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<mods:title id="FCFA920A6D6B4A092DE303CBAEFBEF31">A revision of the Chilean Brachyglutini - Part 7. Revision of Achilia Reitter, 1890: A. cosmoptera species group (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:affiliation id="AE116F5E16E175B508FE610327C6385A">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail: pselaphidae @ yandex. ru</mods:affiliation>
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<emphasis id="E5FC3B6CFFD5FF94E692FE2A869F59B0" bold="true" box="[950,1157,419,443]" italics="true" pageId="5" pageNumber="139">Achilia blanchardi</emphasis>
<bibRefCitation id="B3199A8FFFD5FF94E1A8FE2A873F59B0" author="Raffray A." box="[1164,1317,419,443]" pageId="5" pageNumber="139" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904</bibRefCitation>
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<paragraph id="D737E77EFFD5FF94E6F1FE4A871D59D0" blockId="5.[950,1317,419,476]" box="[981,1287,451,476]" pageId="5" pageNumber="139">
<figureCitation id="4FB3FBFBFFD5FF94E6F1FE4A860359D7" box="[981,1049,451,476]" captionStart="Figs 1-6" captionStartId="8.[153,194,2054,2076]" captionTargetBox="[177,1406,898,2017]" captionTargetId="figure-286@8.[149,1439,882,2039]" captionText="Figs 1-6. Aedeagi of Achilia. (1) A. cosmoptera. (2) A. covidia. (3) A. blanchardi. (4) A. elfridae. (5) A. bifrons. (6) A. temporalis." figureDoi="http://doi.org/10.5281/zenodo.5645110" httpUri="https://zenodo.org/record/5645110/files/figure.png" pageId="5" pageNumber="139">Figs 3</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E100FE4D865B59D7" box="[1060,1089,452,476]" captionStart="Figs 16-22" captionStartId="10.[151,192,2026,2048]" captionTargetBox="[179,1415,984,1976]" captionTargetId="figure-479@10.[153,1438,959,1994]" captionTargetPageId="10" captionText="Figs 16-22. Male antennae of Achilia. (16) A. covidia. (17) A. blanchardi. (18) A. bifrons. (19) A. temporalis. (20) A. pandemica. (21) A. angularis. (22) A. quarantena." figureDoi="http://doi.org/10.5281/zenodo.5639843" httpUri="https://zenodo.org/record/5639843/files/figure.png" pageId="5" pageNumber="139">17</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E168FE4D867359D7" box="[1100,1129,452,476]" captionStart="Figs 23-32" captionStartId="11.[151,192,2024,2046]" captionTargetBox="[188,1424,1432,1985]" captionTargetId="figure-589@11.[151,1436,1428,1996]" captionText="Figs 23-32. Male protibiae (23-25), mesotibiae (26-31) and metatibiae (32) of Achilia. (23, 28) A. elfridae. (24, 30, 32) A. angularis. (25, 31) A. quarantena. (26) A. covidia. (27) A. blanchardi. (29) A. pandemica." figureDoi="http://doi.org/10.5281/zenodo.5645115" httpUri="https://zenodo.org/record/5645115/files/figure.png" pageId="5" pageNumber="139">27</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E150FE4D868B59D7" box="[1140,1169,452,476]" captionStart="Figs 37-42" captionStartId="13.[151,192,2011,2033]" captionTargetBox="[154,1436,257,1939]" captionTargetId="figure-51@13.[151,1437,257,1958]" captionTargetPageId="13" captionText="Figs 37-42. (37, 39, 41) Achilia bifrons. (38, 40, 42) A. blanchardi. Male head in (37-38) dorsal, (39-40) lateral and (41-42) frontal views. Scale bars (200 μm) left for (37, 39, 41) and right for (38, 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5639847" httpUri="https://zenodo.org/record/5639847/files/figure.png" pageId="5" pageNumber="139">38</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E1B8FE4D86A359D7" box="[1180,1209,452,476]" captionStart="Figs 37-42" captionStartId="13.[151,192,2011,2033]" captionTargetBox="[154,1436,257,1939]" captionTargetId="figure-51@13.[151,1437,257,1958]" captionTargetPageId="13" captionText="Figs 37-42. (37, 39, 41) Achilia bifrons. (38, 40, 42) A. blanchardi. Male head in (37-38) dorsal, (39-40) lateral and (41-42) frontal views. Scale bars (200 μm) left for (37, 39, 41) and right for (38, 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5639847" httpUri="https://zenodo.org/record/5639847/files/figure.png" pageId="5" pageNumber="139">40</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E1E0FE4D86FB59D7" box="[1220,1249,452,476]" captionStart="Figs 37-42" captionStartId="13.[151,192,2011,2033]" captionTargetBox="[154,1436,257,1939]" captionTargetId="figure-51@13.[151,1437,257,1958]" captionTargetPageId="13" captionText="Figs 37-42. (37, 39, 41) Achilia bifrons. (38, 40, 42) A. blanchardi. Male head in (37-38) dorsal, (39-40) lateral and (41-42) frontal views. Scale bars (200 μm) left for (37, 39, 41) and right for (38, 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5639847" httpUri="https://zenodo.org/record/5639847/files/figure.png" pageId="5" pageNumber="139">42</figureCitation>
, 
<figureCitation id="4FB3FBFBFFD5FF94E1C8FE4A871D59D0" box="[1260,1287,451,475]" captionStart="Fig" captionStartId="18.[153,188,1968,1990]" captionTargetBox="[888,1379,257,2071]" captionTargetId="figure-493@18.[888,1380,257,2071]" captionTargetPageId="18" captionText="Fig. 63. Distribution map of the Achilia cosmoptera species group. (q blue inverted triangles) A. bifrons. (u green diamonds) A. blanchardi. (J red square) A. cosmoptera. (p blue triangles) A. covidia n. sp." figureDoi="http://doi.org/10.5281/zenodo.5645629" httpUri="https://zenodo.org/record/5645629/files/figure.png" pageId="5" pageNumber="139">63</figureCitation>
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<emphasis id="E5FC3B6CFFD5FF94E61BFE7D86195A03" box="[831,1027,500,520]" italics="true" pageId="5" pageNumber="139">Bryaxis valdiviensis</emphasis>
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, pl. 1 fig. 8 (head and antenna) (nec 
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<bibRefCitation id="B3199A8FFFD5FF94E1AAFD86872B5A2E" author="Blanchard C. E." box="[1166,1329,527,549]" pageId="5" pageNumber="139" pagination="1 - 564" refId="ref16343" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos (pp. 561 - 564). In: Gay C., Historia fisica y politica de Chile, Zoologia. Imprimerie Maulde &amp; Renou, Paris. Vol. 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
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). – 
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<bibRefCitation id="B3199A8FFFD5FF94E072FD8681F55A49" author="Reitter E." pageId="5" pageNumber="139" pagination="321 - 332" refId="ref16938" refString="Reitter E. 1885. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia, Zweiter Theil. Deutsche Entomologische Zeitschrift 29: 321 - 332, pl. II." type="journal article" year="1885">Reitter, 1885: 326</bibRefCitation>
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; 330, pl. 2 fig. 14 (head and antenna).
