treatments-xml/data/7F/31/B5/7F31B537FF93FFFAFE8CFB01B2B8FC96.xml

<document id="31F2CE53AF854C07F1A2660F37C395CD" ID-DOI="10.35249/rche.46.2.20.18" ID-ISSN="0718-8994" ID-Zenodo-Dep="13205002" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722661066298" checkinUser="felipe" docAuthor="Guerrero, Marcelo &amp; Aceituno, Guillermo" docDate="2020" docId="7F31B537FF93FFFAFE8CFB01B2B8FC96" docLanguage="es" docName="RevChilEntomol.46.2.273-290.pdf" docOrigin="Revista Chilena de Entomología (Rev. Chil. Entomol.) 46 (2)" docSource="http://dx.doi.org/10.35249/rche.46.2.20.18" docStyle="DocumentStyle:451321A1F3358405F3D5797D4FC35865.3:RevChilEntomol.2020-.journal_article" docStyleId="451321A1F3358405F3D5797D4FC35865" docStyleName="RevChilEntomol.2020-.journal_article" docStyleVersion="3" docTitle="Gyriosomus maculatus Guerrero y Aceituno 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="278" masterDocId="8308CD4FFF90FFFFFFCEFF9CB03CFFD9" masterDocTitle="Nuevas especies del género Gyriosomus Guérin-Méneville (Coleoptera: Tenebrionidae: Nycteliini) de Chile, y nuevo estatus para Gyriosomus faveopunctatus laevis Kulzer" masterLastPageNumber="290" masterPageNumber="273" pageNumber="276" updateTime="1722700527194" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="695F37F66ED3120D9AAEB1EE25D58D62">Nuevas especies del género Gyriosomus Guérin-Méneville (Coleoptera: Tenebrionidae: Nycteliini) de Chile, y nuevo estatus para Gyriosomus faveopunctatus laevis Kulzer</mods:title>
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<mods:namePart id="4FF00151E3A749447C57A90134ADA3FC">Guerrero, Marcelo</mods:namePart>
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<mods:namePart id="D13966F6593AD84C6C7A362B572E6432">Aceituno, Guillermo</mods:namePart>
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<mods:title id="B99C21B2E1F7217CFED5ACE1B7170990">Revista Chilena de Entomología</mods:title>
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<mods:date id="4977DCB408414A926D5222983531C555">2020</mods:date>
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<mods:title id="B2198134B93EA5F2C122593E4C73D1A4">Rev. Chil. Entomol.</mods:title>
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<subSubSection id="BF8257AAFF93FFFCFE8CFB01B3A9FB6C" box="[322,917,1181,1205]" pageId="3" pageNumber="276" type="nomenclature">
<paragraph id="F7270421FF93FFFCFE8CFB01B3A9FB6C" blockId="3.[322,917,1181,1237]" box="[322,917,1181,1205]" pageId="3" pageNumber="276">
<heading id="AC6FB34DFF93FFFCFE8CFB01B3A9FB6C" box="[322,917,1181,1205]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="276" reason="4">
<taxonomicName id="30987FA2FF93FFFCFE8CFB01B308FB6C" authority="Guerrero y Aceituno" authorityName="Guerrero y Aceituno" authorityYear="2020" box="[322,820,1181,1205]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="276" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="C5ECD833FF93FFFCFE8CFB01B207FB6C" box="[322,571,1181,1205]" italics="true" pageId="3" pageNumber="276">Gyriosomus maculatus</emphasis>
Guerrero y Aceituno
</taxonomicName>
<emphasis id="C5ECD833FF93FFFCFCF4FB01B3A9FB6C" bold="true" box="[826,917,1181,1205]" pageId="3" pageNumber="276">
<taxonomicNameLabel id="DEDF6548FF93FFFCFCF4FB01B3A9FB6C" box="[826,917,1181,1205]" pageId="3" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BF8257AAFF93FFFCFDC1FB21B2F4FB0C" box="[527,712,1213,1237]" pageId="3" pageNumber="276" type="description">
<paragraph id="F7270421FF93FFFCFDC1FB21B2F4FB0C" blockId="3.[322,917,1181,1237]" box="[527,712,1213,1237]" pageId="3" pageNumber="276">
(
<figureCitation id="6FA318A4FF93FFFCFDD9FB21B258FB0C" box="[535,612,1213,1237]" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="3" pageNumber="276">Figs. 2</figureCitation>
A-2H, 9)
</paragraph>
</subSubSection>
<subSubSection id="BF8257AAFF93FFFBFF94FB60B45DFC34" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276" type="diagnosis">
<paragraph id="F7270421FF93FFFCFF94FB60B438FA0C" blockId="3.[90,1149,1276,1493]" pageId="3" pageNumber="276">
<emphasis id="C5ECD833FF93FFFCFF94FB60B0E0FACD" bold="true" box="[90,220,1276,1300]" pageId="3" pageNumber="276">Diagnosis.</emphasis>
Cabeza, pronoto, superficie dorsal de los élitros y pseudopleurón negros brillantes. Faz ventral negra, ligeramente menos brillante que la superficie elitral. Patas con los fémures y tibias negras, tarsos y espolones tibiales marrones oscuros. Antenas negras. Pronoto glabro, convexo, finamente punteado, moderadamente rugoso y realzado en el centro formando una gibosidad; margen anterior casi recto, no visible en vista dorsal (
<figureCitation id="6FA318A4FF93FFFCFB9EFAE1B040FA6C" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="3" pageNumber="276">Fig. 2B</figureCitation>
). Élitros con impresiones tegumentarias grandes y profundas cubiertas de pilosidad blanca. Abdomen con escasos mechones de setas en el primer ventrito (
<figureCitation id="6FA318A4FF93FFFCFC6FFA21B3CBFA0C" box="[929,1015,1469,1493]" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="3" pageNumber="276">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="F7270421FF93FFFBFF94FA60B29CFD6E" blockId="3.[90,1149,1532,1749]" lastBlockId="4.[151,1210,125,695]" lastPageId="4" lastPageNumber="277" pageId="3" pageNumber="276">
<emphasis id="C5ECD833FF93FFFCFF94FA60B0C8F9CD" bold="true" box="[90,244,1532,1556]" pageId="3" pageNumber="276">Descripción.</emphasis>
<typeStatus id="2823BA83FF93FFFCFECCFA61B150F9CC" box="[258,364,1533,1557]" pageId="3" pageNumber="276" type="holotype">Holotipo</typeStatus>
macho. Largo: 
<quantity id="3060A9C4FF93FFFCFDF6FA61B29AF9CD" box="[568,678,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="3" pageNumber="276" unit="mm" value="19.5">19,5 mm</quantity>
; ancho: 
<quantity id="3060A9C4FF93FFFCFCDAFA61B3BEF9CD" box="[788,898,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="276" unit="mm" value="11.5">11,5 mm</quantity>
. 
<emphasis id="C5ECD833FF93FFFCFC5AFA60B3CFF9CD" bold="true" box="[916,1011,1532,1556]" pageId="3" pageNumber="276">Cabeza:</emphasis>
Frente sin mechón de setas en el margen interno del ojo. Antenas extendidas hacia atrás sobrepasan la base del pronoto con los dos últimos segmentos; antenito 11 cubierto con un parche sensorial tomentoso. 
