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<document id="64D809640516F238A05954C9369DA771" ID-CLB-Dataset="29103" ID-DOI="10.11646/zootaxa.4455.2.5" ID-GBIF-Dataset="568a5364-52f2-4feb-b08b-87eb85d4cf77" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457283" ID-ZooBank="8CB311F1-431C-4861-956E-3200329E4F0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539261916473" checkinUser="plazi" docAuthor="Nada, B. &amp; Ballantyne, L. A." docDate="2018" docId="039887E6FFF37A069ECB236EA3666629" docLanguage="en" docName="zootaxa.4455.2.5.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygoluciola dunguna Nada &amp; Ballantyne 2018" docType="treatment" docVersion="7" lastPageNumber="357" masterDocId="FFA1FF9EFFF67A089E5C2657A7076769" masterDocTitle="A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)" masterLastPageNumber="362" masterPageNumber="343" pageNumber="348" updateTime="1698575593942" updateUser="ExternalLinkService">
<mods:mods id="B4C8B37AFCC24380437C14A6D368E0E3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7FF784EA4087C6C06A9E1CCC368BF160">
<mods:title id="97CEB047C6AC289379DBD9827370DDBB">A new species of Pygoluciola Wittmer with unusual abdominal configuration, from lowland dipterocarp forest in Peninsular Malaysia (Coleoptera: Lampyridae: Luciolinae)</mods:title>
</mods:titleInfo>
<mods:name id="77AF0A70ED7F4FFBB3DBD05020F1B49B" type="personal">
<mods:role id="4420DBBBC4AEFFDD27628DB5394F0FA1">
<mods:roleTerm id="AA2D40EA66D251D438A0ABA0792EFC95">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="21E815863920C1B54DCAA28D3930CB1C">Nada, B.</mods:namePart>
</mods:name>
<mods:name id="CA363392EC8C3D8BCDD4CD2ED7127BD3" type="personal">
<mods:role id="EF48AEA288BEB1F4EF3E44F0C45A95D9">
<mods:roleTerm id="508092CE9EAFD714CC5503DC39F0F820">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="658C9A0FFDD39683D16B0692614ED418">Ballantyne, L. A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="36023CFAEC0E3F06E0C5FF677D005F7D">text</mods:typeOfResource>
<mods:relatedItem id="D1D17DEEEDFBDEDF775789DB3BB3F363" type="host">
<mods:titleInfo id="1C35D01BA22F8A06B494B3FAEA123ED5">
<mods:title id="662286070F7705AD66D92160A916EB86">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4846456E0B514EBB253C01DE27A0937E">
<mods:date id="480818A60B12B54C0ED294109F7D2EC8">2018</mods:date>
<mods:detail id="1642D7E0C779D33FFEAAF256FF253071" type="pubDate">
<mods:number id="A188CEED97A0786EE31376FC3AD36272">2018-08-02</mods:number>
</mods:detail>
<mods:detail id="C935B66B4D98C31D6DE61A96B8D63051" type="volume">
<mods:number id="39A25179AC0CFFB282FACDC2AE25788E">4455</mods:number>
</mods:detail>
<mods:detail id="DD37435341C796CCA0E5F5E809A52D2E" type="issue">
<mods:number id="5936FA7A94FB4544FFC2E2A43DAD03F2">2</mods:number>
</mods:detail>
<mods:extent id="4074E14B0D9415B3489647EFD09F3713" unit="page">
<mods:start id="01665E23FB0717829789873B1EC0DF8D">343</mods:start>
<mods:end id="18278F7867629BABF21ED80D2097C254">362</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B86A1A842687D3A07A062ED15A7E81F7">journal article</mods:classification>
<mods:identifier id="F681D563B5E10B0C6C4D16F0532D069C" type="CLB-Dataset">29103</mods:identifier>
<mods:identifier id="71D68FCB0DF359586BF6936110669644" type="DOI">10.11646/zootaxa.4455.2.5</mods:identifier>
<mods:identifier id="A692BFC07C979438AE437F43221F87CB" type="GBIF-Dataset">568a5364-52f2-4feb-b08b-87eb85d4cf77</mods:identifier>
<mods:identifier id="575461DEFBD280975E3DD637B5B1B31F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F31AEF273E7FBBAFF448C28203FBC18C" type="Zenodo-Dep">1457283</mods:identifier>
<mods:identifier id="76D8E468D9499A2F4DBB4D382E8A4926" type="ZooBank">8CB311F1-431C-4861-956E-3200329E4F0F</mods:identifier>
</mods:mods>
<treatment id="039887E6FFF37A069ECB236EA3666629" ID-DOI="http://doi.org/10.5281/zenodo.5959230" ID-GBIF-Taxon="148691255" ID-Zenodo-Dep="5959230" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629" httpUri="http://treatment.plazi.org/id/039887E6FFF37A069ECB236EA3666629" lastPageId="14" lastPageNumber="357" pageId="5" pageNumber="348">
<subSubSection id="C32B657BFFF37A0D9ECB236EA60F621D" pageId="5" pageNumber="348" type="nomenclature">
<paragraph id="8B8E36F0FFF37A0D9ECB236EA545623A" blockId="5.[151,578,1337,1397]" box="[151,578,1337,1363]" pageId="5" pageNumber="348">
<heading id="D0C6819CFFF37A0D9ECB236EA545623A" bold="true" box="[151,578,1337,1363]" fontSize="11" level="1" pageId="5" pageNumber="348" reason="1">
<emphasis id="B945EAE2FFF37A0D9ECB236EA545623A" bold="true" box="[151,578,1337,1363]" pageId="5" pageNumber="348">
<taxonomicName id="4C314D73FFF37A0D9ECB236EA6E2623A" ID-CoL="8N8PY" authority="Nada" authorityName="Nada &amp; Ballantyne" authorityYear="2018" box="[151,485,1337,1363]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="dunguna">
<emphasis id="B945EAE2FFF37A0D9ECB236EA69D623A" bold="true" box="[151,410,1337,1363]" italics="true" pageId="5" pageNumber="348">Pygoluciola dunguna</emphasis>
Nada
</taxonomicName>
sp nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36F0FFF37A0D9ECB230BA60F621D" blockId="5.[151,578,1337,1397]" box="[151,264,1372,1397]" pageId="5" pageNumber="348">Figs 2̄37</paragraph>
</subSubSection>
<subSubSection id="C32B657BFFF37A0E9ECB23CAA35267BC" lastPageId="6" lastPageNumber="349" pageId="5" pageNumber="348" type="materials_examined">
<paragraph id="8B8E36F0FFF37A0D9ECB23CAA5BE62B3" blockId="5.[151,1437,1437,2038]" pageId="5" pageNumber="348">
<materialsCitation id="3B593CADFFF37A0D9ECB23CAA5BE62B3" ID-GBIF-Occurrence="1917075855" collectingDate="2016-07-19" collectionCode="FRIM" collectorName="B. Nada." country="Malaysia" county="Dungun" location="Jengai Forest" municipality="Jengai Forest Reserve" pageId="5" pageNumber="348" specimenCount="2" stateProvince="Terengganu" typeStatus="holotype">
<emphasis id="B945EAE2FFF37A0D9ECB23CAA7DF62DF" bold="true" box="[151,216,1437,1462]" pageId="5" pageNumber="348">
<typeStatus id="548A8852FFF37A0D9ECB23CAA7D362DF" box="[151,212,1437,1462]" pageId="5" pageNumber="348">Type</typeStatus>
.
</emphasis>
<typeStatus id="548A8852FFF37A0D9E8323CAA64062DF" box="[223,327,1437,1462]" pageId="5" pageNumber="348" type="holotype">Holotype</typeStatus>
male.
<emphasis id="B945EAE2FFF37A0D9FC823CAA53662DF" bold="true" box="[404,561,1437,1462]" pageId="5" pageNumber="348">
<collectingCountry id="F3267660FFF37A0D9FC823CAA52B62DF" box="[404,556,1437,1462]" name="Malaysia" pageId="5" pageNumber="348">MALAYSIA</collectingCountry>
.