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<emphasis id="E5FC3B6CFFD5FF94E61BFDC281C95A54" box="[831,979,587,607]" italics="true" pageId="5" pageNumber="139">Achilia clavata</emphasis>
<bibRefCitation id="B3199A8FFFD5FF94E6FEFDC086885A55" author="Raffray A." box="[986,1170,585,607]" pageId="5" pageNumber="139" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904: 137</bibRefCitation>
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, fig. 53 (head and antenna) (synonymized by 
<treatmentCitation id="5629C16FFFD5FF94E113FDEF86EA5A77" author="Jeannel R." box="[1079,1264,614,636]" page="429" pageId="5" pageNumber="139" year="1962">
<bibRefCitation id="B3199A8FFFD5FF94E113FDEF86EA5A77" author="Jeannel R." box="[1079,1264,614,636]" pageId="5" pageNumber="139" pagination="295 - 479" refId="ref16517" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D. &amp; Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 429</bibRefCitation>
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).
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<taxonomicName id="10889CFDFFD5FF94E61BFD0C86AE5A93" ID-CoL="9776" authority="Raffray, 1904: 138" authorityName="Raffray" authorityPageNumber="138" authorityYear="1904" box="[831,1204,643,665]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD5FF94E61BFD0C81EF5A92" box="[831,1013,645,665]" italics="true" pageId="5" pageNumber="139">Achilia blanchardi</emphasis>
<bibRefCitation id="B3199A8FFFD5FF94E6D9FD0A86AE5A93" author="Raffray A." box="[1021,1204,643,665]" pageId="5" pageNumber="139" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904: 138</bibRefCitation>
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(new name for 
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<emphasis id="E5FC3B6CFFD5FF94E077FD0C81E65ABD" italics="true" pageId="5" pageNumber="139">Bryaxis valdiviensis</emphasis>
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). – 
<treatmentCitation id="5629C16FFFD5FF94E188FD29877C5ABD" author="Jeannel R." box="[1196,1382,672,694]" page="426" pageId="5" pageNumber="139" year="1962">
<bibRefCitation id="B3199A8FFFD5FF94E188FD29877C5ABD" author="Jeannel R." box="[1196,1382,672,694]" pageId="5" pageNumber="139" pagination="295 - 479" refId="ref16517" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D. &amp; Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 426</bibRefCitation>
</treatmentCitation>
, 430, figs 195 (head and antenna of male), 196 (head and antenna of female), 197 (aedeagus).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9F92B4F5FFD5FF94E61BFC8E86FD5CCF" pageId="5" pageNumber="139" type="materials_examined">
<paragraph id="D737E77EFFD5FF94E61BFC8E878A5C4A" blockId="5.[831,1437,774,1089]" pageId="5" pageNumber="139">
<emphasis id="E5FC3B6CFFD5FF94E61BFC8E863C5B17" bold="true" box="[831,1062,774,797]" pageId="5" pageNumber="139">
Type material (
<specimenCount id="C18E2CF7FFD5FF94E6FFFC8F86005B17" box="[987,1050,774,796]" pageId="5" pageNumber="139" type="generic">12 ex.</specimenCount>
):
</emphasis>
<materialsCitation id="67E0ED23FFD5FF94E108FC8181845B7E" ID-GBIF-Occurrence="3397993412" collectionCode="MNHN" collectorName="Raffray" country="Chile" determinerName="A. Raffray" pageId="5" pageNumber="139" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectionCode id="B1997FBBFFD5FF94E108FC8186625B16" box="[1068,1144,776,797]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="139">MNHN</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E1A6FC8E86BC5B16" box="[1154,1190,775,797]" pageId="5" pageNumber="139" type="female">1 ♀</specimenCount>
(
<typeStatus id="083359DCFFD5FF94E196FC8E87175B16" box="[1202,1293,775,797]" pageId="5" pageNumber="139" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="10889CFDFFD5FF94E008FC8087945B16" authorityName="Raffray" authorityYear="1904" box="[1324,1422,777,797]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="E5FC3B6CFFD5FF94E008FC8087945B16" box="[1324,1422,777,797]" italics="true" pageId="5" pageNumber="139">A. clavata</emphasis>
</taxonomicName>
); labels verbatim “Museum de Paris; 1917; coll. Raffray / 
<collectingCountry id="AF9FA7EEFFD5FF94E048FCAD87865B31" box="[1388,1436,804,826]" pageId="5" pageNumber="139">Chili</collectingCountry>
/ Type / 
<taxonomicName id="10889CFDFFD5FF94E6BAFCCA81FF5B5C" authorityName="Raffray" authorityYear="1904" box="[926,997,835,855]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="E5FC3B6CFFD5FF94E6BAFCCA81FF5B5C" box="[926,997,835,855]" italics="true" pageId="5" pageNumber="139">clavata</emphasis>
</taxonomicName>
; det. 
<determinerName id="22EB0BBAFFD5FF94E100FCCB86955B5C" box="[1060,1167,833,855]" pageId="5" pageNumber="139">A. Raffray</determinerName>
/ 
<taxonomicName id="10889CFDFFD5FF94E188FCCA86E95B5C" authorityName="Raffray" authorityYear="1904" box="[1196,1267,835,855]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="E5FC3B6CFFD5FF94E188FCCA86E95B5C" box="[1196,1267,835,855]" italics="true" pageId="5" pageNumber="139">clavata</emphasis>
</taxonomicName>
(handwritten by Jeannel)”
</materialsCitation>
. 