<emphasis id="C5ECD833FF93FFFCFE97F9C1B1FCF9AC" bold="true" box="[345,448,1629,1653]" pageId="3" pageNumber="276">Pronoto:</emphasis>
Glabro, con puntuación pequeña homogénea, sólo visible bajo aumento de 10x, moderadamente rugoso; disco convexo, con una gibosidad central inclinada anteriormente, y un surco longitudinal medial profundo que va desde el margen anterior del pronoto hasta la zona media discal; bordes laterales ligeramente curvos; margen anterior delgado y liso, casi recto; margen posterior bisinuado, delgado y liso; margen lateral con una hilera de setas cortas en la ranura formada por el doble margen lateral; ángulos posteriores proyectados hacia atrás, no sobrepasando los húmeros; ancho máximo en la base, 2,3 veces más ancho que largo. Prosterno abovedado, más realzado que el mesosterno, sin apófisis posterior sobresaliente hacia atrás; margen anterior curvo. Mesosterno moderadamente rugoso, con escasas setas en la superficie. Metasterno liso, ligeramente puntuado, con escasas setas. 
<emphasis id="C5ECD833FF94FFFBFD42FEA4B2DFFE8C" bold="true" box="[652,739,312,341]" pageId="4" pageNumber="277">Élitros:</emphasis>
Negros, dorsalmente lisos y brillantes, con abundantes impresiones tegumentarias grandes y profundas, y puntuaciones cubiertas con pilosidad blanca; sutura elitral notoria, no realzada; margen lateral moderadamente grueso, punteado y rugoso, dividido por un surco notorio que se extiende hasta el comienzo del declive elitral, desde donde se vuelve simple, alcanzando el tercio posterior con una notoria banda longitudinal blanca que se interrumpe, la cual alcanza el extremo apical; pseudopleurón liso y brillante, con una banda basal longitudinal pilosa poco notoria e interrumpida que va desde la zona media hasta el extremo apical; epipleura lisa y glabra. 
<emphasis id="C5ECD833FF94FFFBFF59FDA3B0E2FD8E" bold="true" box="[151,222,575,599]" pageId="4" pageNumber="277">Patas:</emphasis>
Fémures pubescente en la superficie ventral, con setas más densas en los profemures; tibias con setas largas y semidecumbentes escasas, más densas en el margen interno de las protibias. 
<emphasis id="C5ECD833FF94FFFBFEF6FDE3B184FD4E" bold="true" box="[312,440,639,663]" pageId="4" pageNumber="277">Abdomen:</emphasis>
Primer ventrito con escasos mechones de setas. 
<emphasis id="C5ECD833FF94FFFBFBCEFDE5B120FD6E" bold="true" pageId="4" pageNumber="277">Órgano genital masculino:</emphasis>
Edeago como en las 
<figureCitation id="6FA318A4FF94FFFBFDDCFD03B262FD6E" box="[530,606,671,695]" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="4" pageNumber="277">Figs. 2</figureCitation>
F-2H.
</paragraph>
<paragraph id="F7270421FF94FFFBFF59FD48B48BFCD4" blockId="4.[151,1209,719,781]" pageId="4" pageNumber="277">
<emphasis id="C5ECD833FF94FFFBFF59FD48B0C1FD34" bold="true" box="[151,253,721,749]" pageId="4" pageNumber="277">
<typeStatus id="2823BA83FF94FFFBFF59FD48B0C8FD35" box="[151,244,724,748]" pageId="4" pageNumber="277" type="alotype">Alotipo</typeStatus>
.
</emphasis>
Largo: 
<quantity id="3060A9C4FF94FFFBFE9BFD48B186FD35" box="[341,442,724,748]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="277" unit="mm" value="25.5">25,5 mm</quantity>
; ancho: 
<quantity id="3060A9C4FF94FFFBFDD6FD48B242FD35" box="[536,638,724,748]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="277" unit="mm" value="14.5">14,5 mm</quantity>
. Élitros con el margen lateral más ancho, con una banda marginal completa en el declive elitral. Primer ventrito abdominal glabro (
<figureCitation id="6FA318A4FF94FFFBFB9CFD69B497FCD4" box="[1106,1195,757,781]" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="4" pageNumber="277">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="F7270421FF94FFFBFF59FCA8B19FFC74" blockId="4.[151,1210,820,941]" pageId="4" pageNumber="277">
<emphasis id="C5ECD833FF94FFFBFF59FCA8B1B6FC94" bold="true" box="[151,394,820,845]" pageId="4" pageNumber="277">
Variación 
<typeStatus id="2823BA83FF94FFFBFEDFFCA9B1B9FC94" box="[273,389,821,845]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
.
</emphasis>
Machos. Largo: 16,0- 
<quantity id="3060A9C4FF94FFFBFD4DFCA9B2D7FC95" box="[643,747,821,845]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="4" pageNumber="277" unit="mm" value="20.5">20,5 mm</quantity>
; ancho: 
<quantity id="3060A9C4FF94FFFBFC84FCA9B3D5FC95" box="[842,1001,821,845]" metricMagnitude="-2" metricUnit="m" metricValue="1.2449999999999999" metricValueMax="1.55" metricValueMin="0.94" pageId="4" pageNumber="277" unit="mm" value="12.45" valueMax="15.5" valueMin="9.4">9,40-15,5 mm</quantity>
. Hembras. Largo: 21,0- 
<quantity id="3060A9C4FF94FFFBFF00FCC9B106FCB5" box="[206,314,853,877]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="4" pageNumber="277" unit="mm" value="25.5">25,5 mm</quantity>
; ancho: 14,0- 
<quantity id="3060A9C4FF94FFFBFE14FCC9B27BFCB5" box="[474,583,853,877]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="277" unit="mm" value="15.5">15,5 mm</quantity>
. El mayor tamaño de las hembras y la forma más angosta de los machos corresponde al dimorfismo sexual caracterÍstico de los integrantes del género 
<taxonomicName id="30987FA2FF94FFFBFED4FC0AB1A0FC74" box="[282,412,918,941]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="genus">
<emphasis id="C5ECD833FF94FFFBFED4FC0AB1A0FC74" box="[282,412,918,941]" italics="true" pageId="4" pageNumber="277">Gyriosomus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F7270421FF94FFFBFF59FC49B45DFC34" blockId="4.[151,1121,981,1006]" box="[151,1121,981,1006]" pageId="4" pageNumber="277">
<emphasis id="C5ECD833FF94FFFBFF59FC49B187FC37" bold="true" box="[151,443,981,1006]" pageId="4" pageNumber="277">Distribución geográfica.</emphasis>
<collectingCountry id="8F8F44B1FF94FFFBFE0CFC49B23FFC34" box="[450,515,981,1005]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
, 