</emphasis>
<collectingRegion id="49F5F812FFF37A0D9C6B23C9A5BC62DC" box="[567,699,1438,1461]" country="Malaysia" name="Terengganu" pageId="5" pageNumber="348">
<emphasis id="B945EAE2FFF37A0D9C6B23C9A5BC62DC" box="[567,699,1438,1461]" italics="true" pageId="5" pageNumber="348">Terengganu</emphasis>
</collectingRegion>
:
<collectingCounty id="62EF4E7CFFF37A0D9C9623C9A42062DF" box="[714,807,1438,1462]" pageId="5" pageNumber="348">Dungun</collectingCounty>
,
<collectingMunicipality id="6BEAAC8AFFF37A0D9D6E23CAA32C62DC" box="[818,1067,1437,1462]" pageId="5" pageNumber="348">
<location id="8EEE602BFFF37A0D9D6E23CAA4C062DC" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9D6E23CAA4C062DC" box="[818,967,1437,1462]" country="Malaysia" county="Dungun" municipality="Jengai Forest Reserve" name="Jengai Forest" pageId="5" pageNumber="348" stateProvince="Terengganu">Jengai Forest</location>
Reserve
</collectingMunicipality>
,
<location id="8EEE602BFFF37A0D9A6B23C9A3D662DF" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9A6B23C9A3D662DF" box="[1079,1233,1438,1462]" country="Malaysia" county="Dungun" municipality="Jengai Forest Reserve" name="Compartment" pageId="5" pageNumber="348" stateProvince="Terengganu">Compartment</location>
40, lat. 4.610939, long. 103.158919
<collectingDate id="EFCBE9D8FFF37A0D9F3F2396A6E262B3" box="[355,485,1473,1498]" pageId="5" pageNumber="348" value="2016-07-19">
<date id="FF8F1030FFF37A0D9F3F2396A6E262B3" box="[355,485,1473,1498]" pageId="5" pageNumber="348" value="2016-07-19">19.vii.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9FAC2395A55362B3" box="[496,596,1473,1498]" pageId="5" pageNumber="348">B. Nada.</collectorName>
(
<collectionCode id="ED20AE35FFF37A0D9C3E2395A5AA62B3" box="[610,685,1474,1498]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36F0FFF37A0E9E9B23B2A35267BC" blockId="5.[151,1437,1437,2038]" lastBlockId="6.[151,1437,151,2025]" lastPageId="6" lastPageNumber="349" pageId="5" pageNumber="348">
<materialsCitation id="3B593CADFFF37A0D9E9B23B2A5A5614B" ID-GBIF-Occurrence="1917075893" collectingDate="2015-06-21" collectionCode="FRIM" collectorName="B. Nada" country="Malaysia" elevation="350" latitude="3.80205" location="Perak" longLatPrecision="1" longitude="101.590965" pageId="5" pageNumber="348" specimenCount="1" stateProvince="Perak" typeStatus="paratype">
<typeStatus id="548A8852FFF37A0D9E9B23B2A6386297" box="[199,319,1509,1534]" pageId="5" pageNumber="348" type="paratype">
<emphasis id="B945EAE2FFF37A0D9E9B23B2A6386297" bold="true" box="[199,319,1509,1534]" pageId="5" pageNumber="348">Paratypes</emphasis>
</typeStatus>
.
<collectingCountry id="F3267660FFF37A0D9F0E23B2A6EF6297" box="[338,488,1509,1534]" name="Malaysia" pageId="5" pageNumber="348">
<emphasis id="B945EAE2FFF37A0D9F0E23B2A6EF6297" bold="true" box="[338,488,1509,1534]" pageId="5" pageNumber="348">MALAYSIA</emphasis>
</collectingCountry>
<emphasis id="B945EAE2FFF37A0D9FA823B2A5476297" box="[500,576,1509,1534]" italics="true" pageId="5" pageNumber="348">
<collectingRegion id="49F5F812FFF37A0D9FA823B2A53D6297" box="[500,570,1509,1534]" country="Malaysia" name="Perak" pageId="5" pageNumber="348">Perak</collectingRegion>
:
</emphasis>
Tanjung Malim, Gunung Liang,
<geoCoordinate id="EE055037FFF37A0D9D9023B2A34D6294" box="[972,1098,1509,1534]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="3.80205">3.80205 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9A0723B2A3F86294" box="[1115,1279,1509,1534]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="101.590965">101.590969 E</geoCoordinate>
,
<elevation id="001CD1C3FFF37A0D9B4C23B2A25B6294" box="[1296,1372,1509,1534]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="348" unit="m" value="350.0">
<quantity id="4CC99B15FFF37A0D9B4C23B2A25B6294" box="[1296,1372,1509,1534]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="348" unit="m" value="350.0">350 m</quantity>
</elevation>
a.s.l,
<collectingDate id="EFCBE9D8FFF37A0D9ECB205EA616614B" box="[151,273,1545,1570]" pageId="5" pageNumber="348" value="2015-06-21">
<date id="FF8F1030FFF37A0D9ECB205EA616614B" box="[151,273,1545,1570]" pageId="5" pageNumber="348" value="2015-06-21">21.vi.2015</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9F41205DA67A614B" box="[285,381,1545,1570]" pageId="5" pageNumber="348">B. Nada</collectorName>
, female (GL350) (
<collectionCode id="ED20AE35FFF37A0D9C13205DA59D614B" box="[591,666,1546,1570]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B593CADFFF37A0D9CF0205EA3FF614B" ID-GBIF-Occurrence="1917075891" box="[684,1272,1545,1570]" collectionCode="ANIC" collectorName="Gombak &amp; J. Buck" country="Malaysia" location="Selangor" pageId="5" pageNumber="348" specimenCount="1" stateProvince="Selangor" typeStatus="paratype">
<collectingRegion id="49F5F812FFF37A0D9CF0205EA416614B" box="[684,785,1545,1570]" country="Malaysia" name="Selangor" pageId="5" pageNumber="348">
<emphasis id="B945EAE2FFF37A0D9CF0205EA416614B" box="[684,785,1545,1570]" italics="true" pageId="5" pageNumber="348">Selangor</emphasis>
</collectingRegion>
:
<collectorName id="26C45326FFF37A0D9D7F205EA484614B" box="[803,899,1545,1570]" pageId="5" pageNumber="348">Gombak</collectorName>
x.4.1965,
<collectorName id="26C45326FFF37A0D9DA0205DA354614B" box="[1020,1107,1545,1570]" pageId="5" pageNumber="348">J. Buck</collectorName>
, male (
<collectionCode id="ED20AE35FFF37A0D9AFA205DA3F7614B" box="[1190,1264,1546,1570]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="5" pageNumber="348">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B593CADFFF37A0D9B58205EA290612C" ID-GBIF-Occurrence="1917075904" collectingDate="2015-01-30" collectionCode="MZUM" collectorName="W. Jusoh" country="Malaysia" latitude="3.233139" location="Thary" longLatPrecision="1" longitude="101.6304" pageId="5" pageNumber="348" specimenCount="1" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9B58205EA277614B" box="[1284,1392,1545,1570]" pageId="5" pageNumber="348" value="2015-01-30">
<date id="FF8F1030FFF37A0D9B58205EA277614B" box="[1284,1392,1545,1570]" pageId="5" pageNumber="348" value="2015-01-30">30.i.2015</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9B21205DA7D0612F" pageId="5" pageNumber="348">W. Jusoh</collectorName>
&amp;
<location id="8EEE602BFFF37A0D9F57207AA648612F" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9F57207AA648612F" box="[267,335,1581,1606]" country="Malaysia" latitude="3.233139" longLatPrecision="1" longitude="101.6304" name="Thary" pageId="5" pageNumber="348">Thary</location>
, male (
<collectionCode id="ED20AE35FFF37A0D9FED2079A508612F" box="[433,527,1582,1606]" pageId="5" pageNumber="348">MZUM</collectionCode>
). Kepong, Forest Research Institute
<collectingCountry id="F3267660FFF37A0D9D93207AA33F612F" box="[975,1080,1581,1606]" name="Malaysia" pageId="5" pageNumber="348">Malaysia</collectingCountry>
,
<geoCoordinate id="EE055037FFF37A0D9A102079A3DB612C" box="[1100,1244,1582,1606]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="3.233139">3.233139 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9AAC207AA290612C" box="[1264,1431,1581,1606]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="101.6304">101.630403 E</geoCoordinate>
</materialsCitation>
,
<date id="FF8F1030FFF37A0D9ECB2006A6176103" box="[151,272,1617,1642]" pageId="5" pageNumber="348" value="2011-06-14">14.vi.2011</date>
, Rover Track,
<quantity id="4CC99B15FFF37A0D9FEE2006A6F16100" box="[434,502,1617,1642]" metricMagnitude="2" metricUnit="m" metricValue="1.71" pageId="5" pageNumber="348" unit="m" value="171.0">171 m</quantity>
a.s.l., O. Khirul-Faizal, male (ADULT8);
<date id="FF8F1030FFF37A0D9D8C2006A34D6103" box="[976,1098,1617,1642]" pageId="5" pageNumber="348" value="2011-06-27">27.vi.2011</date>
, Canopy Walkway Trail, C.N. Nafaruding, male (ADULTCNPY2);
<materialsCitation id="3B593CADFFF37A0D9C2E2022A38F61D8" ID-GBIF-Occurrence="1917075875" collectingDate="2011-07-06" collectionCode="FRIM" collectorName="M. S. Salman" country="Malaysia" elevation="139" latitude="3.55395" location="Waterfall Trail" longLatPrecision="1" longitude="101.734215" pageId="5" pageNumber="348" specimenCount="1" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9C2E2022A5E261E7" box="[626,741,1653,1678]" pageId="5" pageNumber="348" value="2011-07-06">
<date id="FF8F1030FFF37A0D9C2E2022A5E261E7" box="[626,741,1653,1678]" pageId="5" pageNumber="348" value="2011-07-06">6.vii.2011</date>
</collectingDate>
,
<location id="8EEE602BFFF37A0D9D582022A4B961E7" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9D582022A4B961E7" box="[772,958,1653,1678]" country="Malaysia" latitude="3.55395" longLatPrecision="1" longitude="101.734215" name="Waterfall Trail" pageId="5" pageNumber="348">Waterfall Trail</location>
,
<elevation id="001CD1C3FFF37A0D9D812021A33061E4" box="[989,1079,1654,1678]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="5" pageNumber="348" unit="m" value="139.0">
<quantity id="4CC99B15FFF37A0D9D812021A33061E4" box="[989,1079,1654,1678]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="5" pageNumber="348" unit="m" value="139.0">139 m</quantity>
</elevation>
a.s.l,
<collectorName id="26C45326FFF37A0D9AC32021A24061E7" box="[1183,1351,1653,1678]" pageId="5" pageNumber="348">M.S. Salman</collectorName>
, male (ADULTWFALL4) (
<collectionCode id="ED20AE35FFF37A0D9FD820CDA6C861DB" box="[388,463,1690,1714]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
). Kuala Kubu Bharu, Bukit Kutu,
<geoCoordinate id="EE055037FFF37A0D9D0120CEA4DE61D8" box="[861,985,1689,1714]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="3.55395">3.55395 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9DBB20CDA38F61D8" box="[999,1160,1690,1714]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="101.734215">101.734211 E</geoCoordinate>
</materialsCitation>
,
<date id="FF8F1030FFF37A0D9ACA20CEA20061DB" box="[1174,1287,1689,1714]" pageId="5" pageNumber="348" value="2015-05-26">
26.v.