<materialsCitation id="67E0ED23FFD5FF94E68CFCD6879F5BE2" ID-GBIF-Occurrence="3397993368" collectionCode="MNHN" collectorName="Raffray" country="Chile" county="Type" determinerName="A. Raffray" location="Valdivia Prov." municipality="Valdivia Prov." pageId="5" pageNumber="139" specimenCount="1" specimenCount-male="1" stateProvince="Region Los Rios" typeStatus="lectotype">
– SOUTHERN 
<collectingCountry id="AF9FA7EEFFD5FF94E160FCD686945B7F" box="[1092,1166,863,884]" name="Chile" pageId="5" pageNumber="139">CHILE</collectingCountry>
: 
<collectingRegion id="154C299CFFD5FF94E1BDFCD687585B7E" box="[1177,1346,863,885]" country="Chile" name="Los Rios" pageId="5" pageNumber="139">Región Los Ríos</collectingRegion>
: 
<collectingMunicipality id="37537D04FFD5FF94E068FCD6816C5B9A" pageId="5" pageNumber="139">Valdivia Prov.</collectingMunicipality>
: 
<collectionCode id="B1997FBBFFD5FF94E658FCF581D25B9A" box="[892,968,892,913]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="139">MNHN</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E6F5FCF581E95B9A" box="[977,1011,892,913]" pageId="5" pageNumber="139" type="male">1 ♂</specimenCount>
(
<typeStatus id="083359DCFFD5FF94E6DAFCF586425B99" box="[1022,1112,892,914]" pageId="5" pageNumber="139" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="10889CFDFFD5FF94E150FCF786EF5B99" authorityName="Raffray" authorityYear="1904" box="[1140,1269,894,914]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD5FF94E150FCF786EF5B99" box="[1140,1269,894,914]" italics="true" pageId="5" pageNumber="139">A. blanchardi</emphasis>
</taxonomicName>
here designated); labels verbatim “Lectotype / Type / 1880; Chili; Valdivia; leg. Kindermann/ 
<taxonomicName id="10889CFDFFD5FF94E6E3FC31860A5BC7" box="[967,1040,952,972]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="E5FC3B6CFFD5FF94E6E3FC31860A5BC7" box="[967,1040,952,972]" italics="true" pageId="5" pageNumber="139">Bryaxis</emphasis>
</taxonomicName>
; 
<taxonomicName id="10889CFDFFD5FF94E13BFC31868A5BC7" authorityName="Reitter" authorityYear="1883" box="[1055,1168,952,972]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="E5FC3B6CFFD5FF94E13BFC31868A5BC7" box="[1055,1168,952,972]" italics="true" pageId="5" pageNumber="139">valdiviensis</emphasis>
</taxonomicName>
; m. Valdivia / Museum de Paris; 1917; coll. 
<collectorName id="7A7D82A8FFD5FF94E6D4FC5A86235BE2" box="[1008,1081,979,1001]" pageId="5" pageNumber="139">Raffray</collectorName>
/ 
<taxonomicName id="10889CFDFFD5FF94E174FC5C86CF5BE2" authorityName="Raffray" authorityYear="1904" box="[1104,1237,981,1001]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD5FF94E174FC5C86CF5BE2" box="[1104,1237,981,1001]" italics="true" pageId="5" pageNumber="139">A. blanchardi</emphasis>
</taxonomicName>
; 
<determinerName id="22EB0BBAFFD5FF94E1C7FC5A87565BE2" box="[1251,1356,979,1001]" pageId="5" pageNumber="139">A. Raffray</determinerName>
det.”
</materialsCitation>
. 
<materialsCitation id="67E0ED23FFD5FF94E0B4FC5D87975C4A" ID-GBIF-Occurrence="3397993385" collectionCode="MNHN" collectorName="Raffray" country="Chile" determinerName="A. Raffray" pageId="5" pageNumber="139" specimenCount="10" specimenCount-female="9" specimenCount-male="1" stateProvince="Los Rios" typeStatus="paralectotype">
– 
<collectionCode id="B1997FBBFFD5FF94E61BFC7881915C0D" box="[831,907,1009,1030]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="139">MNHN</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E6BDFC7881DB5C0D" box="[921,961,1009,1030]" pageId="5" pageNumber="139" type="male">1 ♂</specimenCount>
and 
<specimenCount id="C18E2CF7FFD5FF94E6D2FC7886045C0D" box="[1014,1054,1009,1030]" pageId="5" pageNumber="139" type="female">9 ♀</specimenCount>
(
<typeStatus id="083359DCFFD5FF94E10AFC7986A65C0D" box="[1070,1212,1008,1030]" pageId="5" pageNumber="139" type="paralectotype">Paralectotypes</typeStatus>
of 
<taxonomicName id="10889CFDFFD5FF94E1C7FC7B87735C0D" authorityName="Raffray" authorityYear="1904" box="[1251,1385,1010,1030]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD5FF94E1C7FC7B87735C0D" box="[1251,1385,1010,1030]" italics="true" pageId="5" pageNumber="139">A. blanchardi</emphasis>
</taxonomicName>
here designated); labels verbatim “Paralectotype / 
<collectingCountry id="AF9FA7EEFFD5FF94E025FB84872B5C28" box="[1281,1329,1037,1059]" pageId="5" pageNumber="139">Chili</collectingCountry>
/ Museum de Paris; 1917; coll. 
<collectorName id="7A7D82A8FFD5FF94E121FBA286545C4A" box="[1029,1102,1067,1089]" pageId="5" pageNumber="139">Raffray</collectorName>
/ 
<taxonomicName id="10889CFDFFD5FF94E145FBA486FE5C4A" authorityName="Raffray" authorityYear="1904" box="[1121,1252,1069,1089]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD5FF94E145FBA486FE5C4A" box="[1121,1252,1069,1089]" italics="true" pageId="5" pageNumber="139">A. blanchardi</emphasis>
</taxonomicName>
; 
<determinerName id="22EB0BBAFFD5FF94E1CBFBA2874C5C4A" box="[1263,1366,1067,1089]" pageId="5" pageNumber="139">A. Raffray</determinerName>
det.”
