880">66</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="F07EDA00FC48E912EB412F72C14D9263">journal article</mods:classification>
<mods:identifier id="BF92D6533B2758197FA7ADD9F26BB8D0" type="CLB-Dataset">53857</mods:identifier>
<mods:identifier id="492A6EB95CD697A056052B6A8A8E815D" type="DOI">10.11646/zootaxa.5089.1.1</mods:identifier>
<mods:identifier id="133E25D951FB16E1FD53541C6B01FEE6" type="GBIF-Dataset">67fbf2b0-b944-485d-90b5-976914d338ad</mods:identifier>
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<mods:identifier id="8E460C4BEBCF7794DF24B07EBA4533A0" type="Zenodo-Dep">5836060</mods:identifier>
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<treatment id="039887EB3020FFEBFF73FB2EF143FC6B" ID-DOI="http://doi.org/10.5281/zenodo.5836072" ID-GBIF-Taxon="209152399" ID-Zenodo-Dep="5836072" LSID="urn:lsid:plazi:treatment:039887EB3020FFEBFF73FB2EF143FC6B" httpUri="http://treatment.plazi.org/id/039887EB3020FFEBFF73FB2EF143FC6B" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="C32B65763020FFEAFF73FB2EF6D6FA84" box="[151,901,1255,1282]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8B8E36FD3020FFEAFF73FB2EF6D6FA84" blockId="6.[151,1358,1255,1320]" box="[151,901,1255,1282]" pageId="6" pageNumber="7">
<heading id="D0C681913020FFEAFF73FB2EF6D6FA84" bold="true" box="[151,901,1255,1282]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="4">
<emphasis id="B945EAEF3020FFEAFF73FB2EF6D6FA84" bold="true" box="[151,901,1255,1282]" pageId="6" pageNumber="7">
<taxonomicName id="4C314D7E3020FFEAFF73FB2EF673FA84" authority="Ortega-Salas &amp; Gonzalez-Soriano" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[151,800,1255,1282]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="akan" status="sp. nov.">
<emphasis id="B945EAEF3020FFEAFF73FB2EF43CFA84" bold="true" box="[151,367,1255,1282]" italics="true" pageId="6" pageNumber="7">Paraphlebia akan</emphasis>
Ortega-Salas &amp; González-Soriano
</taxonomicName>
<taxonomicNameLabel id="A27657943020FFEAFCC3FB21F6D6FA84" box="[807,901,1256,1282]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B65763020FFEAFF73FAC5F6F2FA53" pageId="6" pageNumber="7" type="description">
<paragraph id="8B8E36FD3020FFEAFF73FAC5F01DFAA0" blockId="6.[151,1358,1255,1320]" box="[151,1358,1292,1320]" pageId="6" pageNumber="7">
<figureCitation id="130A2A783020FFEAFF73FAC5F5B0FAA0" box="[151,227,1292,1318]" captionStart="FIGURES 2–11" captionStartId="9.[151,264,1999,2023]" captionTargetBox="[213,1367,251,1955]" captionTargetId="figure-42@9.[200,1387,237,1975]" captionTargetPageId="9" captionText="FIGURES 2–11. Paraphlebia spp. colour pattern of head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5836066" httpUri="https://zenodo.org/record/5836066/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
(head
<specimenCount id="9D37FD743020FFEAFED4FAC4F416FAAE" box="[304,325,1293,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
), 14 (thorax
<specimenCount id="9D37FD743020FFEAFE33FAC4F4BFFAAE" box="[471,492,1293,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
), 63, 83–84 (appendages
<specimenCount id="9D37FD743020FFEAFCE9FAC4F671FAAE" box="[781,802,1293,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
), 118 (posterior lobe of prothorax
<specimenCount id="9D37FD743020FFEAFB43FAC4F1EFFAAE" box="[1191,1212,1293,1320]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
), 150 (map).
