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<document id="A62D6438E9D0E365871A8ABCB76B7EEE" ID-DOI="10.5281/zenodo.193611" ID-GBIF-Dataset="97dd4cba-69c8-4d37-8fe0-72811d79afe3" ID-ISSN="1175-5326" ID-Zenodo-Dep="193611" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460158883150" checkinUser="plazi" docAuthor="Ma, Li, Li, Qiang &amp; Chen, Xue-Xin" docDate="2010" docId="981787F3FF97FFCBE8A6EEFE6670FBF8" docLanguage="en" docName="zt02359p060.pdf" docOrigin="Zootaxa 2359" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Odontopsen shii Ma, Li and Chen, sp. nov." docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="642EFF8BFF97FFCAE831EB0E6742FFAF" masterDocTitle="A rare genus Odontopsen Tsuneki in China (Hymenoptera: Apoidea: Crabronidae), with description of a new species" masterLastPageNumber="60" masterPageNumber="58" pageNumber="58" updateTime="1698235703409" updateUser="plazi">
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<mods:title id="F6BF1BB7F9F3C5C762CB576867A44C68">A rare genus Odontopsen Tsuneki in China (Hymenoptera: Apoidea: Crabronidae), with description of a new species</mods:title>
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<mods:name id="DD9A93F5655E930DF006F7F83DF30DE2" type="personal">
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<mods:namePart id="CD94D0504703E174B13D4C07C96A13C9">Ma, Li</mods:namePart>
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<mods:name id="BFCC8E431E5BF382A0F9293B33FD0C24" type="personal">
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<mods:namePart id="3E6BD161B5C271BCFF7C46DF496655D4">Li, Qiang</mods:namePart>
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<mods:name id="39C4B16B7EF0F82E5E13F950FD1A42A2" type="personal">
<mods:role id="5C9A3BD352E82A91E35CA1E2A7FC7468">
<mods:roleTerm id="2A37C0DB53A94D8B19E17F31052839B9">Author</mods:roleTerm>
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<mods:namePart id="03847B0316FCB81B1C7001C51EB0D043">Chen, Xue-Xin</mods:namePart>
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<mods:typeOfResource id="05AD4E3C4BF1C99A2745C9652DCCEF39">text</mods:typeOfResource>
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<mods:titleInfo id="5ED29E71359E5D8A1136943A566EA30D">
<mods:title id="C55982B996DD564F8916BCADD6020181">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AC37175DB6C2A210B54E537D920B8984">
<mods:date id="9A6EB23B8935DEFF10B39543D782CCA8">2010</mods:date>
<mods:detail id="FA431E4CB19225EA31F784DFA22464F8" type="volume">
<mods:number id="07B44784009AD9B8DD383CDCE5FBC829">2359</mods:number>
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<mods:classification id="ABCA92F41F1A9FBB112B4D1C3070EB8A">journal article</mods:classification>
<mods:identifier id="2FAD7551E0B524DE2FA68F893CB161B9" type="DOI">10.5281/zenodo.193611</mods:identifier>
<mods:identifier id="8E99605C351C99A0436F945A9112D4A1" type="GBIF-Dataset">97dd4cba-69c8-4d37-8fe0-72811d79afe3</mods:identifier>
<mods:identifier id="B73738A8DCC8CBBA57A1FC67BD1A5EB0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="87574B3DBAA393262C833EEDF0AE0D28" type="Zenodo-Dep">193611</mods:identifier>
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<treatment id="981787F3FF97FFCBE8A6EEFE6670FBF8" ID-DOI="http://doi.org/10.5281/zenodo.6210201" ID-GBIF-Taxon="124956115" ID-Zenodo-Dep="6210201" LSID="urn:lsid:plazi:treatment:981787F3FF97FFCBE8A6EEFE6670FBF8" httpUri="http://treatment.plazi.org/id/981787F3FF97FFCBE8A6EEFE6670FBF8" lastPageId="1" lastPageNumber="59" pageId="0" pageNumber="58">
<subSubSection id="58A4656EFF97FFCAE8A6EEFE664EF985" pageId="0" pageNumber="58" type="nomenclature">
<paragraph id="100136E5FF97FFCAE8A6EEFE6539F9A7" blockId="0.[151,635,1520,1578]" box="[151,635,1520,1544]" pageId="0" pageNumber="58">
<heading id="4B498189FF97FFCAE8A6EEFE6539F9A7" bold="true" box="[151,635,1520,1544]" fontSize="10" level="2" pageId="0" pageNumber="58" reason="2">
<emphasis id="22CAEAF7FF97FFCAE8A6EEFE6539F9A7" bold="true" box="[151,635,1520,1544]" pageId="0" pageNumber="58">
<taxonomicName id="D7BE4D66FF97FFCAE8A6EEFE6558F9A7" ID-CoL="48R67" authority="Ma, Li and Chen" authorityName="Ma, Li and Chen" box="[151,538,1520,1544]" class="Insecta" family="Crabronidae" genus="Odontopsen" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="58" phylum="Arthropoda" rank="species" species="shii" status="sp. nov.">
<emphasis id="22CAEAF7FF97FFCAE8A6EEFE660BF9A7" bold="true" box="[151,329,1520,1544]" italics="true" pageId="0" pageNumber="58">Odontopsen shii</emphasis>
,