<collectingRegion id="355CCAC3FF94FFFBFDC0FC49B2C9FC37" box="[526,757,981,1006]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Región de Atacama</collectingRegion>
, Provincia de Huasco (
<figureCitation id="6FA318A4FF94FFFBFBCDFC49B468FC34" box="[1027,1108,981,1005]" captionStart="Figura 15" captionStartId="16.[331,405,1034,1056]" captionText="Figura 15. Distribución de las especies de Gyriosomus en Chile." pageId="4" pageNumber="277">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BF8257AAFF94FFFAFF59FB8AB2B8FC96" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277" type="materials_examined">
<paragraph id="F7270421FF94FFFBFF59FB8AB33DF908" blockId="4.[151,1210,1046,1745]" pageId="4" pageNumber="277">
<emphasis id="C5ECD833FF94FFFBFF59FB8AB17CFBF7" bold="true" box="[151,320,1046,1070]" pageId="4" pageNumber="277">Material tipo.</emphasis>
<materialsCitation id="47F00E7CFF94FFFBFE84FB8AB144FB57" collectingDate="1998-10-15" collectionCode="MNNC" collectorName="Leg. M. Guerrero" country="Chile" determinerName="Guerrero" location="Huasco" pageId="4" pageNumber="277" specimenCount="1" specimenCount-male="1" typeStatus="Holotipo">
<typeStatus id="2823BA83FF94FFFBFE84FB8AB188FBF7" box="[330,436,1046,1070]" pageId="4" pageNumber="277" type="holotype">Holotipo</typeStatus>
macho en 
<collectionCode id="91899CE4FF94FFFBFDF3FB8AB2A4FBF7" box="[573,664,1046,1070]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="4" pageNumber="277">MNNC</collectionCode>
, etiquetado: a) “ 
<collectingCountry id="8F8F44B1FF94FFFBFC91FB8AB39DFBF7" box="[863,929,1046,1070]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
<location id="F24752FAFF94FFFBFC64FB8AB43FFBF4" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFC64FB8AB43FFBF4" box="[938,1027,1046,1069]" country="Chile" name="Huasco" pageId="4" pageNumber="277">Huasco</location>
/ 
<locationDeviation id="E6951D03FF94FFFBFBE6FB8AB0CCFB94" location="Huasco" pageId="4" pageNumber="277">
<quantity id="3060A9C4FF94FFFBFBE6FB8AB44EFBF7" box="[1064,1138,1046,1070]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="277" unit="km" value="13.0">13 km</quantity>
N de Huasco
</locationDeviation>
/ 
<date id="832622E1FF94FFFBFEC1FBAAB1BEFB97" box="[271,386,1078,1102]" pageId="4" pageNumber="277" value="1998-10-15">
<collectingDate id="9362DB09FF94FFFBFEC1FBAAB1BEFB97" box="[271,386,1078,1102]" pageId="4" pageNumber="277" value="1998-10-15">15-X-1998</collectingDate>
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFE6EFBAAB24DFB97" box="[416,625,1078,1102]" pageId="4" pageNumber="277">Leg. M. Guerrero</collectorName>
”, b) “ 
<typeStatus id="2823BA83FF94FFFBFD61FBAAB37DFB97" box="[687,833,1078,1102]" pageId="4" pageNumber="277" type="holotype">HOLOTIPO</typeStatus>
/ 
<taxonomicName id="30987FA2FF94FFFBFCAEFBAAB465FB97" authorityName="Guerrero &amp; Aceituno" authorityYear="2020" box="[864,1113,1078,1102]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="C5ECD833FF94FFFBFCAEFBAAB465FB97" box="[864,1113,1078,1102]" italics="true" pageId="4" pageNumber="277">Gyriosomus maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEDF6548FF94FFFBFBAEFBAAB486FB94" box="[1120,1210,1078,1101]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
/ det. 
<determinerName id="02FBE8E5FF94FFFBFF2AFBCAB173FBB7" box="[228,335,1110,1134]" pageId="4" pageNumber="277">Guerrero</determinerName>
y Aceituno 2020”, c) Genital masculino montado en tarjeta de cartón con sÍmbolo 
<specimenCount id="E19ECFA8FF94FFFBFF35FBEBB133FB48" box="[251,271,1143,1169]" count="1" pageId="4" pageNumber="277" type="male">♂</specimenCount>
impreso
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFE4EFBEAB3D7FB17" collectingDate="1998-10-15" collectionCode="MNNC" collectorName="Leg. M. Guerrero" country="Chile" determinerName="Guerrero" location="Huasco" pageId="4" pageNumber="277" specimenCount="1" specimenCount-female="1" typeStatus="Alotipo">
<typeStatus id="2823BA83FF94FFFBFE4EFBEAB1E6FB57" box="[384,474,1142,1166]" pageId="4" pageNumber="277" type="alotype">Alotipo</typeStatus>
en 
<collectionCode id="91899CE4FF94FFFBFDCCFBEAB261FB57" box="[514,605,1142,1166]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="4" pageNumber="277">MNNC</collectionCode>
, etiquetado: a) “ 
<collectingCountry id="8F8F44B1FF94FFFBFCD6FBEAB366FB57" box="[792,858,1142,1166]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
<location id="F24752FAFF94FFFBFCAEFBEBB385FB57" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFCAEFBEBB385FB57" box="[864,953,1143,1166]" country="Chile" name="Huasco" pageId="4" pageNumber="277">Huasco</location>
/ 
<locationDeviation id="E6951D03FF94FFFBFC1BFBEAB485FB57" box="[981,1209,1142,1166]" location="Huasco" pageId="4" pageNumber="277">
<quantity id="3060A9C4FF94FFFBFC1BFBEAB427FB57" box="[981,1051,1142,1166]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="277" unit="km" value="13.0">13 km</quantity>
N de Huasco
</locationDeviation>
/ 
<date id="832622E1FF94FFFBFF7FFB0AB119FB77" box="[177,293,1174,1198]" pageId="4" pageNumber="277" value="1998-10-15">
<collectingDate id="9362DB09FF94FFFBFF7FFB0AB119FB77" box="[177,293,1174,1198]" pageId="4" pageNumber="277" value="1998-10-15">15-X-1998</collectingDate>
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFE87FB0BB21CFB77" box="[329,544,1174,1199]" pageId="4" pageNumber="277">Leg. M. Guerrero</collectorName>
”, b) “ 
<typeStatus id="2823BA83FF94FFFBFDABFB0AB2DDFB77" box="[613,737,1174,1198]" pageId="4" pageNumber="277" type="alotype">ALOTIPO</typeStatus>
/ 
<taxonomicName id="30987FA2FF94FFFBFCCBFB0BB43EFB77" authorityName="Guerrero &amp; Aceituno" authorityYear="2020" box="[773,1026,1174,1198]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="C5ECD833FF94FFFBFCCBFB0BB43EFB77" box="[773,1026,1174,1198]" italics="true" pageId="4" pageNumber="277">Gyriosomus maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEDF6548FF94FFFBFBC2FB0BB455FB77" box="[1036,1129,1175,1198]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
/ det. 
