plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/B0/87/03B08792DB2ECD09FF6E226D8899FEEF.xml

444 lines
50 KiB
XML
Raw Normal View History

added 03
2024-06-21 12:22:17 +02:00
<document id="CE92AD9DA981320C1D1311F7949ABAD4" ID="10.11646/zootaxa.4221.3.6" ID-CLB-Dataset="37291" ID-DOI="10.5281/zenodo.250320" ID-GBIF-Dataset="d56da6d2-4950-4798-9057-d8875115930b" ID-ISSN="1175-5326" ID-Zenodo-Dep="250320" ID-ZooBank="ABD475CA-CD30-41D0-B9DF-3F6721186781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484727834738" checkinUser="plazi" docAuthor="Trýzna, Miloš" docDate="2017" docId="03B08792DB2ECD09FF6E226D8899FEEF" docLanguage="en" docName="zootaxa.4221.3.6.pdf" docOrigin="Zootaxa 4221 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tophoderes banari Tryzna, sp. nov." docType="treatment" docVersion="8" lastPageNumber="385" masterDocId="FF89FFEADB2FCD01FFF921348D05FFEA" masterDocTitle="Description of a new species of the genus Tophoderes Dejean (Coleoptera: Anthribidae) from east Madagascar, with images of all Madagascan species of the genus" masterLastPageNumber="385" masterPageNumber="377" pageNumber="378" updateTime="1698435854850" updateUser="ExternalLinkService">
<mods:mods id="AFED0918A8DBEA0CF1F6563002586823" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="09AEB12081F116884EE131D03D08523B">
<mods:title id="D72359E5686C5E1C866267EC5F23E921">Description of a new species of the genus Tophoderes Dejean (Coleoptera: Anthribidae) from east Madagascar, with images of all Madagascan species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="AA8202027B05D203419B563ECB315543" type="personal">
<mods:role id="0E9722C4318244B97F3B7AC91E059E8A">
<mods:roleTerm id="D611CF176D2145AAC1E5B428EC38052C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8811FBEEAF79C2AEA6D919EFCCEA80CD">Trýzna, Miloš</mods:namePart>
</mods:name>
<mods:typeOfResource id="9E11FC29BA1AD18CA5EDCDC8963AAF87">text</mods:typeOfResource>
<mods:relatedItem id="132AB1461F628CB90C2DD13D1A093BF5" type="host">
<mods:titleInfo id="D94E116D1B4E40115AB6BF5123FDE669">
<mods:title id="25176A8D61515B4B13914453F3D6EAC6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="251939B26A61C148A6B15198F17C71BD">
<mods:date id="A5F984C434CE14A4A15D5609E26A1970">2017</mods:date>
<mods:detail id="58B16442559E93097A9437A47A5A669A" type="volume">
<mods:number id="ECADD1CCB169EA3C57476564BB471CD9">4221</mods:number>
</mods:detail>
<mods:detail id="F3C0DF74310332E6E9751C393B3C11B7" type="issue">
<mods:number id="921E33314C70E6801D7CE4F4107E0B7A">3</mods:number>
</mods:detail>
<mods:extent id="50CA00F454E436F7A12E6456D701A1F4" unit="page">
<mods:start id="0527C52CFCCEA7E67086D1ADFC40E4CA">377</mods:start>
<mods:end id="A4ADB0C2BB3D2A777BFFCBA540E2BFEF">385</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="07E60EDA4E0946542CF1CE3729CBB447">journal article</mods:classification>
<mods:identifier id="1422EBE0CCA20B3304763FE3EC777FB5" type="CLB-Dataset">37291</mods:identifier>
<mods:identifier id="7FA6ECA973679FDCD1B6E7A33CC5BD09" type="DOI">10.5281/zenodo.250320</mods:identifier>
<mods:identifier id="683A93B98DDA44CE049FA035449A5984" type="GBIF-Dataset">d56da6d2-4950-4798-9057-d8875115930b</mods:identifier>
<mods:identifier id="404229F1AC3489C8726948ABF7EEFAF3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2AED1F349F1046DF79704578EA36169D" type="Zenodo-Dep">250320</mods:identifier>
<mods:identifier id="BC0DFEC98A33A2FED22E604EC442866F" type="ZooBank">ABD475CA-CD30-41D0-B9DF-3F6721186781</mods:identifier>
</mods:mods>
<treatment id="03B08792DB2ECD09FF6E226D8899FEEF" ID-DOI="http://doi.org/10.5281/zenodo.6051455" ID-GBIF-Taxon="127614448" ID-Zenodo-Dep="6051455" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD09FF6E226D8899FEEF" httpUri="http://treatment.plazi.org/id/03B08792DB2ECD09FF6E226D8899FEEF" lastPageId="8" lastPageNumber="385" pageId="1" pageNumber="378">
<subSubSection id="C303650FDB2ECD00FF6E226D8C84FC7E" pageId="1" pageNumber="378" type="nomenclature">
<paragraph id="8BA63684DB2ECD00FF6E226D8F3BFC99" blockId="1.[151,574,857,917]" box="[151,574,857,883]" pageId="1" pageNumber="378">
<heading id="D0EE81E8DB2ECD00FF6E226D8F3BFC99" bold="true" box="[151,574,857,883]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
<emphasis id="B96DEA96DB2ECD00FF6E226D8F3BFC99" bold="true" box="[151,574,857,883]" pageId="1" pageNumber="378">
<taxonomicName id="4C194D07DB2ECD00FF6E226D8CDCFC99" authority="Tryzna" authorityName="Tryzna" box="[151,473,857,883]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB2ECD00FF6E226D8C7CFC99" bold="true" box="[151,377,857,883]" italics="true" pageId="1" pageNumber="378">Tophoderes banari</emphasis>
Trýzna
</taxonomicName>
<taxonomicNameLabel id="A25E57EDDB2ECD00FE19226D8F3BFC99" box="[480,574,857,883]" pageId="1" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA63684DB2ECD00FF6E22488C84FC7E" blockId="1.[151,574,857,917]" box="[151,385,892,917]" pageId="1" pageNumber="378">
(
<figureCitation id="13222A01DB2ECD00FF6622488C0FFC7E" box="[159,266,892,917]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[263,1327,220,1836]" captionTargetId="figure@2.[227,1360,193,1857]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2, Tophoderes banari Trýzna sp. nov.; 1, male holotype, dorsal habitus; 2, female allotype, dorsal habitus; 3 – 4, Tophoderes annulatus Waterhouse, 1875; 3, male, 29 mm, Ankarampotsy, Tantamala; 4, female, 29 mm, Ambositra." httpUri="https://zenodo.org/record/250321/files/figure.png" pageId="1" pageNumber="378">Figs. 1–2</figureCitation>
,