<quantity id="4CC99B15FFF37A0D9A9620CEA20061DB" box="[1226,1287,1689,1714]" metricMagnitude="3" metricUnit="m" metricValue="2.0154" pageId="5" pageNumber="348" unit="m" value="2015.4">2015</quantity>
</date>
, 400 m a.s.l. and
<quantity id="4CC99B15FFF37A0D9E9420EAA61661BC" box="[200,273,1725,1750]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="348" unit="m" value="600.0">600 m</quantity>
a.s.l., B. Nada, male, female (BK400) (BK600) (FRIM).
<materialsCitation id="3B593CADFFF37A0D9DF920E9A5D36077" ID-GBIF-Occurrence="1917075853" collectingDate="2016-05-10" collectorName="S. Muhammad-Jafni &amp; Compartment" country="Malaysia" latitude="4.693225" location="Besul Forest" longLatPrecision="1" longitude="103.167175" municipality="Dungun" pageId="5" pageNumber="348" specimenCount="2" specimenCount-male="2" stateProvince="Terengganu" typeStatus="paratype">
<collectingRegion id="49F5F812FFF37A0D9DF920E9A32E61BC" box="[933,1065,1726,1749]" country="Malaysia" name="Terengganu" pageId="5" pageNumber="348">
<emphasis id="B945EAE2FFF37A0D9DF920E9A32E61BC" box="[933,1065,1726,1749]" italics="true" pageId="5" pageNumber="348">Terengganu</emphasis>
</collectingRegion>
:
<collectingMunicipality id="6BEAAC8AFFF37A0D9A6520E9A39161BF" box="[1081,1174,1726,1750]" pageId="5" pageNumber="348">Dungun</collectingMunicipality>
,
<location id="8EEE602BFFF37A0D9AFF20EAA23561BC" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9AFF20EAA23561BC" box="[1187,1330,1725,1750]" country="Malaysia" latitude="4.693225" longLatPrecision="1" longitude="103.167175" municipality="Dungun" name="Besul Forest" pageId="5" pageNumber="348" stateProvince="Terengganu">Besul Forest</location>
Reserve, Compartment 10, l
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,
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,
<collectingDate id="EFCBE9D8FFF37A0D9CB520B6A45D6193" box="[745,858,1761,1786]" pageId="5" pageNumber="348" value="2016-05-10">
<date id="FF8F1030FFF37A0D9CB520B6A45D6193" box="[745,858,1761,1786]" pageId="5" pageNumber="348" value="2016-05-10">10.v.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9D2D20B5A3646193" box="[881,1123,1761,1786]" pageId="5" pageNumber="348">S. Muhammad-Jafni</collectorName>
,
<specimenCount id="9D37FD79FFF37A0D9A2620B5A3D96193" box="[1146,1246,1761,1786]" pageId="5" pageNumber="348" type="male">2 males</specimenCount>
, female (B10);
<collectorName id="26C45326FFF37A0D9ECB2151A6366077" box="[151,305,1798,1822]" pageId="5" pageNumber="348">Compartment</collectorName>
19, lat.4.63415, long. 103.200553
</materialsCitation>
,
<materialsCitation id="3B593CADFFF37A0D9CBA2152A31C6028" ID-GBIF-Occurrence="1917075854" collectingDate="2016-08-08" collectionCode="FRIM" collectorName="Z. A. Saiful-Azhari" country="Malaysia" latitude="4.548261" location="Jengai Forest" longLatPrecision="1" longitude="102.97988" pageId="5" pageNumber="348" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9CBA2152A4656077" box="[742,866,1797,1822]" pageId="5" pageNumber="348" value="2016-08-08">
<date id="FF8F1030FFF37A0D9CBA2152A4656077" box="[742,866,1797,1822]" pageId="5" pageNumber="348" value="2016-08-08">8.viii.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9D292151A3556077" box="[885,1106,1797,1822]" pageId="5" pageNumber="348">Z.A. Saiful-Azhari</collectorName>
,
<specimenCount id="9D37FD79FFF37A0D9A392151A3C16077" box="[1125,1222,1797,1822]" pageId="5" pageNumber="348" type="male">4 males</specimenCount>
,
<specimenCount id="9D37FD79FFF37A0D9A842151A24B6077" box="[1240,1356,1797,1822]" pageId="5" pageNumber="348" type="female">3 females</specimenCount>
(B19) (
<collectionCode id="ED20AE35FFF37A0D9EC3217DA7ED602B" box="[159,234,1834,1858]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
).
<location id="8EEE602BFFF37A0D9EA3217EA6906028" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9EA3217EA6906028" box="[255,407,1833,1858]" country="Malaysia" latitude="4.548261" longLatPrecision="1" longitude="102.97988" name="Jengai Forest" pageId="5" pageNumber="348" stateProvince="Terengganu">Jengai Forest</location>
Reserve, Compartment 52,
<geoCoordinate id="EE055037FFF37A0D9C83217EA46E6028" box="[735,873,1833,1858]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="4.548261">4.548261 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9D24217DA31C6028" box="[888,1051,1834,1858]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="102.97988">102.979878 E</geoCoordinate>
</materialsCitation>
,
<materialsCitation id="3B593CADFFF37A0D9A75217EA5EC600C" ID-GBIF-Occurrence="1917075892" collectingDate="2016-06-30" collectorName="M. S. Asraf" country="Malaysia" latitude="4.610939" location="Compartment" longLatPrecision="1" longitude="103.15892" pageId="5" pageNumber="348" specimenCount="2" specimenCount-male="2" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9A75217EA3A4602B" box="[1065,1187,1833,1858]" pageId="5" pageNumber="348" value="2016-06-30">
<date id="FF8F1030FFF37A0D9A75217EA3A4602B" box="[1065,1187,1833,1858]" pageId="5" pageNumber="348" value="2016-06-30">30.vi.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9AEE217DA233602B" box="[1202,1332,1833,1858]" pageId="5" pageNumber="348">M.S. Asraf</collectorName>
,
<specimenCount id="9D37FD79FFF37A0D9B1F217DA29B602B" box="[1347,1436,1833,1858]" pageId="5" pageNumber="348" type="male">2 males</specimenCount>
(JI52);
<location id="8EEE602BFFF37A0D9EB52119A684600F" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9EB52119A684600F" box="[233,387,1870,1894]" country="Malaysia" latitude="4.610939" longLatPrecision="1" longitude="103.15892" name="Compartment" pageId="5" pageNumber="348" stateProvince="Terengganu">Compartment</location>
40,
<geoCoordinate id="EE055037FFF37A0D9FEA211AA539600C" box="[438,574,1869,1894]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="4.610939">4.610939 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9C16211AA5EC600C" box="[586,747,1869,1894]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="103.15892">103.158919 E</geoCoordinate>
</materialsCitation>
,
<materialsCitation id="3B593CADFFF37A0D9CAB211AA4BF60E0" ID-GBIF-Occurrence="1917075894" collectingDate="2016-07-19" collectionCode="FRIM" country="Malaysia" latitude="4.8699" location="Jerangau Forest" longLatPrecision="5" longitude="103.04672" pageId="5" pageNumber="348" specimenCount="1" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9CAB211AA47E600F" box="[759,889,1869,1894]" pageId="5" pageNumber="348" value="2016-07-19">
<date id="FF8F1030FFF37A0D9CAB211AA47E600F" box="[759,889,1869,1894]" pageId="5" pageNumber="348" value="2016-07-19">19.vii.2016</date>
</collectingDate>
, R.S. Raja-Ahmad-Bazli, female (JI40) (
<collectionCode id="ED20AE35FFF37A0D9B1A2119A296600F" box="[1350,1425,1870,1894]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
).