</materialsCitation>
.
</paragraph>
<paragraph id="D737E77EFFD5FF94E61BFBDE86FD5CCF" blockId="5.[831,1437,1110,1220]" pageId="5" pageNumber="139">
<emphasis id="E5FC3B6CFFD5FF94E61BFBDE86AC5C67" bold="true" box="[831,1206,1110,1133]" pageId="5" pageNumber="139">
Additional material examined (
<specimenCount id="C18E2CF7FFD5FF94E15DFBDF86B05C67" box="[1145,1194,1110,1132]" pageId="5" pageNumber="139" type="generic">5 ex.</specimenCount>
):
</emphasis>
<materialsCitation id="67E0ED23FFD5FF94E19EFBDE81B15CAC" ID-GBIF-Occurrence="3397993356" collectionCode="FMNH" collectorName="Field Mus. Nat. Hist." location="Orlando Park" pageId="5" pageNumber="139" specimenCount="1" specimenCount-female="1">
<collectionCode id="B1997FBBFFD5FF94E19EFBDE87185C67" box="[1210,1282,1111,1132]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="139" type="Museum">FMNH</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E02EFBDE87365C67" box="[1290,1324,1111,1132]" pageId="5" pageNumber="139" type="female">1 ♀</specimenCount>
; Field Mus. Nat. Hist.; 
<location id="D257B1A5FFD5FF94E696FBFD86205C81" LSID="urn:lsid:plazi:treatment:5F215668FFD5FF97E692FE2A80AD5F10:D257B1A5FFD5FF94E696FBFD86205C81" box="[946,1082,1140,1162]" name="Orlando Park" pageId="5" pageNumber="139">Orlando Park</location>
; 
<taxonomicName id="10889CFDFFD5FF94E16CFBFD871A5C81" authority="Colln." authorityName="Colln." box="[1096,1280,1140,1162]" class="Insecta" family="Pselaphidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="139" phylum="Arthropoda" rank="family">Pselaphidae Colln</taxonomicName>
.; F. C. Fletcher Collection.
</materialsCitation>
<materialsCitation id="67E0ED23FFD5FF94E694FB1B871B5CAC" ID-GBIF-Occurrence="3397993464" box="[944,1281,1169,1191]" collectionCode="MNHN" collectorName="P. Germain" country="Chile" pageId="5" pageNumber="139" specimenCount="1" specimenCount-female="1">
– 
<collectionCode id="B1997FBBFFD5FF94E6E7FB1B86155CAC" box="[963,1039,1170,1191]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="139">MNHN</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E13EFB1886255CAC" box="[1050,1087,1169,1191]" pageId="5" pageNumber="139" type="female">1 ♀</specimenCount>
; 
<collectingCountry id="AF9FA7EEFFD5FF94E169FB18869E5CAC" box="[1101,1156,1169,1191]" name="Chile" pageId="5" pageNumber="139">Chile</collectingCountry>
; 
<collectorName id="7A7D82A8FFD5FF94E1AAFB1B86E75CAC" box="[1166,1277,1169,1191]" pageId="5" pageNumber="139">P. Germain</collectorName>
.
</materialsCitation>
<materialsCitation id="67E0ED23FFD5FF94E023FB1B81925CCF" ID-GBIF-Occurrence="3397993450" collectionCode="HNHM" country="Chile" pageId="5" pageNumber="139" specimenCount="1" specimenCount-male="1">
– 
<collectionCode id="B1997FBBFFD5FF94E03EFB1B877C5CAC" box="[1306,1382,1170,1191]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="139">HNHM</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E055FB18878C5CAC" box="[1393,1430,1169,1191]" pageId="5" pageNumber="139" type="male">1 ♂</specimenCount>
; “ 
<collectingCountry id="AF9FA7EEFFD5FF94E66DFB2781645CCF" box="[841,894,1198,1220]" name="Chile" pageId="5" pageNumber="139">Chili</collectingCountry>
”
</materialsCitation>
. 
<materialsCitation id="67E0ED23FFD5FF94E6B5FB2686FD5CCF" ID-GBIF-Occurrence="3397993364" box="[913,1255,1198,1220]" collectionCode="MNHN" country="Chile" pageId="5" pageNumber="139" specimenCount="2" specimenCount-female="2">
– 
<collectionCode id="B1997FBBFFD5FF94E687FB2681F55CCF" box="[931,1007,1199,1220]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="139">MNHN</collectionCode>
; 
<specimenCount id="C18E2CF7FFD5FF94E6DDFB2686075CCF" box="[1017,1053,1199,1220]" pageId="5" pageNumber="139" type="female">2 ♀</specimenCount>
; “ 
<collectingCountry id="AF9FA7EEFFD5FF94E117FB27867D5CCF" box="[1075,1127,1198,1220]" name="Chile" pageId="5" pageNumber="139">Chili</collectingCountry>
”; Gay; 1849.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9F92B4F5FFD5FF97E61BFB5580FD5A1C" lastPageId="6" lastPageNumber="140" pageId="5" pageNumber="139" type="description">
<paragraph id="D737E77EFFD5FF94E61BFB5581B55D3E" blockId="5.[831,1437,1244,2078]" pageId="5" pageNumber="139">
<emphasis id="E5FC3B6CFFD5FF94E61BFB5581D45CFF" bold="true" box="[831,974,1244,1268]" pageId="5" pageNumber="139">Description:</emphasis>
Body 
<quantity id="10704A9BFFD5FF94E101FB5586DF5CFF" box="[1061,1221,1244,1268]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="5" pageNumber="139" unit="mm" value="1.375" valueMax="1.4" valueMin="1.35">1.35-1.40 mm</quantity>
long, dark brown with reddish elytra; antennae and legs reddish; palpi yellowish.