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF73FA99F62CFAEE" blockId="6.[151,895,1360,1384]" box="[151,895,1360,1384]" pageId="6" pageNumber="7">
<uri id="FFA03AFF3020FFEAFF73FA99F62CFAEE" box="[151,895,1360,1384]" pageId="6" pageNumber="7">
urn:lsid:zoobank.org:act:
<uuid id="FF970C283020FFEAFE6BFA99F62CFAEE" box="[399,895,1360,1384]" pageId="6" pageNumber="7">E52DB127-9AB8-40EF-9BDE-109FF0988EBF</uuid>
</uri>
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF73FA5EF6F2FA53" blockId="6.[151,1437,1430,2033]" pageId="6" pageNumber="7">
<materialsCitation id="3B593CA03020FFEAFF73FA5EF6F2FA53" ID-GBIF-Occurrence="4023503728" collectingDate="1976-08-22" collectionCode="TWD, IBUNAM" collectorName="T. Donnelly" country="Guatemala" county="Rio Hondo" elevation="1700" latitude="15.13" location="Jones" longLatPrecision="76" longitude="-89.53" municipality="Rio Canas" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Zacapa" typeStatus="holotype">
<typeStatus id="548A885F3020FFEAFF73FA5EF450FA37" box="[151,259,1431,1457]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F326766D3020FFEAFEEAFA5EF4E8FA37" box="[270,443,1431,1457]" name="Guatemala" pageId="6" pageNumber="7">GUATEMALA</collectingCountry>
•
<specimenCount id="9D37FD743020FFEAFE37FA5EF4AEFA35" box="[467,509,1431,1459]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
;
<emphasis id="B945EAEF3020FFEAFDE8FA5EF736FA37" bold="true" box="[524,613,1431,1457]" pageId="6" pageNumber="7">
<collectingRegion id="49F5F81F3020FFEAFDE8FA5EF736FA37" box="[524,613,1431,1457]" country="Guatemala" name="Zacapa" pageId="6" pageNumber="7">Zacapa</collectingRegion>
</emphasis>
,
<collectingCounty id="62EF4E713020FFEAFD97FA5EF7A1FA37" box="[627,754,1431,1457]" pageId="6" pageNumber="7">Río Hondo</collectingCounty>
, Trib.
<collectingMunicipality id="6BEAAC873020FFEAFCDFFA5EF6E1FA37" box="[827,946,1431,1457]" pageId="6" pageNumber="7">Río Cañas</collectingMunicipality>
, above
<location id="8EEE60263020FFEAFBECFA5EF119FA37" LSID="urn:lsid:plazi:treatment:039887EB3020FFEBFF73FB2EF143FC6B:8EEE60263020FFEAFBECFA5EF119FA37" box="[1032,1098,1431,1457]" country="Guatemala" county="Rio Hondo" latitude="15.13" longLatPrecision="76" longitude="-89.53" municipality="Rio Canas" name="Jones" pageId="6" pageNumber="7" stateProvince="Zacapa">Jones</location>
; [
<geoCoordinate id="EE05503A3020FFEAFBB8FA5FF19EFA37" box="[1116,1229,1430,1457]" degrees="15.130" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="55" value="15.13">15.130°N</geoCoordinate>
,
<geoCoordinate id="EE05503A3020FFEAFB3DFA5FF001FA37" box="[1241,1362,1430,1457]" degrees="89.530" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="-89.53">89.530°W</geoCoordinate>
];
<quantity id="4CC99B183020FFEAFA80FA5EF5E3FA53" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="m" value="1700.0">
<elevation id="001CD1CE3020FFEAFA80FA5EF5E3FA53" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="7" unit="m" value="1700.0">1700 m</elevation>
</quantity>
;
<date id="FF8F103D3020FFEAFF58FA72F41CFA53" box="[188,335,1467,1493]" pageId="6" pageNumber="7" value="1976-08-22">
<collectingDate id="EFCBE9D53020FFEAFF58FA72F41CFA53" box="[188,335,1467,1493]" pageId="6" pageNumber="7" value="1976-08-22">22 Aug 1976</collectingDate>
</date>
;
<collectorName id="26C4532B3020FFEAFEBFFA72F4B3FA53" box="[347,480,1467,1493]" pageId="6" pageNumber="7">T. Donnelly</collectorName>
leg.;
<collectionCode id="ED20AE383020FFEAFDFAFA72F732FA53" box="[542,609,1467,1493]" pageId="6" pageNumber="7">TWD</collectionCode>
;
<collectionCode id="ED20AE383020FFEAFD89FA72F7B0FA53" box="[621,739,1467,1493]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="6" pageNumber="7" type="University or college">IBUNAM</collectionCode>
:CNIN:OD22717
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B65763020FFEAFF23FA16F706F9C7" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8B8E36FD3020FFEAFF23FA16F706F9C7" blockId="6.[151,1437,1430,2033]" pageId="6" pageNumber="7">