<location id="8EEE602BFFF37A0D9ECB2125A64D60E0" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9ECB2125A64D60E0" box="[151,330,1906,1930]" country="Malaysia" latitude="4.8699" longLatPrecision="5" longitude="103.04672" name="Jerangau Forest" pageId="5" pageNumber="348" stateProvince="Terengganu">Jerangau Forest</location>
Reserve, Compartment 100,
<geoCoordinate id="EE055037FFF37A0D9CC72126A40F60E0" box="[667,776,1905,1930]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="5" value="4.8699">4.8699 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9D4A2126A4BF60E0" box="[790,952,1905,1930]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="103.04672">103.046725 E</geoCoordinate>
</materialsCitation>
,
<materialsCitation id="3B593CADFFF37A0D9D992126A59160C4" ID-GBIF-Occurrence="1917075857" collectingDate="2016-07-20" collectorName="M. S. Asraf" country="Malaysia" latitude="4.862742" location="Compartment" longLatPrecision="1" longitude="103.04864" pageId="5" pageNumber="348" specimenCount="1" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9D992126A34060E3" box="[965,1095,1905,1930]" pageId="5" pageNumber="348" value="2016-07-20">
<date id="FF8F1030FFF37A0D9D992126A34060E3" box="[965,1095,1905,1930]" pageId="5" pageNumber="348" value="2016-07-20">20.vii.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9A092125A3D160E3" box="[1109,1238,1905,1930]" pageId="5" pageNumber="348">M.S. Asraf</collectorName>
, female (JU100);
<location id="8EEE602BFFF37A0D9ECB21C1A63660C7" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9ECB21C1A63660C7" box="[151,305,1942,1966]" country="Malaysia" latitude="4.862742" longLatPrecision="1" longitude="103.04864" name="Compartment" pageId="5" pageNumber="348" stateProvince="Terengganu">Compartment</location>
99,
<geoCoordinate id="EE055037FFF37A0D9F3F21C2A6ED60C4" box="[355,490,1941,1966]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="4.862742">4.862742 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9FAA21C2A59160C4" box="[502,662,1941,1966]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="103.04864">103.048636 E</geoCoordinate>
</materialsCitation>
,
<materialsCitation id="3B593CADFFF37A0D9CFE21C2A30060B8" ID-GBIF-Occurrence="1917075879" collectingDate="2016-08-17" collectionCode="FRIM" collectorName="S. Mohd-Fadhlullah" country="Malaysia" latitude="4.858047" location="Pasir Raja Barat Forest" longLatPrecision="1" longitude="103.00939" pageId="5" pageNumber="348" specimenCount="4" specimenCount-male="4" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9CFE21C2A42B60C7" box="[674,812,1941,1966]" pageId="5" pageNumber="348" value="2016-08-17">
<date id="FF8F1030FFF37A0D9CFE21C2A42B60C7" box="[674,812,1941,1966]" pageId="5" pageNumber="348" value="2016-08-17">17.viii.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9D6421C1A31D60C7" box="[824,1050,1941,1966]" pageId="5" pageNumber="348">S. Mohd-Fadhlullah</collectorName>
,
<specimenCount id="9D37FD79FFF37A0D9A6C21C1A38E60C7" box="[1072,1161,1941,1966]" pageId="5" pageNumber="348" type="male">4 males</specimenCount>
, female (JU99) (
<collectionCode id="ED20AE35FFF37A0D9B1A21C1A29660C7" box="[1350,1425,1942,1966]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
).
<location id="8EEE602BFFF37A0D9ECB21EEA69C60B8" LSID="urn:lsid:plazi:treatment:039887E6FFF37A069ECB236EA3666629:8EEE602BFFF37A0D9ECB21EEA69C60B8" box="[151,411,1977,2002]" country="Malaysia" latitude="4.858047" longLatPrecision="1" longitude="103.00939" name="Pasir Raja Barat Forest" pageId="5" pageNumber="348" stateProvince="Terengganu">Pasir Raja Barat Forest</location>
Reserve, Compartment 87,
<geoCoordinate id="EE055037FFF37A0D9C8921EEA45A60B8" box="[725,861,1977,2002]" direction="north" orientation="latitude" pageId="5" pageNumber="348" precision="1" value="4.858047">4.858047 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF37A0D9D3421EDA30060B8" box="[872,1031,1978,2002]" direction="east" orientation="longitude" pageId="5" pageNumber="348" precision="1" value="103.00939">103.009394 E</geoCoordinate>
</materialsCitation>
,
<date id="FF8F1030FFF37A0D9A4F21EEA38860BB" box="[1043,1167,1977,2002]" pageId="5" pageNumber="348" value="2016-08-02">2.viii.2016</date>
, S. Muhammad-Jafni, 6 males, 2 females,
<materialsCitation id="3B593CADFFF37A0E9F39218AA5FA67C6" ID-GBIF-Occurrence="1917075889" collectingDate="2016-08-22" collectionCode="FRIM" collectorName="Z. A. Saiful-Azhari" country="Malaysia" lastPageId="6" lastPageNumber="349" latitude="4.984031" location="Terengganu" longLatPrecision="1" longitude="102.93217" pageId="5" pageNumber="348" specimenCount="1" specimenCount-female="1" stateProvince="Terengganu" typeStatus="paratype">
<collectingDate id="EFCBE9D8FFF37A0D9F39218AA6E8609F" box="[357,495,2013,2038]" pageId="5" pageNumber="348" value="2016-08-22">
<date id="FF8F1030FFF37A0D9F39218AA6E8609F" box="[357,495,2013,2038]" pageId="5" pageNumber="348" value="2016-08-22">22.viii.2016</date>
</collectingDate>
,
<collectorName id="26C45326FFF37A0D9FA02189A5D4609F" box="[508,723,2013,2038]" pageId="5" pageNumber="348">Z.A. Saiful-Azhari</collectorName>
,
<specimenCount id="9D37FD79FFF37A0D9CBC2189A444609F" box="[736,835,2013,2038]" pageId="5" pageNumber="348" type="female">1 female</specimenCount>
(PB87) (
<collectionCode id="ED20AE35FFF37A0D9DF12189A4FF609F" box="[941,1016,2014,2038]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="5" pageNumber="348">FRIM</collectionCode>
). Hulu
<collectingRegion id="49F5F812FFF37A0D9A112189A3D6609F" box="[1101,1233,2014,2038]" country="Malaysia" name="Terengganu" pageId="5" pageNumber="348">Terengganu</collectingRegion>
Tambahan Forest Reserve, Compartment 27,
<geoCoordinate id="EE055037FFF07A0E9F9726CFA55567C6" box="[459,594,152,176]" direction="north" orientation="latitude" pageId="6" pageNumber="349" precision="1" value="4.984031">4.984031 N</geoCoordinate>
,
<geoCoordinate id="EE055037FFF07A0E9C0126C0A5FA67C6" box="[605,765,151,176]" direction="east" orientation="longitude" pageId="6" pageNumber="349" precision="1" value="102.93217">102.932167 E</geoCoordinate>
</materialsCitation>
,
<date id="FF8F1030FFF07A0E9D5426C0A49567D9" box="[776,914,151,176]" pageId="6" pageNumber="349" value="2016-08-30">30.viii.2016</date>
, Z.A. Saiful-Azhari, 4 males, female (UTT27) (FRIM). All but two specimens (in ANIC and MZUM) are preserved in 70% ethanol.