</paragraph>
<paragraph id="D737E77EFFD5FF97E61BFAB6801A59B3" blockId="5.[831,1437,1244,2078]" lastBlockId="6.[151,756,257,535]" lastPageId="6" lastPageNumber="140" pageId="5" pageNumber="139">
<emphasis id="E5FC3B6CFFD5FF94E61BFAB6816C5D5E" box="[831,886,1343,1365]" italics="true" pageId="5" pageNumber="139">Male</emphasis>
: Head as in 
<figureCitation id="4FB3FBFBFFD5FF94E6D7FAB486BA5D5E" box="[1011,1184,1341,1365]" captionStart="Figs 37-42" captionStartId="13.[151,192,2011,2033]" captionTargetBox="[154,1436,257,1939]" captionTargetId="figure-51@13.[151,1437,257,1958]" captionTargetPageId="13" captionText="Figs 37-42. (37, 39, 41) Achilia bifrons. (38, 40, 42) A. blanchardi. Male head in (37-38) dorsal, (39-40) lateral and (41-42) frontal views. Scale bars (200 μm) left for (37, 39, 41) and right for (38, 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5639847" httpUri="https://zenodo.org/record/5639847/files/figure.png" pageId="5" pageNumber="139">Figs 38, 40 &amp; 42</figureCitation>
, very wide, surface with some very scattered punctures; occipital region raised, frons flattened and separated by large transverse sulcus from large frontal lobe; anterior margin of frontal sulcus slightly raised and pointed at middle. Antennae (
<figureCitation id="4FB3FBFBFFD5FF94E063FA36878E5DDC" box="[1351,1428,1471,1495]" captionStart="Figs 16-22" captionStartId="10.[151,192,2026,2048]" captionTargetBox="[179,1415,984,1976]" captionTargetId="figure-479@10.[153,1438,959,1994]" captionTargetPageId="10" captionText="Figs 16-22. Male antennae of Achilia. (16) A. covidia. (17) A. blanchardi. (18) A. bifrons. (19) A. temporalis. (20) A. pandemica. (21) A. angularis. (22) A. quarantena." figureDoi="http://doi.org/10.5281/zenodo.5639843" httpUri="https://zenodo.org/record/5639843/files/figure.png" pageId="5" pageNumber="139">Fig. 17</figureCitation>
) with scape and pedicel longer than wide; antennomere III distinctly longer than wide; antennomere IV about as long as wide; antennomeres V-VII slightly longer than wide; antennomere VIII as long as wide; antennomere IX wider than VIII and wider than long; antennomere X distinctly wider than long, shorter and wider than IX; antennomere XI elongate, longer than VII-X combined. Metaventrite with deep and wide transverse sulcus on apical twothirds, sulcus with deflected and posteriorly prominent margins forming sort of tubercles. First abdominal sternite slightly raised and flattened at middle; remaining abdominal sternites slightly flattened at middle. Protibiae with distal half slightly swollen; mesotibiae (
<figureCitation id="4FB3FBFBFFD5FF94E065F8EA878F5F70" box="[1345,1429,1891,1915]" captionStart="Figs 23-32" captionStartId="11.[151,192,2024,2046]" captionTargetBox="[188,1424,1432,1985]" captionTargetId="figure-589@11.[151,1436,1428,1996]" captionText="Figs 23-32. Male protibiae (23-25), mesotibiae (26-31) and metatibiae (32) of Achilia. (23, 28) A. elfridae. (24, 30, 32) A. angularis. (25, 31) A. quarantena. (26) A. covidia. (27) A. blanchardi. (29) A. pandemica." figureDoi="http://doi.org/10.5281/zenodo.5645115" httpUri="https://zenodo.org/record/5645115/files/figure.png" pageId="5" pageNumber="139">Fig. 27</figureCitation>
) with distal half swollen and densely pubescent. Aedeagus (
<figureCitation id="4FB3FBFBFFD5FF94E663F82D819D5FB7" box="[839,903,1956,1980]" captionStart="Figs 1-6" captionStartId="8.[153,194,2054,2076]" captionTargetBox="[177,1406,898,2017]" captionTargetId="figure-286@8.[149,1439,882,2039]" captionText="Figs 1-6. Aedeagi of Achilia. (1) A. cosmoptera. (2) A. covidia. (3) A. blanchardi. (4) A. elfridae. (5) A. bifrons. (6) A. temporalis." figureDoi="http://doi.org/10.5281/zenodo.5645110" httpUri="https://zenodo.org/record/5645110/files/figure.png" pageId="5" pageNumber="139">Fig. 3</figureCitation>
) 
<quantity id="10704A9BFFD5FF94E6BCF82D81ED5FB0" box="[920,1015,1956,1980]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="139" unit="mm" value="0.23">0.23 mm</quantity>
long, with dorsal plate subrectangular; dorsal longitudinal struts divergent. Parameres relatively wide with very large and long recurved seta on welldeveloped outer lobe, this seta bifid and very swollen on basal third; apical portion of parameres slightly recurved anteriorly and prolonged laterally at tip; apex of parameres bearing one long ventral medial seta associated with wider and shorter seta. Copulatory pieces consisting of two subequal short sclerites slightly enlarged apically, recurved and more robust basally.
</paragraph>
<paragraph id="D737E77EFFD6FF97E5B3FE4B80FD5A1C" blockId="6.[151,756,257,535]" pageId="6" pageNumber="140">
<emphasis id="E5FC3B6CFFD6FF97E5B3FE4B82FD59D3" box="[151,231,450,472]" italics="true" pageId="6" pageNumber="140">Female</emphasis>
: Similar to male except head not modified; antennae shorter with antennomere XI shorter; metaventrite, abdominal sternites, and legs unmodified.
</paragraph>
</subSubSection>
<subSubSection id="9F92B4F5FFD6FF97E5B3FDA780455A6D" pageId="6" pageNumber="140" type="biology_ecology">
<paragraph id="D737E77EFFD6FF97E5B3FDA780455A6D" blockId="6.[151,755,558,614]" pageId="6" pageNumber="140">
<emphasis id="E5FC3B6CFFD6FF97E5B3FDA7834F5A4D" bold="true" box="[151,341,558,582]" pageId="6" pageNumber="140">Collecting data:</emphasis>
The data in our possession do not allow us to make statements in this regard.