<determinerName id="02FBE8E5FF94FFFBFF59FB2BB13EFB16" box="[151,258,1207,1231]" pageId="4" pageNumber="277">Guerrero</determinerName>
y Aceituno 2020”, c) tarjeta de cartón con sÍmbolo 
<specimenCount id="E19ECFA8FF94FFFBFCA5FB2BB343FB08" box="[875,895,1207,1233]" count="1" pageId="4" pageNumber="277" type="female">♀</specimenCount>
impreso
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFC36FB2AB49EFB36" collectingDate="1991-11-03" collectionCode="CSRR" collectorName="G. Castillo" country="Chile" location="Caldera" pageId="4" pageNumber="277" specimenCount="2" typeStatus="Paratipos">
<typeStatus id="2823BA83FF94FFFBFC36FB2AB454FB17" box="[1016,1128,1206,1230]" pageId="4" pageNumber="277" type="paratype">Paratipos</typeStatus>
(62). 2 
<typeStatus id="2823BA83FF94FFFBFF59FB4BB13BFB36" box="[151,263,1239,1263]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “CHILE- 
<location id="F24752FAFF94FFFBFE67FB4BB23AFB36" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFE67FB4BB23AFB36" box="[425,518,1239,1263]" country="Chile" name="Caldera" pageId="4" pageNumber="277">Caldera</location>
/ El Morro Sur / 
<date id="832622E1FF94FFFBFD10FB4BB370FB36" box="[734,844,1239,1263]" pageId="4" pageNumber="277" value="1991-11-03">
<collectingDate id="9362DB09FF94FFFBFD10FB4BB370FB36" box="[734,844,1239,1263]" pageId="4" pageNumber="277" value="1991-11-03">3-11-1991</collectingDate>
</date>
/ coll: 
<collectorName id="5A6D61F7FF94FFFBFC6CFB4BB41EFB36" box="[930,1058,1239,1263]" pageId="4" pageNumber="277">G. Castillo</collectorName>
” (2 
<collectionCode id="91899CE4FF94FFFBFB9FFB4BB4A6FB36" box="[1105,1178,1239,1263]" pageId="4" pageNumber="277">CSRR</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFB62FB4BB49EFAD6" collectionCode="R, CSRR" country="Chile" county="Caldera" location="Sur de BahIa Inglesa" municipality="Chorrillos" pageId="4" pageNumber="277" specimenCount="1" typeStatus="Paratipos">
1 
<typeStatus id="2823BA83FF94FFFBFF59FB6BB0C7FAD6" box="[151,251,1271,1295]" pageId="4" pageNumber="277" type="paratype">paratipo</typeStatus>
de: “CHILE- 
<collectingCounty id="1E467CADFF94FFFBFE53FB6BB1C6FAD6" box="[413,506,1271,1295]" pageId="4" pageNumber="277">Caldera</collectingCounty>
/ 
<collectingMunicipality id="17439E5BFF94FFFBFDD6FB6BB2B3FAD6" box="[536,655,1271,1295]" pageId="4" pageNumber="277">Chorrillos</collectingMunicipality>
/ 
<location id="F24752FAFF94FFFBFD60FB6BB39CFAD6" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFD60FB6BB39CFAD6" box="[686,928,1265,1295]" country="Chile" county="Caldera" municipality="Chorrillos" name="Sur de BahIa Inglesa" pageId="4" pageNumber="277">Sur de BahÍa Inglesa</location>
/ 
<collectionCode id="91899CE4FF94FFFBFC73FB6BB3EEFAD7" box="[957,978,1271,1294]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="277">R</collectionCode>
. Barrera” (
<collectionCode id="91899CE4FF94FFFBFB9EFB6BB4A6FAD6" box="[1104,1178,1271,1295]" pageId="4" pageNumber="277">CSRR</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFB62FB6BB338FAF6" collectingDate="1961-11" collectionCode="PVGH" collectorName="L. Pena" country="Chile" location="Atacama" pageId="4" pageNumber="277" specimenCount="1" stateProvince="Atacama" typeStatus="Paratipos">
1 
<typeStatus id="2823BA83FF94FFFBFF59FA8BB0C7FAF6" box="[151,251,1303,1327]" pageId="4" pageNumber="277" type="paratype">paratipo</typeStatus>
de: “ 
<collectingRegion id="355CCAC3FF94FFFBFEF7FA8BB199FAF6" box="[313,421,1303,1327]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Atacama</collectingRegion>
/ 
<date id="832622E1FF94FFFBFE08FA8BB21BFAF6" box="[454,551,1303,1327]" pageId="4" pageNumber="277" value="1961-11">
<collectingDate id="9362DB09FF94FFFBFE08FA8BB21BFAF6" box="[454,551,1303,1327]" pageId="4" pageNumber="277" value="1961-11">XI. 1961</collectingDate>
</date>
. 
<collectorName id="5A6D61F7FF94FFFBFDFDFA8BB2B3FAF6" box="[563,655,1303,1327]" pageId="4" pageNumber="277">L. Peña</collectorName>
” (
<collectionCode id="91899CE4FF94FFFBFD67FA8BB2C0FAF6" box="[681,764,1303,1327]" pageId="4" pageNumber="277">PVGH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFCDEFA8BB48AFA96" collectingDate="1997-09-19" collectionCode="PVGH" collectorName="Leg. P. Vidal GH" country="Chile" elevation="300" location="Totoral" pageId="4" pageNumber="277" specimenCount="300" stateProvince="Atacama" typeStatus="Paratipos">
3 
<typeStatus id="2823BA83FF94FFFBFCEBFA8BB3A9FAF6" box="[805,917,1303,1327]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFC1CFA8BB414FAF6" box="[978,1064,1303,1327]" name="Chile" pageId="4" pageNumber="277">CHILE</collectingCountry>
<collectingRegion id="355CCAC3FF94FFFBFBE1FA8BB485FAF6" box="[1071,1209,1303,1327]" country="Chile" name="Atacama" pageId="4" pageNumber="277">ATACAMA</collectingRegion>
/ E 
<location id="F24752FAFF94FFFBFF04FAABB121FA96" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFF04FAABB121FA96" box="[202,285,1335,1359]" country="Chile" name="Totoral" pageId="4" pageNumber="277" stateProvince="Atacama">Totoral</location>
/ 
<date id="832622E1FF94FFFBFEF1FAABB1E7FA96" box="[319,475,1335,1359]" pageId="4" pageNumber="277" value="1997-09-19">
<collectingDate id="9362DB09FF94FFFBFEF1FAABB1E7FA96" box="[319,475,1335,1359]" pageId="4" pageNumber="277" value="1997-09-19">19 Sept. 1997</collectingDate>
</date>
- 
<specimenCount id="E19ECFA8FF94FFFBFE2CFAABB20BFA96" box="[482,567,1335,1359]" count="300" pageId="4" pageNumber="277" type="generic">
<quantity id="3060A9C4FF94FFFBFE2CFAABB20FFA96" box="[482,563,1335,1359]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="277" unit="m" value="300.0">
<elevation id="7CB5E312FF94FFFBFE2CFAABB20FFA96" box="[482,563,1335,1359]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="277" unit="m" value="300.0">300 m</elevation>
</quantity>
.