</paragraph>
</subSubSection>
<subSubSection id="C32B657BFFF07A0E9E9B2688A50B66C4" pageId="6" pageNumber="349" type="diagnosis">
<paragraph id="8B8E36F0FFF07A0E9E9B2688A50B66C4" blockId="6.[151,1437,151,2025]" pageId="6" pageNumber="349">
<emphasis id="B945EAE2FFF07A0E9E9B2688A63D6791" bold="true" box="[199,314,223,248]" pageId="6" pageNumber="349">Diagnosis</emphasis>
. Dorsally pale yellowish brown with diffuse pale brown pronotal markings; ventral abdomen yellow except for white LO in
<date id="FF8F1030FFF07A0E9FFF2753A6C06674" box="[419,455,260,285]" pageId="6" pageNumber="349">V6</date>
, 7, with narrow black posterior margin of
<date id="FF8F1030FFF07A0E9DF72753A4D76674" box="[939,976,260,285]" pageId="6" pageNumber="349">V5</date>
; 9.1¯10.6 mm long. The only species of
<taxonomicName id="4C314D73FFF07A0E9ECB2770A6186629" box="[151,287,295,320]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="B945EAE2FFF07A0E9ECB2770A6186629" box="[151,287,295,320]" italics="true" pageId="6" pageNumber="349">Pygoluciola</emphasis>
</taxonomicName>
having a broad apically truncate MPP which inclines without any bend slightly upwards to engage against the under surface of the very narrow T8, and a yellow ventral abdomen with narrow black posterior margin of
<date id="FF8F1030FFF07A0E9EE42738A7E666E1" box="[184,225,367,392]" pageId="6" pageNumber="349">V5.</date>
Female macropterous and observed in flight; bursa plates in form of two elongated slender hooks which incline anteriorly and posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C32B657BFFF07A069E9B27E0A3666629" lastPageId="14" lastPageNumber="357" pageId="6" pageNumber="349" type="description">
<paragraph id="8B8E36F0FFF07A0E9E9B27E0A266648C" blockId="6.[151,1437,151,2025]" pageId="6" pageNumber="349">
<emphasis id="B945EAE2FFF07A0E9E9B27E0A6B166B9" bold="true" box="[199,438,439,464]" pageId="6" pageNumber="349">Description of male.</emphasis>
9.1
<emphasis id="B945EAE2FFF07A0E9F8327E0A6EA66B9" bold="true" box="[479,493,439,464]" pageId="6" pageNumber="349">–</emphasis>
10.6 mm long. Colour (Figs 2̄9, 14, 16̄18, 25, 26): creamy yellow semitransparent pronotum with single median, and paired reddish markings along the posterior margin, all of which often coincide with retraction of fat body beneath the cuticle (Figs 2, 4, 6¯9); retraction of fat body along anterior margin of pronotum in ethanol preserved specimens allows black head to be visible from beneath and gives the erroneous impression of a dark marking along the pronotum (e.g. Fig. 9); MN and MS brown; elytra semitransparent, very light brown with lateral margins slightly paler (underlying body outline confuses interpretation of elytral colour especially in ethanol preserved specimens where the elytral punctures are dark brown at their bases) (Figs 2, 6); one of the two pinned Gombak males (Fig. 4) pale creamy yellow dorsally, pronotum with faint traces of median ginger brown markings (single median mark and paired areas along posterior margin); head antennae and palpi black (
<figureCitation id="130A2A75FFF07A0E9EC324A0A6266466" box="[159,289,759,784]" captionStart="FIGURES 17, 18" captionStartId="9.[151,264,722,745]" captionTargetBox="[175,1405,193,705]" captionTargetId="figure@9.[175,1411,193,715]" captionTargetPageId="9" captionText="FIGURES 17, 18. Head, antennae and mouthparts, dorsal (17) and left lateral. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457289/files/figure.png" pageId="6" pageNumber="349">Figs 17, 18</figureCitation>
); venter of thorax pale yellow (Figs 3, 6); legs 1, 2 pale brown with black tibiae and tarsi; legs 3 light brown with slightly darker tibiae and black tarsi (BK600 single male with brown tints in middle area of coxae; B19 (1) II has tibiae 3 pale yellow); abdominal ventrites yellow with posterior margin of
<date id="FF8F1030FFF07A0E9A132569A376643E" box="[1103,1137,830,855]" pageId="6" pageNumber="349">V5</date>
narrowly black (Figs 3, 6, 14, 25, 26); LO in
<date id="FF8F1030FFF07A0E9F3A2535A68F6410" box="[358,392,866,889]" pageId="6" pageNumber="349">V7</date>
white, retracted from sides and posterior margin, with white areas at side and area behind LO semitransparent with white fat body visible beneath cuticle (Figs 3, 6, 10, 14), and narrow longitudinal median line on MPP which is brown in the MZUM specimen (Fig. 6); abdominal tergites pale yellowish, semitransparent, T6 slightly darker than rest, T7, 8 slightly paler than preceding tergites (
<figureCitation id="130A2A75FFF07A0E9DC4259BA4EF648C" box="[920,1000,972,997]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Fig. 16</figureCitation>
shows one of two pinned males).
</paragraph>
<paragraph id="8B8E36F0FFF07A0E9E9B25A7A47A6291" blockId="6.[151,1437,151,2025]" pageId="6" pageNumber="349">
Pronotum (Figs 2, 4, 5, 7¯9): width subequal to humeral width; median anterior margin broadly rounded and barely produced in front of the broadly rounded anterolateral corners; lateral margins strongly divergent along anterior 2/3 and posterolateral corners rounded and not projecting beyond median posterior margin; surface along posterior margin coinciding with the brown markings may appear elevated (only clearly visible in pinned specimens Figs 7, 8). Elytron (Figs 2, 4, 5): slightly convex sided; interstitial lines not well defined. Head: (Figs 3, 6, 17, 18, 25, 26) GHW
<date id="FF8F1030FFF07A0E9FF422F6A6CB63D1" box="[424,460,1185,1208]" pageId="6" pageNumber="349">7 x</date>
SIW; ASD &lt;ASW. Mouthparts: apical labial palpomere laterally flattened of the form of a broad triangle with inner edge dentate (up to 7 teeth). Antennae (
<figureCitation id="130A2A75FFF07A0E9D992293A34B63B4" box="[965,1100,1220,1245]" captionStart="FIGURES 17, 18" captionStartId="9.[151,264,722,745]" captionTargetBox="[175,1405,193,705]" captionTargetId="figure@9.[175,1411,193,715]" captionTargetPageId="9" captionText="FIGURES 17, 18. Head, antennae and mouthparts, dorsal (17) and left lateral. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457289/files/figure.png" pageId="6" pageNumber="349">Figs 17, 18</figureCitation>
): length&gt;
<date id="FF8F1030FFF07A0E9A922292A3F363B5" box="[1230,1268,1221,1245]" pageId="6" pageNumber="349">2 x</date>
GHW but not attaining
<date id="FF8F1030FFF07A0E9EA322BFA6256269" box="[255,290,1256,1280]" pageId="6" pageNumber="349">3 x</date>
GHW; scape clavate, all other segments elongate slender about
<date id="FF8F1030FFF07A0E9DAC22BEA3156269" box="[1008,1042,1257,1280]" pageId="6" pageNumber="349">7 x</date>
as long as wide. Abdomen (Figs 3, 6, 10̄16): posterior margin of
<date id="FF8F1030FFF07A0E9FAD235BA514624A" box="[497,531,1292,1315]" pageId="6" pageNumber="349">V7</date>
with elongate apically truncate MPP which curves gently upwards with the apex engaging against the underside of T8 (
<figureCitation id="130A2A75FFF07A0E9C1E2379A593622E" box="[578,660,1326,1351]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Fig. 12</figureCitation>
); dorsally reflexed margins of
<date id="FF8F1030FFF07A0E9DB62378A30B622F" box="[1002,1036,1327,1350]" pageId="6" pageNumber="349">V7</date>
well defined and anterior margin to either side of MPP produced into short rounded pieces (arrowed in
<figureCitation id="130A2A75FFF07A0E9DD62305A4DD6203" box="[906,986,1362,1387]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Fig. 11</figureCitation>
); reflexed margin in two parts on dorsal surface of MPP where the narrow margin between left and right side is visible from beneath as a line in the middle of the MPP (
<figureCitation id="130A2A75FFF07A0E9F7123CEA6D962D8" box="[301,478,1433,1458]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Figs 10, 11, 13</figureCitation>
); much darker markings observed in pinned male (
<figureCitation id="130A2A75FFF07A0E9A7223CEA38562D8" box="[1070,1154,1433,1458]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Fig. 13</figureCitation>
) may be due to age and appear to be accumulated material. T8 with narrow triangular posterior margin, lateral wing like projections and elongate slender anterolateral prolongations that diverge (
<figureCitation id="130A2A75FFF07A0E9D402388A4776291" box="[796,880,1503,1528]" captionStart="FIGURES 10–16" captionStartId="8.[151,264,1912,1935]" captionTargetBox="[208,1361,193,1888]" captionTargetId="figure@8.[208,1361,193,1889]" captionTargetPageId="8" captionText="FIGURES 10–16. (10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="6" pageNumber="349">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8B8E36F0FFF07A0E9E9B2054A51B6080" blockId="6.[151,1437,151,2025]" pageId="6" pageNumber="349">
Aedeagal sheath (
<figureCitation id="130A2A75FFF07A0E9FCF2054A6F76175" box="[403,496,1539,1564]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Figs 22</figureCitation>
̄24): basal ¾ of sheath sternite with narrow sclerotized margins which expand in apical ¼ where they are terminated by a slightly curved, transverse narrow strip of cuticle; to the sides of the apical ¼ (coinciding with the area of expansion of the median strip) there are wider more membranous portions (