</paragraph>
</subSubSection>
<subSubSection id="9F92B4F5FFD6FF97E5B3FDF5834B5ADF" pageId="6" pageNumber="140" type="distribution">
<paragraph id="D737E77EFFD6FF97E5B3FDF5834B5ADF" blockId="6.[151,756,636,725]" pageId="6" pageNumber="140">
<emphasis id="E5FC3B6CFFD6FF97E5B3FDF583375A9F" bold="true" box="[151,301,636,660]" pageId="6" pageNumber="140">Distribution:</emphasis>
<taxonomicName id="10889CFDFFD6FF97E41CFDF6801D5A9E" authorityName="Raffray" authorityYear="1904" box="[312,519,639,661]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E41CFDF6801D5A9E" box="[312,519,639,661]" italics="true" pageId="6" pageNumber="140">Achilia blanchardi</emphasis>
</taxonomicName>
is only known from Valdivia (
<collectingRegion id="154C299CFFD6FF97E421FD1583D65ABE" box="[261,460,668,693]" country="Chile" name="Los Rios" pageId="6" pageNumber="140">Región Los Ríos</collectingRegion>
: Valdivia Prov.) (
<figureCitation id="4FB3FBFBFFD6FF97E7BDFD1580EB5ABF" box="[665,753,668,692]" captionStart="Fig" captionStartId="18.[153,188,1968,1990]" captionTargetBox="[888,1379,257,2071]" captionTargetId="figure-493@18.[888,1380,257,2071]" captionTargetPageId="18" captionText="Fig. 63. Distribution map of the Achilia cosmoptera species group. (q blue inverted triangles) A. bifrons. (u green diamonds) A. blanchardi. (J red square) A. cosmoptera. (p blue triangles) A. covidia n. sp." figureDoi="http://doi.org/10.5281/zenodo.5645629" httpUri="https://zenodo.org/record/5645629/files/figure.png" pageId="6" pageNumber="140">Fig. 63</figureCitation>
: green diamonds).
</paragraph>
</subSubSection>
<subSubSection id="9F92B4F5FFD6FF97E5B3FD6280AD5F10" pageId="6" pageNumber="140" type="discussion">
<paragraph id="D737E77EFFD6FF97E5B3FD62800B5BCA" blockId="6.[151,756,747,1819]" pageId="6" pageNumber="140">
<emphasis id="E5FC3B6CFFD6FF97E5B3FD6283075B08" bold="true" box="[151,285,747,771]" pageId="6" pageNumber="140">Comments:</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E408FD62800F5B08" author="Raffray A." box="[300,533,747,771]" pageId="6" pageNumber="140" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray (1904: 138)</bibRefCitation>
proposed the new name 
<taxonomicName id="10889CFDFFD6FF97E5C3FC8483D85B28" authorityName="Raffray" authorityYear="1904" box="[231,450,781,803]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E5C3FC8483D85B28" box="[231,450,781,803]" italics="true" pageId="6" pageNumber="140">Achilia blanchardi</emphasis>
</taxonomicName>
for 
<taxonomicName id="10889CFDFFD6FF97E72FFC84832A5B48" authority="Reitter, 1883" authorityName="Reitter" authorityYear="1883" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="E5FC3B6CFFD6FF97E72FFC8480EE5B28" box="[523,756,781,803]" italics="true" pageId="6" pageNumber="140">Bryaxis valdiviensis</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E5B3FCA2832A5B48" author="Reitter E." box="[151,304,811,835]" pageId="6" pageNumber="140" pagination="47 - 54" refId="ref16912" refString="Reitter E. 1883. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia. Deutsche Entomologische Zeitschrift 27: 47 - 54, pl. I." type="journal article" year="1883">Reitter, 1883</bibRefCitation>
</taxonomicName>
, described on an unspecified number of specimens from Valdivia, because this name was preoccupied by 
<taxonomicName id="10889CFDFFD6FF97E446FCE582D65BA9" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="E5FC3B6CFFD6FF97E446FCE580725B89" box="[354,616,876,898]" italics="true" pageId="6" pageNumber="140">Pselaphus valdiviensis</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E759FCE382D65BA9" author="Blanchard C. E." pageId="6" pageNumber="140" pagination="1 - 564" refId="ref16343" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos (pp. 561 - 564). In: Gay C., Historia fisica y politica de Chile, Zoologia. Imprimerie Maulde &amp; Renou, Paris. Vol. 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
[now 
<taxonomicName id="10889CFDFFD6FF97E431FC0580A35BA9" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[277,697,906,930]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="E5FC3B6CFFD6FF97E431FC0583F05BA9" box="[277,490,908,930]" italics="true" pageId="6" pageNumber="140">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="B3199A8FFFD6FF97E4DEFC0380AB5BA9" author="Blanchard C. E." box="[506,689,906,930]" pageId="6" pageNumber="140" pagination="1 - 564" refId="ref16343" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos (pp. 561 - 564). In: Gay C., Historia fisica y politica de Chile, Zoologia. Imprimerie Maulde &amp; Renou, Paris. Vol. 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
] (see 
<bibRefCitation id="B3199A8FFFD6FF97E5B3FC2383865BC9" author="Sabella G. &amp; Cuccodoro G. &amp; Kurbatov S. A." box="[151,412,938,962]" pageId="6" pageNumber="140" pagination="129 - 156" refId="ref17106" refString="Sabella G., Cuccodoro G., Kurbatov S. A. 2020. A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae). Revue suisse de Zoologie 127 (1): 129 - 156." type="journal article" year="2020">
Sabella 
<emphasis id="E5FC3B6CFFD6FF97E5C8FC2583015BC9" box="[236,283,940,962]" italics="true" pageId="6" pageNumber="140">et al</emphasis>
., 2020: 140
</bibRefCitation>
, 144-145).