</specimenCount>
/ 27.53.52 s 70.53.32w / 
<collectorName id="5A6D61F7FF94FFFBFC92FAABB415FA96" box="[860,1065,1335,1359]" pageId="4" pageNumber="277">Leg. P. Vidal GH</collectorName>
” (3 
<collectionCode id="91899CE4FF94FFFBFB92FAABB492FA96" box="[1116,1198,1335,1359]" pageId="4" pageNumber="277">PVGH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFF59FACBB2B7FA56" collectingDate="2000-10-15" collectionCode="CGAA" collectorName="J. Mondaca" country="Chile" location="Carrizal Bajo." municipality="Provincia de Huasco" pageId="4" pageNumber="277" specimenCount="1" stateProvince="Atacama" typeStatus="Paratipos">
4 
<typeStatus id="2823BA83FF94FFFBFF61FACBB123FAB6" box="[175,287,1367,1391]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFEACFACBB19AFAB6" box="[354,422,1367,1391]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
, 
<collectingRegion id="355CCAC3FF94FFFBFE7BFACBB29EFAB6" box="[437,674,1367,1391]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Región de Atacama</collectingRegion>
/ 
<collectingMunicipality id="17439E5BFF94FFFBFD09FACBB3FFFAB6" box="[711,963,1367,1391]" pageId="4" pageNumber="277">Provincia de Huasco</collectingMunicipality>
/ 
<location id="F24752FAFF94FFFBFC26FACBB4B0FAB6" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFC26FACBB4B0FAB6" box="[1000,1164,1367,1391]" country="Chile" municipality="Provincia de Huasco" name="Carrizal Bajo." pageId="4" pageNumber="277" stateProvince="Atacama">Carrizal Bajo.</location>
<date id="832622E1FF94FFFBFB58FACBB0DBFA56" pageId="4" pageNumber="277" value="2000-10-15">
<collectingDate id="9362DB09FF94FFFBFB58FACBB0DBFA56" pageId="4" pageNumber="277" value="2000-10-15">15- X-2000</collectingDate>
</date>
/ Leg.: 
<collectorName id="5A6D61F7FF94FFFBFE80FAE4B1EAFA56" box="[334,470,1399,1423]" pageId="4" pageNumber="277">J. Mondaca</collectorName>
E.” (4 
<collectionCode id="91899CE4FF94FFFBFDE0FAEBB2BFFA56" box="[558,643,1399,1423]" pageId="4" pageNumber="277">CGAA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFD56FAE4B48AFA69" collectingDate="2002-10-22" collectionCode="CGAA" country="Chile" location="Carrizal Bajo." municipality="Provincia de Huasco" pageId="4" pageNumber="277" specimenCount="1" stateProvince="Atacama" typeStatus="Paratipos">
7 
<typeStatus id="2823BA83FF94FFFBFD7EFAEBB31CFA56" box="[688,800,1399,1423]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFCAFFAEBB399FA56" box="[865,933,1399,1423]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
, 
<collectingRegion id="355CCAC3FF94FFFBFC7DFAEBB49CFA56" box="[947,1184,1399,1423]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Región de Atacama</collectingRegion>
/ 
<collectingMunicipality id="17439E5BFF94FFFBFF59FA0BB1B6FA76" box="[151,394,1431,1455]" pageId="4" pageNumber="277">Provincia de Huasco</collectingMunicipality>
/ 
<location id="F24752FAFF94FFFBFE69FA0BB27BFA76" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFE69FA0BB27BFA76" box="[423,583,1431,1455]" country="Chile" municipality="Provincia de Huasco" name="Carrizal Bajo." pageId="4" pageNumber="277" stateProvince="Atacama">Carrizal Bajo.</location>
<date id="832622E1FF94FFFBFD83FA04B2FCFA69" box="[589,704,1432,1456]" pageId="4" pageNumber="277" value="2002-10-22">
<collectingDate id="9362DB09FF94FFFBFD83FA04B2FCFA69" box="[589,704,1432,1456]" pageId="4" pageNumber="277" value="2002-10-22">22-X-2002</collectingDate>
</date>
/ Leg.: Guillermo Aceituno A.” (7 
<collectionCode id="91899CE4FF94FFFBFB97FA04B492FA69" box="[1113,1198,1432,1456]" pageId="4" pageNumber="277">CGAA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFF59FA24B24BFA29" collectingDate="1998-10-15" collectionCode="MNNC, IEUMCE, MEUC, CMGG" collectorName="Leg. M. Guerrero" country="Chile" location="Huasco" pageId="4" pageNumber="277" specimenCount="1" typeStatus="Paratipos">
7 
<typeStatus id="2823BA83FF94FFFBFF63FA2BB121FA16" box="[173,285,1463,1487]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFE94FA2BB1A0FA16" box="[346,412,1463,1487]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
<location id="F24752FAFF94FFFBFE6AFA24B1C1FA16" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFE6AFA24B1C1FA16" box="[420,509,1464,1487]" country="Chile" name="Huasco" pageId="4" pageNumber="277">Huasco</location>
/ 
<locationDeviation id="E6951D03FF94FFFBFDD0FA24B337FA16" box="[542,779,1463,1488]" location="Huasco" pageId="4" pageNumber="277">
<quantity id="3060A9C4FF94FFFBFDD0FA24B25BFA16" box="[542,615,1463,1488]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="277" unit="km" value="13.0">13 km</quantity>
N de Huasco
</locationDeviation>
/ 
<date id="832622E1FF94FFFBFCE2FA2BB3A3FA16" box="[812,927,1463,1487]" pageId="4" pageNumber="277" value="1998-10-15">
<collectingDate id="9362DB09FF94FFFBFCE2FA2BB3A3FA16" box="[812,927,1463,1487]" pageId="4" pageNumber="277" value="1998-10-15">15-X-1998</collectingDate>
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFC0EFA24B4ADFA09" box="[960,1169,1464,1488]" pageId="4" pageNumber="277">Leg. M. Guerrero</collectorName>
” (1 
<collectionCode id="91899CE4FF94FFFBFF59FA44B0CEFA29" box="[151,242,1496,1520]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="4" pageNumber="277">MNNC</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFEDDFA44B1BCFA29" box="[275,384,1496,1520]" country="Chile" httpUri="http://grbio.org/cool/85ax-5dji" name="INSTITUTO DE ENTOMOLOGIA UNIVERSIDAD METROPOLITANA DE CIENCIAS DE LA EDUCACION" pageId="4" pageNumber="277" type="Other type of research institution/biorepository">IEUMCE</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFE6FFA44B1C9FA29" box="[417,501,1496,1520]" country="Chile" httpUri="http://grbio.org/cool/mqd5-c2qe" name="Universidad de Chile" pageId="4" pageNumber="277">MEUC</collectionCode>