<figureCitation id="130A2A75FFF07A0E9B67201DA288610A" box="[1339,1423,1610,1635]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Fig. 22</figureCitation>
); tip of sternite terminates in paired narrowed hairy membranous lobes arising at the sides of the transverse strip, and are not clearly visible in
<figureCitation id="130A2A75FFF07A0E9FF120C6A6F861C3" box="[429,511,1681,1706]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Fig. 22</figureCitation>
; anterior margin of tergite shallowly emarginated. Aedeagus (
<figureCitation id="130A2A75FFF07A0E9AE620C6A21F61C3" box="[1210,1304,1681,1706]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Figs 19</figureCitation>
̄21): 2.5 x as long as wide and evenly wide along its entire length; basal piece well defined and in two well cuticularised narrow parts; membranous apical section of LL having short apically acute hair bearing lobe at the outer margin just before origin of the more membranous portion (arrowed in
<figureCitation id="130A2A75FFF07A0E9DC620ACA321607D" box="[922,1062,1787,1812]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Figs 19, 21</figureCitation>
); with dense line of long hairs originating at this lobe and continuing inside their lateral margins (
<figureCitation id="130A2A75FFF07A0E9DD52148A4DB6051" box="[905,988,1823,1848]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Fig. 21</figureCitation>
); membranous portion of LL as wide at base as at apex with both lobes having apices irregularly inclined ventrally (assumed to be an artefact of preparation) (
<figureCitation id="130A2A75FFF07A0E9F6D2131A68B6016" box="[305,396,1894,1919]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Figs 19</figureCitation>
¯21); ML tapering to a rounded narrow apex, much shorter than LL and extending a little beyond the posterior margin of the darker basal portion of the LL which is widely separated for slightly more than half its length in middorsal line; anterior margin of basal portion of LL asymmetrical, being produced on its left side and margin rounded (
<figureCitation id="130A2A75FFF07A0E9FE72187A5176080" box="[443,528,2000,2025]" captionStart="FIGURES 19–24. 19" captionStartId="10.[151,264,1971,1994]" captionTargetBox="[192,1371,195,1899]" captionTargetId="figure@10.[192,1375,195,1934]" captionTargetPageId="10" captionText="FIGURES 19–24. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="6" pageNumber="349">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8B8E36F0FFF17A0F9ECB21C1A40560A2" blockId="7.[151,1435,1942,1996]" pageId="7" pageNumber="350">
<emphasis id="B945EAE2FFF17A0F9ECB21C1A63160C5" bold="true" box="[151,310,1942,1965]" pageId="7" pageNumber="350">FIGURES 2–9</emphasis>
. (2, 3 J152 (1); 4 Gombak pinned male ANIC; 5, 6 Gombak pinned male MZUM; 9 B192)). 2‾6 habitus, 2, 4, 5 dorsal, 3, 6 ventral; 7‾9 pronota dorsal. Scale line is
<quantity id="4CC99B15FFF17A0F9C9D21E1A5F860A2" box="[705,767,1974,1995]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="350" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<caption id="DF4E6678FFFE7A009ECB212FA6BF6082" httpUri="https://zenodo.org/record/1457287/files/figure.png" pageId="8" pageNumber="351" startId="8.[151,264,1912,1935]" targetBox="[208,1361,193,1888]" targetPageId="8">
<paragraph id="8B8E36F0FFFE7A009ECB212FA6BF6082" blockId="8.[151,1436,1912,2027]" pageId="8" pageNumber="351">
<emphasis id="B945EAE2FFFE7A009ECB212FA65260E7" bold="true" box="[151,341,1912,1935]" pageId="8" pageNumber="351">FIGURES 10–16.</emphasis>
(10 Gombak pinned male ANIC; 11‾16 Gombak pinned male MZUM). 10 terminal abdomen from behind, ventral surface uppermost; 11 dorsal surface V7 (rounded projections of reflexed dorsal margin arrowed); 12 right lateral terminal abdomen; 13 dorsal surface MPP only; 14, 16 whole abdomen ventral (14) and dorsal; 15 T8 with lateral ‘wings’ arrowed. Scale line is 1 mm.
</paragraph>
</caption>
<caption id="DF4E6678FFFF7A019ECB2485A37C6580" box="[151,1147,722,745]" httpUri="https://zenodo.org/record/1457289/files/figure.png" pageId="9" pageNumber="352" startId="9.[151,264,722,745]" targetBox="[175,1405,193,705]" targetPageId="9">
<paragraph id="8B8E36F0FFFF7A019ECB2485A37C6580" blockId="9.[151,1147,722,745]" box="[151,1147,722,745]" pageId="9" pageNumber="352">
<emphasis id="B945EAE2FFFF7A019ECB2485A6546581" bold="true" box="[151,339,722,745]" pageId="9" pageNumber="352">FIGURES 17, 18.</emphasis>
Head, antennae and mouthparts, dorsal (17) and left lateral. Scale line is 1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36F0FFFF7A019E9B2547A5CD6248" blockId="9.[151,1437,784,1313]" pageId="9" pageNumber="352">
<emphasis id="B945EAE2FFFF7A019E9B2547A6316440" bold="true" box="[199,310,784,809]" pageId="9" pageNumber="352">Female. (</emphasis>
<figureCitation id="130A2A75FFFF7A019F6A2547A6886440" box="[310,399,784,809]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1936,1959]" captionTargetBox="[215,1371,193,1877]" captionTargetId="figure@11.[208,1379,193,1916]" captionTargetPageId="11" captionText="FIGURES 25–28. Ventral aspect, mating pairs, male to left (25 JI40 (8), 26 B10 (gap 5-6)). 27, 28 dissected abdomen, anterior end to top of page (B19 (2-3)) 27 ventral; 28 dorsal. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457293/files/figure.png" pageId="9" pageNumber="352">Figs 25</figureCitation>
̄37). 9.2̄10.3 mm long. Macropterous and taken in flight. Coloured as for male except for white LO in
<date id="FF8F1030FFFF7A019F742563A64D6424" box="[296,330,820,845]" pageId="9" pageNumber="352">V6</date>
only and
<date id="FF8F1030FFFF7A019FE02562A6E76425" box="[444,480,821,844]" pageId="9" pageNumber="352">V7</date>
, 8 pale coloured;
<date id="FF8F1030FFFF7A019CF72562A5CA6425" box="[683,717,821,844]" pageId="9" pageNumber="352">V7</date>
may have narrow darker marking in anteromedian area and the darker anterior apodeme of
<date id="FF8F1030FFFF7A019F93250EA6F66418" box="[463,497,857,881]" pageId="9" pageNumber="352">V8</date>
may be visible through the transparent cuticle (e.g.
<figureCitation id="130A2A75FFFF7A019A66250FA3976418" box="[1082,1168,856,881]" captionStart="FIGURES 25–28" captionStartId="11.[151,264,1936,1959]" captionTargetBox="[215,1371,193,1877]" captionTargetId="figure@11.[208,1379,193,1916]" captionTargetPageId="11" captionText="FIGURES 25–28. Ventral aspect, mating pairs, male to left (25 JI40 (8), 26 B10 (gap 5-6)). 27, 28 dissected abdomen, anterior end to top of page (B19 (2-3)) 27 ventral; 28 dorsal. Scale line is 1 mm." httpUri="https://zenodo.org/record/1457293/files/figure.png" pageId="9" pageNumber="352">Figs 27</figureCitation>
, 30, 33). Abdomen: LO occupying all of
<date id="FF8F1030FFFF7A019F09252BA67064FC" box="[341,375,892,917]" pageId="9" pageNumber="352">V6</date>
only. Reproductive system differs little from what has been described elsewhere. An elongate vagina leads into a muscular bursa which may or may not be expanded (
<figureCitation id="130A2A75FFFF7A019DE225F7A33B64D0" box="[958,1084,928,953]" captionStart="FIGURES 34–37" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[220,1342,209,1859]" captionTargetId="figure@13.[220,1342,209,1859]" captionTargetPageId="13" captionText="FIGURES 34–37. Female reproductive system (B19 (2-3)). 34 system without ovaries, ovipositor to left and digesting gland to right, from above; 35 as for 34 from side expanded base of spermatheca arrowed; 36 detail bursa and expanded base of spermatheca (arrowed); 37 bursa in area of bursa hooks from above. Figure legend: B = bursa; SDG = spermatophore digesting gland; V = valvifers of the ovipositor. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457295/files/figure.png" pageId="9" pageNumber="352">Figs 36, 37</figureCitation>
show muscular striations). Just behind the bursa is the median oviduct (no median oviduct plates were detected). The bursa bears at its anterior end a thinner walled digesting gland, and towards its posterior end a stalked spermatheca which has an expanded base (
<figureCitation id="130A2A75FFFF7A019EC3225BA623634C" box="[159,292,1036,1061]" captionStart="FIGURES 34–37" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[220,1342,209,1859]" captionTargetId="figure@13.[220,1342,209,1859]" captionTargetPageId="13" captionText="FIGURES 34–37. Female reproductive system (B19 (2-3)). 34 system without ovaries, ovipositor to left and digesting gland to right, from above; 35 as for 34 from side expanded base of spermatheca arrowed; 36 detail bursa and expanded base of spermatheca (arrowed); 37 bursa in area of bursa hooks from above. Figure legend: B = bursa; SDG = spermatophore digesting gland; V = valvifers of the ovipositor. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457295/files/figure.png" pageId="9" pageNumber="352">Figs 35, 36</figureCitation>
). In some cases these may attain similar size (possibly due to immersion in water during dissection) and it may be difficult to determine which is which e.g.