</paragraph>
<paragraph id="D737E77EFFD6FF97E5B3FC4083F45C8B" blockId="6.[151,756,747,1819]" pageId="6" pageNumber="140">
<bibRefCitation id="B3199A8FFFD6FF97E5B3FC40839B5BEA" author="Jeannel R." box="[151,385,969,993]" pageId="6" pageNumber="140" pagination="295 - 479" refId="ref16517" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D. &amp; Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962: 430)</bibRefCitation>
listed 
<taxonomicName id="10889CFDFFD6FF97E4FBFC4282D65C0A" authority="Raffray, 1904" authorityName="Raffray" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="E5FC3B6CFFD6FF97E4FBFC4280975BEA" box="[479,653,971,993]" italics="true" pageId="6" pageNumber="140">Achilia clavata</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E7BAFC4082D65C0A" author="Raffray A." pageId="6" pageNumber="140" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904</bibRefCitation>
</taxonomicName>
as a junior synonym of 
<taxonomicName id="10889CFDFFD6FF97E4F3FC6280705C0A" authorityName="Raffray" authorityYear="1904" box="[471,618,1003,1025]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E4F3FC6280705C0A" box="[471,618,1003,1025]" italics="true" pageId="6" pageNumber="140">A. blanchardi</emphasis>
</taxonomicName>
, but without justifying this action. We have examined the 
<typeStatus id="083359DCFFD6FF97E75DFB8080CC5C2A" box="[633,726,1033,1057]" pageId="6" pageNumber="140" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="10889CFDFFD6FF97E5B3FBA2831C5C4A" authorityName="Raffray" authorityYear="1904" box="[151,262,1067,1089]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="clavata">
<emphasis id="E5FC3B6CFFD6FF97E5B3FBA2831C5C4A" box="[151,262,1067,1089]" italics="true" pageId="6" pageNumber="140">A. clavata</emphasis>
</taxonomicName>
, which is a female resembling in all features those of 
<taxonomicName id="10889CFDFFD6FF97E5DAFBC3838C5C6B" authorityName="Raffray" authorityYear="1904" box="[254,406,1098,1120]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E5DAFBC3838C5C6B" box="[254,406,1098,1120]" italics="true" pageId="6" pageNumber="140">A. blanchardi</emphasis>
</taxonomicName>
, and therefore we confirm the synonymy proposed by Jeannel.
</paragraph>
<paragraph id="D737E77EFFD6FF97E5B3FB01834C5E16" blockId="6.[151,756,747,1819]" pageId="6" pageNumber="140">
<bibRefCitation id="B3199A8FFFD6FF97E5B3FB0183675CAB" author="Jeannel R." box="[151,381,1160,1184]" pageId="6" pageNumber="140" pagination="295 - 479" refId="ref16517" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale (pp. 295 - 479). In: Deboutteville C. D. &amp; Rapoport E. (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962: 430)</bibRefCitation>
stated that the type of 
<taxonomicName id="10889CFDFFD6FF97E783FB0383A85CCB" authority="Raffray, 1904" authorityName="Raffray" authorityYear="1904" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E783FB0383115CCB" italics="true" pageId="6" pageNumber="140">Achilia blanchardi</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E43DFB2E83A85CCB" author="Raffray A." box="[281,434,1191,1216]" pageId="6" pageNumber="140" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904</bibRefCitation>
</taxonomicName>
was housed in the MNHN collections. In the MNHN we found in Raffray’s collection a series of 
<specimenCount id="C18E2CF7FFD6FF97E494FB6E80565CF4" box="[432,588,1255,1279]" pageId="6" pageNumber="140" type="generic">11 specimens</specimenCount>
identified by Raffray as 
<taxonomicName id="10889CFDFFD6FF97E434FA8083C05D14" authorityName="Raffray" authorityYear="1904" box="[272,474,1289,1311]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E434FA8083C05D14" box="[272,474,1289,1311]" italics="true" pageId="6" pageNumber="140">Achilia blanchardi</emphasis>
</taxonomicName>
; the first specimen of the series is a male labeled “Type / 1880; 
<collectingCountry id="AF9FA7EEFFD6FF97E769FAAF809C5D35" box="[589,646,1318,1342]" name="Chile" pageId="6" pageNumber="140">Chili</collectingCountry>
; Valdivia; leg. Kindermann/ 
<taxonomicName id="10889CFDFFD6FF97E442FAC183AD5D55" box="[358,439,1352,1374]" class="Insecta" family="Staphylinidae" genus="Bryaxis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="E5FC3B6CFFD6FF97E442FAC183AD5D55" box="[358,439,1352,1374]" italics="true" pageId="6" pageNumber="140">Bryaxis</emphasis>
</taxonomicName>
; 
<taxonomicName id="10889CFDFFD6FF97E4E8FAC180505D55" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[460,586,1352,1374]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="E5FC3B6CFFD6FF97E4E8FAC180505D55" box="[460,586,1352,1374]" italics="true" pageId="6" pageNumber="140">valdiviensis</emphasis>
</taxonomicName>
; m. Valdivia / Museum de Paris; 1917; coll. Raffray / 
<taxonomicName id="10889CFDFFD6FF97E773FAE180F65D75" authorityName="Raffray" authorityYear="1904" box="[599,748,1384,1406]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E773FAE180F65D75" box="[599,748,1384,1406]" italics="true" pageId="6" pageNumber="140">A. blanchardi</emphasis>
</taxonomicName>
; A. Raffray det.” which we designate here as the 
<typeStatus id="083359DCFFD6FF97E7B6FA0F80EE5D95" box="[658,756,1414,1438]" pageId="6" pageNumber="140" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="10889CFDFFD6FF97E59CFA2180A55DB6" authority="Raffray, 1904" authorityName="Raffray" authorityYear="1904" box="[184,703,1445,1470]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E59CFA2183575DB5" box="[184,333,1448,1470]" italics="true" pageId="6" pageNumber="140">A. blanchardi</emphasis>
<bibRefCitation id="B3199A8FFFD6FF97E473FA2C83F65DB5" author="Raffray A." box="[343,492,1445,1470]" pageId="6" pageNumber="140" pagination="1 - 400" refId="ref16888" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904</bibRefCitation>
(
<bibRefCitation id="B3199A8FFFD6FF97E4DAFA2C80AC5DB6" author="Blanchard C. E." box="[510,694,1445,1469]" pageId="6" pageNumber="140" pagination="1 - 564" refId="ref16343" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos (pp. 561 - 564). In: Gay C., Historia fisica y politica de Chile, Zoologia. Imprimerie Maulde &amp; Renou, Paris. Vol. 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
, and the remaining 
<specimenCount id="C18E2CF7FFD6FF97E41BFA4C83CC5DD6" box="[319,470,1477,1501]" pageId="6" pageNumber="140" type="generic">10 specimens</specimenCount>
(
<specimenCount id="C18E2CF7FFD6FF97E4CBFA4C80265DD6" box="[495,572,1477,1501]" pageId="6" pageNumber="140" type="male">1 male</specimenCount>
and 
<specimenCount id="C18E2CF7FFD6FF97E759FA4C80F75DD6" box="[637,749,1477,1501]" pageId="6" pageNumber="140" type="female">9 females</specimenCount>
) labeled “ 
<collectingCountry id="AF9FA7EEFFD6FF97E5D2FA6C83355DF6" box="[246,303,1509,1533]" name="Chile" pageId="6" pageNumber="140">Chili</collectingCountry>
” without further label data are designated as 
<typeStatus id="083359DCFFD6FF97E590F98C83485E16" box="[180,338,1541,1565]" pageId="6" pageNumber="140" type="paralectotype">paralectotypes</typeStatus>
.