<figureCitation id="13222A01DB2ECD00FEEF22488C54FC7F" box="[278,337,892,917]" captionStart-0="FIGURES 5 – 7" captionStart-1="FIGURES 8 – 10" captionStartId-0="3.[151,264,1615,1638]" captionStartId-1="4.[151,264,1472,1495]" captionTargetBox-0="[177,1398,189,1571]" captionTargetBox-1="[223,1382,191,1428]" captionTargetId-0="figure@3.[153,1433,165,1595]" captionTargetId-1="figure@4.[153,1433,169,1452]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 5 – 7. Tophoderes banari Trýzna sp. nov., male holotype, eadeagus including tegmen; 5, dorsal aspect; 6, ventral aspect; 7, lateral aspect. Scale bar in mm." captionText-1="FIGURES 8 – 10. Tophoderes banari Trýzna sp. nov.; 8, female allotype, hemisternites, ventral aspect; 9, male holotype, pygidium, dorsal aspect; 10, female allotype, pygidium, dorsal aspect. Scale bars in mm." httpUri-0="https://zenodo.org/record/250322/files/figure.png" httpUri-1="https://zenodo.org/record/250323/files/figure.png" pageId="1" pageNumber="378">5–10</figureCitation>
,
<figureCitation id="13222A01DB2ECD00FEA422488C7FFC7E" box="[349,378,892,916]" captionStart="FIGURE 23" captionStartId="7.[151,250,1434,1456]" captionText="FIGURE 23. Distribution of Tophoderes banari Trýzna sp. nov." pageId="1" pageNumber="378">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C303650FDB2ECD00FF6E22F08DD3FB87" pageId="1" pageNumber="378" type="materials_examined">
<paragraph id="8BA63684DB2ECD00FF6E22F08904FC37" blockId="1.[151,1437,964,1997]" box="[151,1025,964,989]" pageId="1" pageNumber="378">
<materialsCitation id="3B713CD9DB2ECD00FF6E22F08904FC37" ID-GBIF-Occurrence="1424788244" box="[151,1025,964,989]" country="Madagascar" elevation="1500" location="Ivohibe" pageId="1" pageNumber="378" specimenCount="1" stateProvince="Fianarantsoa province" typeStatus="holotype">
<emphasis id="B96DEA96DB2ECD00FF6E22F08C30FC37" bold="true" box="[151,309,964,989]" pageId="1" pageNumber="378">
<typeStatus id="54A28826DB2ECD00FF6E22F08DD7FC37" box="[151,210,964,989]" pageId="1" pageNumber="378">Type</typeStatus>
locality.
</emphasis>
East
<collectingCountry id="F30E7614DB2ECD00FE8A22F08CF9FC37" box="[371,508,964,989]" name="Madagascar" pageId="1" pageNumber="378">Madagascar</collectingCountry>
,
<collectingRegion id="49DDF866DB2ECD00FDFE22F08E06FC37" box="[519,771,964,989]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="378">Fianarantsoa province</collectingRegion>
,
<location id="8EC6605FDB2ECD00FCF722F08E63FC37" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD09FF6E226D8899FEEF:8EC6605FDB2ECD00FCF722F08E63FC37" box="[782,870,964,989]" country="Madagascar" name="Ivohibe" pageId="1" pageNumber="378" stateProvince="Fianarantsoa province">Ivohibe</location>
,
<elevation id="0034D1B7DB2ECD00FC8822F08904FC37" box="[881,1025,964,989]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="378" unit="m" value="1500.0">
<quantity id="4CE19B61DB2ECD00FC8822F08EC3FC36" box="[881,966,964,989]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="378" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8BA63684DB2ECD00FF3E22D38DD3FB87" blockId="1.[151,1437,964,1997]" pageId="1" pageNumber="378">
<materialsCitation id="3B713CD9DB2ECD00FF3E22D38F75FBCE" ID-GBIF-Occurrence="1424788242" collectionCode="BSNPC" collectorName="R. Decary" country="Madagascar" elevation="1500" location="Ivohibe" municipality="Madagascar" pageId="1" pageNumber="378" specimenCount="2" stateProvince="FIANARANTSOA PROVINCE" typeStatus="holotype">
<emphasis id="B96DEA96DB2ECD00FF3E22D38EF9FC15" bold="true" box="[199,1020,999,1024]" pageId="1" pageNumber="378">
<typeStatus id="54A28826DB2ECD00FF3E22D38C07FBEA" box="[199,258,999,1024]" pageId="1" pageNumber="378">Type</typeStatus>
material.
<typeStatus id="54A28826DB2ECD00FE7E22D38CF1FBEA" box="[391,500,999,1024]" pageId="1" pageNumber="378" type="holotype">Holotype</typeStatus>
(male):
<collectingCountry id="F30E7614DB2ECD00FD9A22D38E37FBEA" box="[611,818,999,1024]" name="Madagascar" pageId="1" pageNumber="378">MADAGASCAR</collectingCountry>
,
<collectingRegion id="49DDF866DB2ECD00FCBD22D38EF9FC15" box="[836,1020,999,1024]" country="Madagascar" name="Fianarantsoa" pageId="1" pageNumber="378">FIANARANTSOA</collectingRegion>
</emphasis>
PROVINCE: ‘
<collectingMunicipality id="6BC2ACFEDB2ECD00FB6022D38820FBEA" box="[1177,1317,999,1024]" pageId="1" pageNumber="378">Madagascar</collectingMunicipality>
/
<location id="8EC6605FDB2ECD00FABF22D3889EFBEA" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD09FF6E226D8899FEEF:8EC6605FDB2ECD00FABF22D3889EFBEA" box="[1350,1435,999,1024]" country="Madagascar" municipality="Madagascar" name="Ivohibe" pageId="1" pageNumber="378" stateProvince="FIANARANTSOA PROVINCE">Ivohibe</location>
(
<quantity id="4CE19B61DB2ECD00FF6625388DFDFBCE" box="[159,248,1036,1061]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="378" unit="m" value="1500.0">
<elevation id="0034D1B7DB2ECD00FF6625388DFDFBCE" box="[159,248,1036,1061]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="378" unit="m" value="1500.0">1500 m</elevation>
</quantity>
) /
<collectorName id="26EC5352DB2ECD00FEE325398C88FBCF" box="[282,397,1037,1061]" pageId="1" pageNumber="378">R. Decary</collectorName>
1926 [p] (
<collectionCode id="ED08AE41DB2ECD00FDF025388F6DFBCE" box="[521,616,1036,1060]" pageId="1" pageNumber="378">BSNPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CD9DB2ECD00FD8525388C92FBA2" ID-GBIF-Occurrence="1424788245" collectingDate="1976-01" collectionCode="MTDC" collectorName="A. Peyrieras" country="Madagascar" location="Vondrozo" pageId="1" pageNumber="378" specimenCount="1" typeStatus="allotype">
<emphasis id="B96DEA96DB2ECD00FD8525388E50FBCF" bold="true" box="[636,853,1036,1061]" pageId="1" pageNumber="378">
<typeStatus id="54A28826DB2ECD00FD8525388FE4FBCF" box="[636,737,1036,1061]" pageId="1" pageNumber="378" type="allotype">Allotype</typeStatus>
(female):