<docAuthor id="BBD63768FF97FFCAE966EEFE66E1F9A7" box="[343,419,1520,1544]" pageId="0" pageNumber="58">Ma, Li</docAuthor>
and Chen
</taxonomicName>
,
<taxonomicNameLabel id="39F9578CFF97FFCAEA17EEFE6539F9A7" box="[550,635,1520,1544]" pageId="0" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="100136E5FF97FFCAE8A6ED1D664EF985" blockId="0.[151,635,1520,1578]" box="[151,268,1555,1578]" pageId="0" pageNumber="58">
(
<figureCitation id="88852A60FF97FFCAE8AEED1D6641F985" box="[159,259,1555,1578]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Figs. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="58A4656EFF97FFCBE8A6ED566603FD18" lastPageId="1" lastPageNumber="59" pageId="0" pageNumber="58" type="description">
<paragraph id="100136E5FF97FFCAE8A6ED56642BF91A" blockId="0.[151,1437,1624,2029]" pageId="0" pageNumber="58">Female. Body length 8.7 mm. Black; flagellomeres IV–XI yellowish ventrally. Setae on labrum yellowish, dense and long; on mandible yellowish, sparse and short; on lower frons silvery, dense and short; on vertex brown, sparse and long, on mesopleuron and propodeum slightly yellowish, dense and long.</paragraph>
<paragraph id="100136E5FF97FFCBE8F7EDCE6643FF00" blockId="0.[151,1437,1624,2029]" lastBlockId="1.[151,1437,152,1111]" lastPageId="1" lastPageNumber="59" pageId="0" pageNumber="58">
Clypeus broad and polish, gradually impressed toward its middle, which is much lower than the laterals; free margin of clypeus with two broad, round prominences medially; each prominence with oval tubercle near apex, tubercle shiny and markedly elevated (
<figureCitation id="88852A60FF97FFCAE995EC0866A7F8B2" box="[420,485,1798,1821]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Fig. 1</figureCitation>
). Mandible inconspicuously bidentate apically (
<figureCitation id="88852A60FF97FFCAEC33EC086306F8B2" box="[1026,1092,1798,1821]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Fig. 1</figureCitation>
). Interantennal tubercle small, round, conspicuous (
<figureCitation id="88852A60FF97FFCAE9B4EC26668BF890" box="[389,457,1832,1855]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Fig. 1</figureCitation>
). Upper frons with dense minute punctures that are 1× diameter apart. Frontal carina incomplete. Vertex smooth, posterior area deeply emarginated (
<figureCitation id="88852A60FF97FFCAEB0AEC45643EF8CD" box="[827,892,1867,1890]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Fig. 2</figureCitation>
), with dense and very slender transverse striations. Inner orbits in frontal view markedly divergent below (
<figureCitation id="88852A60FF97FFCAEB34EC606405F82A" box="[773,839,1902,1925]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Fig. 1</figureCitation>
). Head in profile with occipital carina produced into conspicuous dentate process, outer margin of process with sparse, sturdy and longitudinal rugae (
<figureCitation id="88852A60FF97FFCAECAFEC9E6246F808" box="[1182,1284,1936,1959]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="0" pageNumber="58">Figs. 3, 4</figureCitation>
). HW: HLD: HLF = 104: 37: 55; TW: EW: EL = 33: 19: 47; POD: OOD: OCD: DEL: DEU = 11: 19: 21: 75: 61; AOD: WAS: IAD = 19: 7: 16. Relative length of flagellomeres I–IV: 20, 14, 13, 12. Length of flagellomere I: width at apex = 20: 4 (
<figureCitation id="88852A60FF96FFCBE882EB9667B1FF00" box="[179,243,152,175]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="1" pageNumber="59">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="100136E5FF96FFCBE8F7EBB4660CFE6B" blockId="1.[151,1437,152,1111]" pageId="1" pageNumber="59">
Pronotum with several short transverse rugae, and apicolateral process (
<figureCitation id="88852A60FF96FFCBEBCFEBB46305FF7E" box="[1022,1095,186,209]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="1" pageNumber="59">Fig. 6</figureCitation>
). Scutum with sparse, minute punctures that are 1–4× diameters apart. Scutellum and metanotum smooth, shiny, and nearly impunctate. Mesopleuron shiny, smooth and impunctate, with very slender, sparse, short oblique transverse striations posteriorly; hypoepimeral area with very slender, dense longitudinal rugae. Propodeal enclosure with shallow depression, and slender, longitudinal rugae; propodeal pad broad, polished; posterior surface of propodeum with very slender, dense, regular, longitudinal rugae. Outer surface of hind tibia with row of yellowish sturdy spines. First recurrent vein of forewing ending in second submarginal cell; second recurrent vein ending in second submarginal cell or interstitial between second and third submarginal cell.