<emphasis id="B945EAEF3020FFEAFF23FA16F41BFA7F" bold="true" box="[199,328,1503,1529]" pageId="6" pageNumber="7">Etymology</emphasis>
. Named
<taxonomicName id="4C314D7E3020FFEAFE55FA16F4B4FA7F" box="[433,487,1503,1529]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3020FFEAFE55FA16F4B4FA7F" box="[433,487,1503,1529]" italics="true" pageId="6" pageNumber="7">akan</emphasis>
</taxonomicName>
/ɑˈkɑn/ (L. noun in apposition), after Akan, the Mayan God of alcoholic beverages and other toxic and hallucinogenic substances. This seems appropriate since this species occurs in the region where the renowned
<collectingRegion id="49F5F81F3020FFEAFED2F9EEF4D4F9C7" box="[310,391,1575,1601]" country="Guatemala" name="Zacapa" pageId="6" pageNumber="7">Zacapa</collectingRegion>
Rum is produced.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763020FFEBFF23F983F7CDFEE3" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="8B8E36FD3020FFEAFF23F983F48DF9E3" blockId="6.[151,1437,1430,2033]" box="[199,478,1610,1637]" pageId="6" pageNumber="7">
<emphasis id="B945EAEF3020FFEAFF23F983F48DF9E3" bold="true" box="[199,478,1610,1637]" pageId="6" pageNumber="7">
Description of
<typeStatus id="548A885F3020FFEAFE93F982F48DF9E3" box="[375,478,1611,1637]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF23F9A6F492F973" blockId="6.[151,1437,1430,2033]" pageId="6" pageNumber="7">
<emphasis id="B945EAEF3020FFEAFF23F9A6F457F90F" box="[199,260,1647,1673]" italics="true" pageId="6" pageNumber="7">Head</emphasis>
. Chiefly black, labrum dark brown to black, anteclypeus dark brown, postclypeus cream over bluish base, with a narrow black line on posterior margin, frons with pale spots anterolateral to antennae, occiput with complete pale occipital bar, eyes dark brown to black; antennae scape black with dark brown distal border, pedicel and flagellum dark brown.
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF23F936F077F8BB" blockId="6.[151,1437,1430,2033]" pageId="6" pageNumber="7">
<emphasis id="B945EAEF3020FFEAFF23F936F44BF89F" box="[199,280,1791,1817]" italics="true" pageId="6" pageNumber="7">Thorax</emphasis>
. Prothorax: chiefly pale on dorsum; middle lobe with black dorsolateral spots; posterior lobe with black lateral spots, see
<figureCitation id="130A2A783020FFEAFEB2F8EAF4E0F8BB" box="[342,435,1827,1853]" captionStart="FIGURES 117–128" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[208,1385,195,1907]" captionTargetId="figure-17@37.[190,1398,181,1919]" captionTargetPageId="37" captionText="FIGURES 117–128. Paraphlebia spp. posterior lobe of prothorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5836120" httpUri="https://zenodo.org/record/5836120/files/figure.png" pageId="6" pageNumber="7">Fig. 118</figureCitation>
for morphology; propleuron black. Pterothorax: see
<figureCitation id="130A2A783020FFEAFBE6F8EAF102F8BB" box="[1026,1105,1827,1853]" captionStart="FIGURES 12–19" captionStartId="24.[151,264,1943,1967]" captionTargetBox="[214,1374,194,1906]" captionTargetId="figure-18@24.[197,1390,181,1919]" captionTargetPageId="24" captionText="FIGURES 12–19. Paraphlebia spp. colour pattern of head and thorax. Head frontal view (12–19); thorax lateral view (14– 19)." figureDoi="http://doi.org/10.5281/zenodo.5836068" httpUri="https://zenodo.org/record/5836068/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
for colour pattern.
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF23F88EF4BBF84B" blockId="6.[151,1437,1430,2033]" pageId="6" pageNumber="7">
<emphasis id="B945EAEF3020FFEAFF23F88EF458F8E6" box="[199,267,1863,1888]" italics="true" pageId="6" pageNumber="7">Wings</emphasis>
. Hyaline, with slight amber tint; FW with 1.0/1.4 post-quadrangular cells, on the left wing, the vein that descends from the subnodus at the first Vx, on the right wing this vein is distal to the first Vx by a distance of twofifths of the second post-quadrangular cell; HW field between CuA and posterior margin lacking supplementary veins, CuA in HW not forked.