, 4 
<collectionCode id="91899CE4FF94FFFBFDD8FA44B253FA29" box="[534,623,1496,1520]" country="Cambodia" httpUri="http://grbio.org/cool/m6jh-aff5" name="Cambodian Molecular Genetics Group" pageId="4" pageNumber="277">CMGG</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFD4CFA44B1DEF9E9" collectionCode="MNNC, IEUMCE, MEUC, PVGH, CMGG, CVMD" collectorName="Leg. P. Vidal &amp; P. Vidal" country="Chile" location="Sept." municipality="Huasco" pageId="4" pageNumber="277" specimenCount="1" stateProvince="Atacama" typeStatus="Paratipos">
21 
<typeStatus id="2823BA83FF94FFFBFD6AFA44B328FA29" box="[676,788,1496,1520]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFC9FFA44B39BFA29" box="[849,935,1496,1520]" name="Chile" pageId="4" pageNumber="277">CHILE</collectingCountry>
W 
<collectingRegion id="355CCAC3FF94FFFBFC16FA44B47DFA29" box="[984,1089,1496,1520]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Atacama</collectingRegion>
/ 
<collectingMunicipality id="17439E5BFF94FFFBFBAEFA44B485FA36" box="[1120,1209,1496,1519]" pageId="4" pageNumber="277">Huasco</collectingMunicipality>
/ 
<date id="832622E1FF94FFFBFF7EFA64B16CF9C9" box="[176,336,1528,1552]" pageId="4" pageNumber="277" value="2000-09-09">
9- 
<location id="F24752FAFF94FFFBFF01FA64B137F9C9" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFF01FA64B137F9C9" box="[207,267,1528,1552]" country="Chile" municipality="Huasco" name="Sept." pageId="4" pageNumber="277" stateProvince="Atacama">Sept.</location>
- 2000
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFEBDFA64B235F9C9" box="[371,521,1528,1552]" pageId="4" pageNumber="277">Leg. P. Vidal</collectorName>
/ colección 
<collectorName id="5A6D61F7FF94FFFBFD6FFA64B2C3F9C9" box="[673,767,1528,1552]" pageId="4" pageNumber="277">P. Vidal</collectorName>
” (3 
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, 2 
<collectionCode id="91899CE4FF94FFFBFC61FA64B420F9C9" box="[943,1052,1528,1552]" country="Chile" httpUri="http://grbio.org/cool/85ax-5dji" name="INSTITUTO DE ENTOMOLOGIA UNIVERSIDAD METROPOLITANA DE CIENCIAS DE LA EDUCACION" pageId="4" pageNumber="277" type="Other type of research institution/biorepository">IEUMCE</collectionCode>
, 2 
<collectionCode id="91899CE4FF94FFFBFB8EFA64B4A8F9C9" box="[1088,1172,1528,1552]" country="Chile" httpUri="http://grbio.org/cool/mqd5-c2qe" name="Universidad de Chile" pageId="4" pageNumber="277">MEUC</collectionCode>
, 11 
<collectionCode id="91899CE4FF94FFFBFF59F984B0DBF9E9" box="[151,231,1560,1584]" pageId="4" pageNumber="277">PVGH</collectionCode>
, 2 
<collectionCode id="91899CE4FF94FFFBFEC7F984B15CF9E9" box="[265,352,1560,1584]" country="Cambodia" httpUri="http://grbio.org/cool/m6jh-aff5" name="Cambodian Molecular Genetics Group" pageId="4" pageNumber="277">CMGG</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFE4FF984B1E5F9E9" box="[385,473,1560,1584]" pageId="4" pageNumber="277">CVMD</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFE23F984B1D0F989" collectingDate="2002-10" collectionCode="CSRR" collectorName="Leg. S. Roitman" country="Chile" location="Huasco" pageId="4" pageNumber="277" specimenCount="1" typeStatus="Paratipos">
3 
<typeStatus id="2823BA83FF94FFFBFDCCF984B24EF9E9" box="[514,626,1560,1584]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFD61F984B2CDF9E9" box="[687,753,1560,1584]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
<location id="F24752FAFF94FFFBFD37F984B36EF9F6" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFD37F984B36EF9F6" box="[761,850,1560,1583]" country="Chile" name="Huasco" pageId="4" pageNumber="277">Huasco</location>
/ 
<locationDeviation id="E6951D03FF94FFFBFCBCF984B410F9F6" box="[882,1068,1560,1584]" location="Huasco" pageId="4" pageNumber="277">
<quantity id="3060A9C4FF94FFFBFCBCF984B391F9E9" box="[882,941,1560,1584]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="277" unit="km" value="7.0">7 km</quantity>
N Huasco
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/ 
<date id="832622E1FF94FFFBFB82F984B485F9E9" box="[1100,1209,1560,1584]" pageId="4" pageNumber="277" value="2002-10">
<collectingDate id="9362DB09FF94FFFBFB82F984B485F9E9" box="[1100,1209,1560,1584]" pageId="4" pageNumber="277" value="2002-10">Oct. 2002</collectingDate>
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFF60F9A5B150F989" box="[174,364,1592,1617]" pageId="4" pageNumber="277">Leg. S. Roitman</collectorName>
” (3 
<collectionCode id="91899CE4FF94FFFBFE55F9A4B1D8F989" box="[411,484,1592,1616]" pageId="4" pageNumber="277">CSRR</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFE38F9A5B439F9A9" collectionCode="R, MNNC, IEUMCE, MEUC, PVGH" collectorName="N. Llanos de Challe &amp; Coleccion P. Vidal" country="Chile" location="Atacama" pageId="4" pageNumber="277" specimenCount="1" stateProvince="Atacama" typeStatus="Paratipos">
7 
<typeStatus id="2823BA83FF94FFFBFDC4F9A4B246F989" box="[522,634,1592,1616]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFD78F9A4B330F989" box="[694,780,1592,1616]" name="Chile" pageId="4" pageNumber="277">CHILE</collectingCountry>
<collectingRegion id="355CCAC3FF94FFFBFCD7F9A4B3BEF989" box="[793,898,1592,1616]" country="Chile" name="Atacama" pageId="4" pageNumber="277">Atacama</collectingRegion>
/ 
<collectionCode id="91899CE4FF94FFFBFC6EF9A5B38AF989" box="[928,950,1593,1616]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="277">R</collectionCode>
. 