</paragraph>
<paragraph id="100136E5FF96FFCBE8F7EADE64D0FD83" blockId="1.[151,1437,152,1111]" pageId="1" pageNumber="59">
Gastral segments polished. Petiole slender, long, smooth, shiny; dorsal surface with a very broad longitudinal carina medially and ventral surface without keel. PL: PW: LTI: WTI: HFL: HTL = 50: 9: 42: 36: 58: 67. Pygidial area broadly triangular, with dense, coarse punctures and with setae (
<figureCitation id="88852A60FF96FFCBEB72E91B64C6FD83" box="[835,900,533,556]" captionStart="FIGURES 1 – 7" captionStartId="1.[151,269,1850,1874]" captionTargetBox="[153,1434,1184,1815]" captionTargetId="figure@1.[151,1436,1165,1826]" captionTargetPageId="1" captionText="FIGURES 1 – 7. Odontopsen shii, Ma, Li and Chen, sp. nov., female. (1) Frontal view of head; (2) dorsal view of head; (3) lateral view of head; (4) ventral view of occipital process; (5) lateral view of antennal flagellomere; (6) dorsal view of pronotum; (7) dorsal view of pygidial plate. Scale bars: Figs. 1 – 5: 1 mm; Fig. 6: 1.48 mm; Fig. 7: 1.89 mm." httpUri="https://zenodo.org/record/193612/files/figure.png" pageId="1" pageNumber="59">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="100136E5FF96FFCBE8F7E936663EFDE0" blockId="1.[151,1437,152,1111]" box="[198,380,568,591]" pageId="1" pageNumber="59">Male: Unknown.</paragraph>
<paragraph id="100136E5FF96FFCBE8F7E9546603FD18" blockId="1.[151,1437,152,1111]" pageId="1" pageNumber="59">
<emphasis id="22CAEAF7FF96FFCBE8F7E95466F3FDDD" bold="true" box="[198,433,602,626]" pageId="1" pageNumber="59">Specimen examined.</emphasis>
<typeStatus id="CF058847FF96FFCBE988E9546562FDDE" box="[441,544,602,625]" pageId="1" pageNumber="59" type="holotype">Holotype</typeStatus>
: female,
<collectingCountry id="68A97675FF96FFCBEAB5E954658AFDDE" box="[644,712,602,625]" name="China" pageId="1" pageNumber="59">China</collectingCountry>
: Tibet, Nyingchi Prefecture, Lulang, 29°30ʹN 94°18ʹE,
<date id="64001025FF96FFCBED12E95462DAFDDE" box="[1315,1432,602,625]" pageId="1" pageNumber="59" value="2007-09-19">19.ix.2007</date>
, coll. Fu-ming Shi, deposited in the Insect Collections of Yunnan Agricultural University, Kunming, Yunnan Province,
<collectingCountry id="68A97675FF96FFCBE8A6E9AE6795FD18" box="[151,215,672,695]" name="China" pageId="1" pageNumber="59">China</collectingCountry>
(YAUK).