</paragraph>
<paragraph id="8B8E36FD3020FFEAFF23F81EF4BDF877" blockId="6.[151,1437,1430,2033]" box="[199,494,2007,2033]" pageId="6" pageNumber="7">Px: FW 34/30, HW 28/27.</paragraph>
<paragraph id="8B8E36FD3021FFEBFF23FF5EF6E1FF53" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FF5EF462FF37" box="[199,305,151,177]" italics="true" pageId="7" pageNumber="8">Abdomen</emphasis>
. Chiefly black; S1 dark brown; S2 with pale ventrolateral horizontal stripes; S3–7 with pale basal rings, on S7 not closing dorsally; S9–10 and cerci pruinose on dorsum.
</paragraph>
<paragraph id="8B8E36FD3021FFEBFF23FF16F7A9FE9B" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FF16F4F9FF7F" box="[199,426,223,249]" italics="true" pageId="7" pageNumber="8">Caudal appendages</emphasis>
. Cerci: see
<figureCitation id="130A2A783021FFEBFDDEFF16F7DEFF7F" box="[570,653,223,249]" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Fig. 64</figureCitation>
for morphology. Paraprocts: rudimentary, superior lobe completely absent; inferior lobe in lateral view smoothly rounded.
</paragraph>
<paragraph id="8B8E36FD3021FFEBFF23FEEEF644FEC7" blockId="7.[151,1437,151,1005]" box="[199,791,294,321]" pageId="7" pageNumber="8">Measurements. Abdomen: 41.5, FW: 34.1, HW: 33.3</paragraph>
<paragraph id="8B8E36FD3021FFEBFF23FE83F7CDFEE3" blockId="7.[151,1437,151,1005]" box="[199,670,330,357]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FE83F49EFEE2" bold="true" box="[199,461,330,356]" pageId="7" pageNumber="8">Description of female.</emphasis>
Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763021FFEBFF23FEA7F1C0FC23" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B8E36FD3021FFEBFF23FEA7F1EFFC07" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FEA7F469FE0E" bold="true" box="[199,314,366,392]" pageId="7" pageNumber="8">Diagnosis</emphasis>
.
<taxonomicName id="4C314D7E3021FFEBFEAEFEA6F745FE0F" box="[330,534,367,393]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFEAEFEA6F745FE0F" box="[330,534,367,393]" italics="true" pageId="7" pageNumber="8">Paraphlebia akan</emphasis>
</taxonomicName>
belongs to the group of species with the mediodorsal flange of the cerci welldeveloped. Three species,
<taxonomicName id="4C314D7E3021FFEBFE24FE5AF761FE2B" authorityName="Ortega-Salas" authorityYear="2022" box="[448,562,403,429]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFE24FE5AF761FE2B" box="[448,562,403,429]" italics="true" pageId="7" pageNumber="8">P. chiarae</emphasis>
</taxonomicName>
,
<emphasis id="B945EAEF3021FFEBFDDBFE5AF7FCFE2B" box="[575,687,403,429]" italics="true" pageId="7" pageNumber="8">
P,
<taxonomicName id="4C314D7E3021FFEBFDBFFE5AF7FCFE2B" authorityName=", Brauer" authorityYear="1871" box="[603,687,403,429]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hyalina">hyalina</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C314D7E3021FFEBFD09FE5AF63DFE2B" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[749,878,403,429]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3021FFEBFD09FE5AF63DFE2B" box="[749,878,403,429]" italics="true" pageId="7" pageNumber="8">P. kukulkan</emphasis>
</taxonomicName>
are similar to
<taxonomicName id="4C314D7E3021FFEBFBF5FE5AF130FE2B" box="[1041,1123,403,429]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFBF5FE5AF130FE2B" box="[1041,1123,403,429]" italics="true" pageId="7" pageNumber="8">P. akan</emphasis>
</taxonomicName>
; however, the distal margin of the cercus has a sclerotized blade-shaped tip (
<figureCitation id="130A2A783021FFEBFD55FE7EF630FE57" box="[689,867,438,465]" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Figs. 63, 65–66</figureCitation>
), while in
<taxonomicName id="4C314D7E3021FFEBFC33FE7EF17BFE57" box="[983,1064,439,465]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFC33FE7EF17BFE57" box="[983,1064,439,465]" italics="true" pageId="7" pageNumber="8">P. akan</emphasis>
</taxonomicName>
the tip is smoothly rounded (
<figureCitation id="130A2A783021FFEBFA8BFE7EF5E4FE70" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Fig. 64</figureCitation>
).