<bibRefCitation id="EFA04B01FFFF7A019D632267A3FC6320" box="[831,1275,1072,1097]" pageId="9" pageNumber="352" refId="ref8493" refString="Ballantyne, L. A. &amp; Lambkin, C. (2006) A phylogenetic reassessment of the rare S. E. Asian firefly genus Pygoluciola Wittmer (Coleoptera: Lampyridae: Luciolinae). Raffles Bulletin of Zoology, 54 (1), 21 - 48." type="journal article">Ballantyne &amp; Lambkin (2006 fig. 37)</bibRefCitation>
indicated the spermatheca and digesting gland of essentially equal size. In this species a large spermatheca was observed but whether this was due to immersion in water or not is not clear. There are two long slender needle shaped bursa hooks projecting either forwards or backwards and lying free in the bursa cavity, with the point of attachment to the bursa wall at their junction (
<figureCitation id="130A2A75FFFF7A019FBE2297A53963B0" box="[482,574,1216,1241]" captionStart="FIGURES 34–37" captionStartId="13.[151,264,1896,1919]" captionTargetBox="[220,1342,209,1859]" captionTargetId="figure@13.[220,1342,209,1859]" captionTargetPageId="13" captionText="FIGURES 34–37. Female reproductive system (B19 (2-3)). 34 system without ovaries, ovipositor to left and digesting gland to right, from above; 35 as for 34 from side expanded base of spermatheca arrowed; 36 detail bursa and expanded base of spermatheca (arrowed); 37 bursa in area of bursa hooks from above. Figure legend: B = bursa; SDG = spermatophore digesting gland; V = valvifers of the ovipositor. Scale lines are 1 mm." httpUri="https://zenodo.org/record/1457295/files/figure.png" pageId="9" pageNumber="352">Figs 34</figureCitation>
̄37). Both curve slightly inwards at their apices but their exact method of functioning within the bursa, and how it relates to a spermatophore, is not entirely clear. The posterior ‘plates’ may ensure the spermatophore is held within the bursa.
</paragraph>
<caption id="DF4E6678FFFF7A019ECB2307A424620E" box="[151,803,1359,1383]" pageId="9" pageNumber="352">
<paragraph id="8B8E36F0FFFF7A019ECB2307A424620E" blockId="9.[151,803,1359,1383]" box="[151,803,1359,1383]" pageId="9" pageNumber="352">
<emphasis id="B945EAE2FFFF7A019ECB2307A60D620E" bold="true" box="[151,266,1360,1383]" pageId="9" pageNumber="352">TABLE 1.</emphasis>
Collection areas for
<taxonomicName id="4C314D73FFFF7A019FB52318A5C1620E" authority="Nada &amp; Ballantyne, 2018" authorityName="Nada &amp; Ballantyne" authorityYear="2018" box="[489,710,1359,1383]" class="Insecta" family="Lampyridae" genus="Pygolucola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="dunguna" status="sp. nov.">
<emphasis id="B945EAE2FFFF7A019FB52318A5C1620E" box="[489,710,1359,1383]" italics="true" pageId="9" pageNumber="352">Pygolucola dunguna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765799FFFF7A019C912307A424620E" box="[717,803,1360,1383]" pageId="9" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8E36F0FFFF7A019EC323D5A35D6024" pageId="9" pageNumber="352">
<table id="F931C450FFFF85F79EC323D5A3B26024" box="[159,1205,1410,1869]" gridcols="3" gridrows="12" pageId="9" pageNumber="352">
<tr id="350134B2FFFF85F79EC323D5A3B262F1" box="[159,1205,1410,1432]" gridrow="0" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC323D5A6E862F1" box="[159,495,1410,1432]" gridcol="0" gridrow="0" pageId="9" pageNumber="352">Location (Compartment no.)</th>
<th id="76D05DCEFFFF85F79D3523D5A4D762F1" box="[873,976,1410,1432]" gridcol="1" gridrow="0" pageId="9" pageNumber="352">Forest ID.</th>
<th id="76D05DCEFFFF85F79A1123D5A3B262F1" box="[1101,1205,1410,1432]" gridcol="2" gridrow="0" pageId="9" pageNumber="352">No. adults</th>
</tr>
<tr id="350134B2FFFF85F79EC323FDA3B262A9" box="[159,1205,1450,1472]" gridrow="1" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC323FDA6E862A9" box="[159,495,1450,1472]" gridcol="0" gridrow="1" pageId="9" pageNumber="352">Bukit Kutu</th>
<td id="76D05DCEFFFF85F79D3523FDA4D762A9" box="[873,976,1450,1472]" gridcol="1" gridrow="1" pageId="9" pageNumber="352">BK400</td>
<td id="76D05DCEFFFF85F79A1123FDA3B262A9" box="[1101,1205,1450,1472]" gridcol="2" gridrow="1" pageId="9" pageNumber="352">1</td>
</tr>
<tr id="350134B2FFFF85F79EC32385A3B26281" box="[159,1205,1490,1512]" gridrow="2" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32385A6E86281" box="[159,495,1490,1512]" gridcol="0" gridrow="2" pageId="9" pageNumber="352">Bukit Kutu</th>
<td id="76D05DCEFFFF85F79D352385A4D76281" box="[873,976,1490,1512]" gridcol="1" gridrow="2" pageId="9" pageNumber="352">BK600</td>
<td id="76D05DCEFFFF85F79A112385A3B26281" box="[1101,1205,1490,1512]" gridcol="2" gridrow="2" pageId="9" pageNumber="352">1</td>
</tr>
<tr id="350134B2FFFF85F79EC323AEA3B26166" box="[159,1205,1529,1551]" gridrow="3" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC323AEA6E86166" box="[159,495,1529,1551]" gridcol="0" gridrow="3" pageId="9" pageNumber="352">Gunung Liang</th>
<td id="76D05DCEFFFF85F79D3523AEA4D76166" box="[873,976,1529,1551]" gridcol="1" gridrow="3" pageId="9" pageNumber="352">GL350</td>
<td id="76D05DCEFFFF85F79A1123AEA3B26166" box="[1101,1205,1529,1551]" gridcol="2" gridrow="3" pageId="9" pageNumber="352">1</td>
</tr>
<tr id="350134B2FFFF85F79EC32076A3B2615E" box="[159,1205,1569,1591]" gridrow="4" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32076A6E8615E" box="[159,495,1569,1591]" gridcol="0" gridrow="4" pageId="9" pageNumber="352">Besul (10)</th>
<td id="76D05DCEFFFF85F79D352076A4D7615E" box="[873,976,1569,1591]" gridcol="1" gridrow="4" pageId="9" pageNumber="352">B10</td>
<td id="76D05DCEFFFF85F79A112076A3B2615E" box="[1101,1205,1569,1591]" gridcol="2" gridrow="4" pageId="9" pageNumber="352">3</td>
</tr>
<tr id="350134B2FFFF85F79EC3201EA3B26136" box="[159,1205,1609,1631]" gridrow="5" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC3201EA6E86136" box="[159,495,1609,1631]" gridcol="0" gridrow="5" pageId="9" pageNumber="352">Besul (19)</th>
<td id="76D05DCEFFFF85F79D35201EA4D76136" box="[873,976,1609,1631]" gridcol="1" gridrow="5" pageId="9" pageNumber="352">B19</td>
<td id="76D05DCEFFFF85F79A11201EA3B26136" box="[1101,1205,1609,1631]" gridcol="2" gridrow="5" pageId="9" pageNumber="352">7</td>
</tr>
<tr id="350134B2FFFF85F79EC32026A3B261EE" box="[159,1205,1649,1671]" gridrow="6" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32026A6E861EE" box="[159,495,1649,1671]" gridcol="0" gridrow="6" pageId="9" pageNumber="352">Hulu Terengganu Tambahan (27)</th>
<td id="76D05DCEFFFF85F79D352026A4D761EE" box="[873,976,1649,1671]" gridcol="1" gridrow="6" pageId="9" pageNumber="352">UTT27</td>
<td id="76D05DCEFFFF85F79A112026A3B261EE" box="[1101,1205,1649,1671]" gridcol="2" gridrow="6" pageId="9" pageNumber="352">5</td>
</tr>
<tr id="350134B2FFFF85F79EC320CFA3B261C7" box="[159,1205,1688,1710]" gridrow="7" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC320CFA6E861C7" box="[159,495,1688,1710]" gridcol="0" gridrow="7" pageId="9" pageNumber="352">Jengai (40)</th>
<td id="76D05DCEFFFF85F79D3520CFA4D761C7" box="[873,976,1688,1710]" gridcol="1" gridrow="7" pageId="9" pageNumber="352">JI40</td>
<td id="76D05DCEFFFF85F79A1120CFA3B261C7" box="[1101,1205,1688,1710]" gridcol="2" gridrow="7" pageId="9" pageNumber="352">2</td>
</tr>
<tr id="350134B2FFFF85F79EC32097A3B261BF" box="[159,1205,1728,1750]" gridrow="8" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32097A6E861BF" box="[159,495,1728,1750]" gridcol="0" gridrow="8" pageId="9" pageNumber="352">Jengai (52)</th>
<td id="76D05DCEFFFF85F79D352097A4D761BF" box="[873,976,1728,1750]" gridcol="1" gridrow="8" pageId="9" pageNumber="352">JI52</td>
<td id="76D05DCEFFFF85F79A112097A3B261BF" box="[1101,1205,1728,1750]" gridcol="2" gridrow="8" pageId="9" pageNumber="352">2</td>
</tr>
<tr id="350134B2FFFF85F79EC320BFA3B26197" box="[159,1205,1768,1790]" gridrow="9" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC320BFA6E86197" box="[159,495,1768,1790]" gridcol="0" gridrow="9" pageId="9" pageNumber="352">Jerangau (99)</th>
<td id="76D05DCEFFFF85F79D3520BFA4D76197" box="[873,976,1768,1790]" gridcol="1" gridrow="9" pageId="9" pageNumber="352">JU99</td>
<td id="76D05DCEFFFF85F79A1120BFA3B26197" box="[1101,1205,1768,1790]" gridcol="2" gridrow="9" pageId="9" pageNumber="352">5</td>
</tr>
<tr id="350134B2FFFF85F79EC32147A3B2604F" box="[159,1205,1808,1830]" gridrow="10" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32147A6E8604F" box="[159,495,1808,1830]" gridcol="0" gridrow="10" pageId="9" pageNumber="352">Jerangau (100)</th>
<td id="76D05DCEFFFF85F79D352147A4D7604F" box="[873,976,1808,1830]" gridcol="1" gridrow="10" pageId="9" pageNumber="352">JU100</td>
<td id="76D05DCEFFFF85F79A112147A3B2604F" box="[1101,1205,1808,1830]" gridcol="2" gridrow="10" pageId="9" pageNumber="352">1</td>
</tr>
<tr id="350134B2FFFF85F79EC32160A3B26024" box="[159,1205,1847,1869]" gridrow="11" pageId="9" pageNumber="352">
<th id="76D05DCEFFFF85F79EC32160A6E86024" box="[159,495,1847,1869]" gridcol="0" gridrow="11" pageId="9" pageNumber="352">Pasir Raja Barat (87)</th>
<td id="76D05DCEFFFF85F79D352160A4D76024" box="[873,976,1847,1869]" gridcol="1" gridrow="11" pageId="9" pageNumber="352">PB87</td>
<td id="76D05DCEFFFF85F79A112160A3B26024" box="[1101,1205,1847,1869]" gridcol="2" gridrow="11" pageId="9" pageNumber="352">9</td>
</tr>
</table>
</paragraph>
<caption id="DF4E6678FFFC7A029ECB21E4A27E6081" httpUri="https://zenodo.org/record/1457291/files/figure.png" pageId="10" pageNumber="353" startId="10.[151,264,1971,1994]" targetBox="[192,1371,195,1899]" targetPageId="10">
<paragraph id="8B8E36F0FFFC7A029ECB21E4A27E6081" blockId="10.[151,1436,1971,2024]" pageId="10" pageNumber="353">
<emphasis id="B945EAE2FFFC7A029ECB21E4A64860A0" bold="true" box="[151,335,1971,1994]" pageId="10" pageNumber="353">FIGURES 19–24</emphasis>
. 19‾21 Aedeagus 19 ventral; 20 dorsal; 21 left lateral (hairy lobes on LL arrowed). 22‾24 aedeagal sheath 22 ventral (position of bulbous apical lobes arrowed); 23 dorsal; 24 right lateral, hairy lobes arrowed. Scale lines are 1 mm.