</paragraph>
</subSubSection>
<subSubSection id="58A4656EFF96FFCBE8F7E9CC66B4FD76" box="[198,502,706,730]" pageId="1" pageNumber="59" type="distribution">
<paragraph id="100136E5FF96FFCBE8F7E9CC66B4FD76" blockId="1.[151,1437,152,1111]" box="[198,502,706,730]" pageId="1" pageNumber="59">
<emphasis id="22CAEAF7FF96FFCBE8F7E9CC661BFD75" bold="true" box="[198,345,706,730]" pageId="1" pageNumber="59">Distribution.</emphasis>
<collectingCountry id="68A97675FF96FFCBE96EE9CC66DDFD76" box="[351,415,706,729]" name="China" pageId="1" pageNumber="59">China</collectingCountry>
(Tibet).
</paragraph>
</subSubSection>
<subSubSection id="58A4656EFF96FFCBE8F7E9EA6670FBF8" pageId="1" pageNumber="59" type="etymology">
<paragraph id="100136E5FF96FFCBE8F7E9EA626DFD53" blockId="1.[151,1437,152,1111]" box="[198,1327,740,764]" pageId="1" pageNumber="59">
<emphasis id="22CAEAF7FF96FFCBE8F7E9EA6605FD53" bold="true" box="[198,327,740,764]" pageId="1" pageNumber="59">Etymology.</emphasis>
The species is named in honour of Prof. Fu-ming Shi, the collector of the
<typeStatus id="CF058847FF96FFCBEC6EE9EB63FEFD53" box="[1119,1212,741,764]" pageId="1" pageNumber="59" type="holotype">holotype</typeStatus>
specimen.
</paragraph>
<paragraph id="100136E5FF96FFCBE8F7E8096626FBBE" blockId="1.[151,1437,152,1111]" pageId="1" pageNumber="59">
<emphasis id="22CAEAF7FF96FFCBE8F7E8096640FCB0" bold="true" box="[198,258,775,799]" pageId="1" pageNumber="59">Note.</emphasis>
This new species shares with the
<typeStatus id="CF058847FF96FFCBEA4FE80665EEFCB0" box="[638,684,776,799]" pageId="1" pageNumber="59">type</typeStatus>
species of
<taxonomicName id="D7BE4D66FF96FFCBEB1AE80664EEFCB0" box="[811,940,776,799]" class="Insecta" family="Crabronidae" genus="Odontopsen" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="22CAEAF7FF96FFCBEB1AE80664EEFCB0" box="[811,940,776,799]" italics="true" pageId="1" pageNumber="59">Odontopsen</emphasis>
</taxonomicName>
with the following distinguishing characters: Frontal carina incomplete below; occipital carina reaching hypostomal carina far from mid-ventral line of head, lower part of carina produced obviously in male but developed into large process in female; pronotum with several oblique lateral carinules but without complete ridge like carina; omaulus curving toward prothorax, without acetabular carina; scrobal sulcus distinct but not deeply impressed, hypoepimeral area weakly reticulate but not markedly bulging; forewing recurrent veins ending at submarginal cells I and II or both at II; hindwing medial vein diverging before cu-a; propodeum largely smooth, only some areas striate; petiole dorsally with median longitudinal carina; female pygidial plate subtriangular.
</paragraph>
<paragraph id="100136E5FF96FFCBE8F7EF136670FBF8" blockId="1.[151,1437,152,1111]" pageId="1" pageNumber="59">
This new species can be distinguished from the
<typeStatus id="CF058847FF96FFCBEAF7EF1365B6FB9B" box="[710,756,1053,1076]" pageId="1" pageNumber="59">type</typeStatus>
species,
<taxonomicName id="D7BE4D66FF96FFCBEB67EF13633CFB9B" authority="Tsuneki, 1964" authorityName="Tsuneki" authorityYear="1964" box="[854,1150,1053,1076]" class="Insecta" family="Crabronidae" genus="Odontopsen" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="59" phylum="Arthropoda" rank="species" species="hanedai">
<emphasis id="22CAEAF7FF96FFCBEB67EF13648EFB9B" box="[854,972,1053,1076]" italics="true" pageId="1" pageNumber="59">O. hanedai</emphasis>
(
<bibRefCitation id="742F4B14FF96FFCBEBEBEF136334FB9B" author="Tsuneki" box="[986,1142,1053,1076]" pageId="1" pageNumber="60" refString="Tsuneki, K. (1964) An interesting new species of the genus Psen captured in Japan with the erection of a new subgenus (Hym.: Sphec.: Pemphredoninae). Insecta Matsumurana, 27, 12 - 15." type="journal article" year="1964">Tsuneki, 1964</bibRefCitation>
)
</taxonomicName>
, by the characters given in the key below.
</paragraph>
</subSubSection>
</treatment>
</document>
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