<taxonomicName id="4C314D7E3021FFEBFF2FFE12F4E1FE73" authorityName=", Brauer" authorityYear="1871" box="[203,434,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hyalina">
<emphasis id="B945EAEF3021FFEBFF2FFE12F4E1FE73" box="[203,434,475,501]" italics="true" pageId="7" pageNumber="8">Paraphlebia hyalina</emphasis>
</taxonomicName>
is set apart from
<taxonomicName id="4C314D7E3021FFEBFD92FE12F79BFE73" box="[630,712,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFD92FE12F79BFE73" box="[630,712,475,501]" italics="true" pageId="7" pageNumber="8">P. akan</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D7E3021FFEBFD32FE12F61BFE73" authorityName="Ortega-Salas" authorityYear="2022" box="[726,840,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFD32FE12F61BFE73" box="[726,840,475,501]" italics="true" pageId="7" pageNumber="8">P. chiarae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C314D7E3021FFEBFC61FE12F155FE73" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[901,1030,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3021FFEBFC61FE12F155FE73" box="[901,1030,475,501]" italics="true" pageId="7" pageNumber="8">P. kukulkan</emphasis>
</taxonomicName>
by having clearly forked paraprocts with well-developed upper and lower branches (
<figureCitation id="130A2A783021FFEBFD25FE36F67DFD9F" box="[705,814,511,537]" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Figs. 62a</figureCitation>
,
<figureCitation id="130A2A783021FFEBFCD8FE36F6D5FD9F" box="[828,902,511,537]" captionStart="FIGURES 79–84" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[218,1357,197,1905]" captionTargetId="figure-17@33.[207,1381,181,1919]" captionTargetPageId="33" captionText="FIGURES 79–84. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view (79–80) W – maximum width of mediodorsal flange, w – maximum width of the distal lobe; dorsal view (81); lateral view (82, 84); posterior view (83)." figureDoi="http://doi.org/10.5281/zenodo.5836104" httpUri="https://zenodo.org/record/5836104/files/figure.png" pageId="7" pageNumber="8">81–82</figureCitation>
). In
<taxonomicName id="4C314D7E3021FFEBFC58FE36F143FD9F" box="[956,1040,511,537]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFC58FE36F143FD9F" box="[956,1040,511,537]" italics="true" pageId="7" pageNumber="8">P. akan</emphasis>
</taxonomicName>
the paraprocts are rudimentary, in lateral view smoothly rounded and with no indication of an upper branch (
<figureCitation id="130A2A783021FFEBFC08FDEAF13EFDBB" box="[1004,1133,547,573]" captionStart="FIGURES 79–84" captionStartId="33.[151,264,1943,1967]" captionTargetBox="[218,1357,197,1905]" captionTargetId="figure-17@33.[207,1381,181,1919]" captionTargetPageId="33" captionText="FIGURES 79–84. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view (79–80) W – maximum width of mediodorsal flange, w – maximum width of the distal lobe; dorsal view (81); lateral view (82, 84); posterior view (83)." figureDoi="http://doi.org/10.5281/zenodo.5836104" httpUri="https://zenodo.org/record/5836104/files/figure.png" pageId="7" pageNumber="8">Fig. 83–84</figureCitation>
).
<taxonomicName id="4C314D7E3021FFEBFB65FDEAF03FFDBB" authorityName="Ortega-Salas" authorityYear="2022" box="[1153,1388,547,573]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFB65FDEAF03FFDBB" box="[1153,1388,547,573]" italics="true" pageId="7" pageNumber="8">Paraphlebia chiarae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D7E3021FFEBFF73FD8EF449FDE7" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[151,282,583,609]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3021FFEBFF73FD8EF449FDE7" box="[151,282,583,609]" italics="true" pageId="7" pageNumber="8">P. kukulkan</emphasis>
</taxonomicName>
are intermediate, having a poorly developed upper branch defined by a shallow groove and a welldeveloped lower branch with an acute projection directed posteriad (
<figureCitation id="130A2A783021FFEBFC4BFDA2F162FD03" box="[943,1073,619,645]" captionStart="FIGURES 85–94" captionStartId="34.[151,264,1943,1967]" captionTargetBox="[214,1372,193,1905]" captionTargetId="figure-18@34.[199,1388,181,1919]" captionTargetPageId="34" captionText="FIGURES 85–94. Paraphlebia spp. caudal appendages lateral view." figureDoi="http://doi.org/10.5281/zenodo.5836110" httpUri="https://zenodo.org/record/5836110/files/figure.png" pageId="7" pageNumber="8">Fig. 93–94</figureCitation>
).