</emphasis>
‘
<collectingCountry id="F30E7614DB2ECD00FC9B25388EEAFBCF" box="[866,1007,1036,1061]" name="Madagascar" pageId="1" pageNumber="378">Madagascar</collectingCountry>
Est /
<location id="8EC6605FDB2ECD00FBCC253889A4FBCF" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD09FF6E226D8899FEEF:8EC6605FDB2ECD00FBCC253889A4FBCF" box="[1077,1185,1036,1061]" country="Madagascar" name="Vondrozo" pageId="1" pageNumber="378">Vondrozo</location>
[p] /
<date id="FFA71044DB2ECD00FB1D25398833FBCF" box="[1252,1334,1036,1061]" pageId="1" pageNumber="378" value="1976-01">
<collectingDate id="EFE3E9ACDB2ECD00FB1D25398833FBCF" box="[1252,1334,1036,1061]" pageId="1" pageNumber="378" value="1976-01">I. 1976</collectingDate>
</date>
[h] //
<collectorName id="26EC5352DB2ECD00FA7925388DFAFBA2" pageId="1" pageNumber="378">A. Peyrieras</collectorName>
[h] (
<collectionCode id="ED08AE41DB2ECD00FECE25048C8BFBA2" box="[311,398,1072,1096]" pageId="1" pageNumber="378">MTDC</collectionCode>
)
</materialsCitation>
. Red label [p] HOLOTYPE / ALLOTYPE /
<emphasis id="B96DEA96DB2ECD00FC68251B8971FBA2" box="[913,1140,1071,1096]" italics="true" pageId="1" pageNumber="378">
<taxonomicName id="4C194D07DB2ECD00FC68251B8917FBA2" box="[913,1042,1071,1096]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">Tophoderes</taxonomicName>
/
<taxonomicName id="4C194D07DB2ECD00FBD0251B8971FBA2" box="[1065,1140,1071,1096]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">banari</taxonomicName>
</emphasis>
<emphasis id="B96DEA96DB2ECD00FB85251B89D3FBA2" bold="true" box="[1148,1238,1071,1096]" pageId="1" pageNumber="378">
<taxonomicNameLabel id="A25E57EDDB2ECD00FB85251B89D3FBA2" box="[1148,1238,1071,1096]" pageId="1" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ M. Trýzna det., 2016.
</paragraph>
</subSubSection>
<subSubSection id="C303650FDB2ECD06FF3E25438FBEF9FE" lastPageId="7" lastPageNumber="384" pageId="1" pageNumber="378" type="description">
<paragraph id="8BA63684DB2ECD00FF3E25438EA2FA82" blockId="1.[151,1437,964,1997]" pageId="1" pageNumber="378">
<emphasis id="B96DEA96DB2ECD00FF3E25438C5CFB7A" bold="true" box="[199,345,1143,1168]" pageId="1" pageNumber="378">Description.</emphasis>
Male
<typeStatus id="54A28826DB2ECD00FE5125438F08FB7A" box="[424,525,1143,1168]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
, (female
<typeStatus id="54A28826DB2ECD00FD8525438FDEFB7A" box="[636,731,1143,1168]" pageId="1" pageNumber="378" type="allotype">allotype</typeStatus>
).
<emphasis id="B96DEA96DB2ECD00FD08254D8E96FB7A" box="[753,915,1145,1168]" italics="true" pageId="1" pageNumber="378">Measurements</emphasis>
(in mm): Total body length—14.78 (15.50). Head: total length—2.48 (2.60); length of rostrum—1.59 (1.59); maximum width of rostrum—1.95 (2.14); length of eye—0.98 (1.07); maximum width across eyes—2.34 (2.55); minimum distance between eyes—1.20 (1.36). Antenna: length of segments: II 0.41 (0.34), III—0.64 (0.59), IV—0.66 (0.50), V—0.52 (0.45), VI—0.45 (0.34), VII—0.43 (0.34), VIII—0.45 (0.36), IX—0.61 (0.55), X—0.45 (0.41), XI—0.52 (0.57). Pronotum: maximum length—3.50 (3.10); width at carina—5.10 (5.80); maximum width—5.10 (5.80); minimum width—2.30 (2.50). Elytra: maximum length—8.80 (9.80); maximum width—6.10 (7.00).
</paragraph>
<paragraph id="8BA63684DB2ECD00FF3E24408DDEFA45" blockId="1.[151,1437,964,1997]" pageId="1" pageNumber="378">
<emphasis id="B96DEA96DB2ECD00FF3E24408C5DFA67" bold="true" box="[199,344,1396,1421]" pageId="1" pageNumber="378">Colouration</emphasis>
of the cuticle of whole body black, only distal part of tarsomere V and claws dark brown in both sexes.
</paragraph>
<paragraph id="8BA63684DB2ECD00FF3E24888E55F8D6" blockId="1.[151,1437,964,1997]" pageId="1" pageNumber="378">
<emphasis id="B96DEA96DB2ECD00FF3E24888C3FFA3F" bold="true" box="[199,314,1468,1493]" pageId="1" pageNumber="378">Vestiture.</emphasis>
<emphasis id="B96DEA96DB2ECD00FEBA24888C85FA3F" box="[323,384,1468,1493]" italics="true" pageId="1" pageNumber="378">Head</emphasis>
predominantly covered with dense whitish appressed setae on dorsal part, this pubescence covering entire dorsal part of rostrum, on vertex formed as wide stripe as wide as interocular width.
<emphasis id="B96DEA96DB2ECD00FB0424D58869FA12" box="[1277,1388,1505,1528]" italics="true" pageId="1" pageNumber="378">Pronotum</emphasis>
and
<emphasis id="B96DEA96DB2ECD00FF6E27308DDCF9F7" box="[151,217,1540,1565]" italics="true" pageId="1" pageNumber="378">elytra</emphasis>
of marmorated appearance.
<emphasis id="B96DEA96DB2ECD00FDEC27318F81F9F6" box="[533,644,1541,1564]" italics="true" pageId="1" pageNumber="378">Pronotum</emphasis>
with two large spots of black setae distinctly bordered by dense stripes of whitish setae in its proximal part. Two black spots in anterior corners of pronotum, rest of pronotum with characteristic pattern formed of black and whitish pubescence, with indistinct tuft of very short black setae on disc. Pronotal declivity black, with narrow central stripe and two wide areas in lateral parts covered with dense whitish setae.
<emphasis id="B96DEA96DB2ECD00FF2627A08C20F947" box="[223,293,1684,1709]" italics="true" pageId="1" pageNumber="378">Elytra</emphasis>
with typical marmorated pattern, whitish pubescence comprises two indistinct transverse stripes in sub-basal and post-median part; these stripes often interrupted in elytral intervals. Particularly on second, fourth and sixth elytral interval, are several indistinct tufts of very short black setae. If such tufts are involved in sub-basal or post-median transverse stripes, they are also formed of whitish setae. Preapical tubercle indistinct, its tip covered with bright whitish setae. Scutellum with sparse whitish setae.