<collectorName id="5A6D61F7FF94FFFBFC74F9A5B49EF989" box="[954,1186,1592,1616]" pageId="4" pageNumber="277">N. Llanos de Challe</collectorName>
/ 
<date id="832622E1FF94FFFBFF59F9C4B123F9A9" box="[151,287,1624,1648]" pageId="4" pageNumber="277" value="2000-11-05">5-Nov. 2000</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFEF5F9C4B233F9A9" box="[315,527,1624,1648]" pageId="4" pageNumber="277">Colección P. Vidal</collectorName>
” (1 
<collectionCode id="91899CE4FF94FFFBFDF4F9C4B2A9F9A9" box="[570,661,1624,1648]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="4" pageNumber="277">MNNC</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFD7FF9C4B322F9A9" box="[689,798,1624,1648]" country="Chile" httpUri="http://grbio.org/cool/85ax-5dji" name="INSTITUTO DE ENTOMOLOGIA UNIVERSIDAD METROPOLITANA DE CIENCIAS DE LA EDUCACION" pageId="4" pageNumber="277" type="Other type of research institution/biorepository">IEUMCE</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFCF5F9C4B3B3F9A9" box="[827,911,1624,1648]" country="Chile" httpUri="http://grbio.org/cool/mqd5-c2qe" name="Universidad de Chile" pageId="4" pageNumber="277">MEUC</collectionCode>
, 4 
<collectionCode id="91899CE4FF94FFFBFC65F9C4B3C1F9A9" box="[939,1021,1624,1648]" pageId="4" pageNumber="277">PVGH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFBC0F9C4B12DF968" collectingDate="2000-11-05" collectionCode="MNNC, IEUMCE, CMGG" collectorName="Leg. M. Guerrero" country="Chile" location="Llanos de Challe" municipality="Huasco" pageId="4" pageNumber="277" specimenCount="1" typeStatus="Paratipos">
6 
<typeStatus id="2823BA83FF94FFFBFBEFF9C4B4ADF9A9" box="[1057,1169,1624,1648]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
de: “ 
<collectingCountry id="8F8F44B1FF94FFFBFF6CF9E4B0D8F949" box="[162,228,1656,1680]" name="Chile" pageId="4" pageNumber="277">Chile</collectingCountry>
<collectingMunicipality id="17439E5BFF94FFFBFF27F9E5B17EF949" box="[233,322,1657,1680]" pageId="4" pageNumber="277">Huasco</collectingMunicipality>
/ 
<location id="F24752FAFF94FFFBFE93F9E4B21CF949" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFE93F9E4B21CF949" box="[349,544,1656,1680]" country="Chile" municipality="Huasco" name="Llanos de Challe" pageId="4" pageNumber="277">Llanos de Challe</location>
/ 
<date id="832622E1FF94FFFBFDF2F9E4B282F949" box="[572,702,1656,1680]" pageId="4" pageNumber="277" value="2000-11-05">
<collectingDate id="9362DB09FF94FFFBFDF2F9E4B282F949" box="[572,702,1656,1680]" pageId="4" pageNumber="277" value="2000-11-05">05-XI- 2000</collectingDate>
</date>
/ 
<collectorName id="5A6D61F7FF94FFFBFD17F9E5B399F949" box="[729,933,1656,1681]" pageId="4" pageNumber="277">Leg. M. Guerrero</collectorName>
” (1 
<collectionCode id="91899CE4FF94FFFBFC1FF9E4B410F949" box="[977,1068,1656,1680]" country="Chile" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="4" pageNumber="277">MNNC</collectionCode>
, 1 
<collectionCode id="91899CE4FF94FFFBFB86F9E4B489F949" box="[1096,1205,1656,1680]" country="Chile" httpUri="http://grbio.org/cool/85ax-5dji" name="INSTITUTO DE ENTOMOLOGIA UNIVERSIDAD METROPOLITANA DE CIENCIAS DE LA EDUCACION" pageId="4" pageNumber="277" type="Other type of research institution/biorepository">IEUMCE</collectionCode>
, 4 
<collectionCode id="91899CE4FF94FFFBFF7EF905B135F968" box="[176,265,1689,1713]" country="Cambodia" httpUri="http://grbio.org/cool/m6jh-aff5" name="Cambodian Molecular Genetics Group" pageId="4" pageNumber="277">CMGG</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="47F00E7CFF94FFFBFEEEF904B2C1F908" country="Chile" determinerName="Guerrero &amp; Aceituno" location="Todos" pageId="4" pageNumber="277" specimenCount="2" typeStatus="Paratipos">
<location id="F24752FAFF94FFFBFEEEF904B15BF969" LSID="urn:lsid:plazi:treatment:7F31B537FF93FFFAFE8CFB01B2B8FC96:F24752FAFF94FFFBFEEEF904B15BF969" box="[288,359,1688,1712]" country="Chile" name="Todos" pageId="4" pageNumber="277">Todos</location>
los 
<typeStatus id="2823BA83FF94FFFBFE51F904B233F969" box="[415,527,1688,1712]" pageId="4" pageNumber="277" type="paratype">paratipos</typeStatus>
portan la siguiente etiqueta: “ 
<typeStatus id="2823BA83FF94FFFBFC49F905B42CF968" box="[903,1040,1689,1713]" pageId="4" pageNumber="277" type="paratype">PARATIPO</typeStatus>
/ 
<taxonomicName id="30987FA2FF94FFFBFBF9F905B13BF908" authorityName="Guerrero &amp; Aceituno" authorityYear="2020" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="277" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="C5ECD833FF94FFFBFBF9F905B13BF908" italics="true" pageId="4" pageNumber="277">Gyriosomus maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEDF6548FF94FFFBFEC3F925B15BF909" box="[269,359,1721,1744]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
/ det. 
<determinerName id="02FBE8E5FF94FFFBFE70F925B215F908" box="[446,553,1721,1745]" pageId="4" pageNumber="277">Guerrero</determinerName>
y 
<determinerName id="02FBE8E5FF94FFFBFD8BF925B28DF908" box="[581,689,1721,1745]" pageId="4" pageNumber="277">Aceituno</determinerName>
2020”
</materialsCitation>
.
</paragraph>
<paragraph id="F7270421FF95FFFAFF95FFE0B3C9FF0D" blockId="5.[91,1149,124,213]" pageId="5" pageNumber="278">
<emphasis id="C5ECD833FF95FFFAFF95FFE0B1B7FF4D" bold="true" box="[91,395,124,148]" pageId="5" pageNumber="278">Otro material estudiado.</emphasis>
<taxonomicName id="30987FA2FF95FFFAFE54FFE1B354FF4C" authority="Guerin-Meneville" authorityName="Guerin-Meneville" box="[410,872,125,149]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C5ECD833FF95FFFAFE54FFE1B2ACFF4D" box="[410,656,125,148]" italics="true" pageId="5" pageNumber="278">Gyriosomus impressus</emphasis>
Guérin-Méneville
</taxonomicName>
(
<figureCitation id="6FA318A4FF95FFFAFCB1FFE1B3ECFF4C" box="[895,976,125,149]" captionStart="Figura 3" captionStartId="10.[152,223,1673,1695]" captionTargetBox="[152,1182,153,1624]" captionTargetId="figure-73@10.[588,796,723,1128]" captionTargetPageId="10" captionText="Figura 3. Gyriosomus impressus Guérin-Méneville.A-C. Sintipo hembra, vistas dorsal, ventral y lateral. D. Detalle del prosterno. E. Etiquetas. F-G. Ejemplar hembra de Cuesta El Gato, Copiapó, vistas dorsal y lateral (escalas: A, B, C y F = 5 mm). Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205008" httpUri="https://zenodo.org/record/13205008/files/figure.png" pageId="5" pageNumber="278">Figs. 3</figureCitation>
A-3E, 3F, 3G). 1 ejemplar sintipo depositado en el 
<collectionCode id="91899CE4FF95FFFAFE31FF01B265FF6D" box="[511,601,157,180]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="278">MNHN</collectionCode>
(
<figureCitation id="6FA318A4FF95FFFAFDA5FF01B28AFF6C" box="[619,694,157,181]" captionStart="Figura 3" captionStartId="10.[152,223,1673,1695]" captionTargetBox="[152,1182,153,1624]" captionTargetId="figure-73@10.[588,796,723,1128]" captionTargetPageId="10" captionText="Figura 3. Gyriosomus impressus Guérin-Méneville.A-C. Sintipo hembra, vistas dorsal, ventral y lateral. D. Detalle del prosterno. E. Etiquetas. F-G. Ejemplar hembra de Cuesta El Gato, Copiapó, vistas dorsal y lateral (escalas: A, B, C y F = 5 mm). Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205008" httpUri="https://zenodo.org/record/13205008/files/figure.png" pageId="5" pageNumber="278">Figs. 3</figureCitation>
A-3E). 1 ejemplar de: 
<collectingCountry id="8F8F44B1FF95FFFAFC78FF01B3C9FF6C" box="[950,1013,157,181]" name="Chile" pageId="5" pageNumber="278">Chile</collectingCountry>
, III Región, Cordillera de Copiapó, 3.400 msnm, Cuesta El Gato, 
<date id="832622E1FF95FFFAFD7CFF20B323FF0D" box="[690,799,188,212]" pageId="5" pageNumber="278" value="2016-01-09">9-01-2016</date>
, Leg. Carlos Vidal.