</paragraph>
<paragraph id="D737E77EFFD6FF97E5B3F9AD80AD5F10" blockId="6.[151,756,747,1819]" pageId="6" pageNumber="140">
Within the 
<taxonomicName id="10889CFDFFD6FF97E42CF9AE83855E36" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[264,415,1575,1597]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="cosmoptera">
<emphasis id="E5FC3B6CFFD6FF97E42CF9AE83855E36" box="[264,415,1575,1597]" italics="true" pageId="6" pageNumber="140">A. cosmoptera</emphasis>
</taxonomicName>
group the males of 
<taxonomicName id="10889CFDFFD6FF97E741F9AE80EE5E36" authorityName="Raffray" authorityYear="1904" box="[613,756,1575,1597]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="E5FC3B6CFFD6FF97E741F9AE80EE5E36" box="[613,756,1575,1597]" italics="true" pageId="6" pageNumber="140">A. blanchardi</emphasis>
</taxonomicName>
are easily distinguished from other species by the shape of the head (
<figureCitation id="4FB3FBFBFFD6FF97E40FF9ED83C75E77" box="[299,477,1636,1660]" captionStart="Figs 37-42" captionStartId="13.[151,192,2011,2033]" captionTargetBox="[154,1436,257,1939]" captionTargetId="figure-51@13.[151,1437,257,1958]" captionTargetPageId="13" captionText="Figs 37-42. (37, 39, 41) Achilia bifrons. (38, 40, 42) A. blanchardi. Male head in (37-38) dorsal, (39-40) lateral and (41-42) frontal views. Scale bars (200 μm) left for (37, 39, 41) and right for (38, 40, 42)." figureDoi="http://doi.org/10.5281/zenodo.5639847" httpUri="https://zenodo.org/record/5639847/files/figure.png" pageId="6" pageNumber="140">Figs 38, 40, 42</figureCitation>
), antennae (
<figureCitation id="4FB3FBFBFFD6FF97E74CF9ED80A35E77" box="[616,697,1636,1660]" captionStart="Figs 16-22" captionStartId="10.[151,192,2026,2048]" captionTargetBox="[179,1415,984,1976]" captionTargetId="figure-479@10.[153,1438,959,1994]" captionTargetPageId="10" captionText="Figs 16-22. Male antennae of Achilia. (16) A. covidia. (17) A. blanchardi. (18) A. bifrons. (19) A. temporalis. (20) A. pandemica. (21) A. angularis. (22) A. quarantena." figureDoi="http://doi.org/10.5281/zenodo.5639843" httpUri="https://zenodo.org/record/5639843/files/figure.png" pageId="6" pageNumber="140">Fig. 17</figureCitation>
) and aedeagus (
<figureCitation id="4FB3FBFBFFD6FF97E42EF90D83545E97" box="[266,334,1668,1692]" captionStart="Figs 1-6" captionStartId="8.[153,194,2054,2076]" captionTargetBox="[177,1406,898,2017]" captionTargetId="figure-286@8.[149,1439,882,2039]" captionText="Figs 1-6. Aedeagi of Achilia. (1) A. cosmoptera. (2) A. covidia. (3) A. blanchardi. (4) A. elfridae. (5) A. bifrons. (6) A. temporalis." figureDoi="http://doi.org/10.5281/zenodo.5645110" httpUri="https://zenodo.org/record/5645110/files/figure.png" pageId="6" pageNumber="140">Fig. 3</figureCitation>
). The females of this species are very similar to those of 
<taxonomicName id="10889CFDFFD6FF97E44EF92F83C75EB7" authorityName="Raffray" authorityYear="1904" box="[362,477,1702,1724]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="elfridae">
<emphasis id="E5FC3B6CFFD6FF97E44EF92F83C75EB7" box="[362,477,1702,1724]" italics="true" pageId="6" pageNumber="140">A. elfridae</emphasis>
</taxonomicName>
, from which they can be distinguished by their wider head with a wider frontal sulcus, and their slightly longer antennomere XI (
<quantity id="10704A9BFFD6FF97E7B2F96A80E95EF1" box="[662,755,1763,1787]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="140" unit="mm" value="0.16">0.16 mm</quantity>
for 
<taxonomicName id="10889CFDFFD6FF97E599F88C83525F10" box="[189,328,1797,1819]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="blanchard">
<emphasis id="E5FC3B6CFFD6FF97E599F88C83525F10" box="[189,328,1797,1819]" italics="true" pageId="6" pageNumber="140">A. blanchard</emphasis>
</taxonomicName>
i vs 
<quantity id="10704A9BFFD6FF97E450F88B80105F11" box="[372,522,1794,1818]" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="6" pageNumber="140" unit="mm" value="0.145" valueMax="0.15" valueMin="0.14">0.14-0.15 mm</quantity>
for 
<taxonomicName id="10889CFDFFD6FF97E712F88C80BD5F10" authorityName="Raffray" authorityYear="1904" box="[566,679,1797,1819]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="140" phylum="Arthropoda" rank="species" species="elfridae">
<emphasis id="E5FC3B6CFFD6FF97E712F88C80BD5F10" box="[566,679,1797,1819]" italics="true" pageId="6" pageNumber="140">A. elfridae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>