</paragraph>
<paragraph id="8BA63684DB2ECD02FF3E267C8DD3F8DA" blockId="1.[151,1437,964,1997]" lastBlockId="3.[151,1437,1708,2021]" lastPageId="3" lastPageNumber="380" pageId="1" pageNumber="378">
<emphasis id="B96DEA96DB2ECD00FF3E267C8C34F8B5" box="[199,305,1864,1887]" italics="true" pageId="1" pageNumber="378">Antennae</emphasis>
with indistinct sparse pubescence, mainly antennomeres VII–VIII covered by longer whitish appressed setae, antennomeres IX–XI (club) with denser, more rusty setae in both sexes.
<emphasis id="B96DEA96DB2ECD00FB7C265989BFF86E" box="[1157,1210,1901,1924]" italics="true" pageId="1" pageNumber="378">Legs</emphasis>
covered with dense setae. All femora with whitish stripe on the distal quarter and whitish tip. Tibiae covered with whitish setae appressed in proximal part and subdecumbent to suberect in distal part; wide stripe in the middle and apical part of tibiae with black setae in both sexes. Tarsomeres predominantly with black setae, only proximal part of tarsomere I and distal part of tarsomere V with whitish setae in both sexes. Venter predominantly with dense blackish or dark brownish pubescence, only lateral part of metasternum and abdominal sternites I–V with mixed sparse whitish setae.
</paragraph>
<caption id="DF66660CDB2DCD03FF6E26628861F861" httpUri="https://zenodo.org/record/250321/files/figure.png" pageId="2" pageNumber="379" targetBox="[263,1327,220,1836]" targetPageId="2">
<paragraph id="8BA63684DB2DCD03FF6E26628861F861" blockId="2.[151,1436,1878,1931]" pageId="2" pageNumber="379">
<emphasis id="B96DEA96DB2DCD03FF6E26628C3EF886" bold="true" box="[151,315,1878,1901]" pageId="2" pageNumber="379">FIGURES 1–4.</emphasis>
1–2,
<taxonomicName id="4C194D07DB2DCD03FE8F26628F86F886" authority="Tryzna" authorityName="Tryzna" box="[374,643,1878,1900]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB2DCD03FE8F26628F30F886" box="[374,565,1878,1900]" italics="true" pageId="2" pageNumber="379">Tophoderes banari</emphasis>
Trýzna
</taxonomicName>
<emphasis id="B96DEA96DB2DCD03FD7226638FD9F886" bold="true" box="[651,732,1879,1900]" pageId="2" pageNumber="379">
<taxonomicNameLabel id="A25E57EDDB2DCD03FD7226638FD9F886" box="[651,732,1879,1900]" pageId="2" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 1, male holotype, dorsal habitus; 2, female allotype, dorsal habitus; 3–4,
<taxonomicName id="4C194D07DB2DCD03FF3026418F6DF861" authority="Waterhouse, 1875" authorityName="Waterhouse" authorityYear="1875" box="[201,616,1909,1931]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B96DEA96DB2DCD03FF3026418CADF861" box="[201,424,1909,1931]" italics="true" pageId="2" pageNumber="379">Tophoderes annulatus</emphasis>
Waterhouse, 1875
</taxonomicName>
; 3, male, 29 mm, Ankarampotsy, Tantamala; 4, female, 29 mm, Ambositra.
</paragraph>
</caption>
<caption id="DF66660CDB2CCD02FF6E277B8F3EF96F" httpUri="https://zenodo.org/record/250322/files/figure.png" pageId="3" pageNumber="380" targetBox="[177,1398,189,1571]" targetPageId="3">
<paragraph id="8BA63684DB2CCD02FF6E277B8F3EF96F" blockId="3.[151,1435,1615,1669]" pageId="3" pageNumber="380">
<emphasis id="B96DEA96DB2CCD02FF6E277B8C39F98F" bold="true" box="[151,316,1615,1638]" pageId="3" pageNumber="380">FIGURES 5–7.</emphasis>
<taxonomicName id="4C194D07DB2CCD02FEBC27648F50F98C" authority="Tryzna" authorityName="Tryzna" box="[325,597,1616,1638]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB2CCD02FEBC27648F03F98C" box="[325,518,1616,1638]" italics="true" pageId="3" pageNumber="380">Tophoderes banari</emphasis>
Trýzna
</taxonomicName>
<emphasis id="B96DEA96DB2CCD02FDA427648FB2F98F" bold="true" box="[605,695,1616,1637]" pageId="3" pageNumber="380">
<taxonomicNameLabel id="A25E57EDDB2CCD02FDA427648FB7F98F" box="[605,690,1616,1637]" pageId="3" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype, eadeagus including tegmen; 5, dorsal aspect; 6, ventral aspect; 7, lateral aspect. Scale bar in mm.
</paragraph>
</caption>
<paragraph id="8BA63684DB2CCD02FF3E260889CBF8BE" blockId="3.[151,1437,1708,2021]" box="[199,1230,1852,1877]" pageId="3" pageNumber="380">
Distal part of prosternum without pair of spots.
<emphasis id="B96DEA96DB2CCD02FD2426088E4DF8BF" box="[733,840,1852,1877]" italics="true" pageId="3" pageNumber="380">Pygidium</emphasis>
covered with sparse whitish setae.
</paragraph>
<paragraph id="8BA63684DB2CCD02FF3E26548990F80F" blockId="3.[151,1437,1708,2021]" pageId="3" pageNumber="380">
<emphasis id="B96DEA96DB2CCD02FF3E26548C3AF893" bold="true" box="[199,319,1888,1913]" pageId="3" pageNumber="380">Structure.</emphasis>
<emphasis id="B96DEA96DB2CCD02FEBE26548C81F893" box="[327,388,1888,1913]" italics="true" pageId="3" pageNumber="380">Head</emphasis>
relatively short, rostrum weakly extended apically, dorsum convex, not depressed in middle, with one fine longitudinal carina reaching between eyes in both sexes. Ratio of rostrum length to maximum width
<quantity id="4CE19B61DB2CCD02FF6E269D8DE1F82B" box="[151,228,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="2.0827999999999998" pageId="3" pageNumber="380" unit="in" value="0.82">0.82 in</quantity>
male,
<quantity id="4CE19B61DB2CCD02FED7269D8C79F82B" box="[302,380,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.8796" pageId="3" pageNumber="380" unit="in" value="0.74">0.74 in</quantity>
female. Eyes spherical, not emarginate, dorsal ocular index
<quantity id="4CE19B61DB2CCD02FBE2269D896DF82B" box="[1051,1128,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.3594" pageId="3" pageNumber="380" unit="in" value="2.11">2.11 in</quantity>
male,
<quantity id="4CE19B61DB2CCD02FB4B269D8805F82B" box="[1202,1280,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.8166" pageId="3" pageNumber="380" unit="in" value="2.29">2.29 in</quantity>
female. Ratio of maximum width across eyes to maximum width of rostrum
<quantity id="4CE19B61DB2CCD02FCAA26F98EA4F80F" box="[851,929,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="3" pageNumber="380" unit="in" value="1.2">1.20 in</quantity>
male,
<quantity id="4CE19B61DB2CCD02FC1526F9893FF80F" box="[1004,1082,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="3.0225999999999997" pageId="3" pageNumber="380" unit="in" value="1.19">1.19 in</quantity>
female.