<taxonomicName id="4C314D7E3021FFEBFBA3FDA2F012FD03" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[1095,1345,619,645]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3021FFEBFBA3FDA2F012FD03" box="[1095,1345,619,645]" italics="true" pageId="7" pageNumber="8">Paraphlebia kukulkan</emphasis>
</taxonomicName>
has the most easily distinguishable cerci, in dorsal view the mediodorsal flange convex and narrowest proximal to half its length (
<figureCitation id="130A2A783021FFEBFF09FD7AF41DFD48" box="[237,334,691,718]" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Fig. 66a</figureCitation>
), while in
<taxonomicName id="4C314D7E3021FFEBFE22FD7AF74BFD4B" box="[454,536,691,717]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFE22FD7AF74BFD4B" box="[454,536,691,717]" italics="true" pageId="7" pageNumber="8">P. akan</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D7E3021FFEBFDCCFD7AF7CBFD4B" authorityName=", Brauer" authorityYear="1871" box="[552,664,691,717]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hyalina">
<emphasis id="B945EAEF3021FFEBFDCCFD7AF7CBFD4B" box="[552,664,691,717]" italics="true" pageId="7" pageNumber="8">P. hyalina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D7E3021FFEBFD34FD7AF611FD4B" authorityName="Ortega-Salas" authorityYear="2022" box="[720,834,691,717]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFD34FD7AF611FD4B" box="[720,834,691,717]" italics="true" pageId="7" pageNumber="8">P. chiarae</emphasis>
</taxonomicName>
each cercus is smoothly curved throughout its length and narrowest at its proximal end (
<figureCitation id="130A2A783021FFEBFDC1FD1EF79AFD77" box="[549,713,726,753]" captionStart="FIGURES 62–66" captionStartId="30.[151,264,1943,1967]" captionTargetBox="[216,1371,196,1905]" captionTargetId="figure-18@30.[205,1382,181,1919]" captionTargetPageId="30" captionText="FIGURES 62–66. Paraphlebia spp. caudal appendages (a) dorsal view; (b) mediodorsal view. W – maximum width of mediodorsal flange, w – maximum width of the distal lobe." figureDoi="http://doi.org/10.5281/zenodo.5836088" httpUri="https://zenodo.org/record/5836088/files/figure.png" pageId="7" pageNumber="8">Figs. 63a–65a</figureCitation>
). The posterior lobe of the prothorax in
<taxonomicName id="4C314D7E3021FFEBFB77FD1EF1B1FD77" box="[1171,1250,727,753]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="akan">
<emphasis id="B945EAEF3021FFEBFB77FD1EF1B1FD77" box="[1171,1250,727,753]" italics="true" pageId="7" pageNumber="8">P akan</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D7E3021FFEBFB16FD1EF036FD77" authorityName="Ortega-Salas" authorityYear="2022" box="[1266,1381,727,753]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFB16FD1EF036FD77" box="[1266,1381,727,753]" italics="true" pageId="7" pageNumber="8">P. chiarae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C314D7E3021FFEBFF73FD32F454FC93" authorityName=", Brauer" authorityYear="1871" box="[151,263,763,789]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hyalina">
<emphasis id="B945EAEF3021FFEBFF73FD32F454FC93" box="[151,263,763,789]" italics="true" pageId="7" pageNumber="8">P. hyalina</emphasis>
</taxonomicName>
is similar in shape with the posterior margin semi-circular and entire (
<figureCitation id="130A2A783021FFEBFBC2FD32F1C1FC93" box="[1062,1170,763,789]" captionStart="FIGURES 105–116" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[205,1381,195,1910]" captionTargetId="figure-18@36.[196,1391,181,1919]" captionTargetPageId="36" captionText="FIGURES 105–116. Paraphlebia spp. posterior lobe of prothorax frontal view (105–107); dorsal view (108–116)." figureDoi="http://doi.org/10.5281/zenodo.5836114" httpUri="https://zenodo.org/record/5836114/files/figure.png" pageId="7" pageNumber="8">Figs. 112</figureCitation>
,