</paragraph>
<paragraph id="F7270421FF95FFFAFF95FF67B242FE4A" blockId="5.[91,1149,251,403]" pageId="5" pageNumber="278">
<emphasis id="C5ECD833FF95FFFAFF95FF67B086FECA" bold="true" box="[91,186,251,275]" pageId="5" pageNumber="278">Hábitat.</emphasis>
Esta especie habita planicies costeras en donde predominan suelos arenosos y pedregosos con presencia de vegetación herbácea y arbustiva caracterÍstica de la costa de la 
<collectingRegion id="355CCAC3FF95FFFAFF95FEA7B101FE8D" box="[91,317,315,340]" country="Chile" name="Atacama" pageId="5" pageNumber="278">Región de Atacama</collectingRegion>
(
<figureCitation id="6FA318A4FF95FFFAFE83FEA7B1A1FE8D" box="[333,413,315,340]" captionStart="Figuras 8-14" captionStartId="15.[91,169,1568,1590]" captionTargetBox="[91,1150,122,1555]" captionTargetPageId="15" captionText="Figuras 8-14. Hábitats de Gyriosomus spp. 8. Chañaral Alto, Provincia de LimarÍ, hábitat de G. resplendens Aceituno y Guerrero sp. nov. 9. Huasco, Provincia de Huasco, hábitat de G. maculatus Guerrero y Aceituno sp. nov. 10. Cuesta El Gato, Provincia de Copiapó, hábitat de G. impressus. 11. Valle Hermoso, Provincia de LimarÍ, hábitat de G. nigrociliatus Guerrero y Aceituno sp. nov. 12. Cordillera de Illapel, Provincia de Choapa, hábitat de G. granocostatus. 13. Tabaqueros, Provincia de LimarÍ, hábitat de G. laevis stat. nov. 14. Cuesta El Espino, Provincia de Choapa, hábitat de G. foveopunctatus." figureDoi="http://doi.org/10.5281/zenodo.13205020" httpUri="https://zenodo.org/record/13205020/files/figure.png" pageId="5" pageNumber="278">Fig. 10</figureCitation>
). Los adultos se refugian bajo plantas que crecen a ras de suelo durante las horas de mayor calor, estando más activos en horas de la mañana y al atardecer, momentos en los cuales se aparean y alimentan.
</paragraph>
<paragraph id="F7270421FF95FFFAFF95FE25B17CFDCB" blockId="5.[91,1149,437,530]" pageId="5" pageNumber="278">
<emphasis id="C5ECD833FF95FFFAFF95FE25B0DFFE0B" bold="true" box="[91,227,441,466]" pageId="5" pageNumber="278">Etimología.</emphasis>
El nombre especÍfico hace referencia a la gran cantidad de impresiones tegumentarias (manchas) blanquecinas presentes en los élitros; del latÍn 
<taxonomicName id="30987FA2FF95FFFAFC79FE46B400FE2B" authorityName="Guerrero &amp; Aceituno" authorityYear="2020" box="[951,1084,474,498]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="maculatus">“maculatus</taxonomicName>
” que significa manchado.
</paragraph>
<paragraph id="F7270421FF95FFFAFF95FDA5B2B8FC96" blockId="5.[91,1150,569,847]" pageId="5" pageNumber="278">
<emphasis id="C5ECD833FF95FFFAFF95FDA5B0C4FD88" bold="true" box="[91,248,569,593]" pageId="5" pageNumber="278">Comentarios.</emphasis>
Esta especie es cercana a 
<taxonomicName id="30987FA2FF95FFFAFDFCFDA6B3C6FD88" authority="Guerin-Meneville" authorityName="Guerin-Meneville" box="[562,1018,569,593]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C5ECD833FF95FFFAFDFCFDA6B31FFD88" box="[562,803,570,593]" italics="true" pageId="5" pageNumber="278">Gyriosomus impressus</emphasis>
Guérin-Méneville
</taxonomicName>
, de la cual se diferencia por la coloración negra brillante. Pronoto con un surco longitudinal medial profundo; gibosidad inclinada anteriormente. Élitros con el margen anterior no observable en vista dorsal, con la superficie lisa y brillante, con abundantes máculas o impresiones tegumentarias y puntuaciones cubiertas con pilosidad blanca en toda la superficie (
<figureCitation id="6FA318A4FF95FFFAFB89FD24B097FD29" captionStart="Figura 2" captionStartId="9.[94,161,1663,1685]" captionTargetBox="[94,557,1164,1557]" captionTargetId="figure-107@9.[91,559,1161,1560]" captionTargetPageId="9" captionText="Figura 2.Gyriosomus maculatus Guerrero yAceituno sp. nov. A.Alotipo hembra, vista dorsal.B-D.Holotipo macho, vistas dorsal, ventral y lateral. E. Detalle del prosterno. F-H. Edeago, vistas dorsal, lateral y ventral (escalas:A, B, C y D= 5 mm; F, G y H= 1 mm).Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205006" httpUri="https://zenodo.org/record/13205006/files/figure.png" pageId="5" pageNumber="278">Figs. 2A, 2B</figureCitation>
). No asÍ en 
<taxonomicName id="30987FA2FF95FFFAFEF8FD44B1FCFD29" box="[310,448,728,752]" class="Insecta" family="Tenebrionidae" genus="Gyriosomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="C5ECD833FF95FFFAFEF8FD44B1FCFD29" box="[310,448,728,752]" italics="true" pageId="5" pageNumber="278">G. impressus</emphasis>
</taxonomicName>
que presenta una coloración negra opaca. Pronoto sin surco longitudinal medial y el margen anterior observable en vista dorsal; gibosidad inclinada hacia atrás. Élitros ligeramente lisos y opacos, con escasas máculas o impresiones tegumentarias sólo visibles hacia los costados (
<figureCitation id="6FA318A4FF95FFFAFE0BFCABB244FC96" box="[453,632,823,847]" captionStart="Figura 3" captionStartId="10.[152,223,1673,1695]" captionTargetBox="[152,1182,153,1624]" captionTargetId="figure-73@10.[588,796,723,1128]" captionTargetPageId="10" captionText="Figura 3. Gyriosomus impressus Guérin-Méneville.A-C. Sintipo hembra, vistas dorsal, ventral y lateral. D. Detalle del prosterno. E. Etiquetas. F-G. Ejemplar hembra de Cuesta El Gato, Copiapó, vistas dorsal y lateral (escalas: A, B, C y F = 5 mm). Abreviaturas: Mgl= Margen lateral, Pt= Prosterno." figureDoi="http://doi.org/10.5281/zenodo.13205008" httpUri="https://zenodo.org/record/13205008/files/figure.png" pageId="5" pageNumber="278">Figs. 3A, 3F, 3G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>