</paragraph>
<caption id="DF66660CDB2BCD05FF6E24F4891CFA1C" httpUri="https://zenodo.org/record/250323/files/figure.png" pageId="4" pageNumber="381" targetBox="[223,1382,191,1428]" targetPageId="4">
<paragraph id="8BA63684DB2BCD05FF6E24F4891CFA1C" blockId="4.[151,1436,1472,1526]" pageId="4" pageNumber="381">
<emphasis id="B96DEA96DB2BCD05FF6E24F48C4EFA3C" bold="true" box="[151,331,1472,1495]" pageId="4" pageNumber="381">FIGURES 8–10.</emphasis>
<taxonomicName id="4C194D07DB2BCD05FEAC24F58F6CFA3D" authority="Tryzna" authorityName="Tryzna" box="[341,617,1473,1495]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB2BCD05FEAC24F58F1DFA3D" box="[341,536,1473,1495]" italics="true" pageId="4" pageNumber="381">Tophoderes banari</emphasis>
Trýzna
</taxonomicName>
<emphasis id="B96DEA96DB2BCD05FD8A24F58FD5FA3C" bold="true" box="[627,720,1473,1494]" pageId="4" pageNumber="381">
<taxonomicNameLabel id="A25E57EDDB2BCD05FD8A24F58FC9FA3C" box="[627,716,1473,1494]" pageId="4" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
;
</emphasis>
8, female allotype, hemisternites, ventral aspect; 9, male holotype, pygidium, dorsal aspect; 10, female allotype, pygidium, dorsal aspect. Scale bars in mm.
</paragraph>
</caption>
<paragraph id="8BA63684DB2BCD05FF3E272A8E4DF80C" blockId="4.[151,1437,1565,2022]" pageId="4" pageNumber="381">
Antennae barely reaching to posterior margin of pronotum in male, females antennae noticeably shorter. Funicle thick, club moderately robust, antennomeres IX and XI as long as wide, X wider than long in both sexes.
<emphasis id="B96DEA96DB2BCD05FF6E27528C03F997" box="[151,262,1638,1661]" italics="true" pageId="4" pageNumber="381">Pronotum</emphasis>
transverse, bell-shaped, widest in proximal part, then gradually narrowing anteriorly. Ratio of its length to width at carina
<quantity id="4CE19B61DB2BCD05FE9A27BD8CB4F948" box="[355,433,1673,1698]" metricMagnitude="-2" metricUnit="m" metricValue="1.7526" pageId="4" pageNumber="381" unit="in" value="0.69">0.69 in</quantity>
male,
<quantity id="4CE19B61DB2BCD05FE0427BD8F49F948" box="[509,588,1673,1698]" metricMagnitude="-2" metricUnit="m" metricValue="1.3462" pageId="4" pageNumber="381" unit="in" value="0.53">0.53 in</quantity>
female. Lateral side rounded, without spine in both sexes. Disc moderately convex, without tubercles. Dorsal transverse carina bisinuate, concave in middle, not interrupted. Lateral carina of pronotum well-developed, long, nearly reaching anterior margin of pronotum, in contact with dorsal transverse carina acute-angled.
<emphasis id="B96DEA96DB2BCD05FE7F27C18CC9F8E4" box="[390,460,1781,1806]" italics="true" pageId="4" pageNumber="381">Elytra</emphasis>
broadly suboval, ratio of the maximum length to maximum width of elytra
<quantity id="4CE19B61DB2BCD05FAB227C28899F8E4" box="[1355,1436,1781,1806]" metricMagnitude="-2" metricUnit="m" metricValue="3.6576" pageId="4" pageNumber="381" unit="in" value="1.44">1.44 in</quantity>
male,
<quantity id="4CE19B61DB2BCD05FF1D262D8C3FF8D8" box="[228,314,1817,1842]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="4" pageNumber="381" unit="in" value="1.4">1.40 in</quantity>
female. Sub-basal tubercles indistinct.
<emphasis id="B96DEA96DB2BCD05FCE2262D8E80F8D8" box="[795,901,1817,1842]" italics="true" pageId="4" pageNumber="381">Abdomen</emphasis>
slightly transverse in both sexes.
<emphasis id="B96DEA96DB2BCD05FAC8262D8899F8D8" box="[1329,1436,1817,1842]" italics="true" pageId="4" pageNumber="381">Pygidium</emphasis>
transverse, ratio of its maximum length to maximum width
<quantity id="4CE19B61DB2BCD05FCBA26098E96F8BC" box="[835,915,1853,1878]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="4" pageNumber="381" unit="in" value="0.75">0.75 in</quantity>
male (
<figureCitation id="13222A01DB2BCD05FC1D2609892FF8BC" box="[996,1066,1853,1878]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1472,1495]" captionTargetBox="[223,1382,191,1428]" captionTargetId="figure@4.[153,1433,169,1452]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Tophoderes banari Trýzna sp. nov.; 8, female allotype, hemisternites, ventral aspect; 9, male holotype, pygidium, dorsal aspect; 10, female allotype, pygidium, dorsal aspect. Scale bars in mm." httpUri="https://zenodo.org/record/250323/files/figure.png" pageId="4" pageNumber="381">Fig. 9</figureCitation>
),
<quantity id="4CE19B61DB2BCD05FBB8260A8994F8BC" box="[1089,1169,1853,1878]" metricMagnitude="-2" metricUnit="m" metricValue="1.8288" pageId="4" pageNumber="381" unit="in" value="0.72">0.72 in</quantity>
female (
<figureCitation id="13222A01DB2BCD05FB0E26098849F8BC" box="[1271,1356,1853,1878]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1472,1495]" captionTargetBox="[223,1382,191,1428]" captionTargetId="figure@4.[153,1433,169,1452]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Tophoderes banari Trýzna sp. nov.; 8, female allotype, hemisternites, ventral aspect; 9, male holotype, pygidium, dorsal aspect; 10, female allotype, pygidium, dorsal aspect. Scale bars in mm." httpUri="https://zenodo.org/record/250323/files/figure.png" pageId="4" pageNumber="381">Fig. 10</figureCitation>
).