<figureCitation id="130A2A783021FFEBFB7AFD32F050FC93" box="[1182,1283,763,789]" captionStart="FIGURES 117–128" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[208,1385,195,1907]" captionTargetId="figure-17@37.[190,1398,181,1919]" captionTargetPageId="37" captionText="FIGURES 117–128. Paraphlebia spp. posterior lobe of prothorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5836120" httpUri="https://zenodo.org/record/5836120/files/figure.png" pageId="7" pageNumber="8">118, 120</figureCitation>
); however, in
<taxonomicName id="4C314D7E3021FFEBFF73FCD6F455FCBF" authorityName="Ortega-Salas" authorityYear="2022" box="[151,262,799,825]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chiarae">
<emphasis id="B945EAEF3021FFEBFF73FCD6F455FCBF" box="[151,262,799,825]" italics="true" pageId="7" pageNumber="8">P chiarae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D7E3021FFEBFEDBFCD6F4E3FCBF" authorityName=", Brauer" authorityYear="1871" box="[319,432,799,825]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hyalina">
<emphasis id="B945EAEF3021FFEBFEDBFCD6F4E3FCBF" box="[319,432,799,825]" italics="true" pageId="7" pageNumber="8">P. hyalina</emphasis>
</taxonomicName>
, depending on the position, it can appear to have a shallow medial notch and is slightly convex towards the outside edges.
<taxonomicName id="4C314D7E3021FFEBFDF2FC8AF65FFCDB" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[534,780,835,861]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3021FFEBFDF2FC8AF65FFCDB" box="[534,780,835,861]" italics="true" pageId="7" pageNumber="8">Paraphlebia kukulkan</emphasis>
</taxonomicName>
differs from the other three species by having subtriangular lateral margins with one straight side at least as long as four-fifths of the lobe width (
<figureCitation id="130A2A783021FFEBFBAAFCAEF1E3FC07" box="[1102,1200,871,897]" captionStart="FIGURES 105–116" captionStartId="36.[151,264,1943,1967]" captionTargetBox="[205,1381,195,1910]" captionTargetId="figure-18@36.[196,1391,181,1919]" captionTargetPageId="36" captionText="FIGURES 105–116. Paraphlebia spp. posterior lobe of prothorax frontal view (105–107); dorsal view (108–116)." figureDoi="http://doi.org/10.5281/zenodo.5836114" httpUri="https://zenodo.org/record/5836114/files/figure.png" pageId="7" pageNumber="8">Fig. 112</figureCitation>
).
</paragraph>
<paragraph id="8B8E36FD3021FFEBFF23FC42F1C0FC23" blockId="7.[151,1437,151,1005]" box="[199,1171,906,933]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FC42F42CFC22" bold="true" box="[199,383,906,933]" pageId="7" pageNumber="8">Natural history</emphasis>
. The
<typeStatus id="548A885F3021FFEBFE24FC42F771FC23" box="[448,546,907,933]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
was collected in a stream within a mountain wet forest.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763021FFEBFF23FC67F143FC6B" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8B8E36FD3021FFEBFF23FC67F143FC6B" blockId="7.[151,1437,151,1005]" pageId="7" pageNumber="8">
<emphasis id="B945EAEF3021FFEBFF23FC67F40AFC4E" bold="true" box="[199,345,942,968]" pageId="7" pageNumber="8">Distribution</emphasis>
. East-Central
<collectingCountry id="F326766D3021FFEBFE1DFC66F724FC4F" box="[505,631,943,969]" name="Guatemala" pageId="7" pageNumber="8">Guatemala</collectingCountry>
: known only from the
<typeStatus id="548A885F3021FFEBFC94FC66F6F3FC4F" box="[880,928,943,969]" pageId="7" pageNumber="8">type</typeStatus>
locality north of
<collectingRegion id="49F5F81F3021FFEBFBBBFC66F1E3FC4F" box="[1119,1200,943,969]" country="Guatemala" name="Zacapa" pageId="7" pageNumber="8">Zacapa</collectingRegion>
on the south slope of the Sierra de las Minas National Park in Río Hondo department at
<quantity id="4CC99B183021FFEBFC64FC1AF68FFC6B" box="[896,988,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="8" unit="m" value="1700.0">1700 m</quantity>
.a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>
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