</paragraph>
</caption>
<caption id="DF4E6678FFFD7A039ECB21C7A46960AF" httpUri="https://zenodo.org/record/1457293/files/figure.png" pageId="11" pageNumber="354" startId="11.[151,264,1936,1959]" targetBox="[215,1371,193,1877]" targetPageId="11">
<paragraph id="8B8E36F0FFFD7A039ECB21C7A46960AF" blockId="11.[151,1436,1936,1990]" pageId="11" pageNumber="354">
<emphasis id="B945EAE2FFFD7A039ECB21C7A65260CF" bold="true" box="[151,341,1936,1959]" pageId="11" pageNumber="354">FIGURES 25–28.</emphasis>
Ventral aspect, mating pairs, male to left (25 JI40 (8), 26 B10 (gap 5-6)). 27, 28 dissected abdomen, anterior end to top of page (B19 (2-3)) 27 ventral; 28 dorsal. Scale line is 1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36F0FFFA7A049ECB21F9A7C1608A" blockId="12.[151,1435,1966,2019]" pageId="12" pageNumber="355">
<emphasis id="B945EAE2FFFA7A049ECB21F9A64860AD" bold="true" box="[151,335,1966,1989]" pageId="12" pageNumber="355">FIGURES 29–33</emphasis>
. Females. (29, 30 PB87 (4); 31 JU100 (6); 32, 33 B19 (2-3)). 29, 31, 32 dorsal; 30 33 ventral. Scale line is
<quantity id="4CC99B15FFFA7A049BD321F8A7C5608A" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="355" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<caption id="DF4E6678FFFB7A059ECB213FA5F260B2" httpUri="https://zenodo.org/record/1457295/files/figure.png" pageId="13" pageNumber="356" startId="13.[151,264,1896,1919]" targetBox="[220,1342,209,1859]" targetPageId="13">
<paragraph id="8B8E36F0FFFB7A059ECB213FA5F260B2" blockId="13.[151,1436,1896,2011]" pageId="13" pageNumber="356">
<emphasis id="B945EAE2FFFB7A059ECB213FA6496017" bold="true" box="[151,334,1896,1919]" pageId="13" pageNumber="356">FIGURES 34–37</emphasis>
. Female reproductive system (B19 (2-3)). 34 system without ovaries, ovipositor to left and digesting gland to right, from above; 35 as for 34 from side expanded base of spermatheca arrowed; 36 detail bursa and expanded base of spermatheca (arrowed); 37 bursa in area of bursa hooks from above. Figure legend: B = bursa; SDG = spermatophore digesting gland; V = valvifers of the ovipositor. Scale lines are 1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36F0FFF87A069E9B26C0A3666629" blockId="14.[151,1437,151,320]" pageId="14" pageNumber="357">
<emphasis id="B945EAE2FFF87A069E9B26C0A6E567D9" bold="true" box="[199,482,151,176]" pageId="14" pageNumber="357">Ecological information.</emphasis>
<taxonomicName id="4C314D73FFF87A069FB126C0A5E667D9" authority="Nada &amp; Ballantyne, 2018" authorityName="Nada &amp; Ballantyne" authorityYear="2018" box="[493,737,151,176]" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="357" phylum="Arthropoda" rank="species" species="dunguna" status="sp. nov.">
<emphasis id="B945EAE2FFF87A069FB126C0A5E667D9" box="[493,737,151,176]" italics="true" pageId="14" pageNumber="357">Pygoluciola dunguna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765799FFF87A069CB026C0A44E67D9" box="[748,841,151,176]" pageId="14" pageNumber="357" rank="species">
<emphasis id="B945EAE2FFF87A069CB026C0A44E67D9" bold="true" box="[748,841,151,176]" pageId="14" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
is present in forested areas where shallow streams flow. The streams were less than
<quantity id="4CC99B15FFF87A069C5226EBA55367BD" box="[526,596,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="357" unit="cm" value="30.0">30 cm</quantity>
deep and 3̄
<quantity id="4CC99B15FFF87A069CBE26EBA42767BD" box="[738,800,188,212]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="357" unit="m" value="10.0">10 m</quantity>
wide. All the streams have small rocks and sandy edges (
<figureCitation id="130A2A75FFF87A069EC32688A7F36791" box="[159,244,223,248]" captionStart="FIGURE 38" captionStartId="14.[151,250,1364,1386]" captionTargetBox="[151,1436,375,1339]" captionTargetId="figure@14.[151,1436,375,1339]" captionTargetPageId="14" captionText="FIGURE 38. Example of a stream in the habitat of P. dunguna. These streams are shallow having small rocks and sandy edges." httpUri="https://zenodo.org/record/1457297/files/figure.png" pageId="14" pageNumber="357">Fig. 38</figureCitation>
). They were found mainly flying among the undergrowth of the forest, 10̄15 meters above ground (
<figureCitation id="130A2A75FFF87A069B292688A7B06675" captionStart="FIGURE 39" captionStartId="15.[151,250,1802,1824]" captionTargetBox="[151,1436,817,1781]" captionTargetId="figure@15.[151,1436,817,1781]" captionTargetPageId="15" captionText="FIGURE 39. The adults of P. dunguna sp. nov. were recorded flying within the undergrowth level of the forest, between 10 and 15 m above ground. Mating pairs were collected on stems and leaves of this vegetation." httpUri="https://zenodo.org/record/1457299/files/figure.png" pageId="14" pageNumber="357">Fig 39</figureCitation>
). On frequent occasions, the females were collected apparently resting on herbaceous plants. Pairs of
<taxonomicName id="4C314D73FFF87A069BDB2752A7FE6629" authority="Nada &amp; Ballantyne, 2018" authorityName="Nada &amp; Ballantyne" authorityYear="2018" class="Insecta" family="Lampyridae" genus="Pygoluciola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="357" phylum="Arthropoda" rank="species" species="dunguna" status="sp. nov.">
<emphasis id="B945EAE2FFF87A069BDB2752A7FE6629" italics="true" pageId="14" pageNumber="357">P. dunguna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765799FFF87A069F5C2770A65E6656" box="[256,345,295,320]" pageId="14" pageNumber="357" rank="species">
<emphasis id="B945EAE2FFF87A069F5C2770A6556629" bold="true" box="[256,338,295,320]" pageId="14" pageNumber="357">sp. nov</emphasis>
.
</taxonomicNameLabel>
were also collected while mating on stems or leaves of undergrowth.
</paragraph>
</subSubSection>
</treatment>
</document>
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