<emphasis id="B96DEA96DB2BCD05FA9B26098C07F890" italics="true" pageId="4" pageNumber="381">Male genitalia.</emphasis>
Aedeagus robust (
<figureCitation id="13222A01DB2BCD05FE2E26558F41F890" box="[471,580,1889,1914]" captionStart="FIGURES 5 – 7" captionStartId="3.[151,264,1615,1638]" captionTargetBox="[177,1398,189,1571]" captionTargetId="figure@3.[153,1433,165,1595]" captionTargetPageId="3" captionText="FIGURES 5 – 7. Tophoderes banari Trýzna sp. nov., male holotype, eadeagus including tegmen; 5, dorsal aspect; 6, ventral aspect; 7, lateral aspect. Scale bar in mm." httpUri="https://zenodo.org/record/250322/files/figure.png" pageId="4" pageNumber="381">Figs. 5–7</figureCitation>
), pedon and tectum wide, both in distal part pointed, apodemes moderately curved. Tegmen robust, its apex wide rounded, without visible setae.
<emphasis id="B96DEA96DB2BCD05FC4626B1898DF874" box="[959,1160,1925,1950]" italics="true" pageId="4" pageNumber="381">Female genitalia.</emphasis>
Hemisternites long and robust (
<figureCitation id="13222A01DB2BCD05FF17269D8C31F82B" box="[238,308,1961,1986]" captionStart="FIGURES 8 – 10" captionStartId="4.[151,264,1472,1495]" captionTargetBox="[223,1382,191,1428]" captionTargetId="figure@4.[153,1433,169,1452]" captionTargetPageId="4" captionText="FIGURES 8 – 10. Tophoderes banari Trýzna sp. nov.; 8, female allotype, hemisternites, ventral aspect; 9, male holotype, pygidium, dorsal aspect; 10, female allotype, pygidium, dorsal aspect. Scale bars in mm." httpUri="https://zenodo.org/record/250323/files/figure.png" pageId="4" pageNumber="381">Fig. 8</figureCitation>
), lateral and median rods very long, median rod reaching to three quarter of length of lateral rod. Toothed plate well developed, apical stylus present but small.
</paragraph>
<caption id="DF66660CDB2ACD04FF6E264E8EAAF807" httpUri="https://zenodo.org/record/250324/files/figure.png" pageId="5" pageNumber="382" targetBox="[158,1403,231,1845]" targetPageId="5">
<paragraph id="8BA63684DB2ACD04FF6E264E8EAAF807" blockId="5.[151,1437,1914,2029]" pageId="5" pageNumber="382">
<emphasis id="B96DEA96DB2ACD04FF6E264E8C7DF87A" bold="true" box="[151,376,1914,1937]" pageId="5" pageNumber="382">FIGURES 11–16. 11,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FE79264F8FDEF87B" authority="(Klug, 1833)" authorityName="Klug" authorityYear="1833" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[384,731,1915,1937]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="frenatus">
<emphasis id="B96DEA96DB2ACD04FE79264F8F4BF87B" box="[384,590,1915,1937]" italics="true" pageId="5" pageNumber="382">Tophoderes frenatus</emphasis>
(Klug, 1833)
</taxonomicName>
, 29 mm, Andasibe-Mantadia Nat. Park, Analamazaotra forest;
<emphasis id="B96DEA96DB2ACD04FA9B264F8887F87A" bold="true" box="[1378,1410,1915,1936]" pageId="5" pageNumber="382">12,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FA7026488CDDF85A" authority="Tryzna &amp; Banar, 2015" authorityName="Tryzna &amp; Banar" authorityYear="2015" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="lidmilae">
<emphasis id="B96DEA96DB2ACD04FA7026488DEFF85A" italics="true" pageId="5" pageNumber="382">T. lidmilae</emphasis>
Trýzna &amp; Baňař, 2015
</taxonomicName>
, 27 mm, Montagne d´Ambre Nat. Park, Ambohitra env.;
<emphasis id="B96DEA96DB2ACD04FBDA26AE8947F845" bold="true" box="[1059,1090,1946,1967]" pageId="5" pageNumber="382">13,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FBB026AF8846F85A" authority="(Klug, 1833)" authorityName="Klug" authorityYear="1833" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[1097,1347,1946,1968]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="B96DEA96DB2ACD04FBB026AF89B3F85A" box="[1097,1206,1946,1968]" italics="true" pageId="5" pageNumber="382">T. funebris</emphasis>
(Klug, 1833)
</taxonomicName>
, 17 mm, Tamatave, forêts d´Alahakato;
<emphasis id="B96DEA96DB2ACD04FE29268C8CEAF827" bold="true" box="[464,495,1976,1997]" pageId="5" pageNumber="382">14,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FE0E268D8E1AF824" authority="Wolfrum, 1959" authorityName="Wolfrum" authorityYear="1959" box="[503,799,1976,1998]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="acarinulus">
<emphasis id="B96DEA96DB2ACD04FE0E268D8F78F824" box="[503,637,1976,1998]" italics="true" pageId="5" pageNumber="382">T. acarinulus</emphasis>
Wolfrum, 1959
</taxonomicName>
, 17 mm, Maroantsetra;
<emphasis id="B96DEA96DB2ACD04FBE8268C8935F827" bold="true" box="[1041,1072,1976,1997]" pageId="5" pageNumber="382">15,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FBCE268D887DF824" authority="Fairmaire, 1888" authorityName="Fairmaire" authorityYear="1888" box="[1079,1400,1976,1998]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="nubeculosus">
<emphasis id="B96DEA96DB2ACD04FBCE268D89C8F824" box="[1079,1229,1976,1998]" italics="true" pageId="5" pageNumber="382">T. nubeculosus</emphasis>
Fairmaire, 1888
</taxonomicName>
, 23 mm, Baie d´Antongil;
<emphasis id="B96DEA96DB2ACD04FE8526E38C9EF806" bold="true" box="[380,411,2007,2028]" pageId="5" pageNumber="382">16,</emphasis>
<taxonomicName id="4C194D07DB2ACD04FE5B26EC8FB5F807" authority="(Klug, 1833)" authorityName="Klug" authorityYear="1833" baseAuthorityName="Klug" baseAuthorityYear="1833" box="[418,688,2007,2029]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="ferrugatus">
<emphasis id="B96DEA96DB2ACD04FE5B26EC8F20F807" box="[418,549,2007,2029]" italics="true" pageId="5" pageNumber="382">T. ferrugatus</emphasis>
(Klug, 1833)
</taxonomicName>
, 23 mm, Tananarivo env.
</paragraph>
</caption>
<caption id="DF66660CDB29CD07FF6E26068CFEF84E" httpUri="https://zenodo.org/record/250325/files/figure.png" pageId="6" pageNumber="383" targetBox="[168,1417,213,1777]" targetPageId="6">
<paragraph id="8BA63684DB29CD07FF6E26068CFEF84E" blockId="6.[151,1437,1842,1956]" pageId="6" pageNumber="383">
<emphasis id="B96DEA96DB29CD07FF6E26068C7EF8A2" bold="true" box="[151,379,1842,1865]" pageId="6" pageNumber="383">FIGURES 17–22. 17,</emphasis>
<taxonomicName id="4C194D07DB29CD07FE7D26078E2EF8A3" authority="Fairmaire, 1901" authorityName="Fairmaire" authorityYear="1901" box="[388,811,1843,1865]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="griseovarius">
<emphasis id="B96DEA96DB29CD07FE7D26078CFDF8A3" box="[388,504,1843,1865]" italics="true" pageId="6" pageNumber="383">Tophoderes</emphasis>
<emphasis id="B96DEA96DB29CD07FE0626008F7BF8A3" box="[511,638,1844,1865]" italics="true" pageId="6" pageNumber="383">griseovarius</emphasis>
Fairmaire, 1901
</taxonomicName>
, 18 mm, Madagascar Sud, vallée d´Ambolo;
<emphasis id="B96DEA96DB29CD07FB062607881BF8A2" bold="true" box="[1279,1310,1843,1864]" pageId="6" pageNumber="383">18,</emphasis>
<taxonomicName id="4C194D07DB29CD07FADF26008C3FF88D" authority="Fairmaire, 1901" authorityName="Fairmaire" authorityYear="1901" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="griseipes">
<emphasis id="B96DEA96DB29CD07FADF26008899F8A3" box="[1318,1436,1844,1865]" italics="true" pageId="6" pageNumber="383">T. griseipes</emphasis>
Fairmaire, 1901
</taxonomicName>
, 24 mm, Anjiro;
<emphasis id="B96DEA96DB29CD07FE1E26658F03F88C" bold="true" box="[487,518,1873,1894]" pageId="6" pageNumber="383">19,</emphasis>
<taxonomicName id="4C194D07DB29CD07FDF526668E14F88D" authority="Alluaud, 1899" authorityName="Alluaud" authorityYear="1899" box="[524,785,1873,1895]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="murinus">
<emphasis id="B96DEA96DB29CD07FDF526668F7DF88D" box="[524,632,1874,1895]" italics="true" pageId="6" pageNumber="383">T. murinus</emphasis>
Alluaud, 1899
</taxonomicName>
, 28 mm, Chaines Anosyennes, Massif nord, Ranomandry;
<emphasis id="B96DEA96DB29CD07FA9D26658886F88C" bold="true" box="[1380,1411,1873,1894]" pageId="6" pageNumber="383">20,</emphasis>
<taxonomicName id="4C194D07DB29CD07FA7026668C70F86C" authority="Jordan, 1895" authorityName="Jordan" authorityYear="1895" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="sikorae">
<emphasis id="B96DEA96DB29CD07FA7026668DE7F86C" italics="true" pageId="6" pageNumber="383">T. sikorae</emphasis>
Jordan, 1895
</taxonomicName>
, 14 mm, Moramanga;
<emphasis id="B96DEA96DB29CD07FD9226448F8FF86F" bold="true" box="[619,650,1904,1925]" pageId="6" pageNumber="383">21,</emphasis>
<taxonomicName id="4C194D07DB29CD07FD6C26458ED8F86C" authority="Wolfrum, 1959" authorityName="Wolfrum" authorityYear="1959" box="[661,989,1904,1926]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="fuscoareatus">
<emphasis id="B96DEA96DB29CD07FD6C26458E36F86C" box="[661,819,1904,1926]" italics="true" pageId="6" pageNumber="383">T. fuscoareatus</emphasis>
Wolfrum, 1959
</taxonomicName>
, 13 mm, Moramanga;
<emphasis id="B96DEA96DB29CD07FB2D264489F6F86F" bold="true" box="[1236,1267,1904,1925]" pageId="6" pageNumber="383">22,</emphasis>
<taxonomicName id="4C194D07DB29CD07FB0426458C19F84E" authority="Jordan, 1895" authorityName="Jordan" authorityYear="1895" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="sinuatocollis">
<emphasis id="B96DEA96DB29CD07FB0426458899F86C" box="[1277,1436,1904,1926]" italics="true" pageId="6" pageNumber="383">T. sinuatocollis</emphasis>
Jordan, 1895
</taxonomicName>
, 16 mm, Moramanga.
</paragraph>
</caption>
<caption id="DF66660CDB28CD06FF6E24AE8E21FA45" box="[151,804,1434,1456]" pageId="7" pageNumber="384">
<paragraph id="8BA63684DB28CD06FF6E24AE8E21FA45" blockId="7.[151,804,1434,1456]" box="[151,804,1434,1456]" pageId="7" pageNumber="384">
<emphasis id="B96DEA96DB28CD06FF6E24AE8C25FA45" bold="true" box="[151,288,1434,1456]" pageId="7" pageNumber="384">FIGURE 23.</emphasis>
Distribution of
<taxonomicName id="4C194D07DB28CD06FE3B24AE8FC8FA5A" authority="Tryzna" authorityName="Tryzna" box="[450,717,1434,1456]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB28CD06FE3B24AE8F33FA5A" box="[450,566,1434,1456]" italics="true" pageId="7" pageNumber="384">Tophoderes</emphasis>
<emphasis id="B96DEA96DB28CD06FDC524AE8F85FA5A" box="[572,640,1434,1456]" italics="true" pageId="7" pageNumber="384">banari</emphasis>
Trýzna
</taxonomicName>
<taxonomicNameLabel id="A25E57EDDB28CD06FD2A24AE8E21FA45" box="[723,804,1434,1455]" pageId="7" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA63684DB28CD06FF3E24E38FBEF9FE" blockId="7.[151,1437,1495,1880]" pageId="7" pageNumber="384">
<emphasis id="B96DEA96DB28CD06FF3E24E38CBAFA1A" bold="true" box="[199,447,1495,1520]" pageId="7" pageNumber="384">Sexual dimorphism.</emphasis>
Male antennae moderately longer, reaching approximately to posterior margin of pronotum, abdominal ventrites slightly flattened.
</paragraph>
</subSubSection>
<subSubSection id="C303650FDB28CD06FF3E272B8CCCF9B6" pageId="7" pageNumber="384" type="etymology">
<paragraph id="8BA63684DB28CD06FF3E272B8CCCF9B6" blockId="7.[151,1437,1495,1880]" pageId="7" pageNumber="384">
<emphasis id="B96DEA96DB28CD06FF3E272B8C4BF9D2" bold="true" box="[199,334,1567,1592]" pageId="7" pageNumber="384">Etymology.</emphasis>
Patronym, dedicated to my friend Petr Baňař, well-known heteropterist and member of many expeditions to
<collectingCountry id="F30E7614DB28CD06FEC527778CC3F9B6" box="[316,454,1603,1628]" name="Madagascar" pageId="7" pageNumber="384">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C303650FDB28CD06FF3E27538C95F8E5" pageId="7" pageNumber="384" type="distribution">
<paragraph id="8BA63684DB28CD06FF3E27538C95F8E5" blockId="7.[151,1437,1495,1880]" pageId="7" pageNumber="384">
<emphasis id="B96DEA96DB28CD06FF3E27538C65F96A" bold="true" box="[199,352,1639,1664]" pageId="7" pageNumber="384">Distribution.</emphasis>
Both known specimens were collected in east
<collectingCountry id="F30E7614DB28CD06FC9427538EF3F96A" box="[877,1014,1639,1664]" name="Madagascar" pageId="7" pageNumber="384">Madagascar</collectingCountry>
,
<collectingRegion id="49DDF866DB28CD06FBF8275389FBF96A" box="[1025,1278,1639,1664]" country="Madagascar" name="Fianarantsoa" pageId="7" pageNumber="384">Fianarantsoa province</collectingRegion>
. The
<typeStatus id="54A28826DB28CD06FAC327538899F96A" box="[1338,1436,1639,1664]" pageId="7" pageNumber="384" type="holotype">holotype</typeStatus>
comes from “Ivohibe,
<quantity id="4CE19B61DB28CD06FE6827BF8CECF94E" box="[401,489,1675,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="384" unit="m" value="1500.0">1500 m</quantity>
”. All of the remaining tropical humid rainforests and dense mountain forests here are extremely significant, because these habitats support animal and plant species with a high rate of endemism. The
<typeStatus id="54A28826DB28CD06FF6E27E78DF4F906" box="[151,241,1747,1772]" pageId="7" pageNumber="384" type="allotype">allotype</typeStatus>
comes from ca
<quantity id="4CE19B61DB28CD06FE5527E78F1FF906" box="[428,538,1747,1772]" metricMagnitude="4" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="7" pageNumber="384" unit="km" value="65.0" valueMax="70.0" valueMin="60.0">60-70 km</quantity>
distant “Vondrozo”, also known as “Vondrozo forest”, which has an elevation from ca
<quantity id="4CE19B61DB28CD06FF0D27C38C89F8E5" box="[244,396,1783,1808]" metricMagnitude="2" metricUnit="m" metricValue="4.7" metricValueMax="6.7" metricValueMin="2.7" pageId="7" pageNumber="384" unit="m" value="470.0" valueMax="670.0" valueMin="270.0">
270 to
<specimenCount id="9D1FFD0DDB28CD06FEBB27C38C89F8E5" box="[322,396,1783,1808]" pageId="7" pageNumber="384" type="generic">670 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C303650FDB28CD09FF3E262F8899FEEF" lastPageId="8" lastPageNumber="385" pageId="7" pageNumber="384" type="diagnosis">
<paragraph id="8BA63684DB28CD06FF3E262F8959F8B2" blockId="7.[151,1437,1495,1880]" pageId="7" pageNumber="384">
<emphasis id="B96DEA96DB28CD06FF3E262F8CD4F8DE" bold="true" box="[199,465,1819,1844]" pageId="7" pageNumber="384">Differential diagnosis.</emphasis>
<taxonomicName id="4C194D07DB28CD06FE22262F8FB5F8DE" box="[475,688,1819,1844]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB28CD06FE22262F8FB5F8DE" box="[475,688,1819,1844]" italics="true" pageId="7" pageNumber="384">Tophoderes banari</emphasis>
</taxonomicName>
<emphasis id="B96DEA96DB28CD06FD43262F8E12F8DE" bold="true" box="[698,791,1819,1844]" pageId="7" pageNumber="384">
<taxonomicNameLabel id="A25E57EDDB28CD06FD43262F8E12F8DE" box="[698,791,1819,1844]" pageId="7" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C194D07DB28CD06FC62262889FAF8DE" authority="Waterhouse, 1875" authorityName="Waterhouse" authorityYear="1875" box="[923,1279,1819,1844]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B96DEA96DB28CD06FC622628892DF8DE" box="[923,1064,1819,1844]" italics="true" pageId="7" pageNumber="384">T. annulatus</emphasis>
Waterhouse,
<quantity id="4CE19B61DB28CD06FB3E262F89FAF8DE" box="[1223,1279,1819,1844]" metricMagnitude="1" metricUnit="m" metricValue="4.7625" pageId="7" pageNumber="384" unit="in" value="1875.0">1875</quantity>
</taxonomicName>
in its similar colour pattern. However, it clearly differs in the characters given in the following key:
</paragraph>
<footnote id="E8022A8ADB28CD06FF6E26B28893F81E" pageId="7" pageNumber="384">
<paragraph id="8BA63684DB28CD06FF6E26B28893F81E" blockId="7.[151,1435,1926,2036]" pageId="7" pageNumber="384">
1. Lateral side of pronotum rounded, without spine in both sexes. Disc of pronotum moderately convex, without tubercles. Pronotum bell-shaped, widest in proximal part. Dorsal transverse carina of pronotum bisinuate, concave in the middle, not interrupted. Sub-basal elytral tubercles indistinct, flattened. Rostrum relatively short, weakly extended apically. Antennomere IX as wide as long, X wider than long in both sexes. Smaller species, ca 15 mm............
<taxonomicName id="4C194D07DB28CD06FBC526EB8846F81E" authority="Tryzna" authorityName="Tryzna" box="[1084,1347,2015,2036]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
<emphasis id="B96DEA96DB28CD06FBC526EB89F1F81E" bold="true" box="[1084,1268,2015,2036]" italics="true" pageId="7" pageNumber="384">Tophoderes banari</emphasis>
Trýzna
</taxonomicName>
<taxonomicNameLabel id="A25E57EDDB28CD06FAB026EB8893F81E" box="[1353,1430,2015,2036]" pageId="7" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</footnote>
<paragraph id="8BA63684DB27CD09FF6E21A38899FEEF" blockId="8.[151,1436,151,261]" pageId="8" pageNumber="385">
- Lateral side of pronotum with distinct spine in both sexes. Disc of pronotum with three tubercles. Pronotum hexagonal, widest in central part. Dorsal transverse carina of pronotum trisinuate, interrupted in middle. Sub-basal elytral tubercles distinct, protruded. Rostrum relatively long, strongly extended apically. Antennomere IX longer than wide, X as wide as long in both sexes. Large-sized species, ca
<quantity id="4CE19B61DB27CD09FDFD21DB8F6EFEEE" box="[516,619,239,261]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="8" pageNumber="385" unit="mm" value="25.0" valueMax="26.0" valueMin="24.0">24–26 mm</quantity>
..........................................
<taxonomicName id="4C194D07DB27CD09FB9421DB8899FEEF" authority="Waterhouse, 1875" authorityName="Waterhouse" authorityYear="1875" box="[1133,1436,239,261]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B96DEA96DB27CD09FB9421DB89ECFEEE" bold="true" box="[1133,1257,239,260]" italics="true" pageId="8" pageNumber="385">T. annulatus</emphasis>
Waterhouse, 1875
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue Copy permalink